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THE 


PROCEEDINGS 


LINNEAN  SOCIETY 


NEW  SOUTH  WALES. 


FOR  TflE   YEAR 


1Q0  2. 


Vol.  XXVII. 


VriXH    THIRTY-XKCREE    I>LA.XES. 


PRIISTED  AND  PUBLISHED  FOR  THE  SOCIETY 

BY 

F.    CUNNINGHAME    &   CO.,    146   PITT    STREET 

AND 

SOLD  BY  THE  SOCIETY. 
1902-03. 


^^7pi- 


SYDNEY : 
F.    CUNNINGHAME    AND    CO.,    PIl^NT 
PITT    STREET. 


LI^T  OF  CONTRIBUTORS  AND  TITLES  OF  PAPERS. 

.r 

'  Andrews,  E.  C,  B.A. —  page 

Preliminary  Note  on  the  Geology  of  the  Queensland 
Coast,  ^YJ^l  References  to  the  Geography  of  the 
Queensland  and  New  South  Wales  Plateau. 
(Wit^nine  Text-figures) M^o 

Baker,  R.  T.,  F.L.S.,  Curator  and  Economic  Botanist, 
Technological  Museum,  Sydney — 

On  Eucalyptus  melanopJdoia,  F.v.M'  and  its  cognate 

Sp^es.    ;(Plate  xi.)  ..        225 

On  a  new  Species  of  Ardisia  from  New  South  Wales. 

(Plate  xvi.) 380 

Contributions  to  a  Knowledge  of  the  Flora  of  Aus- 
tralia.    Part  iv.     (Plates  xxii.-xxiii.)      ...  ...      536 

On   a  new    Species    of    Symjdocos    from    New    South 

Wales.     (Plate  xxviii.)      594 

Betche,  E.,  and  J.  H.  Maipei^j-F.L.S. — 

Notes  from  the  Botanic  Gardens,  Sydney.     No.  8.    ...        55 

Broom,  R.,  M.D.,  B.Sc,  C.M.Z.S.,  Corresponding  Member — 
On  the  Mammalian  and  Reptilian  Vomerine  Bones. 

(Plates  xxiv.-xxvi.)  ...  ...  ...  ...     545 

Cambagk,  R.  H.,  L.S. — 

Notes  on  the  Botany  of  the  Interior  of  New  South 

Wales.     Part  vi.     From  Marsden  to  Narrandera. 

(Plates  viii.-x.)       186 

Notes  on  the  Botany  of  the  Interior  of  New  South 

Wales.     Part    vii.      From    Forbes    to    Bathurst. 

(Plate  xxvii.)  561 

Cheel,  E. — 

Notes  on  Juticus  holoscJuenus,  R.Br.,  and  J.  2)^ismato- 
carpus,  R.Br.;  and  certain  other  New  South 
Wales  Plants  210 


IV.  LIST    OF    CONTRIBUTORS    AND    TITLES    OF    PAPERS. 

Fitzgerald,  VV.  V.,  F.S.Sc,  Lend.,  F.R.H.S.,  Eng.—  page 

Notes  on  some  hitherto  unrecorded  Species  of  Plants 
indigenous  in  the  State  of  Western  Australia. 
{Communicated  hy  J.  H.  Maiden,  F.L.S.,  tf&c.)  ...      241 

Forsyth,  W. — 

Some  Records  of  New  South  Wales  Mosses.     [Title 

only.]  .369 

Froggatt,  Walter  W.,  F.L.S. — 

Notes  on  Australian  Neuroptera  and  their  Life- 
Histories     ...  ...  ...  ...  ...  ...     358 

Hall,  T.  S.,  M.A.— 

On    the   Occurrence   of   Monograptus  in    New   South 

Wales.     (Communicated  hy  John  Mitchell)         ...      654 

Haswell,  W.  a.,  M.A.,  D.Sc,  F.R.S.,  Challis  Professor  of 

Biology,  University  of  Sydney — 
On  a  Gyrocotyle  from  Chimoira  Ogilhyi,  and  on  Gyro- 

cotyle  in  general.     (Plate  vii.)      ...  ...  ...        48 

On  two  remarkable  Sporocysts  occurring  in  Jlytilus 

latus   on    the    Coast   of    New  Zealand.      (Plates 

xix.-xx.)      ...  ...  ...  ...  ...  ...      497 

Hedley,  Charles,  F.L.S. — 

Studies    on    Australian    Mollusca.     Part  vi.      (Plates 

i.-iii.)  7 

Studies   on   Australian   Mollusca.     Part  vii.     (Plates 

xxix.,  xxxi.,  xxxii.,  xxxiii.)  ...  ...  ...      596 

Johnston,  S.  J.,  B.A.,  B.Sc,  Economic  Zoologist,  Techno- 
logical Museum,  Sydney — 
Contributions  to  a  Knowledge  of  Australian  Entozoa. 
No.  ii.  On  a  New  Species  of  Distomum  from  the 
Sawfish  Shark,  Pristiophorus  cirratus,  Lath. 
(Plate  xiii.) 326 


list  of  contributors  and  titles  of  papers.  v. 

Kesteven,  H.  Leighton—  page 

A  Note  on  two  Species  of  Astralium  from  Port  Jackson  2 
Notes  on  Prosobranchiata.     No,  i.  Lotorium.     (Plate 

xvii.)             443 

Notes  on  Prosobranchiata.   No.  ii.  Littorinacea.    (Plate 

XXX.)             620 

Lea,  Arthur  M.,  F.E.S.— 

Revision  of  the  Australian  Curculionidce  belonging  to 

the  Subfamily  Cryptorhynchides.     Part  v.  ...      408 

Lucas,  Thomas  P.,  L.R.C.P.,  Ed.,  L.S.A.,  LoncL,  M.R.C.S., 
Eng.— 
New  Species  of  Queensland  Lepidoptera        ...  ...      246 

JMcAlpink,  D.,  Corresponding  Member — 

Australian  Fungi,  New  or  Unrecorded.     Decades  i.-ii.     373 

MacPherson,  John,  M.A.,  B.Sc,  M.B.,  Ch.M.— 

Ngarrabul    and    other    Aboriginal    Tribes.       Part   i. 

Medical  and  Surgical  Practice      ...  ...  ...      637 

Maiden,  J.  H.,  F.L.S.— 

4. — On  Eucalyptus  Bauer  I  ana,  ^ch.?i,\xQV  ...  ...      214 

5. — On  Eucalyptus  calycogono,  Turcz —  ...  ...      220 

On    a    new    Cryptocarya    from     Lord     Howe    Island, 
together  with  Notes  on  other  Plants  from  that 

Island.     (Plate  XV.)  347 

6. — On  Eucalyptus  bicolor,  A.  Cunn.  ...  ...  ...      516 

7.— On  Eucalyptus  polyantheitios,  ^chuuev.    (Plate  xxi.)     527 

Maiden,  J.  IL,  F.L.S.,  and  E.  Betche — 

Notes  from  the  Botanic  Gardens,  Sydney.      No.  8     ...        55 

Rainbow,  W.  J.,  F.L.S.,  Entomologist  to  the  Australian 
Museum — 
Descriptions   of  some   new  Ara^ieidce  of   New   South 

Wales.     No.  X.     (Plate  xviii.) 484 

8loane,  Thomas  G. — 

A  Revision  of  the  Genus  Xotonomus  (Family  Carabidce; 

Subfamily  Feronini)  ...  ...  ...  ...      252 


VI.  LIST    OF    CONTRIBUTORS    AND    TITLES    OF    PAPERS. 


PAGE 


Smith,  R.  Greig,  M.Sc,  Macleay  Bacteriologist  to  the 
Society — 

The  Gummosis  of  the  Sugar-Caiie  {Bad.  rascularicm, 

Cobb).     (Plates  iv.-v.)       31 

Further  Remarks  upon  the  Mechanism  of  Agglutina- 
tion ...  ...  ...  ...  ...  ...  ...        66 

An  Ascobacterium  from  the  Sugar-Cane,  with  Notes 
upon  the  Nature  of  the  Slime  [Bacterium  sacckari, 
n.  sp.)     (Plate  vi.) 137 

A  Gum  (Levan)  Bacterium  from  a  Saccharine  Exudate 
of  Eucalyptus  K^tuartiana  (^Bacterium  eucalypti^ 
n.  sp.)     (Plate  xii.)  230 

The  Ulcer  Disease  (Black  Ophthalmia?)  of  Rainbow 

Trout  {Micrococcus  pyogenes)        ...  ...  ...      3-52 

The  Bacterial  Origin  of  the  Gums  of  the  Arabin  Group     383 

Turner,  A.  Jefferis,  M.D.,  F.E.S.— 

New  Genera  and  Species  of  Lepidoptera  belonging  to 

the  Family  Noctuidce  ...  ...  ...  ...  7 

Waterhouse,  G.  a.,  B.Sc,  B.E.,  F.E.S.— 

Notes  on  Australian  Rhoyalocera.    Lyccenidce.  Part  i. 

(Plate  xiv.) 331 

Notes  on  Australian  Bhopalocera.    Lyccenidce.    Fart  ii.      648 

Watts,  Rev.  W.  Walter — 

Notes  on  some  New  South  Wales  Hepatics      ..  ...      493 

Watts,  Rev.  W.  Walter,  and  Thomas  Whitelegge, 
F.R.M.S.— 

Whitelegge,  Thomas,  F.R.M.S.,  and  Rev.  W.  Walter 
Watts — 
Census  Muscorum  Australiensium.  A  classified  Cata- 
logue of  the  Frondose  Mosses  of  Australia  and 
Tasmania,  collated  from  available  Publications 
and  Herbaria  Records.  Part  i.  (Issued  separately 
as  a  Supplement  to  Part  3  of  the  Proceedings)  ...      369 


CONTENTS  OF  PROCEEDINGS,  1902. 

PART  I.     (No.   105). 

(Issued  August  22nd,  1902.) 

PAGE 
A  Note  on  two  Species  of    Astralinni  from   Port  Jackson.      By  H. 

Leighton  Kesteven     ...         ...         ...         ...         ...         ...         ...  2 

Studies  on  Australian  Mollusca.     Part  vi.      By  C.  Hedley,  F.L.S. 

(Plates  i.-iii.)      7 

The  Gummosis  of  the  Sugar-Cane  {Bact.  vascularum,  Cobb).     By  R. 

Greig    Smith,   M.Sc,  Macleay   Bacteriologist    to    the   Society. 

(Plates  iv.-v.) 31 

On  a  Gyrocotyle  from  Chinuera  Ogilbyi,  and  on  Gyrocotyle  in  general. 

By  W.  A.   Haswell,  M.A.,  D.Sc,  F.R.S.,  Challis   Professor   of 

Biology,  University  of  Sydney.     (Plate  vii.)  ..  ...         ...         48 

Notes  from  the  Botanic  Gardens,  Sydney.     No,  8.     By  J.  H.  Maiden 

and  E.  Betche ...         55 

Further  Remarks   upon   the    Mechanism   of    Agglutination.     By  R. 

Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the  Society         ...         66 
Note  accompanying  an  exhibit  of  Persoonia  angulata,  R.Br.,  Uncinia 

tenella,  R.Br.,  and  Carex  gracilis,  R.Br.     By  A.  A.  Hamilton  ...         73 
Note  in  Rectification  of  the  Synonymy  of  Eopsaltria  georgiana,  Quoy  & 

Gaim.,  and  E.  gularis,  Quoy  &  Gaim.     By  A.  J.  North,  C.M.Z.S.         74 
New  Genera  and  Species  of   Lepidoptera  belonging  to  the  Family 

Noctuidce.     By  A.  Jefferis  Turner,  M.D.,  F.E.S 77 

An  Ascobacterium  from  the  Sugar-Cane,  with  Notes  upon  the  Nature 

of    the  Slime  {Bacterium  sacchari,  n.sp.)     By  R.  Greig  Smith, 

M.Sc,  Macleay  Bacteriologist  to  the  Society.     (Plate  vi.)  ...       137 

Elections  and  Announcements       ...         ...         ...         ...         30,76 

Notes  and  Exhibits  ...         ..  ...         ...         ...         ...         ...         ...30,73 


PART  II.     (No.  106). 

(Issued  October  17th,  1902.) 


PAGE 


Preliminary  Note  on  the  Geology  of  the  Queensland  Coast,  with 
References  to  the  Geography  of  the  Queensland  and  N.  S.  Wales 
Plateau.     By  E.  C.  Andrews,  B. A.     (With  nine  Text-figures)  ...       146 

Notes  on  the  Botany  of  the  Interior  of  New  South  Wales.     Part  vi. 

•From  Marsden  to  Narrandera.    By  R.  H.  Cambage.    (Plates  viii.-x. )     186 


Vlll.  CONTENTS. 

PART   II.   (continued.) 

PAGE 

Note  on  some  Northern  and  North- Western  Australian  Grass  Finches. 

By  A.  J.  North,  CM. Z.S 207 

Notes  on  Jiuicus  holoscJuenus,  B.Bv.,  ?ind  J.  imsmatocarpus,  R.Br.; 

and  on  certain  other  New  South  Wales  Plants.     By  Edwin  Cheel       210 

4. — On  Eucalyiytus  Baueriana,  ^cho^nev.     By  J.  H.  Maiden 214 

5.— On  Eucalyptus  calycogona,  Tnvcz.     By  J.  H.  Maiden      220 

On  Eucalyptus  melanophloia,  F.v.M.,  and  its  cognate  Species.  By 
R.  T.  Baker,  F.L.S.,  Curator  and  Economic  Botanist,  Technolo- 
gical Museum,  Sydney.     (Plate  xi.) 225 

A  Gum  (Levan)  Bacterium  from  a  Saccharine  Exudate  of  Eucalyptus 
Stuartiana  {Bacterium  eucalypti,  n.sp.).  By  R.  Greig  Smith, 
M.Sc,  Macleay  Bacteriologist  to  the  Society.     (Plate  xii.)  ...       230 

Notes  on  some  hitherto  unrecorded  Species  of  Plants  indigenous  in 
the  State  of  Western  Australia.  By  W.  Y.  Fitzgerald,  F.S.Sc, 
Lond.,  F.R.H.S.,  Eng.    {Communicated  by  J.  H.  Maiden,  F.L.S.)       241 

New   Species   of    Queensland   Lepidoptera.     By   Thomas   P.    Lucas, 

L.R.C.P.,Ed.,  L.S.A.,Lond.,  M.R.C.S.,Eng 246 

A  Revision  of    the   Genus  Nutonomu'i  (Family  Carahidce;  Subfamily 

Feronini).     By  Thomas  G.  Sloane 252 

Elections  and  Announcements       ...  209,  240 

Notes  and  Exhibits 205,237 


page 


PART  III.     (No.   107). 

(Issued  December  loth,  1902.) 

Contributions  to  a  Knowledge  of  Australian  Entozoa.  No.  ii.  On  a 
new  Species  of  Distomum  from  the  Sawfish  Shark,  Pristiojyhorus 
cirratns,  Lath.  By  S.  J.  Johnston,  B.A.,  B.Sc,  Economic 
Zoologist,  Technological  Museum,  Sydney,     (Plate  xiii. )  ...       326 

Notes   on   Australian   Bhopalocera,     Lycccnidce.  Part  i.      By  G.   A. 

Waterhouse,  B.Sc,  B.E.,  F.E.S.     (Plate  xiv.) 331 

On  a  new  Cryptocarya  from  Lord  Howe  Island,  together  with  Notes 

on  other  Plants  from  that  Island.    By  J.  H.  Maiden.    (Plate  xv.)      347 

The  Ulcer  Disease  (Black  Ophthalmia?)  of  Rainbow  Trout  f^J/«'cro- 
coccus  pyogenes).  By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteri- 
ologist to  the  Society    352 


CONTENTS.  IX. 

PART  III.  (continued). 

PAGE 

Notes  on  Australian  Neuroptera  and  their  Life-Histories.    By  Walter 

W.  Froggatt,  F.L.S 358 

Some   Records   of    New   South   Wales   Mosses.      By  W.   Forsyth. 

[Title  only]         3G9 

Census  Muscorum  Australiensium.  A  classified  Catalogue  of  the  Fron- 
dose  Mosses  of  Australia  and  Tasmania,  collated  from  available 
Publications  and  Herbaria  Records,  by  the  Rev.  W.  Walter 
Watts  and  Thomas  Whitelegge,  F.R.M.S.  Part  i.  [Issued 
separately  as  a  Supplement  to  this  Part  of  the  Proceedings]      ...       369 

Australian    Fungi,    New    or    Unrecorded.     Decades   i.-ii.      By    D. 

Mc Alpine,  Corresponding  Member 373 

On  a  new  Species  of  Ardisia  from  New  South  Wales.  By  R.  T. 
Baker,  F.L.S.,  Curator,  Technological  Museum,  Sydney.  (Plate 
xvi.)        380 

The   Bacterial   Origin  of    the   Gums  of    the  Arabin  Group.     By  R. 

Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the  Society       ...       383 

Revision  of  the  Australian  Curcnlionidce  belonging  to  the  Subfamily 

Cnjptorhynchides.     Part  v.     By  Arthur  M.  Lea,  F.E.S.  ...       408 

Notes   on   Prosohranchiata.      No.    i.    Lotorium.     By   H.    Leighton 

Kesteven.     (Plate  xvii.)       ...  443 

Descriptions  of  some  new  Araneidcr.  of    New  South  Wales.     No.  x. 
By  W.    J.  Rainbow,   F.L.S.,    Entomologist  to  the  Australian 
Museum.     (Plate  xviii.)        ...         ...  ...         ...         ...         ...         484 

Elections  and  Announcements      ...  ..  ...         ...         ...  346,  372 

Notes  and  Exhibits 343,370 


PART  IV.     (No.  108). 

(Umed  April  9th   lOOJ.) 

PAGE 

Notes  on  some  New  South  Wales  Hepatics.     By  Rev.  W.   Walter 

Watts        493 

On  two  remarkable  Sporocysts  occurring  in  Mytilus  latrn^,  on  the  Coast 
of  New  Zealand.  By  Professor  W.  A.  Haswell,  M.A.,  D.Sc, 
F.R.S.     (Plates  xix. -XX.)  497 

6. — On  Eucahj2}tus  hicolor,  A.  Cnnn.     By  J.  H.  Maiden  516 

".  —  On  Eucalyptus  jiolyantheiiwfi,  Sch^ner.     By  J.  H.  Maiden.     (Plate 

xxi.)  527 


X.  CONTENTS. 

PART  IV.  (continued). 

PAGE 

Contributions  to  a  Knowledge  of  the  Flora  of  Australia.  Part  iv.  By 
R.  T.  Baker,  F.  L.S.,  Curator  and  Economic  Botanist,  Technolo- 
gical Museum,  Sydney.     (Plates  xxii.-xxiii.) 536 

On  the  Mammalian  and  Reptilian  Vomerine  Bones.  By  R.  Broom, 
M.D.,  B.Sc,  C.M.Z.S.,  Corresponding  Member.  (Plates  xxiv.- 
xxvi.)         545 

Notes  on  the  Botany  of  the  Interior  of  New  South  Wales.     Part  vii. — 

From  Forbes  to  Bathurst.     By  R.  H.  Cambage,     (Plate  xxvii. )    ...     561 

On  a  New  Species  of  SymiJlocos  from  New  South  Wales.  By  R.  T. 
Baker,  F.L.S.,  Curator,  Technological  Museum,  Sydney.  (Plate 
xxviii.)       594 

Studies  on  Australian   Mollusca.     Part  vii.     By  C.    Hebley,  F.L.S. 

(Plates  xxix.,  xxxi.,  xxxii.,  xxxiii.) 596 

Notes  on  Prosohranclnata.       No.  ii.  TAttorinacca.        By  H.  Leighton 

Kesteven.     (Plate  xxx.)  ...         ...  ...          ...  ...  ...     620 

Ngarrabul  and  other  Aboriginal  Tribes,      Part  i.  Medical  and  Surgical 

Practice.     By  John  MacPherson,  M.A  ,  B.Sc,  M.B.,  Ch.M.       ...     6.37 

Notes    on    Australian    Hlwpalocera.     LyccrnidcP  Part  ii.      By  G.   A. 

Waterhoure,  B.Sc,  B.E 648 

On  the  Occurrence  of   Monoriraptua  in  New  South  Wales.     By  T.  S. 

Hall,  M.A.     (Cominunicatcdhy  John  Mitchell)      ...          ...         ...     654 

Notes  and  Exhibits     495,592,656 

Elections  and  Announcements  593 

Donations  and  Exchanges     ...          ...          ...         ...          ..  ...         ...     658 

Title-page  i. 

List  of  Contributors  and  Titles  of  Papers iii. 

Contents  vii 

List  of  Plates xi.. 

Corrigenda        ...         ...          ...          ...         ...          ...         ...         ...         ...     xii. 

List  of  New  Generic  Terms  proposed  in  the  Volume      ...         ...         ...     xii. 

Index 


LIST   OF   PLATES. 

PROCEEDINGS,  1902. 

Plates  I. -III. — Australian  Mollusca. 

Plates  IV. -V. — Bacterium  vai^cnlarnm,  Cobb. 

Plate  VI. — Bacterium  saccliari,  n.sp. 

Plate  VII.  —  Gyrocotyle. 

Plate  VIII. — Camarina  Camhagei,  Baker  (Belah),  Gilganclra,  N.S.W. 

Plate  IX.  -  Geijera parvlflora ,  Lincll.,  (Wilga),  Gilganclra,  N.8.W. 

Plate  X. — Fig.  1.  EticaUjptus  WoolUiana,  Baker. 

Fig.  2.  Eucalyptus  cornea,  Deane  &  Maiden. 
Plate  XI. — Eucalyptus  melanopliloia,  F.v.M. 
Plate  XII, — Bacterium  eucahjpti,  n.sp. 
Plate  XIII. — Bistomum  pristiophori,  n.sp. 
Plate  XIV.  —  Oyyris. 

Plate  XV. — Eheocarpus  sp.,  from  Lord  Howe  Island. 
Plate  XVI. — Ardisia  racemosa,  R.  T.  Baker. 
Plate  XVII. — Lotorium. 
Finite  XYiii. —  Storena  albo7naculata,  n.sp.;    Araneus  liheralis,  n.sp.;    Stepha- 

nopsis  ohtusifrons,  n.sp.;  Cehcnia  distincta,  Cambr. 
Plates  xix-xx. — Sporocysts  from  Mytilns  latus. 

Plate  XXI. — Eucalyptus  poly anthemos,  Schauer  (drawn  from  the  type). 
Plates  xxii. -XXIII. — Fructification  of  Polyporus  mylitt(c,  Cke.  A:  Mass. 
Plate  XXIV. — Mammalian  and  Keptilian  Vomerine  Bones  {Lacerta,Heterouota, 

Tricliosurus,  Miniopterus). 
Plate  XXV. {Amhlystom.a,  Ichthyophis,  Flesiosaurus,  Sphenodon, 

Tropidonotus,  Zootoca). 
Plate  XXVI. {Gomphoynathus,  Ptychognathus,   Testudo,  Ornitho- 

rhynclius). 
Plate  XXVII. — Map  of  New  South  Wales  showing  Botanist's  route. 
Plate  XXVIII. — Symplocos  Bauerleni,  n.sp. 
Plate  XXIX. — Australian  Mollusca. 
Plate  XXX. — Anatomy  of  Pdsella. 
Plates  XXXI. -XXXIII. — Australian  Mollusca. 


CORRIGENDA. 

Page  58,  line  9 — for  E.  puhescem  read  D.  imhe>icens. 

Page  58,  line  18 — for  Piiltenwa  read  Piiltenece. 

Page  251,  line    11 — for  Ceratofornia  read  Ceratoferonia. 

Page  287,  line  23 — for  N.  marginata  read  A\  marginatm. 

Page  444,  line  25 — for  L.  pileare  read  L.  inleari>i. 

Page  464,  in  the  Explanation  of  the  Text-fig.  2 — for  J..  (Septa)  nodiferum 

read  L.  nodiferum. 
Page  466,  last  line — for  Vide  also  P.  [^  L.]fratercnliim  read  T^u/<'also  p.480, 

L.  fraierculum. 
Page  479,  in  the  Explanation  of  Text-fig.  3 — for  Tritonium  f^ijeciosum  read 

Troplion  speciosiim. 


LIST  OF  NEW  GENERIC  TERMS 

Proposed  ix  this  Volume  (1902). 


P.\GE 

PAGE 

J^thalina  (Lepidopt.)   ... 

..     107 

Hyperioiioma  (Coleopt.) 

.     434 

Alapadna  (Lepidopt. )   ... 

..     106 

Icasmn  (Lepidopt.) 

90 

Axioctetn  (Lepidopt.)    ... 

..     105 

LopJioto)iui  (Lepidopt.)... 

.     128 

Axiorata  (Lepidopt.)     ... 

..     120 

Ochthophora  (Lepidopt.) 

89 

Bothynacrum  (Coleopt. ) 

..     437 

PaurophyUa  (Lepidopt.) 

.     124 

liryouiiiiui  (Lepidopt.)  ... 

..     134 

Proscranu  (Lepidopt.). 

.     104 

l)nop]ieropi>i  (Lepidopt. ) 

..     103 

Scleropoides  ( Coleopt. )  . . . 

.     435 

Eccleta  (Lepidopt.) 

86 

Symmolpis  (Lepidopt. ) . . . 

.     127 

Ecpatia  (Lepidopt.) 

..       8.-J 

SyiitJieta  (Lepidopt.) 

84 

Kpitripta  (Lepidopt.)    ... 

123 

TJieiima  (Lepidopt.)      ... 

.     250 

KstJdodora  (Lepidopt. ) . . . 

..     130 

Thelxinoa  (Lepidopt.) 

.     131 

E}idaimonif;)na  ^Lepidopt. ) 

..     247 

Zophochroa  (Lepidopt.) 

.     118 

Hinterois  (Lepidopt.)     ... 

..     121 

OF    THE 

LINNEAN    SOofETY 

OF 


WEDNESDAY,  MARCH  26th,   1902. 


SPECIAL  GENERAL  MEETING. 

The  requisition  upon  which  the  Meeting  was  convened,  signed 
by  sixteen  members,  was  read  by  the  Secretary. 

It  was  moved  by  Mr.  H.  G.  Smith,  and  seconded  by  Mr.  R. 
Etheridge,  Junr., — That  Rule  xii.  be  amended  by  the  insertion 
between  the  word  "President"  and  the  words  "four  Vice- 
Presidents,"  of  the  following  Clause—  "who  shall  hold  office  'for 
not  more  than  one  year  continuously,  but  shall  be  eligible  for 
re-election  after  the  lapse  of  one  j^ear." 

After  some  discussion,  the  motion,  on  being  put  to  the  Meeting, 
was  lost. 

MONTHLY  MEETING. 

Mr.  J.  H.  Maiden,  F.L.S.,  etc.,  President,  in  the  Chair. 

The  Donations  and  Exchanges  received  since  the  previous 
Monthly  Meeting  (27th  November,  1901),  amounting  to  72  Vols., 
185  Parts  or  Nos.,  20  Bulletins,  14  Scientific  Reports,  10  Annual 
(Administrative)  Reports,  20  Pamphlets,  1  Map,  and  6  Miscellanea, 
received  from  115  Societies,  etc.,  and  5  individuals,  were  laid  upon 
the  table. "^ 

*  The  complete  list  for  the  year  will  be  given  in  the  concluding  Part  of 
this  Volume. — [Ed.]. 


A  NOTE  ON  TWO  SPECIES  OF  ASTRALIUM  FROM 
PORT  JACKSON. 

By  H.  Leighton  Kesteven. 

Under  the  name  of  Astralium  tentoriforme^  Jonas,  two  species 
of  that  genus  have,  in  Sydney,  been  united,  'i  hese  two  present 
the  anomaly  of  two  species,  undeniably  distinct,  so  like  one 
another  that  in  some  instances  only  the  operculum  will  show  to 
which  species  a  specimen  belongs.  In  the  neanic  stage,  however, 
the  two  species  are  easily  separable. 

ASTRALIU3I  (Uvaxilla)  fimbriatum,  Lamarck. 

IVochus  fiiiibriatus,  Lamarck,  Anim.  sans  Vert.  ed.  i.,  \\\.,  p.  12, 
1822  :  Delessert,  Recueil,  etc.,  pi.  34,  figs.  6a  and  6,  1841; 
Quoy  &  Gaimard,  Voy.  Astr.  iii.,  p.  229,  pi  61,  fig.  8,  oper- 
culum tigs.  11,  12,  1835  ;  Chenu,  Man.  Conch,  i.,  fig.  2573  ; 
Philippi,  Conch.  Ca)),  ii.,  p.  121,  pi.  20,  fig.  7,  1846. 

Carinidea  Jimbriata,  Swainson,  Proc.  Roy.  Soc.  Van  Diemen's 
Land,  1854,  Vol.  iii.,  p.  39,  pi.  vi.,  figs.  3,  4. 

Calcar  fimbriafAiiiiy  Kiener  &  Fischer,  Coq.  Viv.  Turbinacees,  iii., 
p.  38,  pi.  32,  fig.  2,  1880. 

Astralium  fimbriatum,  Tryon,  Man.  Conch,  x.,  p.  239,  pi.  54,  figs. 
47,  48,  49;   operculum,  pi.  60,  f.  46,  1888. 

Ti'ochus  squamifh-us,  Koch  in  Philippi,  Abild.  i.,  pi.  4,  tig.  9, 
p.  138,  1844. 


BY    H.    LEIGHTON    KESTEVEN. 

Reeve's  figure  of  the  species*^'  is  of  his  A.  pileolum. 


£^iA 


A.  Jimhriatum. 
Fig.  1. — Neanic  stage  and  opercukim  thereof,  and  adult  opercukim. 

Tiyon  regards  A.  squamiferum  as  a  variety  of  A.  Jimhriatum, 
whilst  Fischer  {loc.  cit.)  and  Tate  and  Mayf  consider  the  name 
an  absolute  synonym,  a  decision  which  is  doubtless  correct. 

Other  varieties,  according  to  Tryon,  are  A.  pileolum,  Reeve 
(1842),  and  A.  cucullatum,  Kiener  |  The  former  is  decidedly  a 
distinct  species:  the  latter  will  most  likely  be  found  to  be  distinct 
also,  unless  perhaps  it  is  a  variety  of  A .  teutoriforme. 

This  species  is  not  so  common  in  Port  Jackson  as  the  succeeding. 

AsTRALiUM  (Uvanilla)  tentoriforme,  Jonas. 

Trochus    tentoriforinis,    Jonas,    Zeitsch.    f.    Malak.,    1845,   p.  Q%; 

Philippi,  Conch.   Cab.  ii,,  p.  116,  pi.  20,  fig.  1,  1846;  Reeve, 

Conch.  Icon.  xiii.  pi.  viii.,  fig.  43,  1861. 
Trochus  urvillei,    Philippi,    Conch.    Cab.  ii.   p.  215,    pi.  32,  fig.  4, 

lfe46;  Reeve,  Conch.  Icon,  xiii.,  pi.  ix.,  fig.  46,  1861. 

Trochus  georgianu><,  Quoy,  ^IH.,  fide  Kiener,  »Sp.  Genr.  Trochus, 
pi.  31,  fig.  2,  1880. 

Calcar  tentoriforme,  Kiener  k  Fischer,  Coq.  Viv.  Turbinacees  iii., 
p.  41,  pi.  79,  fig.  2,  1880. 

*  Conch.  Icon.  xiii.  pi.  ix.  fig.  49. 

t  Proc.  Linn.  Soc.  N.S.W.  xxvi.  (2),  1901,  p.  400. 

.tCoq.  Viv.  Turbinacees,  iii.  p.  40,  pi.  32,  fig.  3.  1880. 


TWO    SPECIES    OF    ASTRALIUM    FROM    PORT    JACKSON, 


Astralium  tentoriforme,Tvyoi\,^l?ii\.  Conch,  x.,  p.  240,  pi.  53,  figs. 

41,42,  1888. 
Trochus  ^fimhriaW.s,  Quoy  k  Gaimard,  oj).  cit.,  pi.  61,  fig.  9,  1835. 


A,  tentorifornie. 
Fig.  2. — Neanic  stage. 

Quoy  lV:  Gaimard  {Joe.  cit.)  speak  of  a  variety  "plus  epaisse  et 
plus  pyramidale  "  which  w^as  evidently  the  original  of  their  fig.  9. 
Philippi  quotes  this  figure  as  being  of  his  T.  urvillei.  Fischer 
regards  Philippics  sjDecies  as  the  young  of  a  variety  /3,  7\  georgianus, 
being  presumably  the  adult;  the  figure  of  this  is  the  best  represen- 
tation of  A.  tentoriforme  as  it  is  known  in  Port  Jackson.  The 
species  is  ^ery  common  in  all  the  fresher  reaches  of  Sydney 
Harbour  and  the  coasts  of  New  South  Wales. 

In  literature  these  two  species  are  more  distinct  than  in  life. 
Both  species  are  very  variable.  Specimens  of  A.  tentoriforme 
occur  nearly  as  flat  as  A.  Jimhr latum,  var.  squamiferum,  and 
almost  as  fimbriated  as  var.  cucidlatiim  (see  Tryon's  figs.  52,  50) 
whilst  pyramidal  unornamented  specimens  of  A.  Jimhriatum.  Si,ve 
not  uncommon.  M}'- figures  show  the  differences  between  these 
species  in  their  neanic  stage.  The  best  characters  for  identifica- 
tion of  the  adult  shells  are  offered  b}^  the  base,  and  may  be 
tabulated  as  follows  : — 
A.  fimhriatum. 

(1)  Base  convex. 

(2)  Columella   ending    in    a 

small  tooth 

(3)  Lower    lip    joining    the 

upper  lip  at  the  extreme 
edge. 


A.  tentoriforme 

(1)  Base  concave. 

(2)  Columella  ending  in  a  pro- 

minent tooth. 

(3)  Lower  lip  .set    inside   the 

upper  lip 


BY    H.    LEIGHTON    KESTEVEN. 


5 


As  a  rule  A.  Jimhriahim  is  flatter  than  A.  tentoriforme. 
Although  the  above  characters  are  the  most  constant,  none  of 
them  can  be  implicitly  relied  on.  I  have  seen  A.  fi^nhriatum  with 
a  concave  base  and  A.  tentoriforme  without  any  sign  of  a  tooth. 
There  is,  however,  one  recognition  mark  to  which  one  may  pin 
one's  faith — the  operculum  ;  this  will  be  always  found  reliable. 
The  differences  are  shown  by  my  figures. 


?(^&^- 


O 


A.  fimhriatnm  A.  tentoriforme. 

Fig.  3.— Adult  operculum.  Fig.  4. — Adult  operculum. 

The  radulse  are  almost  exactly  alike;  if  there  is  any  difference  it 
is  that  the  marginals  of  A.  fimhriatum  are  larger  than  those  of 
A.  tentoriforme. 

It  is  more  than  probable  that  both 
species  occur  in  South  Australia  and 
Tasmania,  though  only  A.  fimhriatum 
has  been  recorded  from  there.  2\ 
georgianus  was  obtained  at  King 
George's  Sound,  S.W.  Australia. 

These  two  species  would  seem  to 
present  an  exception  to  the  rule  that 
distinctive  characters  are  inherited 
earlier  in  successive  generations,  for 
by  that  theory  A.  fimhriatum,  should  be 

descended  from  discoidal  ancestors,  whilst  y1.  tentoriforme  should 
come  from  trochiform  ancestors,  and  such  a  divergence  of 
ancestry  would  be  manifested  in  anatomical  characters. 


A.  tentoriforme. 
Fig.  5. — Dentition. 


Postscript. — Two  outline  drawings  of  the  apices  of  Columhella 
seraiconvexa,  Lamarck,  and  C.  australis,  Gaskoin  (figs.  6-7),  shown 


b  TWO    SPECIES    OF    ASTRALIUM    FKOM    PORT    JACKSON. 

me  by  Mr.  C.  Hedley,  illustrate  the  fact  that  they  present  aDother 
exami)le   of,   if    I    may    be    allowed    the    term,   con\ergence    of 


Fig.  0. — C.  andralis.  Fig.  l—C.  semiconvexa. 

development.  Like  the  two  species  discussed  above,  they  are, 
w^hen  adult,  so  alike  that,  devoid  of  their  epidermis,  they  are 
practically  inseparable,  whilst  their  apices  are  quite  different. 
They  are,  however,  easier  to  identify  than  the  two  species  of 
Astralium,  C.  australis,  having  a  sutural  frill  of  epidermis,  which 
is  absent  in  C.  semiconvexa. 

To  Miss  M.  Lodder  I  am  indebted  for  the  opportunity  to 
study  the  growth  of  Astralium  anreum,  Jonas.^  In  the  neanic 
stages  it  is  very  similar  to  A.  fimhriatnm,  Lamarck  [April  30th y 
1902), 


Philippi's  Abbild.,  1844,  Vol.  ii.,  p.  14,  t.  6,  fig.  2. 


STUDIES  ON  AUSTRALIAN  MOLLUSCA. 

Part  VI. 

By  C.  Hedley,  F.L.S. 

(Plates  i.-iii.) 
(Co)iiinued  from  Vol.  xxvi.,  p.  708.) 

BORNIA    FILOSA,  11. sp. 

(Plate  ii.,  figs.  15-17.) 

Shell  small,  thin,  compressed,  subtrigonal,  nearly  as  high  as 
long,  inequilateral.  White.  Sculpture :  numerous  close  fine 
radiating  hair  lines,  which  in  the  young  shell  are  disposed  in 
several  divaricating  bundles;  on  the  older  shell  they  radiate  more 
regularly,  and  are  cut  into  beads  by  concentric  grooves.  Anteri- 
orly and  posteriorly  the  radii  diverge  abruptly  from  the  sculpture 
of  the  median  area.     Length  6,  height  5  mm. 

Hah. — Port  Jackson;  one  valve  adhering  to  an  annelid  tube, 
taken  by  myself  in  Middle  Harbour. 

Type  to  be  presented  to  the  Australian  Museum. 

ROCHEFORTIA    DONACIFOKMIS,   Angas. 

Mysella  donaciformis,  Angas,  P.Z.S.  1878,  p.  863,  pi.  liv.,  f.  13. 

(Plate  i.,  figs.  10-U.) 

Angas  described  and  figured  this  species  from  South  Australia, 
but  his  illustration  has  been  condemned  by  Dall  and  others  as 
unsatisfactory.  A  series  of  drawings  is  therefore  now  given  of 
a  shell  I  o-athered  at  Middle  Harbour,  in  length  6  mm.,  and  in 


»  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

height  5  mm.  The  species  has  ah-eady  been  recorded  from  this 
coast  by  Henn.*  The  generic  title  used  is  adopted  from  Ball's 
investigations.! 

CONGERIA   LUNATA,  n.Sp. 

(Plate  i.,  figs.  1-4.) 

Valves  narrow,  crescentic,  subspiral,  thin.  Posterior  side 
rounded,  divided  by  an  acute  keel  from  the  anterior  which  is 
concave,  and  infolded  to  one-third  of  the  length  of  the  shell,  with 
a  slight  median  byssal  gape.  Colour  white,  rayed  with  purple, 
under  a  thin  gloss}'  brown  epidermis.  Sculpture  :  close  fine 
waved  growth-lines.  Interior  slightly  nacreous.  Septum  reduced 
to  a  narrow  groove,  ending  above  in  a  small  pit.  Anterior 
cardinal  margin  with  two  or  three  oblique  folds.  Ligament 
internal,  running  within  the  dorsal  margin  and  curving  up  in  a 
groove  under  the  umbo.  Umbo  acute,  twisted,  showing  the 
prodissoconch  on  the  summit.  Height  25,  length  IG,  breadth  of 
conjoined  valves  13  mm. 

Hah. — Keppel  Bay,  Queensland;  one  specimen  collected  by  Mr. 
G.  L.  Pilcher  in  1887. 

Type  to  be  presented  to  the  Australian  Museum. 

This  species  adds  a  genus  to  the  Australian  fauna. 

Mactra  parkesiana,  nsp. 

(Plate  i.,  figs.  5-9.) 

Valves  nearly  equilateral,  not  much  longer  than  high,  rather 
compressed,  thin,  rounded  anteriorl}^  and  ventrally,  slightly  pro- 
duced and  angled  postei'iorly.  Colour  white;  surface  smooth  and 
glossy,  under  the  lens  appear  delicate  growth  stride,  faint  radial 
scratches  and  anteriorly  oblique  vermiculate  wrinkles.  Two  sharp 
narrow  ridges  border  the  posterior  dorsal  slope.  Anterior  dorsal 
area  sunken,  but  not  sharply  defined.  The  anterior  lateral  region 
of  the  hinge  of  the  right  valve  presents  two  pockets,  one  abo\e 


*  Henn,P.L.S.N.S.W.  (2),  ix. .  p.  ISO. 
t  Dall,  Trans.  Wagner  Inst.,  iii.,  pt.v.,  1900,  p.  1157. 


HV    C.    HEDLEY.  9 

the  other,  from  the  lower  and  larger  of  which  there  protrudes  a 
slender  tongue  attached  dorsally,  but  free  ventrally.  In  the 
corresponding  position  of  the  left  valve  are  two  pockets  divided 
by  a  septum  which  is  continued  into  the  anterior  arm  of  the 
cardinal.  Length  20,  height  16,  breadth  of  conjoined  valves 
9  mm. 

Hab.  — Port  Jackson:  one  specimen,  containing  the  animal,  was 
found  on  Balmoral  Beach  by  Miss  L.  Parkes,  a  daughter  of  the 
veteran  Australian  statesman,  in  whose  honour  it  is  named. 

Type. — Miss  Parkes  has  generousl}'- presented  her  specimen 
to  the  Australian  Museum. 

This  species  belongs  to  the  Section  Jlaclririida;  the  absence  of 
plicate  sculpture  distinguishes  it  from  other  members  of  the 
group. 

Retusa  nitida,  a.  Adams. 

(Plate  ii.,  fig.  27.) 

Bulla  nitida,  Ad.,  Thes.  Conch,  ii.,  1850,  p.  589,  pi.  125,  f.  127. 

Numerous  specimens  of  a  shell  taken  by  Mr.  J.  Brazier  in 
Torres  Straits  appear  to  be  this  species.  It  is  an  addition  to  the 
Australian  fauna;  indeed,  except  for  a  record  from  the  Persian 
Gulf,"^  it  seems  only  known  from  the  original  locality,  Borneo. 
As  the  original  account  is  rather  unsatisfactory,  I  add  a  figure 
and  description  of  a  Torres  Straits  example. 

Shell  small,  ovate,  rather  thin,  narrowly  umbilicate  above, 
subperforate  below.  Colour  white.  Sculpture:  faint  longitudinal 
oblique  growth-lines  are  crossed  for  the  anterior  and  posterior 
quarters  by  half  a  dozen  lightly  incised  grooves;  the  vertex  also 
radiately  traversed  by  a  few  delicate  raised  threads,  leaving  the 
median  half  of  the  shell  bare  of  ornament.  Aperture  amjDle 
below,  constricted  at  the  waist,  narrow  above  and  rising  in  an 
arch  above  the  vertex.  Columella  broad  and  twisted.  Height 
3*5,  breadth  2-7  mm. 

*  Melvill  &  Standen,  Proc.  Zool.  Soc,  1901,  p.  454. 


10  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

Pyrgulina  perspectiva,  n.sp. 
(Plate  iii.,  fig.  33.) 

Shell  small,  solid,  conical.  Colour  white.  Whorls  five,  besides 
a  prostrate  heterostrophe  apex  of  three  turns,  flattened,  separated 
by  a  deeply  channelled  suture.  Sculpture  :  the  last  whorl  has 
above  a  series  of  about  twenty  straight  oblique  bars,  about  their 
own  breadth  apart,  confluent  at  their  lower  margin  but  not  on 
the  ujjper  one,  both  margins  sinuated  by  the  projecting  ends  of 
the  bars.  Below  these  bars  is  a  deep  groove  succeeded  by  a 
narrow  peripheral  rib.  On  the  base  are  a  few  fine  spiral  threads 
intersected  b}^  growth-lines.  The  hollows  between  the  bars  are 
crossed  Ijy  fine  spiral  threads.  The  series  of  oblique  bars  ascend 
the  spire  ;  the  bars  do  not  correspond  across  the  suture. 
Periphery  of  the  two  topmost  whorls  slightW  constricted.  Base 
somewhat  hollowed.  Aperture  broken  in  the  specimen  studied, 
but  a  prominent  tooth  on  the  columella  remains.  Height  2 •2, 
breadth  1-08  mm. 

Hah. — With  P.  senex:  one  specimen. 

Type  to  be  presented  to  the  Australian  Museum. 

From  those  that  most  resemble  it,  this  species  may  be  dis- 
tinguished by  the  smaller  size  and  striated  interstices. 

Pykgulina  senex,  n.sp. 

(Plate  iii.,  fig.  31.) 

Shell  small,  solid,  subcylindrical.  Colour  white.  Whorls  three 
and  a  half,  besides  an  elevated,  median,  heterostrophe  apex  of 
two  turns,  shouldered  abruptly  and  separated  by  a  deeply 
channelled  suture.  Sculpture :  the  last  whorl  has  above  two 
spiral  keels  joined  at  intervals  by  about  14  short  perpendicular 
bars,  beaded  at.  the  points  of  intersection  and  enclosing  square 
meshes.  Anterior  to  these  are  six  widely  spaced  spiral  keels, 
gradually  diminishing  and  approximating  as  they  recede.  One 
plain  and  the  two  ornate  keels  ascend  the  spire  Delicate 
oblique    growth-lines    occur   in    the    hollows    of    the    prominent 


BY    C.    HEDLEY.  11 

sculpture.  Aperture  ovate,  with  a  parietal  callus  and  a  small 
deeply  seated  plication,  throat  grooved.  Height  1-57,  breadth 
0-8  mm. 

llah. — Ten  miles  south  of  Cape  Sidmouth,  N.  Queensland,  in 
13  fathoms;  two  specimens  (J.  Brazier). 

T3^pe  to  be  presented  to  the  Australian  Museum. 

A  fancied  resemblance  to  the  bald  head  and  hunched  shoulders 
of  an  old  man  suggested  a  name  for  this  shell. 

Pyrgulina  zea,  n.sp. 
(Plate  iii.,  fig.  34.) 

8hell  small,  solid,  conical,  imperforate.  Colour  white.  Whorls 
four,  besides  a  sunken  heterostrophe  apex,  separated  by  deeply 
channelled  sutures.  Sculpture  :  the  body  whorl  has  above  three 
rows  of  beads,  confluent  perpendicularly,  about  18  to  a  row,  set 
about  their  breadth  apart;  anterior  to  these  is  a  deep  groove  and 
a  peripheral  keel,  on  the  base  are  two  smaller  keels.  The  triple 
row  of  beads,  but  not  the  peripheral  keel,  ascend  the  spire.  On 
each  whorl  the  perpendicular  bead  rows  are  set  at  a  slightly 
different  angle  to  the  axis  of  the  shell.  Aperture  slightly  oblique,, 
faintly  grooved  within,  no  parietal  callus,  plication  single,  small 
and  deeply  seated      Height  1-8,  breadth  0*88  mm. 

Hah. — Ten  miles  south  of  Cape  Sidmouth,  N.  Queensland,  in 
13  fathoms;  one  specimen  (J.  Brazier). 

Type  to  be  presented  to  the  Australian  Museum. 

Pyrgulina  umeralis,  n.sp. 
(Pkteiii.,  fig.  32.) 

Shell  small,  very  solid,  imperforate,  elongate-ovate.  Whorls 
three  and  one-half,  besides  the  elevated  lateral  heterostrophe  apex, 
separated  by  a  deeply  channelled  suture.  Colour  white.  Sculp- 
ture :  the  body  whorl  has  al)Ove  a  double  row  of  beads,  each  bead 
confluent  from  above  to  below,  about  sixteen  beads  to  a  row,  set 
less  than  their  breadth  apart  These  are  followed  anteriorly  by 
a  deep  groove  and  a  stout  peripiieral  keel.     Thence  to  the  anterior 


12  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

•extremity  are  four  or  five  grooves  and  keels  diminishing  succes- 
sively. On  the  spire  the  double  row  of  l^eads  and  peripheral  keel 
alone  appear.  Aperture  slightly  oblique,  ovate,  with  a  heavy 
callus  and  single  stout  deep-seated  tooth  on  the  parietal  wall;  the 
throat  traversed  by  six  ridges.     Height  1-72,  breadth  088  mm. 

//«6. —Princess  Charlotte  Bay,  N.  Queensland,  13  fathoms, 
sandy  mud  bottom;  three  specimens  (J.  Brazier). 

Type  to  be  presented  to  the  Australian  Museum. 

Crosse  A  biconica,  n.sp. 
(Plate  ii.,  fig.  24.) 

Shell  minute  but  solid,  biconical,  base  greatly  produced,  peri- 
phery keeled.  Colour  white.  Whorls  four,  somewhat  turreted. 
Sculpture :  first  and  second  whorls  smooth,  third  comparatively 
coarsely  cancellated,  fourth  with  dense  fine  spiral  cords  crossed 
b}^  fainter  growth-lines  which  tend  to  bead  the  interstices.  Base 
two-thirds  of  the  total  height.  Umbilicus  wide  and  deep,  bordered 
by  a  conspicuous  ridge,  and  having  an  elevated  funicle  winding 
within.  Aperture  oblique,  rhomboidal,  channelled  by  the  um- 
bilical ridge,  and  with  a  gutter  at  the  termination  of  the  funicle. 
A  heavy  outstanding  varix  occurs  a  short  distance  behind  the 
aperture.     Height  1-7,  major  diam.  16,  minor  diam.  1  mm. 

Hah. — Torres  Straits,  between  Cape  York  and  New  Guinea; 
one  specimen,  in  company  with  Crossea  striata,  Watson,  dredged 
by  Mr.  J.  Brazier  in  8  fathoms,  on  a  sandy  mud  bottom. 

Type  to  be  presented  to  the  Australian  Museum. 

The  small  size,  produced  base,  and  wide  umbilicus  are  characters 
which  distinguish  this  from  other  Australian  members  of  the 
genus.  Renewed  study  strengthens  the  conviction,  which  I  have 
already  expressed,  that  Crossea  is  unhappily  associated  with  Scala, 
a,nd  that  it  would  be  more  appropriately  ranked  with  Trichotropis 
or  Fossarus. 

Having  examined  the  type  of  Crossea  mimUa,  Petterd,  I  am 
unable  to  agree  either  with  its  author  that  it  is  a  Crossea,  or 


BY    C.    IIEDLEY.  13^ 

with  Tate  and  May^  that  it  is  a  Fossarus.     It  seems  to  me  a 
larval  shell  of  the  Siimsigera  form. 

Crossea  gatliffi,  n.sp. 
(Plate  ii.,  fig.  25.) 

Shell  small,  thin,  globose-conical,  spire  elevated,  gradate,  basal 
funicle  slightly  developed.  Colour  white  C?  bleached).  Whorls 
four.  Sculpture  :  last  whorl  with  fine,  close  spiral  threads  of 
which  every  fourth  is  larger;  at  irregular  intervals  incipient 
varices  traverse  the  whorl;  on  the  penultimate  whorl  the  spiral 
sculpture  is  more  coarse  and  distant.  Umbilicus  narrow,  bordered 
by  a  slight  rib,  which  continuing  to  the  anterior  extremity  is 
there  notched  by  the  pseudocanal.  Aperture  oval,  outer  lip  thin 
and  everted.     Height  1-16,  major  diam.  1-1,  minor  diam.  094  mm. 

Hah. — With  C.  blconica:  one  specimen. 

Type.— To  be  presented  to  the  Australian  Museum. 

This  species  is  named  in  honour  of  m}^  friend  Mr.  J.  H.  Gatliff, 
of  Melbourne.  The  thin  shell,  lack  of  longitudinal  sculpture, 
tabulated  whorls,  and  feeble  basal  funicule  distinguish  it.  C. 
striata^  perhaps  its  nearest  all}-,  is  much  larger,  with  conspicuous 
reticulate  sculpture  and  a  double  umbilical  rib. 

Teinostoma  involuta,  n.sp. 

(Plate  iii.,  fig.  35.) 

Shell  small,  solid,  globose,  spire  slightly  exserted,  umbilicus 
broad  and  deep.  Whorls  3^,  divided  by  a  deeply  channelled 
suture,  rapidly  increasing,  last  whorl  broadening  and  descending 
at  the  aperture.  Sculpture  :  first  whorl  smooth,  the  rest  carrying 
a  row  of  tubercles  along  the  crown  and  basal  edge,  on  the  last 
whorl  about  twenty  above  and  fifteen  below.  The  intervening- 
space  is  obliquely  traversed  by  fine,  close  irregularly  waved  riblets, 
narrower  than  their  smooth  interstices;  these  may  fork  or  end 
abruptly,  but  at  either  end  of  their  course  usually  unite  in  bundles 
of  twos  or  threes  to  form  the  upper  or  lower  tubercle.     Their 


"  Tate  &  May,  these  Proceedings,  1901,  p.  458. 


<^^'^A( 


14  STUDIES    ON    AUSTRALIAN    3I0LLUSCA, 

<ippearance  is  suggestive  of  a  continuous  rib  split  into  riblets  for 
most  of  its  length  but  holding  together  at  top  and  bottom. 
Aperture  very  oblique,  narrowly  ovate,  pointed  above;  peristome 
double  and  massive,  intruding  a  broad  free  lobe  into  the  umbilicus, 
and  clinging  to  the  umbilical  edge  with  a  lesser  lobe.  Umbilicus 
broad,  perspective  smooth  and  excavated  within  the  string  of 
beads  which  bounds  it.  Height  1'3,  major  diam.  1-34,  minor 
diam.  1*2  mm. 

Hah. — Off  Darnley  Island,  Torres  Straits,  in  10-30  fathoms; 
one  specimen  (J.  Brazier). 

Type. — To  be  presented  to  the  Australian  Museum. 

This  species  resembles  certain  forms  of  Liutia  such  as  L.  devexa^ 
Hedley,  from  the  same  region.  The  assignment  of  it  to  Teino- 
stoma  is  governed  by  the  umbilical  tongue,  a  rather  artificial 
character.  As  I  have  previously  indicated,  outlying  forms  of 
Liotia  so  approach  some  species  of  Teinostoynri  that  it  is  difficult 
to  allot  to  each  its  species. 

Liotia  corona,  n.sp. 

(Plate  ii.,  figs.  21-23.) 

Shell  minute,  discoid,  not  nacreous;  spire  sunken,  umbilicus 
wide  and  shallow.  Whorls  three,  last  half-whorl  scarcely  in  con- 
tact with  the  others,  suddenly  and  deeply  deflected.  Colour 
white  C?  bleached).  Sculpture:  last  whorl  ringed  by  16  thick, 
projecting,  distant  ribs  which  fade  above  and  below  at  the  sutures, 
these  ribs  continuing  on  the  suture  for  about  half  a  whorl. 
Interstices  of  the  ribs  faintly  spirally  scratched.  Aperture  very 
oblique,  circular,  fortified  by  a  varix.  Height  028,  major  diam- 
0  82,  minor  diam.  0*66  mm. 

Hah. — With  Crossea  hiconica;  four  specimens. 

Type. --To  be  presented  to  the  Australian  Museum. 

Liotia  incidata,  n.sp. 

(Plate  ii.,  figs.  18-20.) 

Shell  depressedly  globose,  rather  solid,  perforate,  flattened  on 
the  base.     Whorls  three,  margined  at   the  suture,  last  slightly 


BY    C.    HEDLEY.  15 

gibbous  and  descending,  constricted  behind  the  aperture. 
Sculpture :  above  finely  spirally  grooved,  hollowed  at  the  peri- 
phery between  two  keels,  on  the  base  with  radial  riblets  latticed 
by  spiral  threads.  Aperture  subquadrate,  very  oblique,  peristome 
everted,  slightly  thickened.  Height  0  7,  major  diam.  1-14,  minor 
diam.  0-96  mm. 

Hab. — Betw^een  Cape  York  and  New  Guinea,  in  8  fathoms, 
sandy  mud  bottom;  one  specimen  (J.  Brazier). 

Type. — To  be  presented  to  the  Australian  Museum. 

Mecoliotia  spinosa,  n.sp. 

(Plate  ii.,  fig.  26.) 

Shell  minute,  solid,  broader  than  high,  spire  elevated,  base 
flattened.  Colour  white  (?  bleached).  Whorls  four  and  one-half 
in  the  decollate  example  studied.  Sculpture  :  a  strong  projecting 
spiral  keel  runs  down  the  last  three  whorls,  beneath  which  on 
the  body  whorl  is  a  deep  narrow  groove  and  sharp  cord,  succeeded 
by  a  smaller  groove  and  cord.  Radiating  ribs,  of  which  the  last 
whorl  has  fourteen,  commence  at  the  suture,  develop  sharp  points 
on  the  spiral  keel,  partition  the  peripheral  groove  into  a  series  of 
deep  narrow  pits,  bead  the  peripheral  rib  and  disajDpear.  A 
minor  sculpture  of  faint  spiral  threads  is  visible  above  and 
beneath  the  periphery.  Umbilicus  w^de  but  rapidly  narrowing 
above.  Aperture  very  oblique,  circular,  double,  with  a  broadly 
expanded  varix.  Height  0  84,  major  diam.  1,  minor  diam. 
0-82  mm. 

Hah. — Off  PijDon  Islands,  N.  Queensland  ;  9  fathoms,  mud 
bottom;  one  specimen  (J.  Brazier). 

Type. — To  be  presented  to  the  Australian  Museum. 

This  species  is  smaller,  with  a  wider  umbilicus  and  more  prickl}^ 
sculpture,  than  M.  haUigani.'^' 

*  Hedley,  Atoll  of  Funafuti,  1S90,  p.  555. 


16  STUDIES    ON    AUSTRALIAN    iMOLLUSCA, 

Aplustrum  brazieri,  Angas. 

Dlaphana  brazieri,  Angas,  Proc.  Zool.  Soc.  1877,  pp.  175,  189, 
pl.xxvi.,  f.  20. 

(Plate  iii.,  fig.  .36.) 

A  specimen  specifically  identified  by  Mr.  Brazier  is  here  figured. 
It  is  4  mm.  in  length  and  was  obtained  in  sand}'-  mud  off  Watson's 
Bay  in  10  fathoms— a  few  hundred  yards  from  where  the  same 
collector  found  the  type.  If  Mr.  Brazier's  determination  is 
correct,  which  T  have  no  reason  to  doubt,  then  Ano^as'  fio^ure  is 
extremely  bad.  Moreover,  I  conclude  that  the  species  is  wrongly 
placed  in  Diaphana,  and  should  be  transferred  to  Aplustrum. 
Pilsbry,  who  remarked  how  ill  the  species  agreed  with  Diaphana, 
created  a  section  Austro-Jiiaphaiia  for  its  reception,*  which  might 
rank  as  a  subgenus  of  Aplustrum.  A  large  example  of  this  species, 
which  I  have  examined  reached  a  length  of  5  mm. 

Claxculus  denticulatus,  Gray. 

Gra}'  in  King's  Survey  of  the  Intertropical  and  Western  Coasts. 
of  Australia,  ii.,  1827,  Appendix,  p.  479. 

By  modern  wi-iters  this  has  been  treated  as  a  lost  species 
Specimens  were  taken  at  Geraldton,  W.A.,  by  Mr.  A.  U.  Henix 
which  I  recognised  as  the  shell  described  by  Menkef  as  Monodonta 
lupina,  and  afterwards  illustrated  by  Philippi.|  Menke,  who. 
doubtless  had  good  grounds  for  the  belief,  ultimately  considered^ 
that  his  species  was  identical  with  Gray's  Monodonta  denticulata. 
This  withdrawal  of  his  species  escaped  the  notice  of  Prof.  Tate|| 
in  his  revision  of  Menke's  shells.  Trochus  atropurpureus,  Gould, 
and  T.  samoensis,  Hombron  k  Jacquinot,  appear  to  be  synonyms, 
of  Gray's  species. 

*  Pilsbry,  Man.  Conch,  xv.,  1893,  p.  287. 

t  Menke,  Moll.  X.  Holl.  Spec.  1848,  p.  15. 

:;:  Philippi,  Conch.  Cab.  Bd.  ii.,  Abth.  3,  1846,  p.  237,  pi.  30,  f.  3, 

§  Menke,  Zeit.  fiir  Malak.  1844,  p.  58. 

II  Tate,  P.L.S.X.S.W.,  vi.,  1881,  p.  393. 


BY    C.    HEDLEY.  17 

Clathurella  denseplicata,  Dunker. 

Pleurotoma  {Drillia)  denseplicata^  Dunker,  Mai.  Bliitt.  xviii., 
1871,  p.  159:  WeinkaiifF,  Conch.  Cab.  Bd.  iv.,  Abth.  3,  1887, 
p.  107,  pi.  23,  ff.  7,  9. 

This  species,  collected  in  Bass  Straits  by  a  correspondent  of 
the  Godeffroy  Museum,  and  of  which  the  type  is  probably  in  the 
Hamburg  Museum,  has  apparently  been  overlooked  by  Australian 
conchologists.  From  the  figure  and  description  I  judge  this 
species  to  be  identical  with  Clathurella  philomence^  Ten.  Woods, 
over  which  it  has  priority. 

CocHLODESMA  ANGASi,  Crosse  ct  Fisclier. 

On  a  beach  near  the  house  of  my  friend,  Mr.  W.  L.  Mav,  I 
saw  for  the  first  time  a  living  animal  of  this  species.  The 
siphons  are  not  protected  by  a  sheath  like  those  of  Anatina,  and 
can  be  wholly  retracted  within  the  valves.  These  characters 
accord  better  with  Cochlodesma  than  with  Anatina,  to  which 
Angas  (P.Z.S.,  1865,  p.  644)  erroneously  transferred  it. 

Arc  A.  ZEBRA,  Swainson. 
Swainson,  Zool.  Illustrat   2nd  Ser.  iii.,  1833,  pi.  118. 

An  example  of  this  shell  from  Sydney  Harbour  has  l)een 
handed  to  me  by  the  Hon.  J.  Norton,  M.L.C.  This  species  has 
not  to  my  knowledge  been  recorded  from  the  coast  of  New  South 
Wales.  Dr.  Norton's  discovery  was  confirmed  by  Mr.  H.  L. 
Kesteven,  who  again  collected  it  at  the  Inner  South  Head. 

Philobrya  crexatulifera,  Tate. 

Tate,  Trans.  Roy.  Soc.  S.A.  xv.,  1892,  p.  131,  pi.  i.,  if.  11,  Ua. 

This  species  has  not  been  seen  so  far  north  as  this  State.  I 
can  now  announce  that  I  found  it  in  a  cleft  of  the  cliffs  a  mile 
south  of  the  South  Head  Lighthouse,  where  I  took  Zidora  lodder(H, 
Tate  ife  May,  Sirius  badius,  T.  Woods,  Ccecum  aniputatum,  Hedley, 
and  other  rarities.  During  her  recent  visit  to  Sydney  Miss 
Lodder  also  collected  P.  crenatulifera  at  Long  Bay. 


IS 


STUDIES    ON    AUSTKALIAX    MOLLUSCA, 


EuLiMELLA  PULCHRA,  Braziei'. 

Exdimella  imlchra,  Brazier,  P.L.S.N.8.W.  (2),  ix.,  1894,  p.  170, 
pi.  xiv.,  f.  6. 

Eulimella  tricincta,  Tate,  Trans.  Roy.  8oc.  S.A.,  xxii.,  1898^ 
p.  83,  pi.  iv.,  f.  4. 

Although  I  prepared  both  of  the  illustrations  mentioned  I  did 
not  recognise  that  they  ajDplied  to  one  species.  Brazier's  example 
was  young.  On  re-examination  of  authentic  material  I  can  affirm 
their  identity. 

CyclostrExMA  porckllana,  Tate  k  Ma3^ 

Tate  cfe  May,  Trans.  Roy.  Soc.  S.A.  xxiv.,  1900,  p.  102. 

Specimens  of  this  were  collected  in  Sj^dney  Harbour  b}^  Mr. 
H.  L.  Kesteven,  whose  determination  of  them  was  confirmed  by 
Mr.  W.  L.  May. 

Marginella  subbulbosa,  Tate. 

Tate,  Trans.  Roy.  Soc.  S.A.,  i.,  1878,  p.  86. 

This  species  is  a  new  record 
for  ]S^ew  South  Wales.  Mr. 
H.  L.  Kesteven,  who  drew  my 
attention  to  it,  has  kindly 
prepared  sketches  for  the 
accompan3'ing  figures  of  it 
and  M.  stranger.  Tate  and 
May  latel}'  united  these,  but 
the  outline  sketches  here 
M.  suhhitlhosd.  |, resented  will  demonstrate  the  difi'erence  and 
validity  of  each. 

Trophon  goldsteini,  Ten.  Woods. 

Ten.  Woods,  Proc.  Roy.  Soc.  Tas.  1875,  p.  136. 

Mr.  R.  L.  Cherry,  in  finding  this  shell  on  the  beach  at  Catherine 
Hill  Bay,  has  added  another  species  to  the  fauna  of  New  South 
Wales.  " 


21.  stranqci. 


BY    C.    HEDLEY. 


19 


Marginella  mayii,  Tate. 
Tate,  Trans.  Roy.  8oc.  S.A.  xxiv.,  1900,  p.  93. 
Mr.  Brazier  has  shown  me  a  shell  of  which  he  took  a  sinole 

o 

specimen  in  8  fathoms  off  the  Inner  North  Head,  Sydney  Harbour. 
By  comparison  with  an  authentic  specimen  received  from  Mr. 
W.  L.  Ma}''  I  have  identified  this  as  a  dwarf  form  of  M.  mayii. 

Vermetus  caperatus,  Tate  &  May. 

Thylacodes  caperatus,  Tate  cfe  May,  Trans.  Roy.  Soc.  S.A.  xxiv., 
1900,  p.  94. 

During  a  recent  excursion  of  the  Field  Naturalists'  Club  to 
Balmoral  this  species  was  found  by  myself  and  others  for  the  first 
time    in    Australia    in 
plenty  under   stones  at 
half  tide  level. 

The   authors  of    this 
species    state   that   the 
animal      is      unknown. 
Mr.  H.  L  Kesteven  has 
Operculum  and  radula.       j^-^^j^^  prepared  for  me 

the  drawings  for  the  accompanying  illustrations 
of  the  operculum,  radula  and  the  jaw  as  a  whole  and  in  detail. 
The  presence  of  an  operculum  necessitates  the  removal  of  the 
species  from  Thylacodes. 

Bifidaria  macleayi,  Brazier. 

Fnpa  {Vertigo)  tnicleayi^  Brazier,  Proc.  Linn.  Soc.  N.S.W.  i. , 
1876,  p.  110. 

(Plate  iii.,  fig.  39.) 

M}'  drawing  of  this  hitherto  unfigured  species  is  taken  from  an 
authentic  specimen  in  the  Macleay  Museum,  2-5  mm.  in  length. 
It  is  labelled  Barnard  Islands,  No.  iii.,  a  locality  not  named  in 
the  original  description.  The  genus  Bifidaria  was  described  by 
Sterki.* 


Jaw. 


*  Sterki,  The  Nautilus,  vi.,  p.  99,  Jan.  1893. 


20  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

PupiNA  CROSS  EI,  Brazier. 

Pupina  crossei,  Brazier,  Proc.  Linn.  See.  N.S.W.  i.,  1876,  p.  110. 

(Plate  iii.,  fig.  38.) 

One  of  the  type  shells  in  the  Macleay  Museum  is  here 
illustrated.  It  is  from  Palm  Island  (I  understand  the  most 
northern  of  that  archipelago  was  the  only  one  visited  by  the 
Chevert  Expedition),  and  is  7  mm.  in  length. 

Pupina  nitida,  Brazier. 

Fiipiaa  nitida^  Brazier,  Proc.  Linn.  Soc.  N.S.W.  i.,  1876,  p.  136. 

(Plate  iii.,  fig.  37.) 

One  of  the  original  lot  from  Barrow  Island  is  here  figured.  It 
is  9  mm.  in  length.  \ 

Sitala  reedei,  Brazier. 

Helix  {Conidus)  reedei,  Brazier,  Proc.  Linn.  Soc.  N.S.W.  i., 
1876,  p.  101.  Helix  (Conulas)  inaino,  Brazier,  loc.  cit.  Comdiis 
mairio,  Hedley,  op.  cit.  (2),  vi.,  p.  75,  pi.  ix.,  f.  6.  Sitala  maino, 
Hedley,  op.  cit.,  ix.,  p.  384. 

(Plate  iii.,  fig.  45.) 

A  specimen  from  Darnley  Island  (height  2*5,  major  diam.  3*4, 
minor  diam.  3  mm.)  is  here  figured.  I  cannot  distinguish  this 
from  the  New  Guinea  S.  maino,  which  I  therefore  suppress  as  a 
synonym.  Because  it  is  not  of  Latin  form,  the  name  maino  is 
not  as  acceptable  as  reedei. 

Sitala  nepeanensis,  Brazier. 

Helix  (Conidus)  nepeanensis,  Brazier,  Proc.  Linn.  Soc.  N.S.W. 
i.,  1876,  p.  102. 

(Plate  iii.,  figs.  40-42.) 

One  of  the  type  lot  from  Nepean  Island  is  here  figured.  The 
dimensions  of  it  are  :  height  2*6,  major  diam.  3*64,  minor  diam. 
3-48. 


BY    C.    HEDLEY.  21 

SiTALA  DARNLEYENSis,  Brazier. 

Helix  (Oomdufi)  darnleyensis,  Brazier,  Proc.  Linn.  Soc.  N.kS.  W. 
i.,  1876,  p.  102. 

(Plate  iii.,  fig.  43.) 

One  of  the  types  from  Darnley  Island  is  here  drawn.  Its 
dimensions  are  :  height  5,  major  diam.  5-3,  minor  diam.  5  mm. 

SiTALA    BARNARDENSIS,   Brrtzier. 

Hi'lix  {Conulus)  barnardensis,  Brazier,  Proc.  Linn.  Soc.  N.S.W. 
i.,  1876,  p.  102. 

(Plate  iii.,  fig.  4-4.) 

The  single  specimen  taken  by  the  Chevert  Expedition  at 
Barnard  Island,  No.  iii.,  is  here  depicted.  Its  measurements  are: 
height  1*9,  major  diam.  2-35,  minor  diam.  2*1  mm. 

Helix  porti,  Brazier. 
Brazier,  Proc.  Linn.  Soc.  KS.W.  i.,  1876,  p.  104. 

The  single  specimen  from  Albany  Island  is  rather  broken.  It 
seems  to  me  to  be  a  Piipiso^na,  but  not  P.  circumlit2i7n,  Hedley. 

Flammulina  (?)  GRENViLLEi,  Brazicr. 

Helix  (Conulus)  greiivillei,  Brazier,  Proc.  Linn.  Soc.  N.S.W.  i., 
1876,  p.  104. 

(Plate  iii.,  fig.  46.) 

The  single  representative  of  this  species  from  the  Home  Islands 
is  still  preserved  in  the  Macleay  Museum.  Its  size  is  :  height 
1-18,  major  diam.  1*46,  minor  diam.  13  mm.  Brazier  was  in 
error  in  describing  it  as  imperforate,  for  I  find  a  minute  narrow 
perforation.  The  type  is  so  wrapped  in  matted  fungoid  threads 
as  to  obscure  details.  Hence  I  have  been  unable  to  satisfactorily 
make  out  the  lines  of  bristles  described,  and  my  drawing  conveys 
only  the  general  shape.  The  systematic  position  of  the  species 
is  quite  uncertain. 


22  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

With  this  species  I  have  completed  the  work  of  figuring  the 
unfigured  shells  of  the  Chevert  Expedition.  Rhytida  jamesi, 
Brazier  (P.L.S.N.S.W.  i.,  1876,  p.  99)  is  now  regarded  by  its 
author  as  s3^nonymous  with  B.  franklandiensis,  Forbes.  This 
decision,  in  which  I  concur,  makes  a  figure  of  it  needless. 

The  Genus  Ctiloceras. 

(Plate  ii.,  figs.  28,  29,  30.) 

The  name  Ctiloceras  was  proposed  by  Watson*  for  a  remark- 
able shell  taken  in  Torres  Straits.  Whether  the  shell  is  adult, 
and  to  what  group  it  belongs,  are  unknown.  Watson  suggested 
that  it  was  related  to  Ccecnm,  and  yet  included  it  among  Vermetus. 

A  larger  series  than  Watson  studied  has  come  into  my  hands, 
and  it  aj)pears  that  several  species  of  Ctiloceras  occur.  Although 
I  regard  Ctiloceras  as  an  immature  state  comparable  to  Simisigera, 
possibly  the  young  of  species  already  named,  yet  it  seems  con- 
venient to  appl}^  provisional  names  to  the  different  forms. 

For  comparison  a  mutilated  example  of  C.  cyclicitm,  Watson,  is 
shown  at  fig.  28.  It  is  1*6  mm.  in  major  diameter,  and  was 
found  by  Mr.  J.  Brazier  in  Torres  Straits  at  the  depth  of  8 
fathoms. 

Taken  by  the  same  collector  in  the  same  localit}^  is  a  shell 
shown  at  fig.  30  which  measures  1-54  mm.  in  major  diameter. 
It  is  at  once  separable  by  its  clathrate  sculpture.  I  propose  to 
call  it  Ctiloceras  dathratum. 

Another  and  more  advanced  form,  shown  at  fig.  29,  I  name 
Ctiloceras  striatum.  In  this  the  nucleus  is  more  prostrate  than  in" 
the  others,  the  first  spiral  whorl  is  more  tightly  coiled,  leaving  a 
smaller  orifice;  the  radiating  sculpture  is  weak  and  distant;  close 
fine  spiral  grooves  ornament  the  shell.  In  the  example  drawn 
there  are  one  and  one  half  spiral  whorls.  The  aperture  is  rather 
Ijroadly  reflected.  There  is  no  evidence  to  show  whether  this 
feature  indicates  maturit}^  or  the  approach  of  a  third  stage  in 


*  Watson,  Reports  Chall.  Exped.  xv.,  p.  465,  pi.  xxi.  f.  I. 


BY    C.    HEDLEY.  23 

growth.     It  is  1-36  mm.  in  major  diameter,  and  was  taken  })y 
Mr.  Brazier  in  20  fathoms  off  Cape  Grenville,  Queensland. 

On  the  genus  Lippistes. 

In  examining  the  type  shells  of  the  Chevert  Expedition*  I  had 
occasion  to  re^'ise  several  species  referred  to  Trichoti-opis,  and 
concluded  that  the  genus  SejjarcUista  included  four  Australian 
species,  viz.  : — S.  sejyaratista,  Dillwyn,  S.  gracile7ita,  Brazier,  .S'. 
torcidaris,  T.  Woods,  and  S.  gahrieli,  Pritchard  &l  Gatliff. 

A  casual  and  inaccurate  allusion  to  Lippistes  by  Melvill  and 
Standenf  induced  me  to  write  to  Mr.  B.  B.  Woodv/ard  for 
information  on  Lipinstes  and  its  relation  to  Sepai-atista.  Mr. 
Woodward  in  turn  consulted  Mr.  E.  A.  Smith,  and  to  these 
gentlemen  I  am  deeply  indebted  both  for  the  following  informa- 
tion and  for  permission  to  publish  it.     Mr.  Woodward  writes  : — 

^'■Lippistes  is  a  good  genus;  its  type  is  the  Aryonauta  cornu, 
Gmel.,  figured  by  Montfort,|  and  subsequently  and  better  by 
von  Fichtel  (Test.  Micr.,  pi.  i.).  There  are  good  specimens  in 
the  British  Museum  collection,  and  it  is  very  clear. 

"  As  ^''ou  say  there  are  two  Turbo  helicoides,  Gmel. ;  and 
Dillwyn  selected  the  first  and  re-established  Chemnitz's  name  for 
it  of  Tui'ho  separatista;  but  Chemnitz,  not  being  at  that  time 
binomial,  the  species  must  be  attributed  to  Dillwyn. 

"Gray,  in  1847  (P.Z.S.,  p.  136),  apparently  ignorant  of 
Dillwyn's  work,  raised  this  species  to  generic  rank  as  Separatista, 
and,  while  quoting  the  reference  to  Chemnitz  correctly,  misquoted 
Gmelin,  and  gave  as  type,  '  Ttirho  helicina,  Gmelin,'  meaning  of 
course  ^  helicoides.^ 

'•  Then  Adams  (P.Z.S.,  1850,  p.  45),  seemingly  knowing  nothing 
of  Dillwyn's  doings,  and  possibly  recognising  the  double  helicoides, 
rechristened  the  beast  ^Separatista  chemnitzii,^  and  made  some 
observations  on  Lippistes  and  cornu  which  are  absolutely  wide 
of  the  mark. 

"'  Hedley,  Records  Aust.  Museum,  iv.,  1901,  p.  126. 

t  Melvill  &  Standen,  Proc.  Zool.  Soc.  1901,  p.  361. 

X  Montfort,  Conch.  Syst.  ii,,  1810,  pi.  xxxiv.,  p.  217. 


24  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

"  Gray's  surmise  that  Lijypistts  might  be  a  S3^nonym  for  his 
Separatiata  proves  correct,  and  the  former  name  having  priority 
stands,  hence  : — 

''LippiSTES,  Montfort,  1810  [^  Separatista,  Gra}',  1847]. 

"1.  L.  cornu  (GmeL).     Type  [^Separatista  Grayii,  Ad.,  loc.  cit.'] 

(5  carinse). 
•*  2.  L.  seimratista  (Dillwyn).     Type  of  Separatista  (3  carin^e) 
\^  =  Turho  helicoides,  Gmel.,  p.  3598. 
=  Turbo  separatista,  Dillwyn. 
—  Separatista  chemnitzii,  Ad.] 
"  There  are,  I  fancy,  more  species,  but  these  are  for  the  moment 
immaterial." 

In  view  of  this  information  so  kindly  communicated  by  Mr. 
AVoodward,  it  will  be  necessary  to  transfer  the  four  Australian 
species — S.  separatista,  S.  gracileiita,  S.  torcu^aris,  and  >S'.  gahrieli 
— from  Separatista  to  Lippistes.  I  have  noted  some  remarks  b}^ 
Dall  on  Lippistes  (Bull.  Mus.  Comp.  Zool.  xviii.,  1889,  p.  386-7) 
and  Separatista  (Bush,  Trans.  Connect.  Acad,  x.,  1897,  p.  105). 

On  Perry's  Australian  Shells. 
Perry's  work  is  of  particular  importance  to  Australian  natu- 
ralists, because  he  had  access  to  earty  collections  of  Australian 
shells,  especially  to  that  of  Lieut. -Col.  Wm.  Paterson,  who  filled 
the  post  of  Lieut.-Governor  of  New  South  Wales  in  1800-1802, 
and  corresponded*  with  Sir  Joseph  Banks. 

*  Colonel  Paterson  deserves  to  be  remembered  as  one  of  the  earliest 
Australian  conchologists.  He  arrived  in  Sydney  in  1790  as  a  captain  in  the 
New  South  Wales  Corps,  with  a  high  reputation  as  a  traveller  and  naturalist, 
having  already  published  a  work — * '  A  Narrative  of  Four  Journeys  into  the 
Country  of  the  Hottentots  and  Caffraria  in  the  years  1777-8-9,"  dedicated  to 
Sir  Joseph  Banks.  He  had  ample  opportunities  during  two  decades  for 
collecting  in  Australasia.  Besides  various  journeys  in  New  South  Wales,  he 
commanded  the  settlement  at  Norfolk  Island  and  founded  that  at  Port 
Dalrymple  in  Tasmania.  An  affluent  of  the  Hunter  Eiver  is  named  in  his 
honour,  and  Robert  Brown  dedicated  the  genus  Patersonia  to  him.  He 
appears  to  have  corresponded  with  Lamarck,  for  two  Australian  shells  are 
acknowledged — Corbnla  tailensis,  from  a  M.  Patersoon  ;  and  Buccinum 
hrasilkumm,  from  Madame  Paterson. 


BY    C.    HEDLEY.  25 

^The  full  title  of  his  book  is  : — "  Conchology  |  or  the  |  Natural 
History  of  Shells :  |  containing  |  a  new  arrangement  of  the  genera 
and  species,  |  illustrated  by  |  coloured  engravings  |  executed 
from  the  natural  specimens,  |  and  |  including  the  latest  dis- 
coveries I  by  I  George  Perry."     London,  1811. 

The  work  is  post-Linnean,  the  species  are  named  binomially, 
and  are  systematically''  figured  and  described.  In  short,  the 
nomenclature  fulfils  the  requirements  of  modern  zoological  legis- 
lation, and  must  take  precedence  of  younger  names.  The  illus- 
trations vary  in  merit;  some  are  excellent,  others  are  wretched; 
but  most,  if  not  all,  are  recognisable. 

Gray  wrote  in  1829  : — "  I  have  ventured  to  refer  to  this  work, 
as  T  consider  that  it  is  just  that  every  author  should  be  quoted; 
and  this  author  has  anticipated  Lamarck,  Swainson  and  Sowerby 
in  several  species."* 

Gray  put  his  principles  into  practice  by  withdrawing  his 
Cypraea  jyrinceps  in  favour  of  Perry's  prior  Gypraea  valentia. 

Deshayes  did  not  meet  with  Perry's  book  until  he  had  partly 
completed  his  second  edition  of  Lamarck's  Animaux  sans  Verte- 
bres.  But  through  the  latter  part  of  that  work  he  regularly 
quoted  Perry,  and  withdrew  his  species  Fusus  laticostatus  in 
favour  of  Perry's  earlier  name  of  F.  variegatns. 

Sowerby  and  Reeve  were  not  disposed  to  acknowledge  any 
work  which  upset  their  species.  They  ungenerously  denied 
Perry's  right  to  be  cited,  and  assailed  his  nomenclature  with 
hatred  and  abuse. f  So  powerful  and  evil  was  their  influence  that 
Perry's  'Conchology'  has  been  practically  suppressed  by  London 
writers.  Even  the  brothers  Adams,  who  offended  against  modern 
usage  by  adopting  names  from  polynomial  and  pre-Linnean 
writers,  yet  refused  to  admit  Perry's  names. 


*  Gray,  Zool.  Journal,  iv.,  p.  68. 
t  Sowerby,  Zool.  Journal,  iv.,  p.  217:  Reeve,  Conch.  Icon,  iii.,  Cyprsea^ 
Sp.  17,  note;  iv.,  Fusus,  Sp.  33,  note;  and  vii.,  Ranella,  Sp.  47,  note. 


2Q  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

The  restoration  of  certain  of  Perry's  names  is  required  by  the 
rules  of  zoological  nomenclature.  After  a  careful  examination  of 
Perry's  'Conchology,'  and  guided  by  the  references  of  Deshaj'es  and 
others,  I  have  compiled  the  following  list  of  those  Australian 
shells  which  apparently  were  first  described  by  Perry : — 

Monoplex  cornutiis,  Perry  (pi.  iii.,  f.  1),  takes  priority  over 
Triton  exai'atus,  Reeve,  1844. 

Monoplex  formosus,  Perry  (pi.  iii.,  f.  5),  is  recognised  by 
Deshayes  (An.  s.  Vert,  ix.,  p.  635)  as  a  synonym  of  Triton 
clavator,  Lamarck,  1822  {  =  Mitrex  clavator,  Dillwjm,  1817). 
Triton  sinensis,  Reeve,  seems  to  me  to  be  identical, 

Biplex  aust7'alasia,  Perry  (pi.  iv.,  ff.  2,  4),  is  identified  by 
Deshayes  (An.  s.  Vert,  ix.,  p.  542)  as  Ranelia  leucostonia, 
Lamarck,  1822. 

Biiplex  perca^  Perry  (pi.  iv.,  f.  5)  is  given  precedence  by 
Deshaj-es  (An. s.  Vert,  ix.,  p.  556)  over  Ranelia  jndchra,  8owerby, 
1844. 

Biplex  rubicola,  Perr}'-  (pi.  v.,  f.  5),  is  identified  by  Morch  (Cat. 
Yoldi  Coll.,  1852,  p.  106)  as  Ranelia  granifera,  Lamarck,  1822. 

Triplex  frondosa,  Perr}"  (pi.  vi.,  f.  1),  is  quoted  by  Brazier  (Cat. 
Murex,  p.  60)  as  equivalent  to  Murex  australis,  Quoy  tt  Gaimard, 
1832,  and  to  M.  pahniferus,  Sowerby,  1840.  He  overlooked  the 
fact  that  Murex  australis  was  preoccupied  by  Gmelin,  1790. 
Triplex  denudata,  Perry  (pi.  vii.,  f.  2),  seems  to  me  to  be  tlie 
same  species.     I  recommend  the  use  of  Murex  denudata,  Perry. 

Triplex  flavicunda,  Perry  (pi.  vi.,  f.  2),  and  T.  I'ubicunda, 
Perry  (pi.  vi.,  f.  2),  are  marked  by  Deshayes  (An.  s.  Vert,  ix., 
p.  574)  as  synonyms  of  Mnrex  adnstus,  Lamarck,  1822.  Over 
all  these  names  Purpura  scahra,  Martyn  (Univ.  Conch.  1789, 
pi.  113),  has  precedence. 

Sei^ta  parkinsonia,  Perry  (pi.  xiv.,  f.  1),  is  obviously  identical 
with  Triton  fusiforme,  Kiener,  1842. 

Septa  rubicunda,  Perry  (pi.  xiv.,  f.  4),  is  Triton  riodijerum, 
Lamarck,  1822.      Plates  i.-ii.   of  Hunter's  'Historical  Journal' 


BY    C.    HEDLEY.  27 

(1793)  represent  this  species,  but  have  never  Ijefore  been  quoted 
or  identified. 

Voluta  paUersonia,  Perry  (pi.  xvii.,  f.  1),  an  extra-limital 
species,  is  introduced  to  note  that  Deshayes  apparently  erred  in 
considering  (An.  s.  Vert,  ix.,  p.  402)  this  to  be  Cancellaria 
reticulata,  Linn.  Brazier  has  stated  (P.L.S.N.S.W.  [2],  ii.,  p.  996), 
as  I  believe  correctly,  that  this  illustration  represents  F.  nucleus, 
Lamarck.  As,  however,  Lamarck's  preliminary  description  of 
that  species  bears  the  same  date  (1811)  as  Perry's,  there  is  no 
need  to  disturb  the  accepted  nomenclature. 

Cyprcea  valentla,  Perry  (pi.  xxiii.,  f.  2),  is  recognised  by  Gray 
(Zool.  Journ.  iv.,  p.  68)  as  Cyprcea  jyrinceps,  Gray,  1824.  This 
species  has  occurred  at  Warrior  Reef,  Torres  Straits. 

Bulimus  carlnatus,  Perry  (pi.  xxx.,  f.  1),  has  been  recognised 
by  Pilsbry  (Nautilus,  xv.,  p.  8)  as  Cantharidus  peronii,  Philippi, 
1846.  Jay  (Cat.  Shells,  4th  ed.,  1852,  p.  192)  has  wrongly 
identified  this  as  Bulimus  zebra,  Miiller. 

Bulimus  eximius,  Perry  (pi.  xxx.,  f.  2),  has  been  recognised  by 
Pilsbry  (Nautilus,  xv.,  p.  8)  as  Cantharidus  badius,  Wood,  1856. 

Cassidea  labiata,  Peny  (pi.  xxxiv.,  f.  1),  takes  priority  over 
Cassis  achatina,  Lamarck,  1822. 

Mitra  abbatis,  Perry  (pi.  xxxix.,  ff.  2-3),  was  described  by 
Lamarck  in  the  same  year  as  M.  pontificalis.  The  latter  need 
not  be  disturbed. 

Bulla  ferruginosa,  Perry  (pi.  xl.,  f.  2),  is  identified  by  Pilsbry 
(Man.  Conch,  xv.,  p.  389)  as  equivalent  to  Hydatina  albocincta, 
Hoeven  (1839),  but  he  adds  that  its  use  is  barred  by  the  prior 
Bulla  ferruffinea,  Gmelin  (1790). 

A7'anea  triremis,  Perry  (pi.  xlv.,  f.  3),  is  recognised  by  Deshayes 
(An.  s.  Vert,  ix.,  p.  566)  as  Murex  tenuispina,  Lamarck,  1822. 

Pyrida  undidatus,  Perry  (pi.  liv,,  f.  1),  is  Fusus  pyridatus, 
Reeve,  1847. 

Pyrida  australasia,  Perry  (pi.  liv.,  f.  4),  is  identified  by 
Deshayes  (An.  s.   Vert,  ix.,  p.   434)  as  Fasciolar'ia  Jilamentosa, 


28  STUDIES    OX    AUSTRALIAN    MOLLUSCA, 

Lamarck,  1822.     But  it  seems  to  me  to  be  a  variety  of  Fasciolaria 
fusiformis,  Valenciennes  (1840),  whose  name  it  should  replace. 

Pleurotoma  acuta,  Perry  (pi.  li\'.,  f.  5),  is  quoted  by  Deshayes 
(An.  s.  Vert,  ix.,  p.  352)  as  equivalent  to  Pleurotoma  tigrina, 
Lamarck  (1822). 

Tellina  aurea,  Perry  (pi  Iv ,  f.  2)  is  evidently  T.  vulsella, 
Chemnitz,  =  T.  rostrata  of  other  authors  than  Linne,  not  T. 
{Phylloda)  aurea,  Schumacher  (1817).  Since  by  Hanley's  showing 
(Ips.  Linn.  Conch.,  p.  38)  the  Linnean  name  has  been  generally 
misapplied,  and  that  of  Chemnitz  was  not  binomial.  Perry's  name 
should  be  utilised.  I  collected  this  species  at  Dunk  Island, 
Queensland,  last  year. 

Venues  disjecta,  Perry  (pi.  Ivii.,  f.  3),  is  beyond  doubt  identical 
with  Venus  lamellata,  Lamarck  (1818).  This  name  of  Lamarck's 
was  preoccupied  by  Linne  (1767).  The  species  should  be  quoted 
as  Chione  disjecta,  Perry. 


EXPLANATION  OF  PLATES. 

Plate  i. 

Figs.  1-4. — Hinge  and  various  aspects  of  Congeria  lunata,  Hedley. 

Figs.  5-9— Hinge  and  various  aspects  of  3Iacti'a  parkesiana,  Hedley. 

Figs.  10-14. — Hinge  and  various  aspects  of  Rocliefortia  donaciformis ,  Angas. 

Plate  ii. 

Figs.  15-17. — Hinge  and  different  aspects  of  Bornia  filo^a,  Hedley. 

Figs.  18-20. — Various  aspects  of  Liotia  incidata,  Hedley. 

Figs.  21-23. — Various  aspects  of  Liotia  corona,  Hedley. 

Fig.  24. — CroHSea  hiconica,  Hedley. 

Fig.  25.~Cros$ea  gatliffi,  Hedley. 

Fig.  26. — Mecoliotia  spino>^a,  Hedley. 

Fig.  27. — Retusa  nitida,  A.  Adams. 

Fig.  28.  —  Ctiloceras  cycliciuii,  Watson. 

Fig.  29.— C.  striatum,  Hedley. 

Fig.  30  —  C.  clathratnm,  Hedley. 


BY    C.    HEDLEY.  29 


Plate  iii. 
Fig.  31. — Pynjidma  senex,  Hedley. 
Fig.  .3*2. — PiirguUna  umeralis,  Hedley. 
Fig.  33. — Pyrpulina  persjjectiva,  Hedley. 
Fig.  34. — PyrguUna  zea,  Hedley. 
Fig.  35. — Teinostoma  involuta.  Hedley. 
Fig.  36. — Aplustmm  brazier i,  Angas. 
Fig.  37. — Piqnna  nitida,  Brazier. 
Fig.  38. — Pupina  crossei.  Brazier. 
Fig.  39. — Bijidaria  macleayi,  Brazier, 

Figs.  40-42. — Various  aspects  of  Sitala  nepeanensi><,  Brazier. 
Fig.  43. — Sitala  darnleyensis,  Brazier. 
Fig.  44.  —  Sitala  harnarde)ui><f  Brazier. 
Fig.  45. — Sitala  reedei,  Brazier. 
Fig.  46. — Flammulina  (?)  grenvillei,  Brazier. 


30 


NOTES    AND    EXHIBITS. 

Mr.  C.  T.  Musson  sent  for  exhibition  a  remarkable  series  of 
specimens  of  garden  plants  exhibiting  floral  or  vegetati\'e  pro- 
lification,  from  the  Hawkesburj^  Agricultural  College  garden  or 
grounds  at  Richmond.  As  far  as  the  records  go  the  present 
season  is  the  driest  known;  and  the  apparently  correlated  tendency 
on  the  part  of  so  many  species  to  develop  monstrous  groAvths 
exceeds  all  previous  experiences.  Thirty-one  species  were  rei^re- 
sented  in  the  collection  exhibited,  comprising  Lima  Bean,  Lucerne,^ 
Eschscholtzia,  Amarantus  (two  species),  Solanum  (two  species), 
Foxglove,  Pentstemon,  Rose,  Quince,  Dianthus,  Phlox,  Antirrhi- 
num, Salpiglossis,  Petunia,  Geranium,  Mignonette,  Polygonum, 
Squash,  Leptosyne,  Aster,  Cosmos,  Hypochoeris,  Coreopsis,  Gaillar- 
dia,  Zinnia,  Marigold,  Centaurea,  Erigeron,  and  Scabious.  Simi 
lar  examples  of  Zinnia  and  Delphinium  were  also  furnished  by 
Mr.  Henry  Deane  from  his  garden  at  Hunter's  Hill;  and  of 
Campanula  and  Hypochoeris  by  Mr.  H.Selkirk  f  rom  Marrickville 

WEDNESDAY,   APRIL  30th,   1902. 

The  Ordinary  Monthly  Meeting  of  the  Society  was  held  in  the 
Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  AVednesday 
evening,  April  30th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S.,  etc..  President,  in  the  Chair. 

Messrs.  W.  L.  May,  Sandford,  Tasm.;  W.  F.  Petterd,  Laun- 
cestoni  and  A.  Jefferis  Turner,  M.D.,  F.E.S.,  Brisbane,  Q., 
were  elected  Ordinary  Members  of  the  Society. 

ThePresident  announced  that  under  the  provisions  of  Rule  xxv., 
the  Council  had  elected  Professor  David,  B.A.,  F.R.S  ,  F.G.S., 
Mr.  Henry  Deane,  M.A.,  F.L.S.,  Arc,  Dr.  T.  Storie  Dixson,  and  the 
Hon.  James  Norton,  LL.D.,  M.L.C.,  to  ])e  Vice-Presidents;  and 
Mr.  J.  R.  Garland,  M.A.,  (56  EHzabeth  Street),  to  be  Hon. 
Treasurer,  for  the  current  year. 

The  Donations  and  Exchanges  for  the  month,  amounting  to  18 
Vols.,  35  Parts  or  Nos.,  1  Bulletin,  3  Annual  Reports  (Admini- 
strative), 7  Pamphlets,  and  5  Miscellanea,  received  from  41 
Societies,  etc.,  and  2  Individuals,  were  laid  upon  the  table. 


31 


THE  GUMMOSIS  OF  THE  SUGAR-CANE. 

{Bad.  vascularum^  Cobb). 

By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

SociETy. 

(Plates  iv.-v.) 

Gummosis  is  a  disease  pretty  widely  distributed  among  plants,, 
and  is  recognised  by  an  unhealthy  appearance  of  the  plant  as  a 
whole,  by  the  leaves  dying  or  rotting,  and  especially  by  the 
presence  of  a  yellowish  gummy  matter  in  the  vessels.  On  cutting-^ 
across  the  vessels  of  a  badly  diseased  plant,  such  as  the  sugar- 
cane, small  yellow  viscid  drops  of  gum  quickly  gather  on  the  cut 
ends  of  the  vascular  bundles.  In  slightly  diseased  plants  the 
microscope  may  be  necessary  to  demonstrate  the  gum  plugging-^ 
the  vessels.  The  disease  has  been  specially  noted  in  the  sugar- 
cane, the  sugar-beet  and  the  vine  chiefly  on  account  of  the 
economic  importance  of  these  plants,  but  it  is  not  unknown 
among  other  members  of  the  vegetable  kingdom,  as  for  example 
the  fig,  olive,  mulberry,  potato,  carrot,  and  tomato. 

The  c-'ummosis  of  the  vine  has  been  ascribed  to  the  activity  of 
a  bacterium  by  Prillieux  et  Delacroix,"^  but  Mangin,t  and  also 
Ratha}',!  consider  that  the  formation  of  gum  in  the  vessels  by 
bacteria  is  very  problematical.  Indeed,  Mangin  goes  further  and 
says  that  the  bacteria  live  upon  the  gum. 

*  Prillieux  et  Delacroix,  Cent.  f.  Bakt.,  2  Abt.,  i.,  300,  Eef. 

tMangin,  op.  rit.  and  also  ibid.  11.,  621,  Ref. 

JRathay,  ibid,  ii.,  620,  Ref. 


32  THE    GUMMOSIS    OF    THE    SUGAR-CANE, 

The  gummosis  of  the  sugar-beet  has  been  investigated  by  many. 
Among  others,  Arthur  and  Golden  separated  a  bacterium  which, 
when  inoculated  into  sound  beets,  produced  the  disease  in  some 
cases.  Busse"^  also  investigated  the  disease,  and  concluded  from 
his  own  experiments  and  from  those  of  others  that  there  is  a  group 
of  bacteria  which  cause  the  gummosis  of  the  beet.  He  experi- 
mented upon  sound  beets  with  pure  cultures,  and  found  that  after 
a  period  of  growth  the  infected  roots  had  developed  a  greater  or 
less  gummosis.  The  bacteria  that  are  claimed  as  being  the 
exciting  cause  of  gummosis  of  the  beet  are  all  capsulated,  non- 
sporulating  organisms,  and  it  would  appear  that  the  gum  is  the 
capsular  envelope. 

The  Gummosis  of  the  Cane. 

The  bacteriology  of  the  gumming  of  the  sugar-cane  has  been 
investigated  by  Cobb,t  who  invariably  found  microbes  in  the  gum 
of  diseased  stalks.  He  inoculated  the  gum  into  tubes  of  agar  and 
gelatine  and  obtained  bacterial  growths.  The  cultural  characters 
of  the  organism  are  described  as  follows: — "Cultures  on  agar- 
agar  gave  at  first  roundish  colonies  on  the  surface  having  to  the 
unaided  eye  no  structure,  either  radiated  or  concentric.  In  the 
course  of  a  few  weeks  the  yellowish  white  and  somewhat  opal- 
escent growth  had  extended  several  inches  along  the  edge  of  the 
tube  on  the  surface  of  the  agar-agar  and  between  the  tube  and 
the  agar-agar  without  causing  any  liquefaction.  On  gelatine  the 
growth  was  much  slower,  and  remained  circular  and  almost  im- 
perceptibly concentric.  Its  colour,  itc,  were  as  on  agar-agar." 
The  italics  are  in  the  original. 

It  is  unfortunate  that  the  only  information  regarding  the 
growth  characters  which  enables  the  organism  to  be  identified  is 
so   limited.      The    description    would    apply    to  at  least  eleven 

described    non-liquefying    species   of  bacteria.      The    characters, 

f 

*Busse,  Cent.  f.  Bakt.,  2  Abt.,  ill.,  680,  Kef. 
t  Cobb,  Agric.  Gazette  of  N.S.  Wales,  1893. 


BV    11.    GREIG    SMITH.  33 

however,  enabled  Cobb  to  recognise  it  as  a  new  species,  to  w^iicli 
he  gave  the  name  Bac.  vascularum. 

In  an  investigation  into  the  pathogenicity  of  the  organism 
Cobb"^  inoculated  five  apparently  healthy  canes  with  the  gum 
and  retained  one  as  a  control.  Unfortunately  the  control  cane 
was  lost,  and  of  the  others  four  were  gummed  and  one  died.  In 
the  four  stalks  that  were  gummed,  the  gummosis  was  most  marked 
in  the  neighbourhood  of  the  point  of  inoculation.  But  as  all  the 
canes  in  the  stools  developed  gummosis  there  is  the  probability 
that  the  inoculated  canes  would  also  have  become  affected  inde- 
pendently of  the  inoculation.  The  fact  that  gummosis  was  most 
marked  in  the  neighbourhood  of  the  site  of  inoculation  indicates 
that  the  gum  probably  set  up  gummosis;  but  like  the  majorit}^  of 
plant  infection  experiments  the  result  of  Cobb's  experiment  is  not 
decisive.  Even  if  the  experiment  had  been  most  satisfactory  it 
would  onl}^  have  shown  that  cane-gum  could  produce  gummosis, 
and  thus  the  pathogenicit}^  of  the  bacterium  itself  would  remain 
doubtful.  In  all  infection  experiments  the  recognised  and  legiti- 
mate method  is  to  purify  the  bacterium  by  growth  in  plates  of 
artificial  media,  and  after  it  has  been  proved  that  the  bacteria 
are  undoubtedly  pure  by  their  constant  appearances  of  growth, 
etc.,  upon  one  medium,  to  employ  the  pure  culture  for  purposes 
of  inoculation. 

The  Method  of  Research. 

The  cause  of  the  gummosis  of  the  cane,  therefore,  appeared  to 
be  sufficiently  undecided  to  warrant  a  thorough  investigation, 
and  I  decided  to  attack  the  problem  from  the  side  of  the  gum 
rather  than  the  plant— a  method  that  has  not  been  previously 
tried,  and  one  which  promised  to  give  more  conclusive  results 
than  could  be  hoped  for  with  infection  experiments.  If  a  gum 
could  be  produced  in  the  laboratory  from  a  bacterium  isolated 
from  the  cane,  and  if  it  were  identical  with  the  gum  obtained 
from  diseased  canes,  then  the  bacterial  origin  of  the  gum  and  the 


Cobb,  Agric.  Gazette  of  N,S.  Wales,  189i. 


34  THE    GUMMOSIS    OF    THE    SUGAR-CAN K, 

identity  of  the  organism  would  be  placed  beyond  all  doubt.  The 
question,  then,  as  to  the  gum  being  a  pathogenic  secretion  of  the 
plant  itself  would  be  answered  conclusively  in  the  negative. 

The  Isolation  of  the  Organism. 

To  facilitate  the  investigation  the  Colonial  Sugar  Refining  Co. 
offered  me  a  portion  of  their  small  supply  of  cane-gum  which  had 
been  collected  some  years  ago.  Furthermore,  one  of  their  officers, 
Mr.  Morison,  inoculated  tubes  of  media  in  the  cane  fields  with 
gum  exuded  from  diseased  canes.  As,  however,  I  received  several 
specimens  of  gummed  cane,  the  tube  cultures  w^hich  did  not  con- 
tain the  desired  organism  need  not  be  further  noticed.  The 
specimens  of  diseased  cane  were  the  variety  known  as  Rappoe, 
and,  on  being  cut  across,  small  yellow  viscid  drops  exuded  from 
the  ends  of  the  cut  vessels.  By  using  a  lens  of  low  power  (  x  10) 
minute  portions  from  the  centre  of  exuded  drops  of  gum  wei-e 
abstracted  and  inoculated  into  various  media,  which  were  placed 
under  different  conditions  of  culture.  It  need  scarcely  be  said 
that  the  cane  was  cut  and  the  infections  made  with  the  usual 
precautions  as  regards  sterilising  the  outer  surface  of  the  cane, 
knives,  etc. 

Under  anaerobic  conditions  there  was  no  growth  either  in  tluid 
or  on  solid  media.  On  slices  of  cane  and  in  acid  fluids  such  as 
wort  and  cane-juice  no  bacteria  developed.  In  other  neutral 
fluids  the  bacteria  grew  scantily.  On  agar  and  gelatine  in  the 
presence  of  cane  or  fruit  sugar  bacteria  grew  slowly  at  tempera- 
tures of  30°  C.  and  under;  at  37°  and  over  no  growth  coidd  be 
obtained  under  any  condition  whatsoe^'er.  What  appeared  to 
be  the  most  suitable  medium  for  the  grow^th  of  the  gum  bacteria 
was  neutral  cane-juice  gelatine.  The  next  best  media  were 
ordinary  glucose  gelatine  and  slices  of  potato.  From  the  growth 
on  cane  gelatine  a  bacterium  was  separated  in  the  pure  state  by 
repeated  cultivation  upon  plates  of  cane  gelatine,  and  its  cultural 
characters  were  studied  upon  other  media.  The  appearances  of 
the  cultures  are  given  at  the  end  of  this  paper,  and  they  will 
serve  to  identify  the  organism. 


BY    R.    GREIG    SMITH.  35 

Compared  with  other  bacteria  the  organism  is  of  slow  growth. 
On  suitable  solid  media  it  forms  raised  yellowish  patches,  which 
have  the  appearance  and  consistency  of  softened  yellow  bees'-wax, 
and  when  treated  with  water  the  culture  slowly  mixes,  forming 
what  appears  to  be  a  partial  suspension  and  partial  solution. 
In  stroke  culture  on  cane-gelatine  the  growth  collects  in  places 
along  the  stroke,  and  forms  characteristic  hanging  drops  ("  tear- 
drops ").  The  gelatine  is  very  slowly  liquefied,  the  liquefaction 
being  best  seen  by  the  disappearance  of  the  gelatine  under  the 
slime,  which  gradually  collects  as  the  culture  grows,  and  gravitates 
to  the  bottom  of  the  sloping  gelatine. 

The  Bacterial  Gu3i  or  Slime. 
When  a  pure  culture  of  the  bacterium  is  smeared  over  the 
surface  of  neutral  cane-gelatine  or  agar  in  a  suitable  vessel,  and 
incubated,  a  luxuriant  growth  is  obtained  after  the  lapse  of  about 
a  week.  The  culture  is  precisely  similar  in  appearance  and  con- 
sistency to  cane-gum.  Both  have  the  same  soft,  buttery  appear- 
ance and  gummy  consistency,  and  both  slowly  mix  with  water  to 
form  a  pale  yellow  opalescent  solution.  The  opalescence  is  due 
to  the  suspended  bacteria,  from  which  it  is  rather  difficult  to 
separate  without  at  the  same  time  removing  the  gum.  The 
separation  cannot  be  effected  by  filtering  through  paper,  and  the 
usual  coagulating  agents  coagulate  both  gum  and  bacteria. 
Aluminium  hydrate,  for  example,  forms  an  insoluble  compound 
with  the  gum.  An  attempt  was  made  to  clarify  the  suspension 
by  heating  it  up  to  three  atmospheres  in  the  autoclave,  a  pro- 
ceeding which  was  successful  with  another  gum-forming  bacterium, 
but  the  suspension  was  unaltered.  Eventually  the  bacteria  were 
separated  by  filtration  through  porous  porcelain.  A  clear  solu- 
tion of  the  gum  was  thus  obtained,  but  at  the  same  time  it  was 
noted  that  a  considerable  amount  of  the  gum  remained  on  the 
porcelain,  adhering  to  the  bacteria,  even  after  the  suspension  had 
been  boiled  for  some  time  to  diff'use  the  gum.  The  filtered  and 
clear  solution  was  tested  simultaneously  with  an  opalescent 
suspension,  and  it  was  found  that  the  two  behaved  similarly  on 
the  addition  of  the  various  reagents,  from  which  we  must  con- 


36 


THE    GUMMOSIS    OF    THE    SUGAR-CANE, 


elude  that  in  the  suspension  the  bacteria  are  inert,  and  for 
purposes  of  identification  of  the  gum  it  is  unnecessary  to 
separate  them. 

In  testing  the  bacterial  slime  and  the  cane-gum  a  number  of 
reagents  suggested  themselves,  and  the  results  of  the  numerous 
tests  which  identify  the  two  are  shown  in  the  following  table. 
In  considering  the  tests  it  must  be  borne  in  mind  that  the  cane- 
gum  at  my  disposal  was  very  limited,  and  the  solutions  con- 
sequently were  weak,  which  accounts  for  the  absence  of  perceptible 
precipitates  on  the  addition  of  dilute  acetic  acid  and  a  mixture 
of  acetic  and  tannic  acids. 

The  Chemical   Reactioxs  of  the  Gum  fkom  Caxe  and  of  the   Bacterial 
Slime  formed  in  Laboratory  Cultures. 


Cane-Gum. 


Bacterial  Slime. 


Lead  acetate 
Basic  lead  acetate 
Ammoniacal  lead  acetate 
Barium  hydrate 
Calcium  hydrate 
Copper  sulphate 
Ferric  chloride  ... 
Hydrochloric  acid 
Alcohol    .. 


Aluminium  hydrate 
Dilute  acetic  acid 
Glacial     ,,       ,, 
Acid  mercuric  nitrate 


Precipitate. 


Opalescence. 


Precipitate. 


Opalescence. 


Slimy      voluminous  ■  Slimy        voluminous 
ppt.  in  presence  of       ppt.  in  presence  of 


Xanthoproteic  reaction 
Sulphuric  and  phosphotungstic 

acids 
Acetic  and  tannic  acids 
Acetic   acid   and   potas.    ferro- 

cyanide 
Hydrochloric   acid   and   potas. 

mercuric  iodide 
Sodium     hydrate    and    copper 

sulphate 


salts. 
.1  Coagulation. 
.j  No  precipitate. 
.!  Precipitate. 
.1  Ppt.  soluble   in  ex- 

j     cess  ;  no  colour  re- 

I     action. 
.   Colour  reaction. 


Precipitate. 
Opalescence. 

Opalescence. 

Ppt.  on  standing. 


salts. 
Coagulation. 
Opalescence. 
Precipitate. 
Ppt.  soluble  in  excess; 

no  colour  reaction. 

Colour  reaction. 

Precipitate. 
Opalescence  and  pre- 
cipitate. 
Opalescence. 

Precipitate. 


No    biuret    reaction » 
but  ppt. 


No  biuret   reaction 
but  ppt. 

No  reactions  were  obtained  with  KI.^,  AgNO.j,  KOH.  BaCl.^.  NaOH,  cone 
HCl,  picric  acid,  Adamkiewicz  reagent. 


BY    R.    GREIG    SMITH.  37 

The  precipitate  obtained  on  the  addition  of  alcohol  to  a 
suspension  of  the  gum  or  slime  in  water  is  exceedingly 
voluminous  ;  a  precipitate  which  occupies  a  volume  of  about  50 
CO.  becomes,  after  squeezing  out  the  dilute  alcohol  in  a  calico 
strainer,  a  small  pellet  measuring  about  a  quarter  of  a  c.c.  On 
precipitating  the  aqueous  suspension  three  or  four  times  with 
alcohol  an  opalescent  alcoholic  solution  is  obtained,  from  which 
the  gum  can  be  precipitated  by  small  quantities  of  neutral  salts, 
such  as  sodium  chloride.  This  fact,  together  with  the  reactions 
obtained  with  some  of  the  albuminoid  reagents,  suggested  the 
similarity  of  the  gum  to  the  mucins.  That  the  zooglcea  slime  of 
,  bacteria  consists  of  mucin,  or  a  substance  nearly  allied  to  it,  has 
already  been  suggested,  but  there  are  many  points  of  difference 
between  the  bacterial  slime  and  the  mucins.  The  crude  gum 
obtained  by  precipitating  the  cultures  with  alcohol  contains  6'1% 
of  ash  and  3*08%  nitrogen  in  the  ash-free,  dry  substance.  Repeated 
precipitation  with  alcohol,  and  also  filtration  of  the  gum  from  the 
accompanying  bacteria,  would  undoubtedly  lower  this  percentage. 
It  is,  therefore,  apparent  that  the  nitrogen  content  alone  is 
sufficient  to  distinguish  the  slime  from  mucin  or  the  allied 
mucinoids. 

The  identity  of  the  bacterial  slime  with  cane-gum  proves  con- 
clusively that  the  isolated  bacterium  is  the  direct  cause  of  the 
gummosis  of  the  plant,  and  also  that  the  gum  is  no  secretion  of  the 
plant  upon  which  the  bacteria  live  saprophytically.  Since  this 
applies  to  the  gummosis  of  the  sugar-cane,  it  probably  also  applies 
to  the  gummosis  of  the  vine  and  other  plants,  in  which  case 
Mangin  and  Rathaj'-  are  wrong  in  considering  that  the  action  of  the 
bacteria  in  causing  gummosis  of  the  vine  is  very  problematical. 
Cane-gum  is  nitrogenous,  and  when  a  clear  filtered  solution  is 
exposed  to  the  air  it  speedily  produces  a  luxuriant  growth  of 
foreign  organisms.  These  undoubtedly  feed  upon  the  gum,  and 
it  is  possible  that  this  observation  gave  rise  to  the  idea  that  the 
bacteria  in  the  vessels  of  the  plant  live  upon  the  gum. 


38 


THE    GUMMOSIS    OF    THE    SUGAR-CANE, 


The  Conditions  of  Gum-Formation. 

Having  proved  the  relationship  of  the  gum  to  the  bacterium, 
there  remained  the  determination  of  the  conditions  under  which 
the  gum  is  formed.  The  slime  had  been  first  obtained  in 
quantity  by  growing  the  bacteria  upon  cane-juice  agar,  i.e.,  a 
solution  of  4%  of  agar  added  to  an  equal  volume  of  neutralised 
cane-juice;  and  although  the  growth  was  very  good,  yet  there 
appeared  to  be  room  for  improvement.  The  best  medium  is 
undoubtedly  very  faintly  acid  cane-juice  gelatine,  but,  unfor- 
tunately, this  cannot  be  employed  when  the  slime  is  wanted  for 
chemical  investigation,  because  the  medium  is  slowh^  liquefied. 
The  first  experiment  was  made  upon  cane-juice  agar,  and  had  for 
its  object  the  question  of  the  improvement  of  the  agar  by  peptone 
and,  incidentally,  the  eflfect  of  a  temperature  of  30°  as  against 
22°  C.  Previous  tests  had  shown  that  no  growth  occurred  at 
37°.  In  this  and  the  following  experiments  the  numbers 
indicate  the  order  of  merit  of  the  cultures  at  the  time  of  observa- 
tion ;  1  being  the  most  luxuriant  growth,  2  the  next  best,  and 
so  on. 
The  Effect  of  1%  Peptone  on  Cane-agak  Cultures  at  22"^  and  30°  C. 


2  days. 

4  days. 

1%  peptone  at  22°           

30°           

No          „          22°           

30°           

2 
1 
3 
3 

2 
1 
3 
4 

This  experiment  showed  that  more  slime  was  formed  when  the 
cane-agar  contained  1%  of  peptone,  and  when  the  culture  was 
incubated  at  30°.  There,  however,  is  the  doubt  that  1%  peptone 
is  too  much,  and  to  test  this  point  various  quantities  of  a  10% 
peptone  solution  were  added  to  10  c.c.  portions  of  a  faintl}^  acid 
medium  containing  glucose  10%,  sodium  phosphate  0-2%, 
potassium  chloride  0-5%,  agar  2%,  and  the  bacteria  were  inocu- 
lated upon  the  sloped  surface  of  the  portions.  Cane-agar  was  not 
used  on  account  of  the  unknown  nature  of  the  constituents. 


BY    R.    GREIG    SMITH. 


39 


The  Influence  of  Peptone. 


Temperature  =  30°  C 

2  days. 

4  days. 

6  days. 

8  days. 

No  peptone 

0-1  c.c.  10%  solution  =  0'1%      .. 
0-2  „       „           „             0-2%      ... 
O-o  „       ,,           „             0-47%    .. 
0-7  ,,       „          ,,            0-65%    ... 
1-0  ,,       ,,           ,,            0-91%    ... 
2-0  .,       ,,           ,,             1-66%   ... 

5 
4 
4 
.     2 
2 
3 
1 

3 
2 
2 

I 
I 
1 
1 

2 

6 
1 

1 
1 
1 
2 
3 

It  appears  from  these  results  that  about  0-5%  of  peptone  in  the 
medium  is  a  good  quantity  to  employ.  With  a  greater  percen- 
tage the  growth  starts  sooner,  but  as  time  goes  on  it  becomes 
slower,  and  is  ultimately  surpassed  by  the  lower  percentages. 


The  reaction  of  the  medium  best  suited  to  the  bacterium  was 
also  investigated,  and  for  this  purpose  various  quantities  of  10% 
tartaric  acid  and  10%  sodium  carbonate  solutions  were  added  to 
10  c.c.  portions  of  neutral  medium,  containing  saccharose  5%, 
peptone  0-5%,  potassium  phosphate  0-5%,  agar  2%. 


The  Reaction  of  the 

Medium. 

Temperature  =  30°  C. 

2  days. 

4  days. 

6  days. 

8  days. 

10  days. 

Tartaric  acid  0-01%    .. 

1 

1 

1 

1 

1 

„     0-02%    ... 

2 

2 

2 

2 

1 

,,     0-03%    ... 

— 

3 

3 

3 

2 

„     0-05%    ... 

— 

— 

— 

— 

— 

,,     008%    .. 

— 

— 

— 

— 

— 

Sodium  carbonate  0-01% 

— 

4 

6 

7 

8 

0-02% 

— 

5 

1 

8 

— 

„         0-03% 

— 

— 

— 

0-05% 

— 

— 

— 

— 

0-08% 

— 

— 

— 

— 

— 

Neutral  control  tube    . . 

— 

4 

5 

6 

5 

,. 

— 

4 

5 

6 

5 

The  effect  of  slight  differences  of  acid  or  of  alkali  in  the  culture 
media  is  very  pronounced.  In  an  absolutely  neutral  medium  the 
bacteria  grow  very  slowly,  while,  when  the  reaction  is  faintly 
acid,  the  growth  is  quick  aijd  luxurious.      Alkalies  prohibit  the 


40 


THE    GUMMOSIS    OF    THE    SUGAR-CANE, 


multiplication  of  the  microbe.  With  0-02%  the  stroke  was 
slow  to  show  itself,  was  alwaj's  very  scanty,  and  finally,  after 
ten  days'  incubation,  it  dried  up. 

The  Location  of  the  Bacteria. 

Cane-juice  has  an  acidity  equal  to  an  amount  of  tartaric  acid 
varying  from  0*2  to  0-3%,  and  as  the  juice  is  derived  from  the 
tissue  we  can  understand  why  the  bacteria  avoid  these  strongly 
acid  tissues  and  inhabit  the  slightly  acid  or  neutral  fluids  of  the 
vascular  system. 

In  view  of  the  fact  that  in  many  bacterial  diseases  of  plants, 
the  microbes  are  found  only  in  the  vessels,  and  that  the  bacteria 
are  very  sensitive  to  the  reaction  of  the  contents  of  the  vessels, 
it  would  appear  that  the  immunity  of  plants  disease-proof  to 
bacterial  infection  may  depend  upon  a  relativel}'^  greater  acidity 
or  alkalinity  of  the  vascular  contents  as  compared  with  susceptible 
varieties. 


A  peptonised  medium  similar  to  that  used  in  testing  the 
influence  of  peptone  was  employed  in  a  small  test  to  corroborate 
the  general  conclusion  that  had  been  drawm  during  the  work  upon 
the  bacterium  with  regard  to  the  temperature,  and  from  the 
growths  upon  the  agar  it  was  manifest  that  a  temperature  of 
30°  C.  was  best  suited  to  the  organism. 

The  Optimum  Temperature, 


2  days . 

4  days. 

6  days. 

22°    

2 

2 

o 

25^    

3 

2 

2 

30°    

1 

1 

1 

That  the  bacterium  grows  rapidly  at  30°  C.  (86°  F.)  and  not 
at  all  at  37°,  serves  to  explain  in  part  at  least  why  the  disease 
occurs  more  frequently  in  cold,  wet  seasons.  Such  seasons  check 
the  growth  of  the  plant,  the  \  itality  is  lowered,  and  at  the  same 
time  the  bacteria,  being  under  favourable  conditions  of  tempera- 


liY    R.    GREIG    SMITH. 


41 


ture,  grow  rapidly,  and  clog  the  vessels,  on  account  of  which  tlie 
plant  is  unable  to  obtain  its  food  supplies,  and  soon  succumbs. 

One  point  to  be  decided  was  the  influence  of  salts  upon  the 
growth  of  the  bacterium,  and  to  test  this  matter  a  faintly  acid 
medium  containing  peptone  0-5%,  dextrose  10%,  agar  2%,  was 
prepared,  and  about  one-quarter  of  a  gramme  of  salt  was  added  to 
each  10  c.c. 

The  Influence  of  Salts. 


2  days. 

4  days. 

6  days. 

8  days. 

Potassium  nitrate          

3 

1 

2 

2 

,,          monohydrogen  phosphate 
, ,          dihydrogen  phosphate 

, ,          sulphate        . .           

, ,          chloride        

2 

5 
2 
3 

1 

1 

2 
2 

1 

2 
2 

,,          citrate           

1 

2 

2 

,,          sodium  tartrate      

4 

4 

3 

Sodium  chloride 

5 

6 

6 

Ammonium  chloride     

3 

2 

2 

Calcium  chloride 

2 

2 

3 

Magnesium  sulphate     .. 

Check  (no  salts) 

3 
2 

2 

4 

2 
3 

This  experiment  shows  that(l)  the  bacterium  has  a  preference 
for  phosphate  and  is  indifferent  to  the  other  acid  radicles  ;  (2) 
the  acid  potassium  phosphate,  on  account  of  its  acidity,  checked 
the  growth;  (3)  potassium  may  be  replaced  by  calcium,  magnesium 
or  ammonium  ;  (4)  sodium  is  a  distinct  poison. 


The  Poisonous  Action  of  Common  8alt. 

After  noting  the  poisonous  action  of  sodium  salts,  the  idea 
naturally  suggests  itself  that  common  salt  might  be  advantageous 
in  checking  the  gummosis  of  the  cane.  Common  salt,  however, 
is  a  poison  for  most  of  the  higher  plants,  and  might  injure  the 
sugar-cane  if  used  in  doses  sufficiently  great  to  inhibit  the 
growth  of  the  bacterium.  On  enquiry,  I  find  that  healthy  crops 
of  cane  are  grown  in  Fiji  upon  soil  containing  anything  under 
1%  of  common  salt,  although  when  this  amount  is  exceeded    t  h 


42  THE    GUMMOSIS    OF    THE    SUGAR-CANE, 

crops  are  not  so  healthy.  This  would  lead  one  to  expect  favour- 
able results  with  salt  when  applied  to  fields  containing  canes 
liable  to  the  disease. 

I  have  enquired  if  canes  grown  on  the  coast,  and  within  reach 
of  the  salt  spray  become  diseased  to  the  same  extent  as  plants 
oi'own  inland,  and  have  been  informed  by  Mr.  T.  U.  Walton, 
B.8c.,  of  the  Colonial  Sugar  Ptefining  Co.,  who  made  the  necessary 
enquiries,  that  the  majority  of  the  managers  of  the  diiferent  mills 
consider  that  canes  grown  upon  soils  which  are  known  to  be 
pretty  w-ell  impregnated  with  salt  through  being  subject  to  the 
influence  of  tidal  waters,  are  not  less  liable  to  be  affected  with 
gummosis  than  crops  upon  other  soils.  The  opinion  is  that  they 
are  more  prone  to  develop  the  disease,  but  with  regard  to  this  it 
must  be  borne  in  mind  that  such  soils  are  generally  badly  drained, 
and  it  is  to  the  defective  drainage  that  the  prevalence  of  the 
disease  is  ascribed.  Cases  can  be  quoted  in  which  gummosis  has 
been  remedied  by  the  improvement  of  the  land  by  draining  and 
the  planting  of  new  seed  cane. 

In  direct  contradiction  to  the  other  (jflicers,  one  of  the  managers 
thinks  that  it  may  be  safely  said  that  on  lands  adjacent  to  the 
sea  beach,  where  the  soil  is  contaminated  with  salt,  gummosis  has 
not  been  nearly  so  bad  as  in  districts  distant  from  the  sea.  He 
instanced  the  case  of  a  man,  whose  farm  was  on  one  of  the 
islands  in  the  Lakes,  Clarence  River,  being  able  to  grow  a  com- 
paratively sound  crop  of  Mauritius  Ribbon  long  after  this  variet)^ 
of  cane  had  been  abandoned  by  ever3^one  else  on  account  of  its 
tendency  to  develop  gummosis.  A  large  portion  of  his  farm  was 
submerged  by  King  tides. 

The  Colonial  Sugar  Refining  Co.  in  the  following  season  intend 
to  test  the  action  of  common  salt  upon  perfectly  drained  soils. 

The  presence  of  salt  in  the  cane  is  preferable  to  gum,  inasmuch 
as  the  former  collects  in  the  molasses  and  does  not  otherwise 
make  its  presence  evident,  wdiile  the  gum,  besides  diminishing  the 
crop,  interferes  w^ith  the  crystallisation  of  sugar. 

Although  one  might  by  experiment  find  a  substance  that 
would  inhibit  the  disease,  the  best  practice  is,  undoubtedly,  to 


BY    R.    GREIG    SMITH. 


43 


grow  disease-resisting  varieties,  as  has  been  previously  recom- 
mended, not  only  for  gummosis,  but  for  other  diseases.  In  this 
relation  it  may  be  mentioned  that  there  are  one  or  two  well 
known  varieties  which  have  never  been  known  to  develop  gum- 
mosis. The  best  known  of  these  sound  varieties  is  Tanna,  a  stout 
heavy  cane,  hard  in  the  rind  and  containing  a  high  percentage  of 
fibre.  The  sugar  content  is,  however,  not  so  high  as  it  is  in  some 
of  the  varieties  which  are  liable  to  gummosis.  Fields  which  have 
yielded  badly  diseased  crops,  have,  when  planted  with  Tanna, 
produced  perfectly  sound  canes. 

In  order  to  form  a  gum  as  it  undoubtedly  does,  the  bacterium 
probably  requires  a  carbohydrate,  and  although  saccharose  has 
been  employed  and  found  exceedingly  useful,  yet  it  does  not  follow 
that  it  is  necessary,  or  that  saccharose  is  the  best  carbohydrate. 
Other  sugars  might  give  a  better  yield  of  gum.  To  test  this 
question,  about  half  a  gramme  of  various  substances  was  added  to 
10  c.c.  portions  of  a  faintly  acid  medium  containing  peptone 
0-5%,  sodium  phosphate  0*2%,  potassium  chloride  0*5%,  agar  2%, 
and  after  sterilisation  of  the  tubes,  the  sloped  surfaces  were 
inoculated  and  then  incubated. 


The  Influence 

OF  Carbohydrates 

ETC. 

Temperature  =  30° 

2  days. 

4  days. 

6  days. 

Glycerine         

Starch 







Dextrin 







Dextrose           

\ 

2 

3 

Levulose           

I 

1 

Saccharose       

1 

1 

1 

Maltose            

Lactose 







Blank  test        

~ 

— 

— 

This  experiment  showed  that  either  dextrose,  levulose,  or 
saccharose  are  absolutely  necessary  for  the  free  growth  of  the 
organism  and  the  production  of  gum.  The  bacteria  did  not 
grow   in    the    presence    of   the    other    sugars    and  carbonaceous 


44  THE    OUMMOSIS    OF    THE    SUGAR-CANE, 

matters,  from  which  we  infer  that  these  cannot  be  utilised.  Of 
the  three  sugars,  levulose  and  saccharose  are  more  easily  assimi- 
lated than  dextrose. 

The  organism  does  not  secrete  invertase.  This  was  shown  by 
the  practical  absence  of  invert  sugar  in  the  agar  cultures.  In 
testing  inversion  fluid  cultures  are  preferable,  but  as  this  bacteria 
does  not  grow  to  anj^  extent  in  saccharose  fluid  media,  an  agar 
culture  had  to  be  taken.  The  agar  was  melted  and  dissolved  in 
water,  after  which  the  solution  was  treated  with  basic  lead 
acetate,  etc.,  as  is  customary  in  sugar  determinations.  Of  the 
saccharose  in  the  tube  originall}',  onlj''  3%  had  been  inverted  to 
fruit  sugar,  a  quantity  which  might  easily  have  been  produced 
through  the  hydrolytic  action  of  the  small  quantity  of  acid  in  the 
medium. 

The  Most  suitable  Medium. 

From  these  experiments  we  conclude  that  an  excellent  medium 
for  the  growth  of  the  bacterium  and  the  production  of  gum  would 
have  the  following  composition,:  — 

Peptone      ...  ...  ...  ...       0-5 

Saccharose  or  levulose       ...  ...        5  0 

Potassium  phosphate         ...  ...        0*5 

Agar  2-0 

Tap  water 100 

Acidity  of  10  c.c.  =0*14  c.c.  tenth-normal  acid. 
On  this  saccharose  medium  the  bacteria  grew  most  luxuriantly 
.at  the  optimum  temperature  (30°),  and  since  the  organisms 
grow  so  well,  we  cannot  doubt  that  similar  conditions  as  regards 
nutrition,  acidity,  and  temperature  favour  the  development  in 
diseased  canes. 

Bacterium  vascularum,  Cobb. 

Shape,  d:c. — The  organism  appears  as  an  actively  motile,  sh(jrt 
rod,  and  when  stained  and  imbedded  in  balsam  has  an  average 
measurement  of  0*4  :  1  fx.  Carbol  violet  followed  by  dilute 
-iilcohol  produces  the  best  films  ;  fuchsin  stains  the  gum,  which 


BY    K.    GREIG    SMITH.  45 

usually  adheres  more  or  less  to  the  cells  ;  the  blues  stain  but 
feebly.  B}^  using  the  night-blue  method  with  the  scanty 
growth  obtained  on  ordinary  agar  the  flagella  can  be  easily 
stained.  They  are  single  and  terminal.  The  bacteria  are  not 
coloured  by  the  Gram  method  of  staining.  Spores  were  not 
obtained  and  are  probably  never  formed. 

Relations  to  oxygen  and  teinperature. — It  is  a  strong  aerobe^ 
and  grows  best  at  30°  ;  at  37°  there  is  no  growth. 

Ordinary  glucose-gelatine  plate, — The  colonies  develop  slowly. 
Tn  7  days  at  22"  they  are  1  mm.  in  diameter,  and  appear  as 
small,  raised,  viscid  drops.  When  magnified  60-fold  they  appear 
round  and  uniformly  granular,  like  a  thin  yeast  colony ;  the  deep 
colonies  are  like  those  upon  the  surface.  Tn  20  days  the  colonies 
reach  a  diameter  of  4-8  mm,,  and  look  like  drops  of  yellow  bees'- 
wax.  The  medium  shows  no  sign  of  liquefaction,  but  when  the 
colony  is  scraped  or  washed  off  a  pit  is  revealed. 

Ordinary  gelatine  plate. — The  colonies  grow  as  in  the  presence 
of  glucose,  but  much  more  slowly. 

Ordinary  gelatine  stroke. — The  growth  is  scanty,  narrow,  flat, 
and  ivory-white  in  colour.  It  slowlj^  gravitates  to  form  a 
yellowish- white  mass.     The  medium  under  the  stroke  is  depressed. 

Neutral  cane-gelatine  stroke. — There  is  formed  a  characteristic 
convex,  deep  yellow  stroke,  with  waxy  drops  at  intervals,  and  at 
the  base.  The  gelatine  in  contact  with  the  culture  is  slightly 
liquefied  in  three  weeks.  The  colour,  bloom,  and  general  appear- 
ance is  that  of  yellow  wax,  or  of  a  mixture  of  yellow  vaseline  and 
paraffin.  The  "  tear-drop  "  appearance  of  the  stroke  is  character- 
istic. 

Neatral  cane-gelatine  stab. — The  growth  forms  in  the  upper 
portion  of  the  stab  only  in  a  filiform  manner,  and  forms  a  hemi- 
spherical, deep  yellow  nail-head.     No  gas  bubbles  are  produced. 

Ordinary  glucose-gelatine  stab. — As  cane-gelatine,  but  neither 
so  luxuriant  nor  so  deeply  coloured.  ISTo  gas  formation  was 
observed. 

Ordinary  gelatine  stab. — Filiform  growth  in  upper  portion  of 
stab,  with  small,  raised,  white,  glistening  nail-head. 


c 
/- 


fc 


46  THE    GUMMOSIS    OF    THE    SUGAR-CANE. 

Ordinary  luitrletit-agar  stroke. —  A  slow-growing  thin,  yellowish 
white  glistening  stroke. 

Glycerine  niitrient-agar  stroke. — A  thin,  broad,  translucent 
white,  moist,  glistening  growth,  with  turbid  condensed  water. 
The  colour  deepens  to  a  primose-yellow. 

Saccharose  {lOy^Yj'yeptone  {0'iy^)-agar. — A  thin  white  fluid 
growth,  which  gravitates  into  the  condensed  water,  in  which 
there  is  a  yelloAv  sediment. 

Potato. — A  primrose-yellow,  moist,  glistening  growth,  some- 
times raised  and  restricted,  at  other  times  flat,  watery,  and 
spreading  over  the  surface.  Compared  with  agar  or  gelatine, 
the  growth  is  rapid. 

Carrot. — A  raised,  slimy,  yellow  growth,  at  tirst  restricted,  but 
eventually  covers  the  surface  and  gravitates. 

Turnip. — As  on  carrot. 

Sugar-cane. — No  visible  growth. 

Nutrient  houillon. — Slights  turbid  fluid,  with  faint  indol  re- 
action. 

Nitrate  bouillon. — There  is  no  reduction  of  the  nitrate. 

Siveet  ivort. — No  growth. 

Cane-juice. — No  growth. 

Milk. — Unaltered  ;  neutral  reaction. 

The  Affinities  of  the  Bacterium. 

Since  the  organism  does  not  form  colonies  on  gelatine  which 
throw  out  processes  from  the  margin,  it  is  not  jBact.  gummis  to 
which  Comes  ascribes  the  gummosis  of  the  vine.  Bad.  ajni,  said 
to  produce  a  disease  in  celery,  is  larger,  and  grows  more  slowly 
upon  potato,  but  otherwise  there  are  points  of  similarity  in  the 
appearance  of  the  colonies  on  gelatine,  which,  however,  are 
white  as  against  the  j'ellowish  colour  of  Bact.  vasculanim. 
Basse's  bacteria  produce  gas  in  glucose  media,  and  otherwise 
differ.  Bact.  tracheiphilus,  a  bacterium  which  causes  the  wilting 
of  some  of  the  C^icurbitacece  by  plugging  the  vessels,  differs  in 
colour  (white),  and  by  forming  very  ropy  cultures  in  fluid  media, 
especially  when  old.     Arthur  and  Golden  have  described  Bact. 


1?Y    R.    GHEIG    SMITH.  47 

betif,  wJiicli  possibly  causes  gummosis  of  the  sugar  beet,  with  so 
little  detail,  that  it  cannot  be  compared.  The  same  remark 
applies  to  Bad.  mori,  said  by  Boyer  and  Lambert  to  produce  a 
disease  (gummosis  ?)  of  the  mulberry.  Of  the  other  bacteria  not 
associated  with  diseases  of  plants,  there  appear  to  be  none 
described  which  have  the  general  characters  of  this  bacterium,  and 
although  Cobb's  description  is  meagre,  there  is  no  doubt  that  he 
intended  the  name  of  Bac.  vascularutyi  for  this  organism,  which 
he  found  constantly  associated  with  the  gum  of  affected  plants. 


DESCRIPTION  OF  PLATES. 

Plate  iv. 

Fig.  1.  Margin  of  an  impression  taken  from  the  gum  exuded  from  the  cut 
vascular  bundle,  stained  with  fuchsin,  and  partly  decolorised  with  alcohol 
(X  1000). 

Fig.  2.  Film  from  growth  on  cane-gelatine,  stained  with  violet  (  x   1000). 

Fig.  3.  Film  of  growth  on  nutrient  agar,  stained  by  the  night- blue  method 
for  fiagella  (  x   1000). 

Fig.  4.  Colony  on  glucose-gelatine  (  x  80). 

Fig.  5.  Characteristic  "  tear-drop  "  growth  on  cane  gelatine  (  x  J). 

Plate  V. 

Fig.  1.  Section  of  sugar-cane  affected  with  gummosis,  showing  bacteria  in 
large  vessel,  stained  with  dilute  carbol-fuchsin  (  x   500). 

Fig.  2.  Another  section  showing  bacteria  at  the  margin  of  the  contracted 
slime  in  large  vessel  (  x  750). 


48 


ON     A     GYROCOTYLE     FROM     GHIMJ^RA     OGILBYI, 
AND    ON    GYROCOTYLE    IN    GENERAL, 

By  W,  a.  Haswell,   M.A.,  D.Sc,  F.R.8.,  Challis  Pkofessok 
OF  Biology,  University  of  Sydney. 

(Plate  vii.) 

The  Monozoic  Cestodes  of  the  genus  Gyrocotyle  ( Ampliiptyches) 
are,  in  the  adult  condition,  parasites  exclusively  of  fishes  of  the 
order  Holocephali,  and  have  been  found  not  only  in  the  northern 
Cki?ncera  monstrosa,  but  in  the  southern  Callorhynchus  antarcticus 
and  C.  argenteus.  It  is,  therefore,  not  a  matter  for  surprise  that 
the  new  species  of  the  former  genus  trawled  by  the  "  Thetis  "  in 
1898,  and  described  b}'  Mr.  E.  R.  Waite  under  the  name  of  C. 
Oyilbyi,*  proves  to  be  the  host  of  a  member  of  this  group.  A 
few  months  ago  Mr.  Waite  sent  me  two  specimens  in  alcohol  of 
a  species  of  Gyrocotyle,  taken  from  a  specimen  of  C/mnchra 
Ogilhyi,  recenth^  received  by  the  Australian  Museum  from  Manlv. 

In  comparing  these  specimens  with  the  published  accounts  of 
G.  2triia  and  G.  rugosa,  and  with  a  preserved  specimen  and 
sections  of  the  latter,  a  difficulty  occurs  which  has  to  be  dealt 
with  at  the  outset.  Contrary  opinions  are  held  b}^  difterent 
observers  as  to  the  orientation  of  the  animal.  By  Wagener  (8), 
followed  by  Diesing,  Olsson,  Monticelli  (5, 6),  Braun  (2,  3),  and 
others,  the  end  Ijearing  the  peculiar  plicated  sti-ucture  termed  the 
funnel  is  looked  upon  as  posterior.  By  Baldwin  Spencer  (7),  and 
l)y  Lijnnberg  (4),  on  the  other  hand,  the  funnel  end  is  regarded  as 


*  E.    11.    Waite,    "Scientitic    Results   of    the    Trawling   Expedition    of 
H.M.C.S.  '  Thetis."*     Australian  Museum,  Memoir  iv.,  1899. 


BY    W.    A.    HASWELL.  49 

anterior.  It  will  be  iiecessar}-,  therefore,  to  decide  which  of 
these  opposed  views  is  to  be  adhered  to  in  making  this  comparison. 

Spencer's  contribution  to  the  subject  was  published  at  least  a 
3^ear  before  Lonnberg's;  but  the  latter  reached  his  conclusion 
independently,  and  adduces  several  reasons  of  some  weight  in 
support  of  his  position.  He  alleges,  in  the  first  jDlace,  that  the 
animal  creeps  in  the  direction  of  the  end  which  bears  the  funnel. 
On  this  point  I  am  unable  to  make  any  statement,  as  I  have  not 
had  an  opportunity  of  observing  living  specimens.  But,  in  the 
absence  of  other  evidence,  it  is  worthy  of  note  that  Wagener 
states  "Die  Schwanzkrause  bleibt  meist  ruhig  wahrend  der  Kopf- 
theil  des  Thieres  sich  langsam  bewegt." 

The  direction  of  the  spinules  on  the  surface  Lunnberg  regards 
as  affording  further  evidence  in  favour  of  his  view.  If  the  end 
with  the  funnel  is  posterior,  then  these  spinules  lie  with  their 
points  directed  forwards.  Further  support  Lonnberg  conceives 
to  be  derivable  from  the  structure  of  the  nervous  system,  nerve- 
cells  being  more  numerous  in  the  commissure  at  the  end  which 
he  looks  upon  as  anterior.  Monticelli  (5),  however,  states  that  the 
contrar}^  is  the  case. 

But  a  comparison  of  the  reproductive  apparatus  of  Gyrocotyle 
with  that  of  allied  forms  appears  to  me  to  place  it  beyond  a  doubt 
that  the  sucker  end  is  that  which  corresponds  to  the  end  usually 
regarded  as  anterior.  If  we  take  the  contrary  view  it  is 
impossible  to  trace  any  correspondence  in  the  arrangement  of  the 
parts  :  whereas  a  general  unity  of  plan  at  once  becomes  apparent 
on  the  position  becoming  reversed.  When  we  bear  in  mind  that, 
though  Gyrocotoyle  is  unsegmented,  it  possesses  a  number  of 
highly  specialised  Cestode  features  in  its  adult  structure,  and  is 
characterised,  moreover,  by  the  occurrence  in  its  life-history  of 
the  peculiar  and  characteristically  Cestode  hexacanth  larva,  the 
necessity  for  tracing  such  a  correspondence  becomes  manifest. 
The  end  which  bears  the  sucker  is  seen,  as  a  result  of  such  a 

*  Whether  or  not  this  cori-esponcis  to  the  anterior  end  in  other  classes  of 
Flat-worms  is  a  question  outside  the  limited  scope  of  the  present  paper. 
4 


50  ON  A  GYROCOTYLE  FROM  C HIM. ERA, 

comparison,  to  correspond  to  the  scolex  end*  in  the  segmented 
Cestodes,  and  to  that  at  which  the  mobile  organ  is  situated  in 
Caryophyllaeus.  •'' 

Lonnberg  has  shown  that  G.  urna  is  subject  to  verj'  remark- 
able variations  in  form.  In  its  normal  condition  it  has  the  lateral 
borders  and  the  characteristic  posterior  funnel-like  organ  thrown 
into  a  number  of  complicated  plaits.  But  in  some  examples  the 
plaits  of  the  lateral  borders,  or  of  the  funnel,  or  of  both,  may  be 
entirely  absent;  and  in  some  the  funnel  becomes  replaced  b}^  a  long 
cylindrical  tube.  This  modification  of  the  funnel  has  not  been 
observed  in  G.  rugosa  ;  but  living  specimens  of  that  form  have 
not  been  studied  :  the  degree  of  folding  of  the  lateral  l)orders 
has  been  observed  to  be  subject  to  variation 

In  view  of  these  variations  it  becomes  difficult  to  fix  upon  any 
definite  and  constant  differences  between  the  form  from  Chimccra 
monstrosa  and  that  from  CaUorhynchus  antarcticus.  Braun 
suggests  that  the  position  of  the  reproductive  apertures  may 
afford  a  means  of  distinguishing  between  them. 

Spencer's  account  of  these  apertures  in  G.  rugosa  is  somewhat 
confusing  owing  to  certain  inconsistencies  in  the  text,  and  has  to 
be  interpreted  with  the  aid  of  the  figures.  He  describes  the 
presence  on  the  "left"  side  ventrally  of  a  well-marked  somewhat 
conical  and  muscular  papilla.  The  male  aperture  is  "  on  the 
ventral  side  just  at  the  base  of  the  papilla,  towards  the  middle 
line."  This  statement  is  not  borne  out  by  an  examination  of  the 
only  specimen  of  G.  rugosa  at  my  command  at  present,  in  which 
the  aperture  is  situated  at  the  extremity  of  the  papilla,  and  is 
not  consistent  with  Spencer's  own  figures,  one  of  which  (Plate  ii., 
fig.  1)  represents  th".  opening  in  question  as  situated  on  the  mar- 
gin or  approximately  so,  and  another  (fig.    4)   shows  it   on   the 


*  That  the  rejection  of  Hinnberg's  and  Spencer's  views  on  the  orientation 
of  Gyrocotyle  appears  also  to  lead,  as  Benham  (1)  points  out,  to  a  reversal  of 
the  usual  view  with  regard  to  the  position  of  the  sucker  in  AmphUina  need 
only  be  mentioned  here. 


BY    W.    A.    HAS  WELL.  51 

ventral  surface  some  little  distance  from  the  margin,  but  mani- 
festly on  the  summit  of  the  papilla. 

If  we  accept  Lonnberg's  account  of  the  position  of  these 
apertures  in  G.  urna,  there  would  appear  to  be  a  sufficiently 
strongly-marked  difterence  in  this  respect  between  that  species 
and  G.  ruyosa.  In  the  former  the  male  aperture  is  ventral, 
situated  towards  the  middle  line,  and  the  vaginal  aperture  is 
always  in  front  of  it  on  the  dorsal  surface.  In  the  latter  the 
male  aperture  is  ventral,  but  marginal  or  sub-marginal;  and  the 
female  dorsal,  somewhat  to  the  right  of  the  middle,  and  a  little 
behind  the  male.  A  further  external  difference  between  the  two 
forms  concerns  the  cirrus. 

In  the  specimen  of  G.  ruyosa  in  my  possession  (figs.  3,  4  and  5) 
this  is  a  prominent  process,  conical  in  shape,  its  base  situated  on 
the  ventral  side  between  the  middle  line  and  the  right  margin  of 
the  body.  It  is  strongly  curved  round  towards  the  dorsal  side, 
lying  in  a  deep  notch  in  the  right  border,  its  apex  actually 
projecting  on  the  dorsal  side  of  the  body.  From  the  appearance 
presented  it  is  obvious  that  the  specimen  has  been  killed  with  the 
parts  approximately  in  the  position  for  self-impregnation,  the 
apex  of  the  penis  being  approximated  towards  the  vaginal 
aperture  on  the  dorsal  side.  There  is  no  evidence  of  the 
occurrence  of  any  such  arrangement  in  G.  urna.  On  the  contrary, 
L'Jnnberg  states  very  positively  "Da  die  beiden  Genitaloffnungen 
auf  verschiedenen  Korperfliichen  liegen  ist  natiirlich  Selbstbe- 
frachtung  ganz  unmoglich  bei  diesen  Cestoden,  und  auch 
Wechselkreuzung  sehr  schwierig  "  (4,  p.  40). 

The  parasite  of  Chimcera  Ogilhyi  (fig.  1)  is  more  nearly  allied 
to  G.  urna  than  to  G.  rugosa.  How  far  the  slight  differences 
between  it  and  the  northern  species  are  of  a  constant  character 
can  only  be  determined  by  comparison  of  large  series ;  in  t)ie 
meantime  I  refer  to  it  as  G.  nigrosetosa. 

The  two  specimens,  which  are  both  large  (about  70  mm.  in 
length),  resemble  one  another  in  most  respects.  The}^  both  have 
richly  plaited  borders  and  a  "funnel"  with  greatly  folded  walls. 
The  anterior  sucker  resembles  that  of  G.  urna. 


52  ON  A  GYROCOTYLE  FEOM  CHIM.flRA, 

The  spinules  (fig.  6)  are  dark  brown  or  black.  Each  is  a  stout 
cylindrical  rod  swollen  towards  the  middle,  and  with  the  proximal 
extremity  slightly  constricted  off  as  a.  rounded  knob.  They  con- 
form in  shape  exactl}'-  to  the  description  of  those  of  G.  iirna  given 
b}'  Lonnberg.  If  we  are  to  judge  from  Spencer's  figure  the 
spinules  of  G.  rugosa  are  of  simpler  shape.  They  are  mainly 
confined  to  the  dorsal  surface,  but  at  the  anterior  end  at  the  sides 
of  the  sucker  and  on  the  stalk  of  the  funnel  a  number  extend  on 
to  the  ventral  surface.  Over  the  rest  of  the  ventral  surface  are 
scattered  a  very  few  ;  these  are  all  much  smaller  than  those  on 
the  dorsal  surface.  There  are  very  few  on  the  lateral  plaits,  and 
none  on  tho  funnel.  As  in  G.  urna  and  G.  rugosa,  they  all  lie 
with  their  apices  directed  forwards  or  forwards  and  outwards. 
The  longest  are  0*3  mm.  in  length. 

On  the  right  border,  not  far  from  the  anterior  extremity  (fig.  2), 
in  both  specimens,  is  a  well-marked  notch.  On  the  dorsal  aspect, 
close  to  this  is  the  conspicuous  aperture  of  the  vagina  (9).  The 
male  aperture  is  on  the  ventral  surface,  a  little  to  the  right  of  the 
middle  line,  and  slightl}"  behind  the  vaginal  opening.  There  is 
no  cirrus  distinguishable  in  either  specimen.  The  ejaculatory 
duct  has  a  chitinous  internal  lining  beset  with  fine  denticles  or 
spinules  as  in  most  Cestodes.  These  appear  to  be  absent  in  G. 
rugosa,  but  are  present  in  G.  arna.* 

The  specimens  were  not  in  good  condition  for  minute  investi- 
gation, and  I  have  not  attempted  more  than  a  general  examina- 
tion of  the  structure  of  the  remainder  of  the  reproductive 
apparatus.  So  far  as  my  results  go  the}"  correspond  with  what 
has  been  described  by  Lonnberg  and  by  Spencer.  As  Lonnberg 
has  stated,  the  continuation  backwards  of  the  vagina  to  the 
receptaculum  seminis  is  a  very  fine  tube  which  runs  on  the  ventral 
side  of  the  uterus,  and  not  on  the  dorsal  as  represented  by  Spencer 
in  his  coloured  diagram.  The  main  vitelline  ducts  also,  which  are 
represented  in  the  same  figure  on  the  dorsal  side  of  the  ovary, 
are  in  reality  ventral  in  position. 

*  Obviously  what  Lonnberg  (4,  p.  38)  refers  to  as  cilia  are  of  this  nature. 


BY    W.    A.    IIASWELL.  53 

The  eggs  (fig.  7)  are  elliptical,  about  0-08  mm.  in  long  diameter. 
They  are  thus  larger  than  those  of  most  segmented  Cestodes,  l3ut 
considerably  smaller  than  those  of  G.  ruyosa,  which  measure 
0*1  mm.,  and  are  the  largest  Cestode  eggs  the  dimensions  of  which 
have  been  recorded.  In  G.  nigrosetosa  a  circular  area  of  the  shell 
at  one  pole  is  much  thinner  than  the  rest,  and  is  probably 
differentiated  as  an  operculum.  At  the  opposite  pole  there  is 
occasionally,  though  by  no  means  frequently,  a  filament,  usually 
very  short.  Coalescence  of  two  eggs  by  fusion  between  their 
shells  is  by  no  means  rare.  No  hooked  embryos  are  present  in 
either  of  the  specimens. 

In  my  specimen  of  G.  rugosa  the  eggs  differ  from  those  of  G. 
nigrosetosa  not  only  in  size,  but  in  the  appearance  of  the  shell, 
which  does  not  appear  to  be  provided  with  an  operculum. 
Hexacanth  embryos  are  abundant,  and  are  remarkable  for  their 
large  size  and  the  comparatively  colossal  dimensions  of  their 
hooks.  These  are  disposed  in  pairs  like  those  of  hexacanth 
embryos  in  general.  In  G.  uima  Lonnberg  failed  to  find  any 
hooks,  and  this  circumstance,  when  we  take  into  account  the  fact 
that  the  observer  in  question  had  abundance  of  material  at  his 
disposal,  would  seem  to  jDoint  to  an  important  difference  between 
the  two  species  with  regard  to  the  structure  of  the  embryo. 


LITEEATUEE. 

1. — Benham,  W.  B.  —  "  A  Treatise  on  Zoology,"  edited  by  E.  Ray  Lankester. 

Part  iv.      "  The  Platyhelmia,  Mesozoa  and  Nemertini  "  (1901). 
2. — Braun,  M. —  "  Gyrocotyle,  Amphiptyches  u.  Verwandte."       Central- 

blatt  f.  Bakteriologie  u.  Parasitenkunde.     vi.  Bd.  (1889). 
3. "Vermes"    of    Bronn's    "  Klassen    u.     Ordnungen    des 

Thierreiehs." 
4. — LoENXBEKG,  EixAR — "  Anatomlsche  Studien  liber  Skandinavische  Ces- 

toden."      Kongl.    Svenska    Vetenskaps-Akademiens    Handlingar, 

Bd.  24. 
5.  —  MoNTicELLi,  F.  8.— "Sul  sistema  nervosa  dell'  Amphiptyches  urna." 

Zool.  Anz.,  1889. 
6, "  Appunti  sui  Cestodaria."     Atti  della  E.  Accad.  dell  Scienze 

lis.  e  nat.  di  Napoli,  1892. 


54  ON  A  GRYOCOTYLE  FEOM  CHIM^ERA. 

7. — Spencer,  W.    BALD^YIX— "  The    Anatomy   of    Amphiptyches    urna." 

Trans.  E.S.  Vict.,  1889. 
8. — Wagener,  E,   G. — -"  Ueber  einen  neuen  in  der  Chimcera  monstrosa 

gefundenen   Eingeweidewurm    Amphiptyches    urna,    Grube    und 

Wagener."     Miiller's  Archiv  f.  Anat.  Physiol.  Jahrg.  1852. 
9.— Will,  H. — "Anatomie  von  Caryophyllseus  mutabilis  Rud."      Zeitschr. 

wiss.  Zool.,  Ivi.  Bd.  (1893). 


EXPLANATION  OF  PLATE  VII. 

Fig.  1. — GyrocotyU  nigrosetosa,  natural  size. 

Fig.  2. — Gyrocotyle  nigrosetosa,  anterior  end,  ventral  aspect,  magnified; 
te.,  lobes  of  testis;  -s.,  sucker;  «..,  uterus;  vit.,  lobes  of  vitelline  glands;  J, 
opening  of  ejaculatory  duct;    $  ,  opening  of  vagina. 

Fig.  3. — Gyrocotyle  rugosa,  dorsal  surface  of  the  anterior  region  (extremity 
with  sucker  cut  off);  c,  cirrus;  va.,  opening  of  vagina. 

jpig.  4, — The  same  specimen,  ventral  surface. 

Fig.  5. — Outline  of  a  transverse  section  through  the  specimen  of  Gyrocotyle 
rugosa  represented  in  figs.  3  and  4,  in  the  region  of  the  cirrus;  v. ,  ventral 
surface;  d.,  dorsal  surface;  e.d.,  ejaculatory  duct  in  terminal  part  of  cirrus; 
c,  cirrus. 

Fig,  6, — Spinules  of  Gyrocotyle  nigrosetosa. 

Fig.  7. — Egg  of  Gyrocotyle  nigrosetosa. 


50 


N0TE8  FROM  THE  BOTAN^IC  GARDENS,  SYDNEY. 

No.  8. 
By  J.  H.  Maidex  and  E.  Betche. 
STERCULIACE^. 
KekaudrExVIa  Hillii,  F.v.M.  {Seringea  Hillii,  F.v.M,,  m  Census.) 

Jennings  (J.  L.  Boorman;  October,  1901).  A  new  locality  for 
a  rare  plant.  According  to  the  collector  it  is  a  shrub  about  6  to 
8  feet  high,  with  large  blue  flowers  (not  purplish  as  described  in 
Moore  and  Betche's  'Handbook  of  the  Flora  of  New  South  Wales,' 
from  old  specimens),  growing  on  dry  ridges  amongst  boulders, 
about  half  a  mile  from  the  Queensland  border. 

The  gQiiu^  Kei-audrenia  \^  confined  to  Australia  with  the  excep- 
tion of  a  single  species  which  occurs  in  Madagascar.  This 
Madagascar  plant  is,  according  to  Mueller's  'Census,'  identical 
with  K.  Hillii.  It  seems  extraordinary  that  a  rather  local 
Australian  plant  should  recur  alone  in  Madagascar,  but  we  have 
no  means  of  verifying  Mueller's  statement.  We  observe  (Proc. 
Linn.  Soc.  xx.,  104)  that  Madagascar  also  possesses  one  species 
of  RuUngia  (also  a  Sterculiaceous  genus),  all  the  other  Rulingias 
being  natives  of  Australia. 

RUTACE.a:. 

AsTEROLASiA  CORREIFOLIA,  Benth.,  var.   MOLLIS  (Syn.  A.  7noUis, 

Benth.) 

Warrumbungle  Ranges  (W.  Forsyth;  October,  1899  and  1901). 

The  specimens  collected  by  Mr.  Forsyth  in  the  Warrumbungle 
Ranges  diifer  from  Bentham's  description  of  A.  mollis,  made  from 
specimens  collected  by  Eraser  more  than  half  a  century  previously 
in  the  same  locality  (called  Arbuthnot  Range  at  that  time),  in  the 
lobed  ovarium  and  in  the  long  pedicels.     The  discrepancy  in  the 


56  NOTES    FROM    THE    BOTANIC    GARDENS,   NO.  VIII., 

length  of  pedicels  is  easily  explained  b}^  the  fact  that  Bentham 
had  seen  no  fruits,  and  the  pedicels  in  this  species  are  very 
variable  in  length,  and  lengthen  out  with  the  maturing  fruit.  In 
some  of  last  year's  fruiting-specimens  from  the  Warrumbungle 
Ranges,  the  pedicels  are  nearly  1  inch  long,  while  some  specimens, 
collected  two  j^ears  previously  in  the  same  locality,  have  the 
young  flowers  on  pedicels  often  under  J  of  an  inch  long.  The 
discrepancy  in  the  shape  of  the  ovarium  (Bentham  describes 
it  as  "rounded  at  the  top")  cannot  be  explained  by  simple 
variation,  and  it  is  difficult  to  believe  that  a  trained  observer  like 
Bentham  should  have  made  such  a  mistake.  Perhaps  Bentham 
had  such  young  flowers  under  observation  that  the  lobes  were 
scarcely  formed.  The  ripe  carpels  of  Mr.  Forsyth's  specimens 
are  beaked  like  those  of  A.  correifolia,  which  proves  that  the 
young  carpels  in  the  ovarium  are  also  lobed. 

Unfortunately  the  type  specimens  are  not  in  Australia,  so  that 
we  can  only  surmise  that  Fraser's  and  Forsyth's  specimens  are 
identical,  Avithout  being  absolutely  certain. 

Mueller  unites  A.  coi'i'ei folia,  Benth.,  and  A.  MueUeri,  Benth., 
under  the  name  of  Eriostemon  correifolius.  We  propose  now  to 
go  a  step  further  and  include  A.  mollis  in  the  variations  of  A. 
correifolia.  Bentham  separates  A.  mollis  from  A.  correifolia  and 
Muelleri  on  account  of  the  lobed  stigma  and  the  rounded  ovarium; 
but  we  find  both  very  unreliable  characters  in  this  section  of  the 
genus.  In  A.  mollis  the  stigma  is  distinctly  lobed,  while  scarcely 
lobed  in  A.  correifolia;  Ijut  in  A.  Muelleri  the  stigma  seems  so 
variable  that  we  can  onl}'  describe  it  as  "  more  or  less  distinctly, 
lobed  or  almost  entire."  The  true  A.  correifolia  is  so  variable  in 
shape  and  size  of  leaves,  indumentum,  and  length  of  pedicels  that 
both  A.  Muelleri  and  A.  mollis  cannot  be  maintained  as  separate 
species  if  the  character  of  stigma  and  ovarium  are  proved  to  be 
variable. 

The  chief  characteristics  of  the  three  forms  of  A.  correifolia 
are  : — 

Normal    Form. — Leaves    glabrous    above.       Floivers    white. 
Stigma  entire,  or  nearly  so. 


BY    J.    H.    MAIDEN    AND    E.    BETCHE.  57 

Var.  MuELLERi. — Leaves  tomeiitose  on  both  sides.  Floivers 
yellow.     Stigma  more  or  less  lobed,  or  nearly  entire. 

Var.  MOLLIS. — Leaves  tomentose  on  both  sides.  FloicerswhitG. 
Stigma  distinctly  lobed. 

Phebalium  Ralstoni,  Benth. 

Warrumbungle  Ranges  (W.  Forsyth;  October,  1901). 

Previously  recorded  only  from  the  Southern  Dividing  Range. 
The  Warrumbungle  Ranges  specimens  are  unfortunately  only  in 
fruit,  but  they  resemble  so  closely  Dr.  Leichhardt's  type-specimens 
from  Castle  Rock  Mountain  (a  locality  whose  situation  we  do  not 
know)  that  we  do  not  hesitate  to  refer  them  to  this  species.  The}'- 
differ  from  Bentham's  description  in  the  'Flora  Australiensis '  in 
the  prominent  beak  of  the  ripe  carpels  and  in  the  stellate 
tomentum  on  the  young  branches,  which  occurs  sparingly  also  on 
the  underside  or  the  young  lea\'e8. 

RHAMNACE^. 

Stenanthemum  Scortechinii,  F.v.M.   {Crifptandra  Scortechinii, 
F.V.M.,  in  Census.) 

Jennings  (J.  L.  Boorman ;  October,  1901V  A  new  locality 
for  a  rare  plant.  This  shrub  grovv^s  5  to  6  feet  high  in  rich 
alluvial  soil  on  the  side  of  a  creek.  The  Jennings  specimens 
are  entirely  identical  with  Mueller's  type-specimens  from  the 
Severn  River.  Other  Xew  South  Wales  localities,  representing 
a  rather  narrow-leaved  form,  are  : — Head  of  Maeleay  River,  open 
sandy  ground  (Carron;  between  1860  and  1870);  Braidwood(W. 
Bauerlen;  1888);  Bermagui  (W.  Bauerlen,  1889);  Apsley  Falls, 
Walcha  district  (E.  Betche;  December,  1898). 

SAPINDACE^. 
D0D0NJ5A  peduncularis,  Lindl.,  var.  hirsuta,  var.nov. 

Jennings  (J.  L.  Boorman;  October,  1901).  A  very  handsome 
bushy  shrub  attaining  about  2  feet  in  height,  with  short  broad 
cuneate,  rather  crowded  leaves,  deeply  3-toothed  at  the  truncate 
end;  growing  in  rich  alluvial  soil  on  the  banks  of  a  creek.     It 


58  NOTES    FROM    THE    130TAXIC    GARDENS,    NO.  VIII., 

differs  from  the  type  chiefly  in  the  absence  of  all  viscidity;  and 
in  the  short  spreading  hairs,  which  densely  cover  the  whole  plant, 
except  the  old  branches.  The  capsules  attain  fully  J  of  an  inch 
in  breadth.  Male  flowers  not  seen.  Though  Bentham  describes 
the  species  as  glabrous,  we  find  the  tendenc}'-  to  hairiness  in 
several  specimens,  amongst  others  in  Mitchell's  specimens  collected 
at  Mt.  Pluto  near  the  Warrego  River,  and  described  by  Lindle}' 
in  "Mitchell's  'Tropical  Australia,'  p.  342,  as  E.  j)uhescenii.  Ben- 
tham's  remark  in  the  'Flora  A^ustraliensis'  in  regard  to  D.piihescens, 
"  the  supposed  pubescence  is  apparently  a  mistake,"  shows  that  he 
cannot  have  seen  the  type  specimens,  for  Major  Mitchell's  speci- 
mens are  distinctly  hairy,  though  viscid,  sparse-leaved,  and  %vith 
all  the  characteristics  of  a  plant  grown  in  dry  country,  very 
different  in  appearance  from  the  luxuriant  hirsute  form  found  by 
Mr.  Boorman  at  Jennings. 

LEGUMINOS^. 

OXYLOBIUM    PULTEN^A,   DC. 

Hastings  River  (W.  Best,  communicated  by  W.  Forsyth  ; 
August,  1901).     Most  northern  locality  recorded. 

PULTEN.EA    PLUMOSA,   Sieb. 

Bombala  (J.  H.  Maiden;  November,  1901). 

A  rather  rare  PultencEa,  confined  to  New  South  Wales. 
Bentham  gives  its  habitat  as  "Port  Jackson  to  the  Blue  Mountains  " 
Since  then  man}^  additional  localities  have  been  discovered  within 
the  borders  of  New  South  Wales.  The  most  northern  locality . 
known  to  us  is  Apsley  Falls,  in  the  Walcha  district,  and  its 
southern  limit  has  now  been  extended  to  Bombala. 

PSORALEA    ERIANTIIA,   Benth. 

Paldrumatta  Bore,  on  sandy  ridges  (P.  Corbett;  October,  1901). 

A  rare  plant  in  New  South  Wales,  though  with  extensive  range 
in  Central  Australia.  A  sparingly  branched  erect  shrub  about 
3  to  4  feet  high,  densely  covered  with  a  soft  tomentum  of  white 
hairs,  especially  long  on  the  calyces.     The  flowers  are  white,  with 


BY    .T.    H.     MAIDEN    AND    E.    BETCHE.  59^ 

a  dark  almost  black  keel,  the  standard  and  wings  often  (not 
always)  striped  with  blue.  The  Paldrumatta  Bore  specimens  are 
much  more  hairy  than  Mitchell's  prostrate  type-specimens  from 
the  Balonne  River,  Queensland. 

Acacia  rhigiophylla.  F.v.M. 

AVest  Wyalong  (R.  H.  Cambage;   September,  1900). 

New  for  New  South  Wales.  Previously  only  recorded  from  the 
desert  on  the  Lower  Murray  River  towards  Mt.  Barker  Range 
in  South  Australia. 

Acacia  harpophylla,  F.v.M, 

Marsden  (R.  H.  Cambage;  September,  1900). 

The  trees  attain  a  height  of  20  to  30  feet,  according  to  Mr. 
Cambage,  and  cover  about  5  acres,  3|  miles  west  of  Marsden. 
The  most  southern  locality  hitherto  recorded  is  Coolabah. 

HALORAGEa:. 
Myriophyllum  pedunculatum,  Hook.  f. 

Mt.  Wilson  (Jesse  Gregson  ;  December,  1901,  in  flower; 
February,  1902,  in  fruit). 

A  southern  species,  recorded  in  these  Proceedings  in  April, 
1899,  as  "new  for  New  South  AVales  "  from  Mt.  Kosicusko,  but 
never  found,  or  at  all  events  recorded,  north  of  this  station.  The 
floating  specimens  from  Mt.  Wilson  have  long  and  slender  stems 
with  leaves  above  1  inch  long;  specimens  growing  in  mud  are 
shorter,  and  resemble  much  more  the  Tasmanian  specimen  figured 
by  J.  D.  Hooker  in  his  'Flora  of  Tasmania'  (Vol.  i.  t.  23  b.).  The 
male  flowers  are  very  shortly  pedunculate  or  sessile;  the  carpels 
are  only  sparingly  tuberculate. 

MYRTACEiE. 

Baeckea  diffusa,  Sieb. 

Port   Macquarie   (G.    R.   Brown  ;   February,  1897);   Hastings 
River  (W.  Best,  communicated  b}^  W.  Forsyth;  August,  1901). 
Most  northern  locality  recorded. 


60  NOTES    FROM    THE    BOTANIC    GARDENS,    NO.  YIIL, 

UMBELLIFER^. 
ACTINOTLS    FORSYTHII,   Sp.llOV. 

Blacklieath,  Blue  Mountains  (W.  Forsytli;  February,  1902). 

An  ascending  much-branched  annual  or  biennial  attaining 
about  1  foot  in  height,  with  somewhat  striate,  terete,  rather  wir}^ 
stems,  sparingly  covered  with  appressed  hairs.  Leaves  ^  to  1 
inch  long,  glabrous  on  both  sides  or  nearly  so,  generally  twice 
ternately  divided,  the  segments  often  again  lobed  or  toothed,  the 
ultimate  lobes  tending  to  linear-lanceolate,  acute,  with  recurved 
or  thickened  margins,  the  radical  ones  and  lower  stem-leaves  on 
petioles  often  above  1  inch  long,  the  upper  ones  gradually  more 
sessile  and  less  divided.  Umbels  on  long  slender  peduncles. 
Involucre  radiating  to  I  inch  in  diameter  in  the  largest  umbels 
seen;  the  bracts  lanceolate,  acute,  about  12  in  number,  densely 
hairy  inside  with  long  white  silky  hairs,  nearly  glabrous  outside, 
often  brownish  and  conspicuously  1-  or  3-veined.  Flowers 
numerous,  on  slender  glabrous  pedicels,  a  few  of  the  outer  ones 
often  male,  without  any  ovarium;  rarely  the  males  are  in  several 
rows  at  the  circumference.  Calyx  limb  truncate  or  sinuate-lobed, 
glabrous.  Petals  small,  spathulate,  on  slender  claws.  Stamens 
on  slender  filaments  with  violet  anthers.  Fruit  densel}^  ciliate 
on  the  edges  with  long  white  hairs,  the  flat  sides  glabrous  and 
dark  brown  when  ripe,  with  a  prominent  rib  along  the  centre. 

In  affinities  it  is  more  closely  allied  to  the  beautiful  West 
Australian  A.  leucocephalus  than  to  an}^  of  the  East  Australian 
species  of  Actinotus,  though  its  general  ajDpearance  is  more  like  a 
large  flowered  form  of  our  modest  A.  minor.  It  is  the  only  East 
Australian  Actinotus  with  petals,  and  has  also  a  truncate  calyx 
in  common  with  A.  leucocephalus,  but  differs  from  this  species 
chiefly  in  the  fruit,  apart  from  the  more  striking  but  less  essential 
difference  of  size,  length  of  involucral  bracts,  and  indumentum. 
Like  A.  Gihhonsii  it  has  a  peculiar  fruit  with  densely  ciliate  edges 
and  glabrous  carinate  sides,  but  nothing  else  in  common  with  this 
species. 


BY    J.    H.    MAIDEX    AXD    E.    BETCHE.  61 

COMPOSITE. 

Olearia  lepidophylla,  Benth. 

Emmaville  (J.  L.  Boorman;  October,  1901). 

The  most  northern  locality  recorded.  A  common  plant  in  the 
southern  colonies  and  southern  districts  of  New  South  \yales; 
the  present  note  brings  its  range  up  to  the  Queensland  border. 

Olearia  adenophora,  F.v.M. 

At  the  Gap,  near  Cobborah  (W.  Forsyth;  October,  1901). 
The  most  northern   locality  for  a  plant   rare  in  New  South 
Wales,  though  more  common  in  Victoria. 

Cassinia  leptocephala,  F.v.M. 

AVarrumbungie  Ranges  (W.  Forsyth;  October,  1901). 

The  locality  "Port  Jackson"  given  in  Bentham's  'Flora  Aus- 
traliensis '  seems  to  be  a  mistake.  Mueller  gives  the  very  vague 
locality  "Nova  Austro-Cambria "  in  his  original  description 
(Fragm.  iii ,  138),  though  he  gives  "Port  Jackson"  on  his 
herbarium  label.  This  herbarium  label  probably  misled  Bentham, 
and  the  mistake  was  copied  by  Dr.  Woolls  in  his  '  Plants  of  the 
NeighlDOurhood  of  Sydney,'  and  in  Moore  and  Betche's  'Handbook 
of  the  Flora  of  New  South  Wales.'  We  have  never  heard  of  this 
species  having  been  collected  in  the  Port  Jackson  district,  and  as 
this  district  is  naturally  the  best  explored  one  in  the  State,  it  is 
scarcely  probable  that  such  a  prominent  plant  should  have  escaped 
notice;  besides  the  whole  appearance  of  the  plant  is  that  of  an 
inhabitant  of  dry  mountain  ranges. 

Other  authenticated  localities  of  the  species  in  the  Herbarium 
are  : — Moonan  Brook,  Upper  Hunter;  and  Upper  Hastings  River 
(ascent  to  tableland;  J.  H.  Maiden). 

EPACRIDEiE. 

Leucopogon  exolasius,  F.v.M. 

Woronora  River  (E.  Cheel;  October,  1901). 
The  only  locality  recorded   is  "  near  Camden,"  where  it  was 
collected  by  Dr.  Leichhardt  in  1843.     It  is  perhaps  too  closely 


•62  NOTES    FROM    THE    BOTANIC    GARDENS,    NO.  VIII.  , 

ullied  to  L.  setf'ger,  R.Br.,  to  be  separated  from  it  as  a  species, 
but  we  would  like  to  further  study  the  variations  of  that  species 
before  proposing  a  change. 

MoNOTOCA  LEDiFOLiA,  A.  Cunn. 

Woronora  River  (E.  Betche  ;  Januar\^,  1894:  E.  Oheel ; 
October,  1901). 

Previously  only  recorded  from  the  Blue  Mountains,  but 
apparently  not  uncommon  at  the  head  waters  of  the  coastal  rivers 
in  the  Port  Jackson  sandstone  district. 

SOLANE.S:. 
Anthocbrcis  scabrella,  Benth. 

Wallangarra  (J.  L.  Boorman;  October,  1901), 

A  rare  plant,  previously  onh^  recorded  from  a  single  locality, 
viz.,  "Nepean  River  in  the  Blue  Mountains,"  In  the  Wallangarra 
localit}^  it  is  a  much  spreading  and  intricately  branched  shrub  not 
a,bove  2  feet  high,  growing  on  sparsely  timbered  dry  rocky  hill 
sides.  The  peduncles  are  shorter  than  in  the  type  and  often 
apparently  axillary,  though  in  reality  terminal  on  very  short 
branchlets  in  the  axils  of  the  leaves.  In  the  Nepean  River 
locality  (J.  H.  ]\[aiden  has  collected  it  at  Erskine  Creek,  Nepean 
River,  10  miles  from  Penrith)  the  shrub  is  about  3  to  4  feet  high, 
with  considerably  longer  filiform  peduncles. 

VERBENACEJE. 
Gmelina  Leichhardtji,  F.v.  M. 

Botanic  Gardens,  Sydney  (cultivated,  April,  1902). 

This  is  the  first  time  that  we  have  noticed  ripe  fruits  on  the 
tree  in  the  Gardens;  and  as  the  fruits  are  not  described  in  the 
^ Flora  Australiensis'  we  give  a  short  description  of  them: — Fruits 
of  a  dull  mauve,  almost  blue  colour,  somewhat  depressed-globular, 
nearly  1  inch  in  diameter;  always  provided  with  the  persistent 
flattened  out  and  enlarged  calyx. 


BY    J.    H.    MAIDEN    AND    E.    BETCHE.  63 

LAURINEJE. 

LiTSBA  RETICULATA,  Beufcli.  [Tefyrantheva  reticulata^  Meissn.). 

Wyong  (J.  L.  Boorman;  February,  1900). 

Recorded  in  the  'Flora  Australiensis'  only  from  Queensland. 

It  has  been  long  known  to  be  not  uncommon  in  the  northern 
brush-forests  of  New  South  Wales  (see  a  note  by  one  of  us  in 
Agric.  Gazette  of  N.S.  Wales,  Vol.  v.  [1894],  p.  826),  and  now 
its  range  has  been  traced  as  far  south  as  Wyong. 

PR0TEACE5:. 
Banksia  ericifolia,  Linn.  f. 

Byron  Bay  (A.  H.  Hammond;  April,  1902). 

Most  northern  locality  recorded  for  New  South  Wales,  the 
Hastings  River  being  hitherto  the  most  northerly  record  for  this 
State.  It  is,  however,  admitted  by  F.  M.  Bailey  in  his  'Queensland 
Flora,'  with  the  note  :  "  Recorded  for  Queensland  by  F.v.M. 
without  locality." 

THYMELE^. 

PixMELEA  LiGUSTRiNA,  Labill,  var.   GLABRA,  vai'.nov. 

Kiandra  District  (fruiting  specimens ;  E.  Betche,  February, 
1897),  Kiandra  District  (flowering  specimens ;  W.  Forsyth, 
December,  1901). 

A  small  bushy  shrub  about  18  inches  high,  with  crowded  erect 
oblong  leaves  from  5  to  8  lines  long.  Flower-heads  nodding, 
with  6  to  8  large  involucral  bracts  glabrous  inside,  and  of  a 
purplish  cast  when  in  fruit.  Receptacle  glabrous.  Persistent 
base  of  the  calyx  glabrous  from  the  first,  on  short  glabrous  stalks; 
calyx-tube  glabrous,  the  lobes  generally  sparingly  hairy. 

Bentham's  type-specimens  of  var.  macrosteyiay  collected  by 
Waterhouse  in  sandy  scrub  in  Kangaroo  Island,  differ  from  the 
Kiandra  specimens  only  in  the  hairy  receptacle,  and  possibly  in 
the  size  of  the  shrub,,  which  is  not  stated  by  Waterhouse. 
Perhaps  our  new  variety  should  be  a  distinct  species,  but  we  are 
not  sure  whether  the  absence  of  hairs  on  the  involucral  bracts 


64  NOTES    FROM    THE    BOTAXIC    GARDENS,    NO.  VIII., 

and  receptacle  is  constant.  The  alpine  forms  of  var.  hypei-icina 
have  frequently  coloured  bracts  and  a  tendency  to  nodding  flower, 
heads,  and  approach  the  new  Kiandra  form  closely  in  habit. 

URTICACEiE. 
Ficus  Hbnneana,  Miq. 

National  Park,  Port  Hacking  (M.  Bell:  February,  1900:  J.  L.. 
Boorman;  January,  1902). 

New  for  New  South  Wales.  It  seems  very  strange  that  a 
Ficus  described  from  specimens  collected  on  the  barren  Booby 
Island  in  Torres  Strait  (about  10°  S.  lat.j,  and  not  recorded 
further  south  than  Rockingham  Bay,  Queensland,  should  recur 
again  in  Port  Hacking,  near  Sydney;  but  we  have  no  doubt  about 
the  correctness  of  the  identification.  The  Port  Hacking  speci- 
mens differ  from  the  small  specimen  of  Henne's  type  from  Booby 
Island  only  in  the  somewhat  smaller  size  of  the  leaves. 

According  to  notes  by  Messrs.  Bell  and  Boorman,  the  largest 
of  the  few  trees,  growing  near  "  Wentworth's  Hut "  in  the 
National  Park,  attains  a  trunk  diameter  of  about  5  feet,  with 
gnarled  spreading  branches,  and  is  probably  considerably  over 
100  years  old.  The  leaves  are  deciduous,  but  the  trees  remain 
bare  only  a  few  weeks  in  the  year  before  the  appearance  of  the 
new  leaves.  The  ripe  receptacles  are  reddish,  spotted  with  white.. 
Amongst  the  New  South  Wales  species  of  Ficus,  it  is  most  nearly 
allied  to  F.  Cnnnhighamii,  Miq.,  with  which  it  has  also  the 
deciduous  leaves  in  common,  but  from  which  it  is  easil}^  dis- 
tinguished by  the  larger  and  pedunculate  receptacles,  and  by  the 

shape  of  the  leaves. 

CYPERACEiE. 

CvPERUS  LAEViGATUS,  Linn. 

Manning  River  (E.  Cheel:  December,  1899). 

New  for  New  South  AVales.  An  almost  cosmopolitan  species 
found  chiefly  in  maritime  districts,  and  recorded  in  Mueller's 
'Second  Census'  only  from  West  Australia,  although  we  have- 
South  Australian  specimens  from  Port  Lincoln  and  Mt.  Lynd- 
liurst. 


BY    J.    H.    MAIDEN    AND    E.    BETCHE.  65 

The  Manning  River  specimens  belong  to  the  form  with  very 
few  (1  to  3)  pale  coloured  spikelets,  and  closely  resemble  some 
European  specimens;  they  are  less  rigid  than  all  West  Australian 
specimens  we  have  seen,  with  nuts  sometimes  nearly  as  long  as 
the  glumes. 

XJncinia  tenella,  R.Br. 

Katoomba,  Blue  Mountains  (A.  A.  Hamilton;  December,  1901); 
Mt.  Wilson  (Jesse  Gregson;  April,  1902). 

The  Australian  species  of  Uncinia  are  all  southern  plants, 
abounding  in  the  alpine  districts  of  Victoria  and  Tasmania. 

Mt.  Wilson  is  the  most  northern  localit}^  recorded  for  the 
genus  in  Australia,  with  the  exception  of  Uncinia  dehilior,  F.v.M., 
from  summit  of  Mt.  Gower,  Lord  Howe  Island. 


Xote. — It  is  to  be  understood  that  the  localities  given  in  this 
series  of  papers  are  in  New  South  Wales,  unless  otherwise  stated. 


66 


FURTHER  REMARKS  UPON  THE  MECHANISM  OF 
AGGLUTINATION. 

By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

Society. 

Two  years  ago,  at  the  April  meeting  of  this  Society,  I  read  a 
paper*  upon  the  mechanism  of  agglutination,  in  which  I  reviewed 
the  theories  regarding  the  phenomenon  which  had  been  published 
up  to  that  time.  I  also  gave,  as  the  result  of  my  work  and 
experience,  the  hypothesis  that  the  agglutination  of  bacteria  is 
due  to  the  formation  of  a  chemical  precipitate  upon  the  bacteria 
through  the  interaction  of  the  agglutinating  substance  of  the 
active  serum  with  the  agglutinable  substance  secreted  by  the 
bacteria.  The  chemical  precipitate  upon  the  bacteria  is  flocculated 
by  the  salts  of  the  serum  or  of  the  bouillon,  and  the  bacteria  are 
carried  with  the  precipitate  mechanically.  My  hypothesis  differed 
from  that  of  Nicolle  and  of  Paltauf  by  taking  into  account  the 
function  of  the  salts  and  the  mechanical  part  played  by  the 
organisms. 

Since  the  publication  of  my  paper  two  authors  have  published 
papers  detailing  experiments  which  show  the  function  of  salts  in 
promoting  agglutination,  but  they  miss  the  point  that  it  is  the 
precijntate  on  the  bacteria  that  is  agglutinated,  clumped,  or 
flocculated  by  the  salts. 

I  was  the  first  to  emphasise  the  necessary  role  of  the  salts  in 
explaining  the  mechanism  of  the  process;  and  although  Bordet 

*  Proc.  Linn.  Soc.  N.S.W.  xxv.,  75.  Referat  in  Cent.  f.  Bakt.  1  Abt.  xxx.. 
209. 


BY    R.    GREIG    SMITH.  67 

had  published  an  experiment  showing  how  common  salt  was 
necessary  in  causing  an  agglutination,  he  failed  to  accentuate  the 
part  played  by  salts.  Although  most  writers  upon  the  subject 
agree  in  considering  agglutination  as  a  chemical  phenomenon, 
they  do  not  understand,  or  at  least  fail  to  indicate,  the  steps  in 
the  process.  Were  it  otherwise,  the  recently  published  papers 
upon  the  role  of  salts  in  agglutination  would  probably  not  have 
been  written. 

The  first  of  these  papers  is  by  Joos,'^'  who,  after  enumerating 
his  experiments,  concluded  that  — 

1.  No  agglutination  takes  place  when  the  agglutinating  sub- 

stance acts  upon  the  agglutinable  substance  in  the  absence 
of  common  salt. 

2.  Agglutination  always  occurs  when  the  three  substances  are 

together,  viz.,  the  agglutinating  substance,  the  agglutin- 
able substance  and  salt. 

3.  In  the  absence  of  salt  the  agglutinating  substance  is  quickly 

combined  with  the  agglutinable  substance  of  the  bacteria. 
This  combination  alters  their  vitality  in  no  way. 

4.  There  is  a  close  relation  between  the  relative  amounts  of 

substances  which  together  bring  about  the  phenomenon  of 
agglutination  and  of  the  obtained  quantity  of  agglutinated 
substance. 

5.  Salt  plays  an  active  role  in  the  phenomenon. 

6.  Salt  enters  into  the  combination  of  the  agglutinating-  and 

agglutinable  substance. 

7.  Agglutination  can  occur  in  a  salt-free  medium  if  the  bacteria 

contain  salt. 

8.  The  "physical  theory"  of  Bordet  is  untenable. 

These  conclusions  of  Joos  simply  bear  out  the  contention  that 
agglutination  depends  upon  the  flocculating  action  of  a  particular 
salt,  viz.,  common  salt,  for  the  reason  that  this  was  the  only  salt 
with  which  he  experimented. 

"  Joos,  Zeit.  f.  Hygiene,  xxxvi.,  422. 


68  MECHANISM    OF    AGGLUTINATIOX, 

A  year  afterwards  Friedberger"^  published  a  paper  in  which  he 
showed  that  other  salts  were  as  useful  as  common  salt  in  causing 
agglutination.     He  concluded  that : — 

1.  Agglutination  does  not  occur  in  the  complete   absence  of 

crystalline  substances. 

2.  Of  these  substances  the  inorganic  salts  are  the  most  active 

and  they  differ  among  themselves  in  their  activity. 

3.  The  rapidity  of  the  onset  of  the  agglutination  of  dialysed 

cultures  depends  upon  the  salt  content  of  the  bacterial 
suspension. 

4.  The  rapidity  of  the  onset  of  the  agglutination  of  a  bacterial 

emulsion  depends  upon  its  salt  content. 

5.  The  action  of  salts  in  agglutination  is  not  chemical. 

Thus  the  function  of  salts  in  general  in  causing  agglutination 
is  emphasised.  Exception  must,  however,  be  taken  to  the  5th 
item,  which  is  rather  startling.  Friedberger  does  not  attempt  to 
explain  the  mechanism  of  the  process,  and  that  he  cannot  under- 
stand the  part  played  by  the  salts  is  made  evident  by  his  sweeping 
assertion.  By  chemical,  he  means  that  the  salts  do  not  chemically 
combine  with  the  albumen  of  the  cell,  an  idea  which  has  never 
been  advanced  with  an}^  positiveness.  Bordet,  perhaps  by  con- 
sidering that  Kraus'  precipitate  did  not  play  a  part  in  agglutina- 
tion, suggested  the  idea  of  an  enzyme  altering  an  albuminoid 
constituent  of  the  microbic  cell;  but  beyond  this  inference  there 
is  nothing  to  justify  the  assumption  that  coagulation  of  the  cell 
albuminoids  may  occur.  That  the  bacteria  simply  become  immo- 
bilised is  well  known.  Were  a  portion  of  their  albuminoid  matter 
coagulated,  probably  something  more  than  immobilisation  would 
ensue.  Joos  confirmed  the  fact  that  agglutination  does  not  alter 
the  vitality  of  the  bacteria. 

In  a  later  paper  Joos,t  after  pointing  out  that  Fried berger's 
researches  strengthened  his  own  former  conclusions,  finally  pro- 

*  Friedberger,  Cent.  f.  Bakt.  xxx.,  330. 
t  Joos,  Cent.  f.  Bakt.  1  Abt.  xxx.,  853. 


BY    R.    GREIG    SMITH.  69 

pounded  the  following  as  an  explanation  of  the  mechanism : — 

The  phenomenon  of  agglutination  is  made  up  of  two  quite 

distinct  phases.     In  the  first  the  specific  substances  combine 

with  the  salts.     The  second  is  that  in  which  the  microbes 

modified  by  this  combination  unite  into  floccules. 

The  first  phase  depends  upon  a  chemical  combination 
analogous  to  the  formation  of  double  salts,  or  of  certain 
additive  compounds.  It  does  not  depend  upon  a  molecular 
attraction  [Juxtapposition]  as  in  dyeing  or  porosity  pheno- 
mena. 

The  second  phase  is  entirely  similar  to  a  chemical  precipi- 
tate, and   does  not  depend  exclusively  upon  a  disturbance 
of    the    molecular    equilibrium    of    the    mixture,   which    is 
produced  by  the  solution  of  the  salt. 
These  statements  are  vague,  and  we  do  not  gather  from  them 
whether   the  individual    specific    substances   (agglutinating  and 
agglutinable)  unite  with  the  salts  and  then  combine,  or  combine 
and  then  unite — an  important  point.     Nor  do  we  see  how  the 
microbes  are  modified  by  this  combination  of  salt  and  specific 
substance.     Nor  is  the  relation  between  the  microbes  and  the 
combined  specific  substance  defined.     Although  this  is  an  attempt 
to  explain  the  mechanism  by  bringing  the  phenomenon  into  line 
with  the  formation  of  chemical  precipitates,  the  mechanism  itself 
is  still  unexplained.     That  the  phenomenon  is  different  from  that 
which  occurs  in  staining  fibres,  etc.,  may  be  open  to  doubt,  for 
Knecht  (Ber.  1902,  35,  1022)  has   shown   that  dyeing   depends 
upon  the  formation  of  lakes  between  the  colour  and  certain  con- 
stituents of  wool  and  silk. 

On  the  whole  there  is  a  tendency  in  these  papers  to  consider 
the  phenomenon  in  a  manner  that  is  too  narrow  and  too  indefinite 
to  be  scientific.  That  agglutination  is  similar  to  coagulation  and 
precipitation  appears  to  be  admitted  by  most  writers,  e.g.,  Bordet, 
Duclaux,  Nicolle,  Paltauf,  Kohler  and  Joos;  but  the  mechanism 
of  coagulation  does  not  appear  to  be  understood,  or  Joos  would 
probably  not  have  emphasised  the  role  of  common  salt  in  agglu- 
tination, and  Bordet  would  have  mentioned  the  function  of  calcium 


/U  MECHANISM    OF    AGGLUTINATION, 

salts  in  the  coagulation  of  casein,  to  which  he  likened  agglutina- 
tion. 

I  have  already^  explained  the  mechanism  of  flocculation,  coagu- 
lation or  precipitation  by  the  hydrate  theory  of  solution,  and 
although  I  understand  that  it  has  also  been  explained  by  the 
ionisation  theor}',  yet  I  was,  and  still  am,  not  satisfied  with  the 
latter  theorj^  as  being  a  good  one  for  the  purpose. 

The  mechanism  of  agglutination  is  explained  in  the  following- 
manner  by  the  hydrate  theory  of  solution.  The  unknown  agglu- 
tinating substance  of  the  serum  and  the  obscure  agglutinable 
product  of  microbic  metabolism  are  each  dissolved  in  their 
respective  fluids  b}'' reason  of  the  "  residual  affinities  "  which  they 
possess  for  loose  water  molecules,  (In  explanation  of  these  terms 
it  may  be  said  that  residual  affinit}^  is  a  chemical  affinity  of  a 
weak  kind,  and  that  a  water  solution  consists  of  molecular 
aggregates  and  loose  water  molecules.)  Each  molecule  of  the 
active  substances  is  the  centre  of  a  system,  and  is  surrounded  b}^  a 
zone  of  water  molecules.  The  solutions  are  in  equilibrium  until 
they  are  mixed,  when  the  active  substances  combine,  the  result 
of  chemical  affinity.  Whether  or  not  a  decomposition  also  takes 
place  we  are  unable  to  say,  and  at  present  this  question  may  be 
ignored.  The  combined  or  double  molecule  now  beomes  the  centre 
of  a  new  water  system,  but  its  residual  affinity  for  the  loose  water 
molecules  is  weaker  than  the  residual  affinity  of  the  saline  matter 
in  the  fluid.  The  origin  of  the  saline  matter  is  immaterial.  The 
salts  attract  the  water  molecules,  and  withdraw  them  from  the 
influence  of  the  double  molecule.  The  residual  affinities  of  the 
double  molecules  are  thus  free,  and  a  kind  of  molecular  tension 
is  produced.  The  double  molecules  are  then  attracted  to  one 
another  and  become  molecular  aggregates,  which  gradually  increase 
in  complexity,  while  the  equilibriums  between  aggregate  and  solu- 
tion are  constantly  altering.  Eventuall}''  the  aggregates  grow  so 
massive  that  they  become  visible  as  a  precipitate.  This  is  Kraus' 
precipitate.     When  substances  separate  out  from  solution  they 

*  Jour.  Soc.  Chem.  Ind.  xvii.,  117. 


BY    R.    GREIG    SMITH.  71 

tend  to  form  upon  small  solid  particles  just  as  rain-drops  gather 
around  dust  particles,  crystals  form  around  string,  and  even 
precipitates  adhere  to  glass.  Thus  Kraus'  precipitate  forms  upon 
the  flagella  and  upon  the  bacterial  cell.  It  forms  upon  the 
bacteria  the  more  readily  since  the  capsule  is  saturated  with  the 
agglutinable  substance  which  slowly  oozes  through  the  membrane. 
But  the  building  up  of  the  molecular  aggregates  still  proceeds, 
and  the  precipitated  films  surrounding  the  bacteria  are  attracted 
to  one  another,  so  that  small  clumps  are  formed  and  agglutination 
is  made  manifest,  because  we  can  see  the  comparatively  large 
bacteria.     The  thin  precipitated  films  elude  our  observation. 

Although  the  residual  affinity  alone  has  been  used  to  explain 
the  phenomenon,  yet  the  base  of  the  flocculating  salt  over  and 
above  its  attraction  for  the  loose  water  molecules  plays  a  part  in 
combining  with  the  molecular  aggregates.  The  base  partly 
attracts  the  Avater  molecules,  and  partly  replaces  them.  Even 
some  of  the  acid  radicles  may  in  some  instances  be  carried  down 
(attracted)  with  the  precipitate  as  in  the  coagulation  of  slimes 
and  mucin  bodies  by  salts  in  alcoholic  solution. 

Agglutination  is  a  chemical  phenomenon,  and  the  natural  laws 
which  govern  it  are  those  which  obtain  in  the  precipitation  of 
chemical  compounds,  the  coagulation  of  precipitates,  the  floccula- 
tion  of  clay  particles,  and  even  the  solution  and  diffusion  of 
substances  in  water.  The  bodies  of  the  bacteria  have  tended  to 
obscure  the  mechanism  of  the  process,  a  thing  which  should  not 
have  been  done  in  view  of  what  Nicolle  did  when  he  agglutinated 
particles  of  talc  suspended  in  filtered  typhoid  cultures. 

A  paper  has  recently  been  written  by  Harrison"^  upon  the 
agglutinating  substance  of  typhoid  bacteria.  He  took  typhoid 
bacteria  which  agglutinated  readily  with  active  sera,  and  treated 
them  with  with  pyocyanase,  an  enzyme  which  has  the  jDower  of 
dissolving  the  outer  membranes  of  bacteria.  After  washing  the 
membraneless  bacteria  with  water,  he  found  that  they  were 
not  now  capable  of  being  agglutinated      From  this  he  concluded 

*  Harrison,  Cent.  f.  Bakt.  1  Abt.  xxx.,  115. 


72  MECHANISM    OF    AGGLUTINATION. 

that  the  agglutinable  substance  existed  "  entirely  in  the  outer 
layers  of  the  bacilli." 

Jochmann,'^'  in  abstracting  a  paper  by  Kohler,  remarked  that 
Harrison  had  in  a  very  clear  manner  proved  Nicolle's  assumption 
that  the  agglutinable  substance  is  found  in  the  outer  membranes 
of  the  bacteria.  With  this  I  do  not  agree.  The  function  of  a 
cell  membrane  is  to  retain  the  cell  contents,  and  the  membrane  of 
the  typhoid  bacterium  retains  the  agglutinable  substance  to  a 
considerable  extent.  That  it  slowly  diffuses  through  is  admitted 
from  the  scanty  formation  of  Kraus'  precipitate  in  younger 
cultures,  and  the  more  copious  precipitation  in  older  cultures. 
But  if  we  remove  the  capsule  or  membrane  there  is  no  restraint 
put  upon  the  diffusion  of  the  agglutinable  substance  from  within 
the  cell.  Harrison's  experiments  simply  show  that  by  removing 
the  membrane  or  capsule  one  of  the  slowly  diffusible  constituents 
of  the  cell  can  be  quickly  removed,  a  circumstance  which  no  one 
would  have  doubted.  They  do  not  prove  that  the  agglutinable 
substance  is  in  the  surface  layers  of  the  cell.  One  might  remove 
the  covering  from  a  bee-hive,  and  with  the  same  amount  of  reason 
say  that  the  bees  which  swarm  out  were  in  the  covering,  or  when 
an  egg  is  broken  that  all  the  fluid  contents  were  originally  in  the 
substance  of  the  shell.  No  !  The  agglutinable  substance  saturates 
the  bacterium  as  a  whole ;  there  is  no  evidence  to  show  that  it  is 
otherwise. 


Jochmann,  Cent.  f.  Bakt.  1  Abt.  xxx.  585. 


73 


NOTES    AND    EXHIBITS. 

Mr.  A.  A.  Hamilton  exhibited  specimens  of  some  interesting- 
plants  as  follows : — 

Persoonia  angulata,  R.Br. — Lawson  (A.  A.  Hamilton;  Novem- 
ber, 1901).  Flowers  erect,  solitary  in  the  axils  of  the  leaves,  one 
of  the  floral  leaves  reduced  to  a  bract  supporting  each  flower; 
corolla  slightly  pubescent,  tipped  with  conspicuous  points;  pedicels 
and  bracts  pubescent;  ovulary  glabrous;  branches  angular  (a 
character  not  common  in  the  genus).  Described  from  specimens 
collected  by  A.  Cunningham  on  the  Blue  Mountains,  without 
flowers  or  fruits,  Bentham  says  of  these  specimens  (B.FL,  v., 
4:02)  : — "  Evidently  distinct  from  all  other  species  and  most  pro- 
bably a  Persoonia,  but  it  must  remain  doubtful  until  flowering 
specimens  have  been  examined."  It  is  interesting  to  note  that 
Bentham  placed  this  species  (provisionally)  in  the  section  Amhly- 
ayithera,  for  an  examination  of  the  flowers  shows  this  to  be  its 
correct  position.  Specimens  were  submitted  to  Mr.  Maiden,  who 
has  confirmed  the  identification  of  the  species.  Mr.  Maiden  had 
collected  flowering  specimens  of  this  plant  at  Woodford. 

Uncinia  tenella,  R.Br. — Katoomba  (A.  A.  Hamilton;  Decem- 
ber, 1901).  Recorded  in  the  '  Flora  Australiensis' from  Victoria 
and  Tasmania;  and  in  the  'Handbook  of  the  Flora  of  N.S.W.' 
from  the  southern  parts  of  the  colony  in  forests.  Mr.  Maiden 
had  also  kindly  confirmed  the  determination  of  this  species. 

Carex gracilis,  R.Br. — South  Clifton  (A.  A.  Hamilton;  January, 
1902).  Recorded  in  both  the  'Flora  Australiensis '  and  the 
'  Handbook  of  the  Flora  of  N.S  W.,'  from  the  Grose  River  north 
to  Queensland.  The  identification  of  this  species  had  been  con- 
firmed by  Mr.  Maiden. 


74  NOTES    AND    EXHIBITS. 

Mr.  Whitelegge  exhibited  specimens  of  U^ispongia  Illatvarrce, 
Whitelegge,  which  he  had  recently  obtained  alive  under  rock 
ledges  on  the  coast.  These  examples  are  much  finer  in  texture 
and  more  compact  than  the  type;  and  are  considered  to  be  at 
least  equal  in  value  to  any  toilet  sponge  imported. 

Professor  Haswell  showed  a  mounted  specimen  of  Gyrocotyle 
nigrosetosa,  sp.n.,  in  illustration  of  his  paper. 

Mr.  Froggatt  exhibited  a  beautiful  collection  of  Coleoptera, 
comprising  about  400  species,  brought  together  by  him  during  a 
few  days'  visit  to  Southern  Queensland  and  the  Northern  Rivers 
of  New  South  Wales,  in  October  last. 

Mr.  R.  Greig  Smith  exhibited  a  series  of  cultures,  various 
chemical  products,  and  sections  under  the  microscope,  illustrating 
the  life-history  and  characteristics  of  Bacterium  vascular um, 
Cobb. 

Dr.  MacPherson  exhibited  an  aboriginal  stone  chisel  weighing 
40oz.,  turned  up  by  the  plough  at  Beaufort,  near  Glen  Innes; 
such  implements  were  used  by  the  Ngarrabul  tribe  in  shaping 
the  handles  of  shields  which  were  made  from  the  wood  of  the 
Black  Kurrajong  (Sterculia  diver  si  folia).  Also  an  example  of  a 
stone  axe  for  use  without  a  handle,  obtained  30  years  ago  from 
the  Yukumbul  tribe  at  Bukulla  in  the  Inverell  District. 

Dr.  Norton  exhibited  an  extraordinary  instance  of  prolification 
of  the  inflorescence  in  a  cultivated  specimen  of  Nicotiana  sp. 

Mr.  North  reported  that  he  had  received  for  examination 
two  birds  shot  by  Mr.  Tom  Carter  on  the  23rd  February,  1902, 
in  a  dense  patch  of  mangroves  at  Point  Cloates,  N.W.  Australia. 
One  is  an  adult  specimen  of  Eopsaltria  pulverulenta,  Bonap., 
( =  Eopsaltria  leucura,  Gould),  and  the  other  the  skin  of  a  young 
bird  referable  to  Eo2Jsaltriagularis,QuoyetGa,imard(= Eopsaltria 
griseogularis,  Gould).  The  latter  specimen  is  unfortunately  much 
mutilated,  the  tip  of  the  upper  mandible,  most  of  the  quills, 
upper  tail-coverts,  and  all  but  two  tail-feathers  being  blown  away, 


NOTES    AND    EXHIBITS.  75 

owing  to  Mr.  Carter's  inability  to  get  far  enough  away  from  the 
bird  without  losing  sight  of  it.  Mr.  George  Masters  also  found 
this  species  inhabiting  the  mangroves  at  Port  Lincoln,  S.A., 
during  the  latter  part  of  1 865.  Count  Salvadori  places  Eo])saliHa 
'pulverulent a  in  the  genus  Foncilodryas ;  if  rightly  so,  then  E. 
georgiana  must  also  be  included  in  it.  The  type  of  Eopsaltria 
gularis  was  figured  and  described  by  MM.  Quoy  &Gaimardfrom 
a  specimen  collected  at  King  George's  Sound,  W.A.,  during  the 
voyage  of  the  "  Astrolabe."  Subsequently  Gould  described  and 
figured  the  same  species  under  the  name  of  E.  griseogularis,  from 
a  specimen  obtained  at  Swan  River.  In  his  Handbook,*  Gould 
erroneously  placed  Muscicapa  georgiana,  Quoy  &  Gaimard, 
as  a  synonym  of  his  Eopsaltria  griseogularis,  and  also  erred  in 
placing  Muscicapa  gularis,  Quoy  &  Gaimard,  as  a  synomym  of 
Eopsaltria  leitcog aster,  Gould.  Dr.  Gadowf  has  similarly  con- 
fused the  synonymy  of  the  two  species,  and  he  erroneously 
describes  Eopsaltria  gularis  under  the  name  of  Eopsaltria 
georgiana,  and  the  latter  species  under  the  name  of  the  former. 
The  correct  synonymy  of  these  two  sj^ecies  as  regards  the  works 
referred  to  is  as  follows  : — 

Eopsaltria  geokgiana 

White-bellied  Robin. 

Gole-mouche  georgien,  Quoy  et  Gaim.,  Voy.  de  I'Astrol.,  Atlas^ 
pi.  3,  fig.  4. 

Muscicapa  georgiana,  Quoy  et  Gaim.,  Voy.  de  I'Astrol.,  Zool.  i. 
p.  175(1830). 

Eopsaltria  leucogaster,  Gould,  P.Z.S.  1846,  p.  19;   Birds  Austr. 
fol.  iii.  pi.  13  (1848);   Handbk.  Birds  Austr.  i.  p.  296  (1865). 

Eopsaltria  gularis  (nee  Quoy  et  Gaim.),  Gadow,  Cat.  Birds  Brit. 
Mus.  viii.  p.l81(1883). 


*  Handbook  of  the  Birds  of  Australia,  i.  p.  294(1865). 
t  Catalogue  of  Birds  in  the  Brit.  Museum,  viii.  p.  178(1883). 


76  NOTES    AND    EXHIBITS. 

EOPSALTRIA    GULARIS. 

Grey-breasted  Robin. 

Gohe-mouche   a  gorge   hlanclie,  Quoy  et  Gaim.,  Yoy.  de  I'Astrol. 

Atlas,  pi.  4,  fig.  1. 
Muscicapa  gidaris,  Quoy  et   Gaim.,  Voy.  de  I'Astrol.,  Zool.   i. 

p.  176  (1830). 
Eo27saltria griseogularis,  Gould,  P.Z.S.  1837,  p.  144;  Birds  Aiistr. 

fol.  iii.  pi.  12  (1848);   Handbk.  Birds  Austr.  i.  p.  294  (1865). 
Eopsaltria  georgiana   (nee   Quoy  et   Gaim.),  Gadow,  Oat.  Birds 

Brit.  Mus.  viii.  p.l78(1883). 


WEDNESDAY,  MAY  28th,  1902. 


The  Ordinary  Monthly  IMeeting  of  the  Society  was  held  in  the 
Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening.  May  28th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S.,  &c.,  President,  in  the  Chair. 

Mr.  William  P.  F.  Dorph,  M.R.A.S.,  A.J.S.  Bank,  Inverell, 
N.S.W.,  was  elected  an  Ordinary  Member  of  the  Society. 

The  President  referred  to  the  death  of  Mr.  J.  B.  Garland,  a 
Member  and  the  only  son  of  the  Society's  esteemed  Honorary 
Treasurer,  on  23rd  inst.  It  was  resolved  that  an  expression  of 
the  Society's  sympathy  should  be  conveyed  to  Mr.  James  R. 
Garland  and  family. 

The  Donations  and  Exchanges  for  the  month,  amounting  to  9 
Vols.,  61  Parts  or  Nos.,  5  Bulletins,  1  Annual  Report  (Admini- 
strative), 2  Pamphlets,  and  2  Miscellanea,  received  from  48 
Societies,  cfec,  and  2  Individuals,  were  laid  upon  the  table. 


77 


NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA 
BELONGING  TO  THE  FAMILY  NOCTUIDuE. 

By  a.  Jefferis  Turner,   M.D.,  F.E.S. 

The  Noctnidcti  of  Australia  have  been  brought,  by  the  labours  of 
a  series  of  entomologists,  into  such  a  state  of  confusion  that  it  has 
been  impossible  for  any  local  worker  unable  to  examine  the  t3^pes  to 
identify  any  of  the  more  obscure  species  with  any  degree  of  confi- 
dence. To  attempt  to  describe  new  species  under  these  conditions 
would  have  been  only  to  add  to  the  existing  confusion.  The  only 
remedy  for  this  state  of  affairs  is  the  revision  by  an  observer 
working  with  real  scientific  method  of  the  whole  group,  including 
a  critical  examination  of  the  types,  more  especially  of  those  in 
the  collection  of  the  British  Museum.  This  much-needed  work 
is  at  present  being  carried  out  by  Sir  George  Hampson  greatly 
to  the  benefit  of  workers  at  this  group  of  insects  in  all  parts  of 
the  world.  To  him  I  am  particularly  indebted  for  his  generous 
assistance  in  examining  and  classifying  the  whole  of  my  Austra- 
lian collection  within  the  limits  of  this  family.  Upon  this 
foundation  I  have  been  able  to  write  the  present  paper.  The 
sub-families  and  generic  identifications  are  those  made  by  him, 
with  a  few  exceptions,  some  of  which  are  indicated  in  the  text. 

Altogether  some  ninety  new  species  are  here  described. 

I  am  particularly  indebted  to  Mr.  F.  P.  Dodd  for  many  new 
species  from  Northern  Queensland,  and  for  a  still  larger  number 
not  yet  recorded  as  Australian;  to  Mr.  P.  Illidge  for  a  few  species 
from  Southern  Queensland  otherwise  unknown  to  me;  and  to  Mr. 
G.  Lyell  for  all  that  I  know  of  the  Victorian  species. 

J  have  been  at  some  pains  to  identify  the  species  described  in 
various  papers  by  Dr.  T.  P.  Lucas.     Many  I  have  so  far  failed  alto- 


78 


NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 


gether  to  identify,  some  no  doubt  because  the  species  are  unknown 
to  me,  but  others  probably  because  they  can  only  be  identified  by  an 
examination  of  the  types.  The  following  list  of  the  species  that 
I  have  succeeded  in  identifying  may  prove  of  use  to  others.  The 
second  column  contains  the  genera  and  species  to  which  they  are 
referred  by  Sir  G.  Hampson. 


Leucania  fumata 


Ophiodes  parcimacida. 

(misprinted  parcemacula) 
Fodina  gloriosa 


P.L.S.N.S.W.  1889. 

Canthylidia  moribunda.  Gn. 

P.L.S.N.S.W.  1891. 

Ophiusa  pa7'Gimactda,  Luc. 


F.  stola,  Gn 


Leucania  laheculis 
L.  linearis 
Agrotis  enunciatus 
Hypogramma  distincta 
Thalpochares  dioidens 
Zethes  conscripta 
Z.  chordoplioides 
Thermesia  tenehrica 
Herminia  delicata 


.R.S.Q.  1891. 

L.  abdominalis,  Wlk. 
L.  venalba,  Moore 
Spodoptera  utnbraticula,  Wlk. 
Gadirtha  pidchra^  Butl. 
Eublemma  vagusana,  F.v.R. 
Sarcopteron  conscriptta,  Luc. 
Episparis  chordophoides,  Luc. 
Opliyx  ochroptera,  Gn. 
Nodaria  anuatalrs,  Wlk. 


P.L.S.N 

Gharicha  sanguinaia 
Luperina  ditata 
Prionop/hora.  torquesaria 
Holocryptis  phasianura 
Grammodes  excellens 
Grammodes  pidcherrbna 

Grammodes  divaricata 

G.  pallens 

Herminia  semicircular  is 


,S.W.  1892. 

Criofrontia  sanguinata,  Luc. 
Acronycta  ditata^  Luc. 
Prionophora  torquesaria,  Luc. 
Holocryptis  phasianura,  Luc. 
Gram>modes  excellens,  Luc. 
G.  pulcherrima,  Luc. 

(G.  minor,  Swin.,  is  a  synonym) 
Ophiusa  constricta,  Butl. 
0.  gravaia,  Gn. 
Adrapsa  sem>icircularis,  Luc. 


BY    A.    JEFFERIS    TURNER. 


79 


P.L.S.N.S.W.  1893 

Pantydia  cajnstrata 
Homodes  violacea 
Westermannia  ocellata 
Jlelipotis    collustrans 
Sympis  parkeri 


Pantydia  cajnstrata,  Luc. 
Micraeschits  ohlatarhis^  Wlk. 
Westermannia  conchidia,  Butl. 
Ercheia  cyllaria^  Cram. 
Sympis  parkeri,  Luc. 


P.R.S.Q.   1894. 


Thalpochares  pectorora 


Madojje  curvata 


Madope  leucocrossa 


Leucania  sejoulchralis 
Bryophila  exquisita 


Imleanga  Jluviatilis 


Euhlemma  semirufa,  Hmps. 

(As  I  have  not  seen  the  refer- 
ence, this  name  may  be  a  synonym) 
Euhlemma  curvata,  Luc. 

P.R.S.Q.  1899. 

Corgatha  Jiguralis,  Wlk. 

P.R.S.Q.  1900. 

L.  irregularis,  Wlk. 
Blenina  lichenopa,  Meyr. 
(A  variable  species) 

P.R.S.Q.  1901. 

Mecodina  Jluviatilis,  Luc. 

N0CTUID5:. 


Agrotinae. 

Cantiiylidia  tenuistria,  n.sp. 

(J 9-  32-33  mm.  Head  and  thorax  pinkish-grey.  Antenna? 
ochreous-whitish.  Abdomen  whitish-ochreous.  Legs  pinkish; 
posterior  tarsi  ochreous-whitish.  Forewings  elongate-triangular; 
costa  slightly  arched,  apex  rounded,  hind  margin  slightly  oblique, 
slightly  rounded ;  pinkish-grey,  towards  termen  ochreous-tinged ; 
costal  edge  whitish;  a  narrow  whitish  streak  from  before  middle  of 
disc  to  middle  of  termen;  four  similar  but  shorter  streaks  between 
and  parallel  to  veins,  two  above  and  two  below  the  preceding: 
cilia   pinkish-grey.      Hindwings  with    termen    gently  rounded  ; 


80         NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

whitish  ;    towards  termen  more   or  less   suffused  with  fuscous ; 
sometimes  with  a  few  pinkish  scales  along  veins;  cilia  whitish. 
Brisbane,  Q.;  four  specimens,  in  January. 

Agrotis  hypochalchis,  n.sp. 

(J.  34  mm.  Head,  thorax,  and  abdomen  brownish-grey. 
Antennae  fuscous  ;  towards  base  brownish-grey.  Legs  grey  ; 
femora  and  coxae  densely  hair}^  Forewings  elongate-triangular; 
costa  almost  straight,  apex  rounded,  termen  rounded,  scarcely 
oblique;  brownish-grey;  traces  of  blackish  dotted  lines  near  base, 
and  transversel}-^  across  disc  at  ^;  a  minute  black  dot  in  disc 
beneath  costa  at  4;  reniform  conspicuous,  dark  reddish-brown, 
posteriorly  narrowly  edged  with  whitish;  two  parallel  fine  dotted 
lines  from  costa  at  |  to  inner  margin  at  f ,  outwardly  curved  in 
disc;  a  sinuate  subterminal  dark  shade;  a  fine  interrupted  blackish 
line  on  termen:  cilia  fuscous.  Hind  wings  with  termen  slightly 
sinuate;  dark  grey;  cilia  grey,  with  a  darker  basal  line.  Under- 
surface  of  both  wings  greenish-ochreous  over  anterior  §,  thence 
fuscous. 

Brisbane,  Q.;  one  specimen,  in  October:  83'dney,  N.S.W.;  one 
specimen  (Coll.  Lyell). 

The  colouring  of  undersurface  is  peculiar  and  characteristic. 
It  is  closely  allied  to  Agrotis  subnigra,  Wlk. 

Mamestrinse. 

Leucania  polysticha,  n.sp. 

^.  40  mm.  Head  and  palpi  ochreous-whitish  irrorated  with 
fuscous.  Antennae  ochreous-whitish.  Thorax  pale  brownish- 
ochreous  mixed  with  fuscous;  a  few  conspicuous  black-tipped 
scales  near  inner  border  of  tegulae.  Abdomen  grey.  Legs 
ochreous-whitish  irrorated  with  blackish.  Forewings  elongate- 
triangular,  costa  straight  except  near  apex,  apex  round-pointed, 
hind  margin  rounded,  moderately  oblique;  brown-whitish  with 
whitish  streaks  on  veins  and  numerous  fuscous  streaks  between 
veins;  a  stronger  fuscous  streak  from  base  to  bej'^ond  cell,  inter- 


BY    A.    JEFFERIS    TURNER.  81 

sected  by  median  vein;  a  blackish  dot  close  beneath  centre  of 
this  streak;  a  row  of  minute  blackish  dots  from  costa  at  |,  first 
outwardly  and  then  strongly  inwardly  curved  to  inner  margin  at 
f ;  a  second  similar  line  from  costa  at  f,  connected  with  preceding 
by  some  dots  on  costa,  similarl}'-  but  more  gently  curved  to  inner 
margin  at  |;  a  row  of  fuscous  dots  on  termen,  being  the  termina- 
tions of  the  intervenal  streaks  :  cilia  fuscous,  opposite  veins 
narrowly  interrupted  with  whitish.  Hindwings  with  termen 
rounded,  indented  beneath  apex;  white;  a  very  slight  fuscous 
suffusion,  and  a  series  of  blackish  dots  on  apical  half  of  termen; 
cilia  white. 

Brisbane,  Q.;  one  specimen,  in  February;  a  second  specimen 
(S)  in  Coll.  Illidge. 

Leucania  melanopasta,  n.sp. 

(J.  30  mm.  Head  whitish.  Palpi  whitish,  irrorated  with  fus- 
cous. AntenncB  ochreous-whitish.  Thorax  w^hitish,  sparsely 
irrorated  with  black.  Abdomen  whitish.  Legs  whitish,  sparsely 
irrorated  with  dark  fuscous;  external  spurs  dark  fuscous.  Fore- 
wings  elongate,  costa  straight  except  near  apex,  apex  round- 
pointed,  hindmargin  rounded,  moderately  oblique :  ochreous- 
whitish,  mixed  with  whitish-grey  and  sparsely  irrorated  with 
black,  the  black  scales  being  most  numerous  near  base;  a  whitish- 
grey  longitudinal  streak  from  base  to  termen;  a  faintly  marked 
series  of  grey  dots  forming  a  line  from  costa  at  4,  first  outwardly 
and  then  inwardly  curved  to  inner  margin  about  |;  a  series  of 
terminal  dots  tending  to  be  prolonged  into  disc  as  stout  streaks 
between  veins  :  cilia  whitish,  mixed  with  fuscous.  Hindwings 
with  termen  rounded,  indented  beneath  apex;  whitish;  a  very 
faint  fuscous  suffusion  and  a  series  of  dark  fuscous  dots  alone 
termen;  cilia  w^iitish. 

Brisbane,  Q. ;  one  specimen,  in  April. 

Caradrininae. 
EuPLEXiA  callisina,  n.sp. 
9.   18  mm.     Head  dark  fuscous,  mixed  wdth  pale  ochreous  and 
whitish.     Palpi  dark  fuscous  with  a  few  ochreous-whitish  scales; 


£ 


82  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

apex  of  second  and  third  joints  and  anterior  border  of  second  joint 
whitish-ochreous.  Antennae  wbitish-ochreous,  obscurely  annu- 
lated  with  pale  fuscous;  beneath  ochreous-fuscous.  Thorax  dark 
fuscous,  mixed  with  ochreous-brown  and  whitish.  Abdomen 
dark  grey;  crests  as  thorax.  Legs  dark  fuscous,  irrorated  and 
annulated  with  whitish-ochreous.  Forewings  elongate-triangular; 
costa  gently  rounded,  apex  round-pointed,  termen  rounded,  crenu- 
late,  slightly  oblique;  dark  fuscous,  mixed  with  ochreous-brown 
and  whitish,  the  centre  of  disc  being  darker;  veins  incompletely 
outlined  in  white  and  ochreous;  orbicular  represented  by  an  oval 
ring  of  mixed  white  and  ochreous  scales;  reniform  similar  but 
larger  and  biconcave ;  basal  irroration  tending  to  form  several 
darker  and  lighter  transverse  lines;  a  dark  fuscous  transverse 
median  shade;  two  closely  approximated  pale  lines  from  costa  at 
J,  first  outwardly,  then  inM^ardly  curved  to  inner  margin  at  J;  a 
fine  ochreous  subterminal  line  preceded  by  a  series  of  wedge- 
shaped  black  spots;  beyond  this  is  a  terminal  band  of  whitish 
irroration;  an  interrupted  black  hindmarginal  line  :  cilia  ochreous 
at  base,  then  a  fine  black  line,  then  fuscous  mixed  with  ochreous- 
whitish.  Hindwings  with  termen  rounded,  wavy;  dark  grey, 
inclining  to  fuscous;  cilia  whitish-ochreous  with  a  dark  fuscous 
line  at  J,  on  inner  margin  grey. 

Gisborne,  Vic;  one  specimen,  in  January,  taken  by  Mr.  G. 
Lyell. 

A  very  fine  and  distinct  species. 

EUPLEXIA    LEUCOSTIGMA,   n.Sp. 

^.  34  mm.  Head,  palpi  and  thorax  fuscous,  mixed  with  pale 
brown  and  whitish.  Antennae  fuscous,  in  male  simple,  with  very 
short  ciliations  (i).  Abdomen  fuscous,  mixed  with  whitish ; 
crests  dark  fuscous.  Legs  dark  fuscous  mixed  with  whitish  ; 
posterior  pair  mostly  whitish;  anterior  tarsi  dark  fuscous  annu- 
lated with  whitish.  Forewings  elongate-triangular,  costa  gently 
arched,  apex  round-pointed,  termen  rounded,  wavy  oblique;  grey 
mixed  with  brownish  and  whitish;  a  broad  oblique  dark  grey 
median  shade,  not  reaching  inner  margin,  and  containing  a  fine 


BY    A.    JEFFERIS    TURNER.  83 

oval  black  ring,  which  represents  the  orbicular;  reniform  ante- 
riorly brownish,  posteriorly  white,  incompletely  outlined  with 
black,  and  connected  with  costa  by  a  white  spot;  claviform  repre- 
sented by  a  short  thick  black  streak;  a  fine  dentate  black  line 
near  base;  a  second  outwardly  curved  dentate  line  before  orbicu- 
lar; a  fine  dentate  line  from  costa  at  |,  first  outwardly  then 
inwardly  curved  to  inner  margin  at  |;  a  fine  black  terminal  line: 
cilia  fuscous  mixed  with  whitish.  Hindmargin  with  termen 
rounded,  wav3^;dark  fuscous;  toward  base  whitish;  with  a  fuscous 
crescentic  mark  at  end  of  cell,  and  a  dark  curved  median  line; 
cilia  fuscous  mixed  with  whitish. 

Gisborne,  Vic;  one  specimen,  in  January,  taken  by  Mr.  G. 
Lyell. 

EUPLEXIA    POLYCMETA,  n.Sp. 

(J.  36  mm.  Head  and  palpi  grey,  irrorated  with  black. 
Antennae  dark  fuscous;  in  male  shortly  pectinate  (1),  near  apex 
serrate.  Thorax  dark  fuscous,  with  a  strong  bifid  posterior  crest. 
Abdomen  grey ;  crests  dark  fuscous.  Legs  whitish,  pinkish- 
tinged,  irrorated  with  dark  fuscous.  Forewings  rather  broadly 
triangular,  costa  nearly  straight,  apex  round-pointed,  termen 
crenate,  well  rounded,  slightly  oblique;  grey,  with  whitish  and 
brownish  irrorations  and  black  lines;  two  black  lines'  from  costa 
near  base  to  median  vein;  two  linos  from  costa  at  J  to  inner 
margin  at  J;  disc  internal  to  this  densely  irrorated  with  whitish; 
orbicular  and  reniform  whitish-grey,  slenderl}''  outlined  with* 
black,  and  containing  each  a  darker  grey  circle;  a  dentate  line, 
very  fine  except  on  costa,  from  costa  at  J  to  middle  of  inner 
margin,  passing  between  orbicular  and  reniform;  two  parallel 
dentate  lines  describing  a  sigmoid  curve  from  costa  at  §  to  inner 
margin  at  f ;  this  is  followed  near  inner  margin  by  a  crescentic 
blackish  shade,  and  above  this  by  a  narrow  whitish  irro ration; 
inner  marginal  portion  of  mid-disc  and  a  broad  submarginal  band 
suffused  with  brownish;  a  series  of  black  intervenal  black  streaks 
cross  this  band  to  termen;  a  whitish  irroration  at  tornus;  a  fine 
black  terminal  line  :  cilia  fuscous,  irrorated  with  whitish.      Hind- 


84  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

wings  with  termen  crenate,  rounded;  dark  fuscous;  cilia  whitish, 
with  a  fuscous  median  Hne.  Hind  wings  beneath  whitish,  with  a 
dark  fuscous  discal  dot  and  a  very  broad  dark  fuscous  terminal 
band. 

Lawson,  Blue  Mts.,  N.S.W;  one  specimen, 

Elusa  oenolopha,  n.sp. 

(^9.  26-29  mm.  Head,  palpi,  and  thorax  reddish-brown  tinged 
with  purple.  Antennae  ochreous-fuscous,  in  male  unipectinate, 
toward  apex  simple;  pectinations  long  (6)  on  outer  side  of  antennal 
stalk.  Abdomen  ochreous-brown;  crests  purplish;  tuft  in  male 
pale  ochreous.  Legs  brownish-ochreous,  tibise  purplish-tinged; 
posterior  tibise  in  male  with  a  large  dense  tuft  of  long  purjDle 
hairs  on  dorsal  surface.  Forewings  elongate-triangular,  costa 
moderately  arched,  apex  round-pointed,  termen  slightly  rounded? 
oblique;  reddish-brown  with  some  faintl}-  marked  fuscous  lines; 
orbicular  obsolete  or  represented  by  a  minute  whitish  dot;  reni_ 
form  grey,  narrowly  edged  with  whitish,  constricted  above  middle, 
or  divided  into  two  unequal  segments;  a  faint  dentate  line  from 
costa  at  ^  to  inner  margin  at  J;  traces  of  a  median  and  two  posterior 
lines:  cilia  reddish-brown.  Hind  wings  with  termen  rounded: 
grey:  cilia  whitish,  purplish-tinged. 

Brisbane,  Q.;  two  specimens. 

S  Y  N  T  H  E  T  A,    n.g. 

•  Face  flat.  Palpi  moderate,  ascending,  not  nearly  reaching 
vertex;  second  joint  loosely  scaled;  terminal  joint  short,  smooth- 
scaled,  porrect.  Tongue  well  developed.  Antennae  of  male 
shortly  laminate,  apex  simple,  ciliated.  Thorax  with  a  small 
posterior  crest.  Abdomen  with  small  dorsal  crests.  Posterior 
tibife  with  two  pairs  of  spurs.  Hindwings  with  vein  8  abruptly 
diverging  from  cell  at  base. 

Beside  the  type,  described  below,  I  refer  Mnmestra  nigerrima, 
Gn.,  to  this  genus. 

From  Magttsa,  Wlk.,  to  which  it  is  closely  allied,  and  also  from 
the  genus  described  below,  it  may  be  distinguished  by  the  palpi 


BY    A.    JEFFERIS    TURNER.  85 

not  reaching  vertex.  All  three  genera  differ  in  the  structure  of 
vein  8  of  hind  wings  from  Frodenia,  Gn.,  and  Spodoptera,  Gn.,  in 
which  vein  8  diverges  very  gradually  from  cell. 

Synth  ETA  xylitis,  n.sp. 

(J.  34  mm.  Head  and  palpi  ochreous-whitish;  face  with  an 
indistinct  blackish  transverse  line;  palpi  with  a  few  blackish 
scales.  Antennse  grey.  Thorax  whitish  mixed  with  brownish 
and  dark  fuscous;  collar  ochreous-whitish  with  a  fine  transverse 
dark  fuscous  line.  Abdomen  grey;  crests  formed  by  loose  whitish- 
grey  hairs;  an  ochreous-brown  spot  on  dorsum  of  terminal  segment; 
tuft  whitish-ochreous,  towards  apex  pinkish-tinged.  Legs  whitish 
mixed  with  pinkish  and  dark  fuscous.  Forewings  elongate- 
triangular,  costa  slightly  rounded,  apex  rounded,  termen  rounded, 
slightly  oblique;  pale  brownish  mixed  with  whitish,  markings 
dark  fuscous;  a  short  oblique  line  from  costa  near  base  to  fold; 
followed  by  a  number  of  short  costal  transverse  strigulee;  orbicular 
represented  by  a  pale  circular  area  preceded  and  followed  by  a 
dark  fuscous  spot;  reniform  faintl}^  outlined,  its  lower  end  white; 
immediately  succeeding  reniform  is  a  fine  incomplete  wavy  trans- 
verse line;  traces  of  a  subterminal  line;  a  well  marked  interrupted 
terminal  line  :  cilia  pale  brown-whitish,  with  a  fine  interrupted 
dark  fuscous  median  line.  Hindwings  with  termen  rounded, 
slightly  wavy;  dark  fuscous,  towards  base  paler;  cilia  whitish,  at 
apex  fuscous. 

Townsville,  Q.;  in  March;  one  specimen  received  from  Mr.  F. 
P.  Dodd. 

E  c  P  A  T  I  A,   n.g. 

Face  flat.  Palpi  ascending,  reaching  vertex ;  second  joint 
loosely  scaled;  terminal  joint  stout,  ascending.  Thorax  with  a 
strong  posterior  crest.  Abdomen  crested,  crests  strongly  developed 
on  4th  and  5th  segments.  Forewings  broadly  triangular.  Hind- 
wings  with  vein  8  abruptly  diverging  from  cell  at  base. 

Allied  to  Magusa,  Wlk.,  from  which  it  differs  in  the  broadly 
triangular  forewings,  crested  thorax,  and  more  strongly  crested 
abdomen.     Type  Catephia  didcistriga,  Wlk.     The  resemblance  of 


86         NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

this  species  to  Catephia  is  merely  superficial,  the  structure  of  vein 
5  of  hind  wings  being  altogether  different. 

Caradrina  chromoneura,  n.sp. 

(^.  29  mm.  Head  dark  fuscous  mixed  with  whitish.  Pa.lpi 
white  with  a  few  dark  fuscous  scales;  basal  J  of  second  joint  dark 
fuscous.  Antennae  fuscous;  in  male  simple.  Thorax  dark  fuscous 
mixed  with  whitish  and  ochreous,  the  ochreous  scales  being 
especially  developed  on  collar.  Abdomen  ochreous-f uscous.  Legs 
dark  fuscous,  annulated  with  whitish.  Forewings  narrowly  elon- 
gate-triangular, costa  gently  arched,  apex  round-pointed,  termen 
bowed  between  veins  3  and  4,  oblique;  white,  irrorated  sparsely 
with  dark  fuscous,  more  densely  in  centre  of  disc;  orbicular  and 
claviform  obsolete;  reniform  obscurely  indicated  by  an  oblique 
whitish  loop  from  costa;  an  incomplete  black  line  near  base;  a 
finely  dentate  black  line  from  costa  at  \  obliquely  to  inner  margin 
at  f ;  a  third  black  dentate  line  from  costa  at  f  curved  first  out- 
wardly then  inwardly  to  inner  margin  at  j ;  base,  an  incomplete 
transverse  line  before  \^  and  most  of  veins  outlined  with  ochreous; 
a  blackish  suffusion  anterior  and  posterior  to  reniform;  a  series 
of  wedge-shaped  blackish  terminal  dots :  cilia  dark  fuscous  barred 
with  white.  Hindwings  with  termen  slightly  sinuate;  dark  grey, 
towards  base  paler;  cilia  dark  grey  barred  with  white,  on  inner 
margin  whitish. 

Ulverstone,  Tas.;  one  specimen  (Coll.  Lyell). 

E  c  c  L  E  T  A,   n.g. 

Head  loosely  scaled.  Face  not  projecting.  Eyes  smooth. 
Tongue  well  developed.  Palpi  rather  large,  upwardly  curved, 
reaching  base  of  antennae,  second  joint  loosely  scaled  towards 
apex,  terminal  joint  well  developed,  with  appressed  scales. 
Antennae  in  male  with  long  pectinations  (10),  shorter  towards 
apex,  apical  jy  simple.  Thorax  with  a  slight  posterior  crest. 
Abdomen  smooth.  Posterior  tibiae  with  two  pairs  of  spurs. 
Forewings  with  vein  2  from  middle  of  cell,  3  from  angle,  6 
separate,  7,  8,  9  stalked,  10  connected  by  a  bar  with  7,  8,  9. 
Hindwings  normal. 


BY    A.    .TEFFERIS    TURNER.  87 

ECCLETA  XUTHOPHANES,  11. Sp. 

(J9.  28-30  mm.  Head,  palpi,  antennjie,  and  thorax  dark  fuscous. 
Abdomen  grey.  Legs  dark  fuscous,  mixed  with  ochreous-whitish. 
Forewings  elongate-triangular,  costa  moderately  arched,  apex 
rounded,  hindmargin  rounded,  slightly  oblique  ;  dark  fuscous, 
with  a  few  scattered  paler  scales;  orbicular  represented  by  a 
whitish  dot;  reniform  by  a  minute  whitish  crescentic  mark;  traces 
of  a  transverse  blackish  line  at  J,  and  of  an  outwardly  curved  line 
from  costa  at  J  to  inner  margin  at  f ;  a  whitish  subterminal  line: 
cilia  dark  fuscous.  Hind  wings  with  termen  rounded,  slightly 
sinuate  beneath  apex;  coppery-red;  a  terminal  band  and  cilia 
fuscous;  inner  marginal  cilia  whitish-ochreous. 

Towns ville  and  Stradbroke  Island,  Q.;  three  specimens. 

EuteliansB. 

Ingura  cyanodes,  n.sp. 

(J9.  28-32  mm.  Head,  palpi,  and  antennae  grey,  purplish- 
tinged.  Thorax  bluish  or  purplish-grey;  anterior  edge  of  collar 
ochreons-brown;  a  transverse  dark  fuscous  line  immediately  behind 
collar.  Abdomen  grey,  ochreous-tinged;  second,  third,  and  fourth 
segments  edged  posteriorly  with  black;  beneath  whitish,  mixed 
with  pale  ochreous-brown;  with  a  lateral  row  of  black  dots,  nnd 
three  median  dark  fuscous  lines.  Legs  dark  fuscous;  posterior 
pair  white,  irrorated  with  dark  fuscous.  Forewings  narrow- 
elongate,  in  female  rather  broader,  costa  slightly  sigmoid,  apex 
round-pointed,  termen  obliquely  rounded;  whitish-grey,  somewhat 
bluish-tinged;  towards  costa  grey;  an  outwardly  curved  oblique 
black  line  from  inner  margin  at  ^,  preceded  b}'  a  small  white 
suffusion,  and  succeeded  by  an  ochreous-fuscous  suffusion ;  a 
second  outwardly  curved  fine  black  line  from  middle  of  inner 
margin  very  obliquely  outwards  to  near  termen,  where  it  curves 
abruptly  upwards  and  backwards  towards  costa  at  |;  a  minute 
black  dot  before  middle  of  disc;  succeeded  by  two  fine  grey  lines 
from  inner  margin  to  centre  of  disc,  parallel  to  second  black  line; 


88  NEW    GKNKRA    AND    SPECIES    OF    LEPIDOPTERA, 

second  black  line  is  preceded  in  its  lower  half  by  a  white  line, 
absent  in  female;  and  succeeded  towards  tornus  by  some  ochreous 
and  purplish  suffusions;  a  very  fine  crenate  black  line  from  angle 
of  second  line  toward  tornus;  a  dark  fuscous  subterminal  line, 
and  a  fine  dark  fuscous  line  on  termen  :  cilia  grey,  basal  half 
mixed  with  whitish,  bluish-tinged  scales.  Hind  wings  with 
termen  rounded,  slightly  waved;  fuscous;  towards  base  whitish; 
veins  darker  fuscous;  an  elongate  white  mark  at  tornus;  blackish 
subterminal  and  terminal  lines  ;  cilia  grey,  bases  whitish,  at 
tornus  wholly  whitish. 

Townsville,  Q. ;  in  December  and  February;  two  specimens 
from  Mr.  F.  P.  Dodd,  who  informs  me  that  they  were  bred  from 
larvae  feedino:  on  Malaisia  tortuosa. 


'& 


EUTELIA    OXYLOPHA,  H.sp. 

^^.  29  mm.  Head,  palpi,  antennae  and  thorax  brownish-grey; 
thorax  with  a  sharp  median  crest  throughout.  Abdomen  brownish- 
grey,  with  two  or  three  median  dark  fuscous  dots  posteriorly. 
Legs  dark  grey,  with  obscure  whitish  annulations.  Forewings 
elongate-triangular,  costa  straight,  apex  round-pointed,  termen 
sharpl}^  angled  on  vein  3,  oblique ;  brownish-groy ;  a  minute 
blackish  dot  beneath  costa  at  4;  followed  by  four  faintly  indicated, 
waved,  transverse,  darker  lines;  three  minute  whitish  dots  on 
apical  part  of  costa;  a  row  of  dark  fuscous  terminal  dots :  cilia 
fuscous.  Hindwings  with  termen  gently  rounded,  crcnulate  ; 
fuscous;  towards  base  pale;  three  dark  fuscous  dots  on  inner 
margin  above  tornus ;  a  narrow  fuscous  terminal  line ;  cilia 
fuscous. 

Has  a  general  resemblance  to  Eutelia  dilatrix,  Gn.,  but  may  be 
readily  distinguished  by  the  ridge-like  crest  of  thorax,  which 
extends  right  to  anterior  edge;  and  also  by  the  absence  of  the 
ante-apical  costal  triangle  on  the  forewings  of  the  latter  species, 
which  has  a  posterior  thoracic  crest  only. 

Brisbane,  Q.;  two  specimens  taken  by  Mr.  11.  Illidge. 


BY    A.    JEFFERIS    TURNKR.  89 

Sarrothripinae. 
Blesina  metachrysa,  n.sp. 

(J.  40  mm.  Head  and  palpi  white,  irrorated  with  dark  fuscous. 
Antennae  dark  fuscous,  with  a  few  whitish  scales;  in  male  slightly- 
serrate,  shortly  ciliated  (J).  Thorax  fuscous,  irrorated  with 
whitish  and  ochreous-brown  scales;  central  area  paler.  Abdomen 
deep  ochreous-yellow ;  terminal  segment  fuscous;  tuft  whitish- 
grey;  dorsal  crests  whitish,  irrorated  with  ochreous-brown  and 
whitish.  Legs  white,  irrorated  with  fuscous.  Forewings  oblong, 
costa  arched  at  base,  then  straight,  apex  rectangular,  termen 
slightly  rounded,  hardly  oblique;  dark  grey,  with  whitish  and 
ochreous-brown  irroration,  and  black  lines  ;  a  small  tuft  of 
ochreous-yellow  hairs  on  base  of  inner  margin  ;  an  inwardly 
oblique,  acutely  dentate  line  from  costa  at  1;  a  broad,  nearly 
straight  streak,  interrupted  in  disc  from  costa  at  ^  to  tornus; 
this  gives  off  in  disc  a  fine  irregularly  dentate  line  to  inner 
margin  beyond  middle;  a  fine  irregularly  dentate  line  from  mid- 
costa,  curved  first  outwardly  then  inwardly  to  inner  margin  at  f ; 
a  finely  dentate  subterminal  line,  interrupted  by  the  oblique 
streak;  terminal  part  of  disc  densely  irrorated  with  whitish;  a 
fine  fuscous  crenulate  terminal  line :  cilia  white,  mixed  with  dark 
fuscous  towards  base,  and  with  a  dark  fuscous  median  line. 
Hindwings  with  termen  rounded,  crenulate;  deep  ochreous-yellow; 
a  broad  dark  fuscous  terminal  band,  narrowing  to  a  point  at 
tornus;  cilia  white  opposite  veins  2  to  4,  above  this  dark  fuscous, 
beneath  deep  ochreous-yellow. 

Townsville,  Q.;  in  February;  two  specimens  received  from  Mr. 
F.  P.  Dodd. 

OCHTHOPHORA,  n.g. 

Face  flat,  with  a  fringe  of  projecting  scales  on  forehead.  Palpi 
rather  large,  ascending;  terminal  joint  nearly  as  long  as  second, 
stout,  obtuse.  Antennae  in  male  serrate.  Tongue  rudimentary. 
Thorax  with  a  large  undivided  posterior  crest.  Abdomen  smooth. 
Posterior  tibiae  with  two  pairs  of  spurs.  Forewings  with  vein  2 
from  f ;  7,  8,  9  stalked,  10  connected  by  a  bar  with  8,  9  beyond 


90         NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

7.     Hindwings  with  4  and  5  short-stalked,  8  connected  with  cell 
before  middle. 

Sir  George  Hampson  referred  the  type  species  to  Gadirtha^ 
Wlk.,  but  the  dense  posterior  thoracic  crest  seems  a  sufficient 
distinction. 

OCHTHOPHORA    SERICINA,  n.sp. 

^.  40  mm.  Head  grey;  lower  I  of  face  whitish.  Palpi  whitish; 
posterior  surface  grey.  Antennae  grey.  Thorax  gre}-,  mixed 
with  brownish;  a  fine  transverse  black  line  near  base  of  crest. 
Abdomen  grey.  Legs  grey;  posterior  pair  whitish.  Forewings 
oblong,  costa  strongly  arched  at  base,  thence  straight  to  near 
apex,  apex  rounded,  termen  slightly  rounded,  hardly  oblique; 
glossy  grey;  many  wavy  whitish-grey  lines  from  costa  becoming 
lost  in  disc;  three  dark  fuscous  dots  in  a  triangle  close  to  base;  a 
small  dark  fuscous  irroration  at  base  of  inner  margin;  a  fine 
strongly  waved  black  line  from  costa  at  \  to  inner  margin  at  |^; 
orbicular  dark  grey,  slenderly  outlined  with  whitish;  reniform 
small,  dark  grey;  a  fine  acutely  dentate  black  line  from  mid-costa, 
obliquely  outwards,  describing  a  complete  loop  in  disc,  then 
returning  to  form  a  second  loop,  which  encloses  reniform,  and 
ending  in  inner  margin  at  j;  a  wavy  whitish-grey  sabterminal 
line;  a  row  of  faintly  darker  terminal  dots:  cilia  grey.  Hind- 
wings  with  termen  rounded,  indented  on  vein  5;  white;  a  fuscous 
terminal  band  from  apex,  narrowing  and  not  quite  reaching  tornus 
cilia  grey,  towards  tornus  white. 

Brisbane,  Q.;  one  specimen  in  December. 

I  c  A  s  M  A,  n.g. 

Face  flat,  forehead  with  a  projecting  ridge  of  scales.  Palpi; 
long,  rough-haired  ;  second  joint  somewhat  ascending ;  terminal 
joint  porrect  or  slightly  descending,  longer  than  second,  obtuse. 
Tongue  present.  Thorax  with  a  small  posterior  crest.  Abdomen 
with  a  smooth  patch  of  scales  on  first  segment,  representing  a 
dorsal  crest.  Posterior  tibite  with  two  pairs  of  spurs.  Forewings 
with  veins  7,  8,  9  stalked;  10  connected  b}^  a  bar  with  8,  9  beyond 


BY    A.    JEFFERIS    TURNER.  91 

7.      Hindwings  with  vein  5  absent,  8  connected  with  cell  before 
middle. 

The  peculiar  palpi  distinguish  this  from  Hypothripa,  Hmps. 
They  appear  to  resemble  those  of  Arachnognatha^  Hmps.,  which, 
however,  has  the  thorax  smooth,  and  vein  5  of  hindwings  present. 

ICASMA    MINUTUM,  n.sp. 

^.  14  mm.  Head,  palpi,  and  thorax  grey,  mixed  with  whitish^ 
lower  half  of  face  white.  Antennae  dark  grey.  Abdomen  grey; 
first  segment  with  a  small  patch  of  dark  fuscous  and  whitish 
scales.  Legs  whitish-grey.  Forewings  oblong,  posteriorly  dilated, 
costa  strongly  arched  at  base,  thence  straight  to  near  apex,  apex 
rounded,  termen  slightly  rounded,  slightly  oblique;  grey,  mixed 
with  whitish,  lines  dark  fuscous;  two  short  lines  from  costa  near 
base;  a  suffused  line  from  costa  at  4  to  inner  margin  at  \\  two. 
parallel  closely  approximated  lines  from  costa  at  4  to  before 
middle  of  inner  margin;  beyond  this  is  an  obscure  whitish  spot 
representing  the  reniform;  a  fine  wavy  line  from  costa  at  f,  first 
outwardly,  then  inwardl}'  curved  to  inner  margin  at  |;  a  fine 
dentate  subterminal  line,  followed  by  a  fine  whitish  line:  cilia 
grey.  Hindwings  with  termen  wavy,  indented  between  veins  4 
and  6;  grey;  cilia  grey. 

A  very  small  and  inconspicuous  species.  The  type  is  not  in 
the  best  condition,  but  the  generic  characters  should  be  suflicient 
for  identification. 

Brisbane,  Q.;  one  specimen,  in  July. 

Plotheia  rhythmopis,  n.sp. 

2.  26mm.  Head  grey.  Palpi  grey;  beneath  whitish;  terminal 
joint  dark  fuscous  at  base  on  upper  surface.  Antennae  ochreous- 
f  uscous.  Abdomen  grey.  Legs  grey ;  posterior  pair  whitish. 
Forewings  oval,  posteriorly  strongh'-  dilated,  costa  moderately 
arched,  apex  rounded,  termen  very  obliquely  rounded;  grey;  a 
wavy  blackish  line  from  costa  at  ^  to  inner  margin  at  1;  a  second 
similar  line  from  costa  at  f ,  obtusely  angulated  in  disc,  to  inner 
margin  at  4;  the  disc  between  these  lines  is  largely  suff'used  with 


92         NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

brownish-fuscous,  containing  two  suffused  dark  fuscous  spots 
representing  orbicular  and  reniform,  and  a  dark  fuscous  line  from 
middle  of  costavery  obliquely  outwards  to  near  second  black  line, 
and  continued  parallel  and  close  to  this  line  to  inner  margin;  two 
faint  lines  of  brownish  suffusion  from  tornus,  and  from  termen 
above  tornus;  a  series  of  dark  fuscous  terminal  dots:  cilia  grey. 
Hindwings  with  termen  rounded;  gre}^- whitish;  on  termen  grey; 
cilia  whitish -grey. 

Brisbane,  Q.;  one  specimen. 

Sarrothripa  symmicta,  n.sp. 

(J.  23  mm,  Head,  palpi,  antennae,  and  thorax  grey,  slightly 
ochreous-tinged.  Abdomen  grey.  Legs  whitish;  anterior  pair 
mixed  with  dark  fuscous.  Forewings  elongate-oblong,  somewhat 
dilated  posteriorly,  costa  strongly  arched  at  base,  thence  slightly 
concave  to  near  apex,  apex  rounded,  termen  rounded,  slightly 
oblique;  grey,  slightly  ochreous-tinged,  markings  dark  fuscous;  a 
short  transverse  streak  from  base  of  costa;  two  or  three  lines  from 
basal  part  of  costa,  lost  in  disc;  a  suffused  spot  on  fold;  a  fine  line 
from  mid-costa  angulated  first  outwards  then  inwards  near  costa, 
thence  outwardly  curved,  bent  obliquely  outwards  near  inner 
margin  to  margin  of  |;  on  this  rests  a  wedge-shaped  spot  in  mid 
disc;  two  fine  closely  approximated  parallel  lines  from  costa  at  i 
first  inwardly  then  outwardly  curved  to  before  tornus;  a  sub- 
terminal  series  of  dots  best  marked  towards  tornus;  some  minute 
terminal  dots:  cilia  whitish.  Hindwings  with  termen  rounded, 
indented  beneath  apex;  grey;  cilia  grey- whitish. 

Brisbane,  Q.;  one  specimen. 

Superficially  rather  like  S.  tnetaspilella,  Wlk.,  but  may  be 
readily  distinguished  by  the  double  posterior  line. 

Dendrothripa  clopaea,  n.sp. 

9.  24  mm.  Head  grey,  mixed  with  whitish.  Palpi  white, 
mixed  with  fuscous  and  black.  Antennae  grey.  Tliorax  grey, 
mixed  with  whitish  and  black.     Abdomen  ochreous-grey,  with  a 


BY    A.    JEFFERIS    TURNER.  93 

small  basal  crest  of  grey  mixed  with  whitish.  Legs  whitish, 
anterior  pair  dark  fuscous  anteriorl}^;  middle  pair  with  a  few 
dark  fuscous  scales  Forewings  elongate-oblong,  posteriorly 
dilated,  costa  strongly  "arched  at  base,  thence  slightly,  apex  round- 
pointed,  hindmargin  slightly  rounded,  slightly  oblique;  grey  mixed 
with  fuscous  and  whitish,  lines  black;  an  outwardly  curved  line 
from  costa  near  base  not  quite  reaching  inner  margin;  a  second 
line  from  costa  at  J,  obscurely  angulated  near  costa,  thence 
straight  or  slightly  sigmoid  to  middle  of  inner  margin;  a  third 
line  from  middle  of  costa  very  obliquely  outwards  forming  a  strong 
obtuse  projection  in  disc,  and  ending  in  inner  margin  at  |;  reni- 
form  obscurely  indicated  by  a  small  ochreous-brown  suffusion 
between  second  and  third  lines;  a  row  of  black  terminal  dots: 
cilia  grey,  mixed  with  dark  fuscous  and  whitish.  Hindwings 
with  termen  rounded,  slightly  sigmoid  beneath  apex;  gre}^;  cilia 
grey,  apices  whitish. 

Brisbane,  Q.;  one  specimen,  in  November. 

Dendrothripa  baeota,  n.sp. 

(J.  12-13  mm.  Head,  palpi,  antennae,  and  thorax  grey.  Abdo- 
men ochreous-grey;  with  a  small  grey  basal  crest.  Legs  grey; 
posterior  pair  whitish.  Forewings  elongate-triangular,  costa 
gently  arched,  apex  round-pointed,  termen  slightty  rounded, 
slightly  oblique;  whitish-grey;  a  broad  fuscous  suffusion  on  base 
of  costa,  and  another  at  -I;  a  fine  fuscous  line  from  costa  before 
middle,  and  another  beyond  middle,  first  obscurely  continued  to 
middle  of  inner  margin,  second  outwardly  curved,  twice  obtusely 
dentate  in  disc,  to  inner  margin  at  f ;  second  line  immediately 
preceded  by  a  minute  black  discal  dot;  a  third  suffusion  on  costa 
at  |;  a  faintly  marked  fuscous  subterminal  line;  a  series  of  black 
terminal  dots  :  cilia  whitish-grey.  Hindwings  with  termen 
rounded,  slightly  sinuate  beneath  apex;  whitish,  veins  and  termen 
grey;  cilia  whitish. 

Townsville,  Q.;  in  January;  two  specimens  received  from  Mr. 
F.  P.  Dodd. 


94  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

AcontiansB. 
Maurilia  iocephala,  n.sp. 

9.  25  mm.  Head,  palpi,  and  thorax  pale  ochreous-brown  with 
purple  reflections.  Antennae  fuscous.  Abdomen  whitish-grey. 
Legs  purplish-grey  mixed  with  white;  posterior  pair  mostly  white. 
Forewings  elongate-triangular,  costa  strongly  arched,  apex  rect- 
angular, termen  straight,  not  oblique ;  pale  ochreous-brown, 
towards  termen  whitish-grey,  irrorated  with  purple-grey  scales; 
lines  purple-grey;  first  from  costa  at  ^  to  middle  of  inner  margin; 
second  from  costa  at  f,  wavy,  to  inner  margin  at  J;  preceded  by 
an  obscure  discal  dot;  a  subterminal  and  a  terminal  sei'ies  of  dark 
fuscous  dots:  cilia  purple-grey,  apices  pale  pink.  Hindwings 
with  termen  rounded,  slightly  sigmoid  beneath  apex;  whitish,  a 
narrow  terminal  fuscous  suffusion;  cilia  whitish. 

Townsville,  Q.;  in  March;  one  specimen  received  from  Mi-.  F. 
P.  Dodd. 

ACONTIA    ExMBOLOSCIA,   n.Sp. 

5.  35  mm.  Head  white,  posteriorly  yellowish;  face  and  palpi 
reddish.  Antennae  grey,  towards  base  whitish.  Thorax  yellow, 
with  a  reddish-brown  central  streak.  Abdomen  orange-ochreous. 
Legs  reddish-brown  ;  anterior  femora  whitish ;  posterior  pair 
whitish,  tinged  with  reddish.  Forewings  elongate-triangular, 
apex  rounded,  termen  straight,  rounded  near  tornus,  not  oblique; 
yellow;  a  wedge-shaped  fuscous  suffusion  having  its  apex  at  base 
of  wing,  its  base  extending  whole  length  of  termen,  obscuring  the 
lines,  which  nevertheless  can  mostly  be  traced  through  it:  lines 
and  dorsal  margin  reddish-brown;  a  line  from  costa  at  -^  outwardly 
oblique,  forming  an  acute  angle  in  disc^  thence  inwardly  oblique 
to  fold;  a  median  discal  dot  just  above  suffusion;  a  line  from  costa 
at  f  obliquely  outwards,  forming  an  acute  angle  near  to  termen, 
thence  inwardly  oblique  to  mid-clorsum,  interrupted  before  dorsum; 
a  line  from  costa  near  apex  to  angle  of  posterior  line;  a  short  line 
from  dorsum  at  i  obliquely  outwards  to  suffusion :  cilia  fuscous. 
Hindwings  with  termen  rounded;  orange-ochreous;  cilia  grey,  on 
inner  margin  orange-ochreous. 


BY    A.    JEFFERIS    TURNER.  95 

Towiisville,  Q.;  in  February;  two  specimens  received  from  Mr. 
F.  P.  Dodd,  and  I  have  seen  others. 
Allied  to  A.  transversa,  Gn. 

Homopterinae. 

Ophiusa  trophidota,  n.sp. 

(J9.  63-65  mm.  Head  and  palpi  brown.  Antennae  whitish. 
Thorax  brown,  a  fine  whitish  transverse  line  before  apex,  similar 
lines  across  patagia.  Abdomen  grey.  Legs  grey.  Forewings 
triangular,  costa  moderately  arched,  more  strongly  before  apex, 
apex  acute,  terraen  sinuate  beneath  apex,  bowed  on  vein  4, 
oblique;  brown;  a  straight  ochreous-whitish  line  from  costa  near 
base  to  median  vein;  a  straight,  slightly  waved,  deep  brown  line, 
margined  with  ochreous-whitish  from  costa  at  J  to  dorsum  at  f ; 
a  deep  brown  median  fascia  bounded  anteriorly  by  a  line,  from  4 
costa  to  f  dorsum,  posteriorly  by  an  ochreous-whitish  line  from 
costa  at  f,  angled  in  disc,  thence  slightly  inwardly  curved  to 
dorsum  at  J;  three  whitish  dots  on  costa  beyond  this;  a  small 
dark  fuscous  subapical  dot ;  a  terminal  series  of  minute  dark 
fuscous  dots  between  veins:  cilia  brownish,  with  a  pale  basal  line. 
Hindwings  with  termen  rounded;  fuscous;  a  whitish  line  from 
inner  margin  near  tornus  obliquely  towards  mid-costa,  lost  in 
disc;  a  similar  line  from  tornus  to  opposite  mid-termen;  a  narrow 
whitish  suffusion  on  termen,  with  a  row  of  minute  dark  fuscous 
•dots  between  veins;  cilia  grey,  with  a  whitish  basal  line. 

Brisbane,  Q.;  three  specimens. 

Noctuinae. 
Crioa  nycterina,  n.sp. 

^.  38  mm.  Face  and  palpi  dark  fuscous,  mixed  with  ochreous- 
whitish;  lower  edge  of  frontal  projection  and  a  line  behind  eyes 
pinkish.  Antenna?  pale  ochreous-fuscous,  partly  annulated  with 
blackish.  Thorax  dark  fuscous,  with  a  moderate  central  crest. 
Abdomen  grey,  mixed  with  ochreous-whitish;  a  dark  fuscous  spot 
on  dorsum  at  base.  Legs  dark  fuscous,  irrorated  and  annulated 
with  whitish.      Forewings  elongate  triangular,  costa  very  slightly 


96         NEW  GKNERA  AND  SPECIES  OF  LEPIDOPTERA, 

arched,  apex  rather  acute,  hindmargin  slightl}^  rounded,  wavy, 
very  oblique;  dark  fuscous,  suffused  with  brown,  with  a  few 
scattered  whitish  scales,  markings  black;  an  acutely  angulated 
line  from  costa  near  base  as  far  as  fold:  a  similar  line  traceable 
with  diificulty  from  costa  at  ^  to  inner  margin  at  -|;  an  oblique 
blackish  line  in  disc  followed  by  a  whitish  line  representing 
reniform;  a  line  from  middle  of  costa  very  obliquely  outwards, 
then  bent  parallel  to  termen,  and  again  bent  backwards  beneath 
reniform,  and  thence  acutely  dentate  to  inner  margin  at  §;  three 
or  four  white  dots  on  apical  fourth  of  costa;  a  blackish  subterminal 
shade  posteriorly  dentate;  a  fine  terminal  line  interrupted  by 
whitish  dots,  which  are  connected  by  fine  blackish  lines  with  sub- 
terminal  shade:  cilia  dark  fuscous,  apices  dotted  with  whitish. 
Hindwings  with  termen  rounded,  w^avy ;  fuscous ;  basal  half 
ochreous-whitish  ;  cilia  whitish,  barred  with  fuscous.  Under 
surface  of  fore-  and  hindwings  with  crescentic  fuscous  discal  spots 
and  fuscous  terminal  bands. 

Brisbane,  Q. ;  one  specimen. 

Closely  allied  to  Crioa  acronyctina,  Butl.,  but  fore  wings  much 
darker,  abdominal  crest  less  developed,  and  hindwings  sharply 
divided  between  ochreous-whitish  and  fuscous. 

Crioa  leptochroma,  n.sp. 

(J9.  30  mm.  Head  whitish,  mixed  with  pale  greenish-fuscous; 
face  whitish.  Palpi  whitish,  mixed  with  a  few  dark  fuscous 
scales;  external  surface  of  second  joint  dark  fuscous  at  base. 
Antennae  grey;  towards  base  whitish,  with  a  few  dark  fuscous 
transverse  bars.  Thorax  with  a  strong  erect  crest  extending 
from  middle  to  posterior  edge:  whitish,  mixed  with  brownish; 
posterior  aspect  of  crest  irrorated  with  dark  fuscous.  Abdomen 
whitish-ochreous,  mixed  with  grey.  Legs  dark  fuscous,  mixed 
with  whitish;  tarsi  annulated  with  whitish;  posterior  pair  mostly 
whitish.  Forewings  elongate-triangular,  costa  moderately  arched, 
apex  round-pointed,  hindmargin  rounded,  obliciue;  white,  suffused 
with  fuscous  and  greenish;  a  fine  irregularly  dentate  line  from 


BY    A.    JEFFERIS    TURNER.  97 

costa  at  J  to  inner  margin  at  |;  reniform  represented  by  a  fine 
dark  fuscous  circular  I'ing,  enclosing  a  small  pale  fuscous  spot; 
second  line  from  costa  at  |  obliquely  outwards,  wavy,  curved  in  disc, 
and  ending  in  inner  margin  at  §;  this  is  succeeded  by  a  greenish 
line,  a  fuscous  shade,  and  a  greenish  subterminal  shade;  these  are 
interrupted  by  whitish  veins  dotted  with  dark  fuscous  :  cilia 
whitish-grey,  bases  grey,  interrupted  with  whitish  opposite  veins. 
Hindwings  with  termen  rounded,  slightly  wavy;  pale  j^ellowish; 
a  broad  fuscous  terminal  band  narrowing  to  tornus;  above  tornus 
is  an  elongate  white  spot,  interrupted  by  a  few  dark  fuscous  scales; 
cilia  ochreous- whitish,  with  a  fine  median  grey  line;  at  tornus 
dark  fuscous. 

Gympie,  Q.;  in  January;  two  specimens  received  from  Mr.  R. 
Illidge. 

CORULA    OCTOGESIMA,   n.sp. 

(J.  29  mm.  Head,  palpi,  and  thorax  whitish-grey,  irrorated 
Avith  darker  grey;  second  joint  of  palpi  with  appressed  scales. 
Antennae  ochreous-fuscous,  towards  base  whitish-grey;  in  male 
with  fine  bristles  (1)  and  short  ciliations  (|)  near  base,  thence 
broadly  laminate.  Abdomen  ochreous-whitish.  Legs  whitish, 
irrorated  with  grey.  Forewings  elongate,  posteriorly  dilated, 
costa  slightly  arched,  apex  rounded,  termen  sinuate,  scarcely 
oblique;  gre}'',  mixed  with  whitish,  especially  towards  base  and 
termen,  with  a  few  scattered  ochreous-brown  scales,  markings 
dark  fuscous;  a  suffused  spot  near  base  of  costa  representing  basal 
line;  a  transverse  line  from  costa  at  ^  to  dorsum  at  |,  rather 
broadly  suffused  near  costa  ;  two  adjacent  fine  circular  rings 
arranged  transversely  before  middle  of  disc,  and  a  third  reniform 
ring  beyond  centre;  each  ring  contains  a  dark  fuscous  centre,  and 
the  whole  form  the  figure  80;  a  fine  line  from  costa  at  |,  at  first 
outwards,  then  parallel  to  termen  and  waved  to  dorsum  at  |;  an 
interrupted,  thick,  wavy  line  at  f;  a  fine  subterminal  line :  cilia 
whitish.  Hindwings  with  termen  sinuate ;  whitish ;  faintly 
suft'used  with  grey  towards  termen;  cilia  whitish. 

Brisbane,  Q.;  one  specimen. 


98  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

CoRULA   DISPILA,  11. Sp. 

(J.  27  mm.  Head  whitish-grey.  Palpi  whitish,  external  sur- 
face irrorated  with  dark  fuscous;  second  joint  with  long  spreading 
scales  at  apex.  Antennse  grey;  in  malew4th  a  pair  of  bristles  on 
each  joint  ("i)  and  evenly  ciliated  (1),  towards  apex  simple. 
Thorax  grey,  mixed  with  whitish  and  dark  fuscous.  Abdomen 
grey.  Legs  grey;  posterior  pair  white:  anterior  tarsi  dark  fuscous, 
aniiulated  with  white.  Forewings  elongate-oblong,  costa  strongly 
arched  near  base,  thence  nearl}^  straight,  apex  rectangular,  termen 
slightly  rounded,  hardly  oblique;  pale  grey,  mixed  with  white, 
markings  dark  fuscous;  a  fine  angulated  basal  line  from  costa  to 
dorsum;  a  line  from  costa  at  J  to  dorsum  at  J;  a  squarish  ring 
enclosing  a  whitish  area  represents  the  orbicular,  a  similar  more 
elongate  dark-centred  ring  the  reniform;  a  line  from  costa  beyond 
middle,  bent  outwards  at  a  right  angle  beneath  costa,  and  twice 
again  in  disc,  so  as  to  partly  enclose  reniform,  thence  to  dorsum 
at  J;  an  interrupted  acutely  dentate  transverse  line  at  f,  ending 
in  a  dark  fuscous  suffusion  at  tornus;  a  black  interrupted  sub- 
terminal  line,  and  a  fine  black  terminal  line:  cilia  whitish. 
Hind  wings  with  termen  nearl}^  straight  to  vein  2,  thence  curved, 
slightly  wavy;  grey,  towards  base  paler;  cilia  whitish. 

Brisbane,  Q.;  in  November;  one  specimen. 

COKULA    METABLETA,   n.sp. 

(^9-  24-28  mm.  Head  grey,  sometimes  brownish.  Palpi  grey. 
Antennae  grey;  in  male  shortly  laminate,  moderately  ciliated  (1), 
with  longer  bristles  (3).  Thorax  grey,  mixed  with  fuscous. 
Abdomen  grey;  rarely  pinkish-tinged.  Legs  grey,  irrorated,  and 
tarsi  annulated,  with  dark  fuscous.  Forewings  elongate-triangular, 
costa  moderately  and  evenl}^  arched,  apex  round-pointed,  termen 
rounded,  slightly  obli(j[ue;  gre}',  mixed  with  whitish-grey;  lines 
dark  fuscous,  varying  in  intensity  in  different  specimens  :  an 
outwardly  curved  line  near  base,  usually  distinct  only  near  costa; 
an  outwardly  curved  line  from  I  costa  to  J  dorsum,  sometimes 
nearly  obsolete,  sometimes  thickened  and  conspicuous;  orbicular 


BY    A.    JEFFERIS    TURNER.  99 

and  reniform  spots  whitish-grey,  with  fuscous  margins  and  centres, 
sometimes  very  distinct,  sometimes  partly  obsolete;  a  posterior 
outwardly  curved  line,  with  two  slightly  developed  obtuse  denta- 
tions from  §  costa  to  |  dorsum;  followed  by  a  fuscous  shade 
mixed  with  brownish,  sometimes  obsolete;  an  interrupted  blackish 
subterminal  line:  cilia  grey.  Hindwings  with  termen  sigmoid; 
grey,  rarely  brown-pinkish;  with  a  broad  fuscous  terminal  band- 
cilia  whitish-grey,  rarely  pinkish-tinged. 

Brisbane,  Q.;  four  specimens  taken  by  Mr.  R.  lUidge. 

A  Variable  species.  It  may  be  distinguished  from  Corula 
dispila,  which  it  closely  resembles,  by  the  proportionately  longer 
forewings,  with  costa  not  incurved  towards  base,  and  with 
differently  shaped  posterior  line,  the  more  sigmoid  termen  of 
hindwings,  and  the  laminate  antennae  of  the  male. 

Corula  mesospila,  n.sp. 

(J9.  21-24  mm.  Head,  palpi,  and  thorax  whitish,  with  a  very 
few  scattered  dark  fuscous  scales;  second  joint  of  palpi  with 
appressed  scales.  Antennae  fuscous;  in  male  with  long  bristles 
(3),  absent  towards  apex,  and  even  ciliations  (1).  Abdomen 
whitish-grey,  or  ochreous-whitish.  Legs  whitish,  irrorated  with 
dark  fuscous;  posterior  pair  whitish;  anterior  tibiae  annulated 
with  dark  fuscous.  Forewings  elongate-triangular,  costa  gently 
arched,  apex  rather  acute,  termen  slightly  rounded,  slightly  oblique; 
whitish,  mixed  with  pale  ochreous-brown,  with  a  few  scattered 
black  scales,  markings  black;  a  very  short  streak  from  base;  a 
line  from  costa  near  base  reaching  as  far  as  fold;  a  line  from  costa 
atito  dorsum  at  ^;  orbicular  and  reniform  spots  faintly  outlined, 
the  latter  sometimes  dark-centred;  a  posterior  line  from  costa 
beyond  middle  obliquely  outwards,  then  sharply  bent  round  so  as 
to  form  an  obtuse  projection,  below  which  it  is  slightly  inwardly 
curved  to  dorsum  at  |;  a  whitish  line  incompletely  margined  with 
blackish  dots  at  |^;  an  interrupted  subterminal,  and  a  fine  terminal 
line:  cilia  whitish,  bases  mixed  with  grey.  Hindwings  with 
termen  nearly  straight  to  vein  2,  there  strongly  curved;  grey; 
cilia  whitish. 


100  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

Brisbane,  Q.;  in  September;  three  specimens. 

Allied  to  the  preceding,  from  which  it  may  be  distinguished 
by  the  different  shape,  brownish  irroration,  and  differently  formed 
lines  of  fore  wings. 

CORULA    STENOPHYLLA,   n.Sp. 

X.  22  mm.  Head  and  palpi  grey;  second  joint  of  palpi  with 
appressed  scales.  Antennae  grey;  male  with  long  bristles  (3), 
absent  towards  apex,  and  even  ciliations  (1).  Thorax  grey,  mixed 
with  whitish-grey;  some  dark  fuscous  scales  at  base  of  posterior 
crest.  Abdomen  ochreous-whitish.  Legs  gre}',  mixed  with 
whitish ;  posterior  pair  whitish.  Forewings  narrow-elongate, 
costa  very  slightly  arched,  apex  round-pointed,  termen  slightly 
rounded,  moderately  oblique;  grey,  mixed  with  whitish,  with  a 
small  amount  of  pale  brownish  suffusion  in  disc,  lines  dark  fuscous; 
a  faint  line  from  costa  near  base  barely  reaching  fold;  a  better 
marked  line  from  costa  at  ^  to  fold;  a  slightl}^  wavy  line  from 
costa  at  J  to  dorsum  at  J,  preceded  by  a  whitish  line;  a  line  from 
costa  at  |,  slightly  wavy,  curved  slightly  outwards  in  disc,  then 
straight  to  dorsum  at  |,  followed  by  a  whitish  line;  an  irregular 
anteriorly  dentate  blackish  line  at  i,  followed  by  a  whitish  line; 
orbicular  and  reniform  obsolete;  three  fuscous  dots  on  apical 
third  of  costa;  a  blackish  subterminal  and  a  fine  terminal  line: 
cilia  whitish  grey.  Hind  wings  with  termen  sigmoid;  ochreous- 
whitish;  a  terminal  grey  band  not  reaching  tornus;  cilia  whitish. 

Brisbane,  Q.;  in  December;  one  specimen. 

CORULA    EREMNA,  U.sp. 

(J.  23  mm.  Head  and  palpi  dark  grey,  irrorated  with  dark 
fuscous;  second  joint  of  palpi  much  dilated  with  loose  spreading 
scales  towards  apex.  Antennae  fuscous;  in  male  with  long  bristles 
(3),  absent  towards  apex,  and  even  ciliations  (^).  Thorax  dark 
grey,  irrorated  with  dark  fuscous.  Abdomen  grey.  Legs  grey, 
annulated  with  dark  fuscous;  posterior  pair  whitish,  with  a  few 
irrey  scales.  Forewings  elongate-triangular,  costa  slightly  arched, 
apex  round-pointed,  termen  rounded,  moderatel}'  oblique;  grey, 


BY    A.    JEFFERIS    TURNEK.  101 

irrorated  with  dark  fuscous,  witli  a  few  whitish  scales,  markings 
dark  fuscous  or  black;  a  dot  on  base  of  dorsum;  an  inwardly 
curved  line  from  costa  near  base  to  fold;  a  straight  line  from  costa 
at  J  to  dorsum  at  J;  orbicular  and  reniform  spots  faintly  outlined; 
a  line  from  costa  beyond  middle  very  obliquely  outwards,  bent 
twice  at  right  angles  in  disc  to  form  a  projection  enclosing 
reniform,  thence  inwardly  curved  to  dorsum  at  |;  an  oblique  line 
from  near  apex,  much  angulated  in  disc,  to  tornus;  an  interrupted 
subterminal,  and  a  fine  terminal  line :  cilia  grey,  apices  whitish- 
grey,  Hindwings  with  termen  slightly  wavy,  straight  to  vein  3; 
gre}'',  towards  base  whitish;  cilia  whitish. 
Killarney,  Q.;  in  October;  one  specimen. 

CORULA    ABEB^A,   n.sp. 

^9.  30-34  mm.  Head  whitish,  irrorated  with  pale  brownish- 
ochreous.  Palpi  whitish,  irrorated  with  pale  brownish-ochreous 
and  dark  fuscous;  terminal  joint  mostly  dark  fuscous;  second  joint 
with  appressed  scales.  Antennae  whitish-grey;  in  female  whitish- 
ochreous;  in  male  with  long  bristles  (4),  absent  towards  apex,  and 
even  ciliations  (IJ)-  Thorax  whitish,  irrorated  with  pale 
brownish-ochreous,  a  few  dark  fuscous  scales  posteriorly.  Abdo- 
men whitish,  suffused  with  pale  brownish-ochreous,  somewhat 
pinkish-tinged.  Legs  whitish,  mixed  with  pale  brownish-ochreous 
and  dark  fuscous.  Fore  wings  elongate-triangular,  costa  very 
slightly  arched,  apex  rectangular,  termen  slightly  sigmoid,  scarcely 
oblique;  white,  irrorated  with  whitish-grey  and  pale  brownish- 
ochreous,  markings  dark  fuscous;  a  minute  dot  at  base  of  dorsum; 
an  acutely  outwardly  angled  line  from  costa  near  base  to  fold;  an 
oblique  wavy  line  from  costa  at  ^  to  dorsum  at  J;  spots  obsolete, 
orbicular  represented  by  a  small  tuft  of  scales;  an  outwardly 
oblique  line  from  mid-costa,  lost  in  disc;  a  wavy  line  from  costa 
at  I  forming  a  moderate  double  projection  in  disc,  then  inwardly 
curved  to  dorsum  at  f ;  a  series  of  elongated  submarginal  black 
dots  ending  in  a  spot  above  tornus;  a  greyish  subterminal  line;  a 
row  of  black  dots  on  termen:  cilia  white,  with  a  few  grey  scales. 
Hindwings  with  termen  rounded,  slightly  sigmoid  beneath  apex; 


& 


102  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

grey,  towards  base  ochreous-whitish;  cilia  whitish-grey,  on  dorsum 
whitish  ochreous. 

Brisbane,  Q. ;  two  specimens,  and  I  huve  since  seen  others 
which  show  considerable  variation. 

ACANTHOLIPES    TOXICA,  n.sp. 

$.  20  mm.  Head,  palpi,  and  thorax  reddish-brown.  Antennae 
pale  brownish,  obscurely  annulated  with  dark  fuscous.  Abdomen 
rey ;  towards  base  mixed  with  brown.  Legs  whitish-brown, 
irrorated  with  dark  fuscous;  posterior  pair  whitish,  with  dark 
fuscous  spurs;  anterior  tarsi  dark  fuscous,  annulated  with  whitish. 
Forewings  elongate-triangular,  costa  scarcely  arched,  apex  acute, 
termen  concave  beneath  apex,  strongly  outwardly  bowed  between 
\'eins  3  and  4;  reddish-brown,  with  a  few  fuscous  scales;  several 
dark  fuscous  and  whitish  dots  on  costal  edge;  a  minute  white  dot 
margined  with  dark  fuscous  in  disc  before  middle;  reniform  white: 
cilia  reddish-brown,  apices  whitish.  Hindwings  with  termen 
rounded;  dark  gre}^;  cilia  pale  reddish-brown,  apices  whitish. 

Brisbane,  Q.;  one  specimen. 

Dahlia  capnobela,  n.sp. 

9.  27-29  mm.  Head,  thorax,  and  abdomen  ochreous-brown. 
Face  and  palpi  dark  fuscous.  Antennas  whitish-ochreous,  obscurely 
annulated  with  fuscous.  Legs  dark  fuscous;  posterior  pair  whitish- 
ochreous,  mixed  with  fuscous.  Forewings  rather  broadly  triangular, 
costa  slightly  arched  at  base,  thence  straight,  apex  rounded,  termen 
rounded,  slightly  oblique;  ochreous-brown;  a  fine  angulated  dark 
fuscous  line  from  costa  at  i  to  dorsum  at  ^,  with  an  obtuse  outward 
projection  belowmiddle,  a  dark-fuscous  discal  dot,  sometimes  white- 
centred  just  over  projection;  a  white  dot  with  fine  dark-fuscous 
edge  near  centre  of  disc;  a  fine  acutel}^  dentate  dark-fuscous  line 
from  costa  at  |  to  dorsum  at  |;  an  angulated  fuscous  shade 
followed  by  a  pale  line  at  4;  a  series  of  blackish  terminal  dots: 
cilia  ochreous-brown.  Hindwings  with  termen  rounded;  colour 
and  markings  as  forewings,  but  basal  line  wanting,  onl}"  one 
discal  dot,  and  that  not  white-centred. 

Brisbane,  Q.;  two  specimens. 


BY    A.    JEFFERIS    TURNER.  103 

Hyperlopha  aridela,  n.sp. 

$.  52  mm.  Head  ochreous-grey,  with  a  fine  anteriorly- angled 
white  line  between  antenna?.  Palpi  pale  grey,  with  a  few 
scattered  black  scales.  Antennae  white,  towards  apex  grey. 
Thorax  pale  grey,  anteriorly  ochreous-tinged,  sparsely  irrorated 
with  black  scales.  Abdomen  whitish-grey,  along  centre  ochreous- 
tinged,  with  sparsely  scattered  black  scales.  Legs  pale  grey,  with 
sparsely  scattered  black  scales.  Forewings  triangular,  costa  rather 
strongly  arched,  apex  rectangular,  termen  angled  on  vein  3,  above 
angle  straight,  beneath  wavy  and  very  oblique  to  tornus;  pale 
grey,  somewhat  purplish-tinged,  with  sparsely  scattered  black 
scales;  lines  faint,  ochreous-grey;  first  from  i  costa  to  J  dorsum; 
second  from  I  costa  to  1  dorsum;  third  from  J  costa  obliquely 
outwards,  angled  beneath  costa,  thence  wavy  to  dorsum  at  |;  a 
subterminal  shade,  containing  towards  dorsum  three  snow-w^hite 
spots  arranged  in  line,  margined  with  dark  brown,  and  jDOsteriorly 
with  an  outer  margin  of  whitish-ochreous,  followed  by  a  small 
dark  fuscous  suffusion:  cilia  pale  grey,  beneath  angle  fuscous- 
brown.  Hind  wings  with  termen  rounded;  dark  fuscous,  basal 
half  pale  ochreous;  an  elongate-oval  dull  white  blotch  from  tornus 
to  opposite  mid-termen;  cilia  white. 

Brisbane,  Q.;  in  December;  two  specimens. 

Dnopheropis,  n.g. 

Face  with  small  projecting  cone  of  hairs.  Palpi  moderate, 
obliquely  ascending,  scarcely  reaching  vertex ;  second  joint 
thickened  with  loosely  appressed  hairs  above  and  beneath ; 
terminal  joint  short,  stout,  obtuse,  partly  concealed.  Tongue 
present.  Antennae  of  male  with  a  double  row  of  pectinations 
extending  to  apex.  Thorax  with  a  vertical  anterior  crest. 
Abdomen  with  two  tufts  of  hairs  on  dorsum  of  first  segment,  and 
a  small  crest  of  loose  hairs  on  second  segment.  Legs  normal. 
Forewings  with  vein  2  from  §,  3  from  near  angle,  6  from  upper 
angle,  7,  8,  9  stalked,  10  connected  by  a  bar  with  7,  8,  9  to  form 
areole,  7  arising  from  8,  9  well  beyond  areole.     Hindwings  with 


104        NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

3,  -4  approximated  at  base,  5  bent  towards  4  at  base,  6,  7  connate, 

8  anastomosing  with  cell  near  base. 

Sir  G.  Hampson  was  unable  to  refer  the  type  to  any  descriljed 

genus. 

Dnopheropis  scoTiEA,  n.sp. 

(J.  23  mm.  Head  fuscous.  Palpi  dark  fuscous.  Antenna" 
grey;  in  male  with  long  pectinations  (8-9).  Thorax  dark  fuscous, 
collar  paler.  Abdomen*  whitish-gre3^  Legs  dark  grey.  Fore- 
wings  elongate-triangular,  costa  moderately  arched,  apex  rectan- 
gular, termen  straight,  rounded  beneath,  scarcely  oblique:  dark 
fuscous,  markings  black;  a  fair-sized  dot  in  disc  at  |,  and  another 
in  middle,  the  latter  containing  some  whitish-ochreous  scales;  a 
row  of  .subterminal  dots,  forming  an  outwardly  curved  line, 
obsolete  towards  margins :  cilia  dark  fuscous.  Hindwings  broad, 
termen  rounded,  slightly  indented  beneath  apex;  whitish;  cilia 
whitish. 

Brisbane,  Q.;  one  specimen  taken  by  Mr.  R.  Illidge. 

Proscrana,  n.g. 

Face  flat.  Palpi  stout,  moderately  long,  porrected ;  second 
joint  loose-haired;  terminal  joint  short,  stout,  obtuse.  Antennte 
bipectinated  in  both  sexes  nearly  to  apex;  each  pectination  ending 
in  or  bearing  a  long  bristle.  Tongue  well  developed.  Thorax 
and  abdomen  not  crested.  Fore  wings  with  vein  2  from  |  or  |, 
3  from  angle,  6  from  upper  angle  of  cell,  7,  8,  9  stalked,  10  con- 
nected by  a  Ijar  with  8,  9  opposite  origin  of  7.  Hindwings 
normal. 

Proscrana  tephropis,  n.sp. 

-J^.  25-32  mm.  Head  and  palpi  whitish,  mixed  with  grey. 
Antennae  white,  with  a  few  dark  fuscous  scales;  pectinations  dark 
fuscous;  in  male  8,  in  female  4.  Thorax  whitish,  mixed  with 
grey  anteriorly.  Abdomen  whitish.  Legs  whitish-grey,  irrorated 
with  grey;  anterior  femora  and  tibiae  clothed  with  long  hair  in 
both  sexes.  Forewings  elongate-triangular,  costa  arched  at  base, 
thence  straight  to  near  apex  where  it  is  abruptly  arched,  apex 


BY    A.    JEFFERIS    TURNER.  105 

pointed,  termen  aiigulated  on  ^  ein  4,  slightly  concave  above  angle 
and  slightly  sinuate  beneath;  whitish-grey;  a  whitish  streak  along 
costa,  narrowing  to  a  point  at  f ,  irrorated  with  a  few  dark  fuscous 
scales;  in  female  a  faint  posterior  line  from  a  dark  fuscous  dot  on 
costa  at  |-  to  dorsum  at  J,  obsolete  in  male:  cilia  whitish,  slightly 
pinkish-tinged,  barred  with  dark  fuscous,  bases  dark  fuscous. 
Hind  wings  with  termen  rounded,  whitish-grey;  cilia  whitish. 

Townsville,  Q.;  in  January;  four  bred  specimens  received  from 
Mr.  F.  P.  Dodd,  who  informs  me  that  the  larvae  feed  on  the  young- 
foliage  of  IScyphiphora  hydrojyhylacea. 

Proscrana  poliophylla,  n.sp. 

9.  36  mm.  Head  and  palpi  whitish-grey,  irrorated  with  grey. 
Antennae  whitish-gre}';  pectinations  in  female  1^;  a  long  bristle 
arises  from  about  the  middle  of  the  posterior  aspect  of  each  pectina- 
tion. Thorax  and  abdomen  whitish-grey.  Legs  whitish-grey. 
Forewings  triangular,  costa  moderately  arched,  apex  acute,  termen 
angulated  on  vein  4,  above  angulation  concave,  beneath  straight; 
whitish-grey ;  a  posterior  grey  line  from  costa  at  |  becoming 
obsolete  in  disc:  cilia  whitish-grey,  mixed  with  fuscous.  Hind- 
wings  with  termen  bowed  on  vein  4;  whitish-grey;  cilia  whitish, 
with  several  dark  fuscous  interruptions. 

Warwick,  Q.  ;  in  October,  one  specimen.  When  taken  it 
carried  the  wings  erect,  and  had  a  very  Geometrid  appearance. 

Very  similar  to  the  preceding,  but  shape  of  forewings  is 
diiferent,  posterior  line  is  nearer  base,  and  antennie  and  forelegs 
also  differ. 

A  X  I  0  c  T  E  T  A,  n.g. 

Face  with  rounded  projection.  Palpi  long  (4-5);  second  joint 
obliquely  ascending,  reaching  level  of  vertex,  clothed  beneath 
with  long  scales  ;  terminal  joint  well  developed,  down-curved, 
obtuse.  Tongue  well  developed.  Thorax  and  abdomen  without 
crests.  Posterior  tibiae  with  two  pairs  of  spurs.  Forewings  with 
termen  excavated  beneath  apex  and  acutely  angled  on  vein  4; 
vein  2  from  |,  3  from  before  angle,  6  from  below  upper  angle  of 
cell,  7,  8,  9  stalked,  10  anastomosing  with  8,  9  from  just  beyond 


106  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

7  to  near  bifurcation.     Hindwings  with  vein  5  well  developed, 
ai^proximated  to  4  at  base. 

AXIOCTETA   OENOPLEX,   n.sp. 

9.  28  mm.  Head  purplish-gre3^  Palpi  purplish-grey,  with  a 
few  dark  fuscous  scales.  Antennae  grey.  Thorax  reddish-brown, 
anteriorly  purplish-grey.  Abdomen  vinous-purple,  beneath 
ochreous-whitish.  Legs  purple-grey,  irrorated  with  dark  fuscous; 
posterior  pair  ochreous-whitish  above.  Forewings  triangular, 
costa  slightly  arched,  apex  acute,  falcate,  termen  excavated  beneath 
apex,  acutely  angled  on  vein  4;  reddish-brown,  towards  termen 
suffused  with  ochreous-grey;  a  pair  of  faint  fuscous  dots  arranged 
transversely  before  middle  of  disc;  a  subterminal  row  of  similnr 
dots:  cilia  purple-fuscous,  apices  white.  Hindwings  with  termen 
rounded;  vinous-purple,  towards  base  and  costa  ochreous-whitish; 
a  subterminal  row  of  fuscous  dots;  cilia  purple-whitish. 

Brisbane,  Q.;  in  April,  one  specimen. 

A  L  A  P  A  D  N  A,   n.g. 

Face  with  projecting  scales.  Palpi  long  (4-5),  porrect;  second 
joint  very  long,  expanded  above  by  long  rough  scales;  terminal  joint 
slender,  short,  obtuse.  Tongue  present.  Antennae  in  male 
ciliated.  Thorax  smooth.  Abdomen  with  a  small  dorsal  crest. 
Posterior  tibiae  with  two  pairs  of  spurs.  Forewings  with  vein  2 
from  |,  3  from  before  angle,  6  from  well  before  upper  angle  of 
cell,  7  from  upper  angle,  8,  9  stalked,  10  absent.  Hindwings 
with  3  and  4  long-stalked,  5  well  developed,  approximated  to  3, 
4  at  base. 

Alapadna  pauropis,  n.sp. 

(J9-  15-17  mm.  Head  and  palpi  ochreous-grey,  with  a  few 
fuscous  scales.  Antennae  whitish-ochreous.  Thorax  ochreous- 
brownish,  with  a  dark  fuscous  posterior  spot.  Abdomen  whitish- 
ochreous;  crest  dark  fuscous.  Legs  dark  fuscous,  mixed  with 
ochreous-whitish;  posterior  pair  mostly  ochreous-whitish.  Fore- 
wings elongate-triangular,  costa  gently  arched,  apex  round- 
pointed,    termen    rounded,    oblique ;    ochreous-brown,    markings 


BY    A.    JEFFERIS    TURNEK.  107 

darker  brown,  somewhcat  variable;  a  basal  Kiie  from  i  costa  to  \ 
dorsum;  a  suffused  median  line,  sometimes  expanded  into  a  broad 
fascia,  immediately  followed  by  a  more  or  less  "crescentic  dark- 
edged  white  discal  dot;  a  suffused  angulated  subterminal  line, 
sometimes  preceded  by  one  or  two  finer  lines;  a  row  of  small  dark 
fuscous  terminal  dots :  cilia  ochreous-brown,  irrorated  with  dark 
brown.  Hind  wings  with  termen  rounded;  whitish  or  grey;  a 
crescentic  mark  followed  by  a  curved  median  line  grey  ;  an 
interrupted  grey  terminal  line;  cilia  ochreous-whitisli  or  grey. 
Brisbane,  Q.;  from  September  to  December,  not  uncommon. 

PrONOPIIORA    TRITOSTICHA,   n.Sp. 

(J.  22  mm.  Head  pale  brownish-ochreous,  sides  white.  Palpi 
pale  brownish-ochreous;  apex  of  second  joint  and  terminal  joint 
white.  Antennse  whitish;  in  male  bipectinate  (2|),  towards  apex 
simple,  with  a  long  bristle  arising  from  stalk  behind  each  pecti- 
nation. Thorax  whitish ;  collar  and  bases  of  patagia  pale 
brownish-ochreous.  Abdomen  whitish.  Legs  whitish;  anterior 
pair  grey  on  inner  and  lower  surfaces.  Fore  wings  narrow  elon- 
gate ;  costa  gently  arched,  apex  rounded,  termen  obliquely 
rounded;  pale  brownish-ochreous,  with  three  shining  snow-white 
longitudinal  streaks;  first  from  base  to  apex,  narrowly  margined 
with  black  beneath,  near  termen  the  marginal  line  separates  from 
streak;  second  from  disc  at  ^,  anteriorly  acute,  to  termen  at  J; 
third  from  base  along  fold  not  quite  reaching  tornus;  a  row  of 
black  dots  close  to  termen:  cilia  ochreous- whitish.  Hindwing 
with  termen  sinuate ;  white,  with  a  scanty  grey  suffusion  ;  a 
narrow  grey  terminal  line;  cilia  white. 

Townsville  and  Bundaberg,  Q.;  two  specimens. 

^  T  H  A  L  I  N  A,   n.g. 

Face  with  projecting  scales.  Palpi  long  (3-4);  second  joint 
with  spreading  loose  scales  above  and  below,  obliquely  upturned; 
terminal  joint  well  developed,  stout,  obtuse,  porrect.  Tongue 
present.  Thorax  smooth.  Abdomen  with  a  basal  crest.  Posterior 
tibite  w^ith  two  pairs  of  spurs.     Fore  wings  with  vein  2  from  |,  3 


108  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

from  f,  4  from  angle,  6  from  below  upper  angle  of  cell,  7  short- 
stalked  with  8,  9,  10  connected  by  a  bar  with  8,  9  opposite  7. 
Hindwings  with  5  well  developed,  very  closely  approximated  to  4 
at  base,  8  anastomosing  with  cell  near  base  only. 

-^THALINA    ASAPHES,  n.sp. 

2'  19mm.  Head,  palpi,  and  thorax  fuscous.  Antennae  fuscous, 
with  obscure  blackish  annulations.  Abdomen  fuscous  ;  basal 
crest  dark  fuscous.  Legs  fuscous;  posterior  pair  and  middle  tarsi 
whitish  ;  anterior  tarsi  anniilated  with  whitish.  Forewings 
elongate-triangular,  costa  rather  strongly  arched,  apex  rounded, 
termen  slightly  rounded,  scarcely  oblique  ;  fuscous  strigulated 
witii  darker  fuscous;  costal  edge  paler;  a  suffused  discal  dot 
beneath  ^  of  costa;  an  incomplete  line  of  white  dots  from  disc 
beneath  costa  at  4,  angulated  outwards,  then  incurved  to  dorsum 
at  §;  near  dorsum  at  ^  and  before  dorsal  part  of  this  line  are 
some  red  scales;  an  interrupted  dark  fuscous  terminal  line,  near 
which  are  some  red  scales:  cilia  fuscous,  with  a  row  of  basal 
whitish  dots.  Hindwings  with  termen  rounded;  grey;  cilia  grey; 
bases  whitish. 

Card  well,  Q.;  in  August,  one  specimen. 

Eakias  ochrophylla,  n.sp. 

(J9.  18-21  mm.  Head  and  palpi  whitish,  faintly  yellowish- 
tinged.  Antennae  whitish,  towards  apex  grey.  Thorax  pale 
yellow.  Abdomen  whitish.  Legs  whitish.  Forewings  elongate, 
costa  moderately  arched,  apex  round-pointed,  termen  scarcely 
rounded,  oblique;  pale  yellow,  greenish-tinged,  sometimes  partly 
suffused  with  pale  grey  between  veins;  sometimes  two  ol^lique 
lines  of  deeper  colour  in  terminal  part  of  disc:  cilia  pale  yellow. 
Hindwings  with  termen  rounded;  whitish;  cilia  whitish. 

Birchip,  Vic;  in  November;  two  specimens,  in  Coll.  Lyell. 

Erastrianse. 

COSMOPHILA    PSAMATHODES,   U.sp. 

(J.  38  mm.  Head,  thorax,  and  abdomen  pale  brownish-ochreous. 
Palpi    pale    brownish-ochreous,    irrorated    with    reddish-brown. 


BY    A.    JEFFERIS    TURNER.  109 

Antennse  ochreous-fuscous,  towards  base  paler;  in  male  minutely 
ciliated  (J)  with  longer  bristles  (f).  Legs  whitish-ochreous, 
irrorated  with  reddish-brown.  Fore  wings  elongate-triangular, 
costa  nearly  straight,  apex  acute,  termen  wav}'-,  acutely  angled  on 
vein  4,  excavated  above  and  below  angle,  oblique:  pale  brownish- 
ochreous;  a  darker  line  from  costa  at  ^  to  dorsum  at  I;  immedi- 
ately followed  beneath  costa  by  a  minute  white  discal  dot;  a 
similar  line,  better  marked,  from  costa  at  |,  angulated  outwards 
close  to  costa,  and  continued  as  an  inwardly  oblique  incurved  line 
to  dorsum  at|^;  a  minute  blackish  discal  dot  beneath  costa  before 
second  line;  a  faint  subterminal  shade:  cilia  ochreous-brown, 
apices  whitish,  interrupted  by  fuscous.  Hindwings  with  termen 
wav}^,  rounded ;  ochreous,  suffused  with  reddish-brown  ;  cilia 
whitish-ochreous,  obscurel}'  barred  with  reddish-brown. 
Stradbroke  Island,  Q.;  one  specimen. 

Erastria  bryistis,  n.sp. 

(J.  30  mm.  Head  pale  green,  posterior  margin,  face,  and  palpi 
brown.  Antennae  grey,  towards  base  paler;  in  male  minutely 
ciliated  (J).  Thorax  with  a  rounded  posterior  crest ;  brown, 
mixed  with  pale  green.  Abdomen  with  fourth  and  fifth  segments 
slightly  crested;  ochreous-whitish,  irrorated  with  dark  fuscous. 
Legs  ochreous-whitish  irrorated,  and  tarsi  annulated  with  dark 
fuscous.  Fore  wings  elongate-triangular,  costa  gently  arched, 
apex  rounded,  termen  rounded,  slightly  oblique  ;  pale  green, 
central  part  of  disc  except  near  base  brown;  lines  dark  fuscous; 
incomplete  fine  angulated  basal  and  subbasal  lines  ;  a  fine 
angulated  outwardly-curved  line  from  costa  at  J  to  dorsum  at  J; 
this  is  followed  by  a  broad  fuscous  sufFusioti  from  costa  before 
middle  obliquely  to  tornus;  a  finely  dentate  posterior  line  obsolete 
towards  costa;  a  broad  dark  fuscous  longitudinal  streak  aboAe 
dorsum  interrupted  at  transverse  lines:  a  fine  interrupted  terminal 
line;  orbicular  and  reniform  distinctly  outlined:  cilia  greenish- 
white,  barred  with  brown.  Hindwings  with  termen  wavy, 
rounded;  fuscous,  towards  base  paler;  a  darker  discal  dot,  and 


110  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

posterior  line  immediately  followed  by  a  whitish  line;  an  inter- 
rupted white  line  close  to  termen;  cilia  fuscous,  apices  whitish. 
Brisbane,  Q.;  one  specimen  received  from  Mr.  F.  P.  Dodd. 

Erastroides  lichnomima,  n.sp. 

(J.  26  mm.  Head  and  thorax  white,  mixed  with  dark  fuscous 
and  green ;  thorax  with  a  small  posterior  crest.  Palpi  dark 
fuscous,  terminal  joint  and  apex  of  second  joint  white.  Antennse 
whitish-ochreous,  with  a  fe\v^  dark  fuscous  scales;  in  male  simple. 
Abdomen  dark  fuscous,  sides  and  apex  whitish-brown  [crests 
probably  denuded].  Legs  dark  fuscous,  irrorated  and  annulated 
with  white.  Forewings  narrow  elongate-triangular,  costa  nearly 
straight,  apex  round -pointed,  termen  rounded,  oblique;  whitish, 
irrorated  with  greenish  and  dark  fuscous,  lines  dark  fuscous;  a 
line  from  costa  near  base  to  fold;  a  dentate  line  from  costa  at  j 
to  dorsum  at  4 ;  a  whitish  mark  from  mid-costa,  enclosing  some 
dark  fuscous  scales,  and  edged  in  disc  with  dark  fuscous  to 
represent  reniform;  a  longitudinal  streak  in  terminal  half  of  disc 
interrupted  by  reniform;  a  dentate  line  from  costa  at  |  to  dorsum 
at  |-  slightly  outwardly  curved ;  a  whitish  subterminal  line 
followed  by  some  dark  streaks  between  veins;  veins  in  terminal 
area  outlined  in  greenish :  cilia  dark  fuscous,  mixed  with  white. 
Hind  wings  with  termen  rounded;  grey;  cilia  grey,  with  a  few 
whitish  scales. 

Brisbane,  Q.;  in  March,  one  specimen. 

HOMODES    ERIZESTA,   n.Sp. 

^.  22-23  mm.  Head,  palpi,  thorax,  and  abdomen  reddish- 
brown,  tinged  with  purple;  thorax  smooth;  abdomen  crested  on 
3rd  and  4th  segments,  crests  dark  purple-fuscous,  with  metallic 
lustre.  AntennLie  ochreous-brown;  in  male  slightly  serrate,  e\  enly 
ciliated  (1).  Legs  reddish-brown;  posterior  pair  whitish-ochreous. 
Forewings  elongate-triangular,  costa  nearly  straight,  strongly 
curved  near  apex,  apex  rounded,  termen  rounded,  oblique ; 
reddish-brown,  tinged  with  purple,  alternating  in  fine  regular 
strige  with  whitish;  a  line  from  costa  at  J  to  dorsum  at  J,  a  second 


BY    A.    JEFFERIS    TURNER.  HI 

from  costa  at  |  to  dorsum  at  f,  and  a  third,  better  marked,  from 
costa  at  i  outwardly  curved  to  dorsum  at  4,  slightly  darker  in 
shade  and  interspersed  with  shining  metallic  scales;  a  roundish 
ochreous  suJBfusion  in  costal  part  of  disc  before  posterior  line,  and 
a  broad  ochreous  suffused  line  from  beneath  costa  at  ^  to  tornus; 
a  subterminal  row  of  purplish  dots  containing  metallic  scales;  an 
ochreous  suffusion  along  termen  ;  two  fine  parallel  interrupted 
black  terminal  lines  :  cilia  reddish-purple.  Hindwings  with 
termen  rounded;  colour  and  markings  as  forewings;  inner  marginal 
cilia  whitish-ochreous. 

Townsville  and   Geraldton,  Q.;   from  November  to  January: 
several  specimens  received  from  Mr.  F.  P.  Dodd. 

HyELA    ARC40TYPA,  n.sp. 

(J9.  25-26  mm.  Head  and  thorax  grey,  irrorated  with  white. 
Palpi  fuscous,  with  a  few  whitish  scales.  Antennae  whitish- 
ochreous  ;  in  male  serrate,  ciliated  (1).  Abdomen  whitish, 
irrorated  with  whitish-grey.  Legs  grey,  irrorated  with  whitish. 
Forewings  elongate-triangular,  costa  straight,  apex  round-pointed, 
termen  bowed,  oblique;  grey,  mixed  with,  whitish  and  dark  fuscous, 
markings  clear  white;  a  short  suffused  line  from  base;  an  oblique 
line  from  beneath  i  of  costa  to  f  of  dorsum;  an  oblique  oblong 
spot,  edged  with  dark  fuscous,  representing  orbicular;  an  hour- 
glass-shaped spot  with  grey  centre  and  dark  fuscous  margins 
representing  reniform,  sometimes  a  longitudinal  streak  from  its 
lower  extremity  is  produced  towards  base,  beneath  and  touching- 
orbicular;  a  sigmoid  subterminal  line  from  costa  at  f  to  dorsum 
at  §,  followed  by  a  short  inwardly  oblique  streak  from  costa;  two 
subterminal  fuscous  suffusions,  one  above  and  one  beneath  centre 
of  disc,  follow  this  line;  a  dark  fuscous  terminal  line  immediately 
preceded  by  a  whitish  line:  cilia  grey,  ajDices  white.  Hindwings 
with  termen  rounded,  wavy  beneath  apex;  grey;  cilia  whitish, 
with  a  grey  basal  line. 

Brisbane  and  Warwick,  Q.;  two  specimens,  that  from  the  latter 
locality  in  October. 


112  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

Hyela  thapsina,  n.sp. 

(J.  18-20  mm.  Head  and  palpi  ochreous-yellow.  Antennae 
grey;  in  male  simple.  Thorax  with  a  small  rounded  posterior 
crest  ;  brownish-fuscous  ;  collar  and  patagia  ochreous-yellow. 
Abdomen  grey;  tuft  brownish-ochreous.  Legs  ochreous;  anterior 
tibiae  and  tarsi,  and  middle  tarsi  fuscous,  annulated  narrowly 
with  whitish-ochreous.  Forewings  elongate,  posteriorly  dilated, 
costa  straight,  apex  round-pointed,  termen  slightly  rounded, 
slightly  oblique  ;  ochreous-yellow  ;  a  broad  brownish-fuscous 
terminal  fascia,  anterior  edge  wavy  from  |-  costa  to  |  dorsum  : 
cilia  brownish-fuscous.  Hindwings  with  termen  rounded,  slightly 
indented  beneath  apex;  dark  grey,  near  base  pale  ochreous;  cilia 
pale  ochreous,  with  a  grey  basal  line. 

Townsville,  Q.;  in  February  and  March;  two  specimens  received 
from  INIr.  F.  P.  Dodd. 

Though  Sir  George  Hampson  considers  this  a  new  species,  I  am 
inclined  to  consider  it  a  variety  of  Tarache  detrita,  ButL 

Raparna  cyclostigma,  n.sp. 

(J9.  22-21  mm.  Head,  palpi,  and  thorax  gre}'.  AntennjB 
dark  fuscous;  in  male  minutely  ciliated  (\).  Abdomen  gre}', 
with  some  fuscous  scales.  Legs  fuscous,  irrorated,  and  tarsi 
annulated  with  whitish.  Forewings  elongate-triangular,  apex 
tolerably  pointed,  termen  rounded,  oblique;  grey;  black  dots  on 
costa  near  base  and  at  |,  sometimes  obsolete;  a  fine  wavy  grey 
line  from  second  dot  to  dorsum  at  ^;  orbicular  and  reniform  finely 
outlined;  a  fine  whitish  line  from  costa  at  |,  at  first  obliquely 
outwards,  forming  a  rounded  projection  in  disc,  thence  inwardly 
oblique  to  dorsum  at  ^;  on  the  anterior  margin  of  this  line  are  a 
variably  developed  series  of  black  spots,  sometimes  obsolete  ; 
veins  towards  termen  more  or  less  distinctly  outlined  with  ochreous- 
whitish  ;  a  dentate  grey  subterminal  shade  ;  a  row  of  black 
terminal  dots :  cilia  grey.  Hindwings  with  termen  scarcely 
rounded;  grey;  a  short  dark  fuscous  line  from  i  of  inner  margin; 
and  a  second  fainter  line  from  tornus;  a  series  of  terminal  black 
dots;  cilia  grey. 


BY    A.    JEFFERIS    TURNER.  113 

Townsville  and  Brisbane,  Q. ;  from  December  to  April;  five 
specimens. 

Allied  to  Raparna  nebulosa,  Moore,  from  which  it  may  be 
distinguished  by  the  rounded  projection  of  posterior  line.  It  is 
a  highly  variable  species,  no  two  of  my  specimens  being  alike.  Two 
deserve  mention — («)  with  an  elongate  black  fascia  preceding- 
posterior  line  and  obliterating  reniform;  (6)  without  black  spots, 
but  with  posterior  |  of  forewings  suffused  with  fuscous. 

CORGATHA    DICHIOXISTIS,   n.Sp. 

$.  14-18  mm.  Head,  thorax,  and  abdomen  brownish-grey. 
Palpi  brownish-grey,  irrorated  with  dark  fuscous.  Antennae 
whitish-ochreous,  towards  base  sometimes  annulated  with  dark 
fuscous;  in  male  simple,  ciliated  (2).  Legs  whitish-ochreous; 
anterior  tibi?e  grey.  Forewings  elongate-triangular,  costa  straight, 
slightly  arched  near  apex,  apex  pointed,  termen  angled  on  vein 
3,  slightly  concave  above  and  below  angle;  ochreous-grey,  irrorated 
with  metallic  scales;  an  oval  white  spot  on  costa  at  \,  and  a 
rounded  white  spot  before  |-;  a  transversely  elongate  small  discal 
spot  beneath  costa  before  middle,  dark  fuscous;  a  faint  wavj'  line 
from  costa  at  |  to  dorsum  at  |;  some  small  black  terminal  dots: 
cilia  dull  purplish,  apices  whitish.  Hindwings  with  termen 
rounded;  colour,  posterior  line,  terminal  dots,  and  cilia  as  fore- 
wings. 

Townsville  and  Brisbane,  Q.;  in  May  and  March;  two  specimens. 

CORGATHA    DIPYRA,    n.sp. 

(J9  20  mm.  Head  ochreous-grey;  face  and  palpi  dull  reddish. 
Antenna;  grey,  near  base  reddish;  in  male  simple,  ciliated  (1). 
Thoi-ax  ochreous-gre}',  with  metallic  reflections.  Abdomen  brown, 
mixed  with  dark  fuscous.  Legs  pale  ochreous  ;  anterior  arid 
middle  femora  and  tibise  fuscous.  Forewings  elongate-triangular, 
costa  straight,  apex  acute,  termen  angulated  on  vein  3,  slightly 
concave  above  and  below  angle;  reddish-purplish-grey;  an  inter- 
rupted dark  fuscous  suffusion  along  costa,  the  scales  having  a 
bluish-metallic  lustre;  a  few  similar  metallic  scales  scattered  in 


114  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

disc;  a  fuscous  discal  spot  beneath  mid-costa ;  a  slender  dentate 
fuscous  posterior  line  from  costa  at  4  to  dorsum  at  §;  termen  with 
a  narrow  dark  fuscous  suffusion  except  at  angles:  cilia  reddish- 
purple,  with  a  white  median  line  and  rosy  apices;  these  are 
interrupted  by  dark  fuscous  at  angles.  Hindwings  with  termen 
slightly  rounded;  colour  and  markings  as  forevvings,  but  posterior 
line  better  marked  and  double,  and  a  fine  subterminal  line. 

Brisbane,  Q.;  in  October  and  November;  two  specimens. 

This  species  and  the  preceding  are  allied  to  Corgatha  drosera^ 
Meyr. 

Metachrostis  epichroma,  n.sp. 

(J.  21  mm.  Head,  palpi,  and  thorax  dark  gvQy.  Antenna? 
fuscous;  in  male  simple,  minutel}-  ciliated  {\).  Abdomen  pale 
ochreous,  irrorated  with  grey.  Legs  grey,  irrorated  with  whitish; 
posterior  pair  mostly  whitish.  Forewings  elongate-triangular, 
costa  very  slightly  arched,  apex  round-pointed,  termen  slightly 
rounded,  oblique;  grey,  mixed  towards  base  with  whitish,  towards 
termen  with  pale  brownish,  markings  dark  fuscous;  two  fine  wavy 
parallel  lines  from  costa  at  J  to  dorsum  at  \;  a  broad  median 
fascia,  narrower  towards  costa,  preceded  by  a  minute  discal  dot, 
its  anterior  edge  straight,  posterior  edge  angled  in  disc,  towards 
costa  suffused;  two  fine  parallel  lines  from  costa  at  |  obliquely 
outwards,  forming  a  rounded  projection  in  disc,  thence  to  dorsum 
at  f ,  preceded  by  a  double  discal  dot;  a  fine  crenate  subterminal 
line;  a  fine  terminal  line:  cilia  grey,  with  a  few  whitish  scales. 
Hindwings  with  termen  slightly  sinuate;  deep  yellow,  with  a 
l)road  dark  fuscous  terminal  band;  cilia  fuscous. 

Brisbane,  Q.;  one  specimen. 

The  forewings  are  very  similarly  marked  to  those  of  Metachrostis 
punctigera,  Wlk. 

Metachrostis  camptozona,  n.sp. 

^.  23  mm.  Head  brownish-gre}'.  Palpi  brownish-grey,  irro- 
rated with  dark  fuscous.  Antennje  dark  fuscous,  basal  joint 
whitish,  purple-tinged.  Thorax  brownish-grey,  apices  of  patagia 
fuscous.      Abdomen   fuscous,    mixed   with   whitish.       Legs   dark 


BY    A.    JfiFFERlS    TURNER.  115 

fuscous,  annulated  with  whitish ;  posterior  pair,  except  tarsi, 
whitish.  Forewings  elongate-triangular,  costa  nearly  straight, 
termen  rounded,  oblique;  brownish-grey,  slightly  purplish-tinged, 
markings  fuscous;  a  spot  on  costa  near  base,  and  another  beneath 
it  on  fold;  a  fine  dentate  line  from  \  costa  to  ^  dorsum,  near 
dorsum  preceded  by  a  similar  parallel  line;  a  broad  median  fascia 
from  costa  before  middle  obliquely  outwards,  abruptly  bent  in 
disc,  and  continued  as  three  wavy  parallel  lines  to  dorsum  beyond 
middle;  a  dot  on  costa  beyond  middle,  with  another  beneath  it  in 
disc;  a  straight  ochreous-brown  shade  preceded  by  a  darker  line 
from  costa  at  4  to  before  tornus;  a  terminal  deep  brown  shade, 
and  some  dark  fuscous  terminal  dots  :  cilia  fuscous,  mixed  with 
brownish-grey  and  whitish.  Hindwings  with  termen  rounded, 
veins  3  and  4  connate;  dark  grey;  cilia  dark  grey,  barred  with 
whitish. 

Brisbane,  Q.;  one  specimen  taken  by  Mr.  R.  lUidge. 

]MlCRAESCHUS    DIACAUSTUS,    n.sp. 

(J.  16  mm.  Head,  palpi,  and  thorax  fuscous-brown.  Antenna3 
fuscous;  in  male  simple,  minutely  ciliated  (J).  Abdomen  pale 
reddish-brown.  Legs  brownish-fuscous,  mixed  with  whitish,  tarsi 
dark  fuscous,  annulated  with  whitish.  Forewings  elongate- 
triangular,  costa  strongly  arched,  apex  round-pointed,  termen 
moderately  angled  on  vein  4,  above  angle  concave,  beneath 
straight,  pale  reddish-brown;  a  paler  spot  on  costa  at  J,  from 
which  proceeds  a  fine  dark  dentate  line  to  dorsum  at  J;  a  white 
dark-edged  dot  in  middle  of  disc;  a  second  costal  pale  spot  at  |, 
from  which  a  dark  line  proceeds  obliquely  outwards,  forming  a 
marked  projection  in  disc,  and  continued  as  a  dentate  line  to 
dorsum  at  f ;  a  dark  terminal  line :  cilia  whitish-ochreous,  inter- 
rupted by  fuscous-brown.  Hindwings  with  termen  rounded ; 
ochreous-whitish,  suffused  with  reddish-brown  ;  cilia  whitish- 
ochreous.  On  underside  hindwings  and  margins  of  forewings  are 
suffused  with  dark  fuscous. 

Brisbane,  Q.;  one  specimen. 

The  type  is  not  in  the  best  condition. 


116        NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 
E  U  B  L  E  M  M  A,  Hb. 

Veins  3  and  4  of  the  hind  wing  are  frequently  stalked  in  this 
genus.  There  is  also  considerable  variation  in  the  scaling  of  the 
palpi,  and  length  of  the  terminal  joint;  but  these  differences  do 
not  appear  to  call  for  generic  subdivision. 

EUBLEMMA    GLAUCOCHKOA,    n.sp. 

(J,  19  mm.  Head,  palpi,  thorax,  and  abdomen  whitish-gre}-. 
Palpi  rough-scaled,  terminal  joint  short,  reaching  vertex. 
Antennae  gre}' :  in  male  simple,  ciliated  (1).  Legs  whitish; 
anterior  and  middle  pairs  irrorated  with  grey.  Forewings  elon- 
gate-triangular, costa  straight,  apex  rounded,  termen  rounded, 
oblique:  with  a  tuft  of  long  scales  on  dorsal  margin  before  middle; 
pale  grey,  greenish-tinged,  towards  costa  and  termen  sparsely 
irrorated  with  dark  fuscous;  lines  whitish;  first  from  costa  at  ^ 
to  dorsum  at  i,  slightly  outwardly  curved:  second  from  mid-costa 
to  mid  dorsum  with  an  obtuse  median  outward  projection;  third 
from  costa  at  f  to  dorsum  at  f  with  a  prominent  acute  median 
projection:  cilia  whitish,  Hindwings  with  termen  rounded; 
veins  3  and  4  stalked;  grey,  towards  base  whitish;  cilia  whitish. 

Townsville,  Q.;  in  August;  one  specimen  received  from  Mr.  F. 
P.  Dodd. 

EUBLEMMA    SPHRAGIDOTA,  n.Sp. 

(J.  14  mm.  Head,  thorax,  and  abdomen  whitish.  Palpi 
purplish-grey,  terminal  joint  whitish;  second  joint  with  short 
appressed  scales,  terminal  joint  short,  obtuse,  Imrely  reaching 
vertex.  Antennae  whitish;  in  male  simple,  ciliated  (1).  Legs 
purplish-grey,  irrorated,  and  tarsi  annulated  with  white;  posterior 
pair  white.  Forewings  elongate-triangular,  costa  nearly  straight, 
apex  round-pointed,  termen  rounded,  oblique;  \vhitish-grey;  a 
broad  transverse  reddish-brown  fascia  before  middle,  its  anterior 
edge  narrowh'  dark  fuscous,  wavy,  its  posterior  edge  angled  out- 
wardl}-;  just  at  angle  is  a  large  oblong  dark  fuscous  discal  spot; 
a  fine  wavy  grey  posterior  line  from  costa  at  §  to  dorsum  at  f , 
near  costa  reddish-brown;  a  small  reddish-brown  mark  inclosino- 


BY    A.    JEPFERIS    TURNER.  117 

a  whitish  clot  on  costa  before  apex;  a  faint  grey  subterminal 
shade:  cilia  whitish-grey.  Hindwings  with  termen  slightly 
rounded;  veins  3  and  4  stalked;  whitish-grey,  with  traces  of  two 
lines  of  deeper  grey  beyond  middle;  cilia  whitish-grej'-. 

Townsville,  Q.;  in  October;  one  specimen  received  from  Mr.  F. 
P.  Dodd. 

EUBLEMMA    EURYNIPHA,  n.sp. 

(J.  18  mm.  Head  whitish-ochreous.  Palpi  fuscous;  short, 
porrect  (?),  second  joint  roughly  scaled,  terminal  joint  short, 
obtuse.  [Antennae  broken.]  Thorax  fuscous.  Abdomen  whitish- 
ochreous.  Legs  fuscous,  annulated  with  whitish;  posterior  pair, 
except  tarsi,  whitish-ochreous.  Forewings  narrow  elongate- 
triangular,  costa  straight,  except  close  to  base  and  apex,  apex 
tolerably  pointed,  termen  straight,  oblique;  snow-white;  costal 
edge  narrowly  dark  fuscous;  a  dark  fuscous  basal  fascia  limited 
by  a  line  from  J  costa  to  J  dorsum;  a  dark  fuscous  subterminal 
fascia,  anterior  edge  wavy  from  4  costa  to  4  dorsum,  posterior 
edge  touching  termen  only  at  tornus;  termen  white,  with  a  series 
of  dark  fuscous  dots:  cilia  fuscous,  mixed  with  whitish.  Hind- 
wings  with  termen  rounded;  veins  3  and  4  separate;  whitish- 
ochreous,  with  a  broad  grey  terminal  suffusion;  cilia  whitish- 
ochreous. 

The  apparently  porrect  palpi  may  be  accidental,  as  the  head 
of  the  type  has  been  injured. 

Birchip,  Vic;  in  January,  one  specimen  in  Coll.  Lyell. 

EUBLEMMA    RUFIPUNCTA,  n.Sp. 

(J.  10-14  mm.  Head,  palpi,  thorax,  and  abdomen  grey.  Palpi 
with  second  joint  rough-scaled,  terminal  joint  minute,  not  reach- 
ing vertex.  Antennae  ochreous-grey;  in  male  simple,  ciliated  (2). 
Legs  grey ;  posterior  pair  whitish-grey.  Forewings  elongate- 
triangular,  costa  slightly  arched,  apex  round-pointed,  termen 
obliquely  rounded;  grey,  sometimes  purplish-tinged;  with  straight, 
oblique  darker  lines,  first  from  costa  at  J  to  dorsum  at  J,  second 
from  mid-costa  to  dorsum  at  |,  third  from  costa  near  apex  to 
tornus;  the  second  line  is  interrupted  at  J  from  costa  by  a  reddish 


118  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

dot ;  a  fine  terminal  line :  cilia  purplish-grey,  irrorated  with 
whitish.  Hind  wings  with  termen  slightly  rounded;  veins  3  and 
4  stalked;  grey,  a  darker  transverse  line  at  J;  a  pale  siibterminal 
line  from  inner  margin  not  reaching  costa;  cilia  grey. 

Townsville,  Q.;  in  July  and  September;  two  specimens  received 
from  Mr.  F.  P.  Dodd. 

EUBLEMMA    LOZOSTROPHA,   n.sp. 

9.  12-13  mm.  Head  and  palpi  whitish-ochreous,  vertex  some- 
times white;  palpi  short,  not  reaching  vertex,  second  joint  loosely 
scaled.  Antennae  grey,  towards  base  whitish-ochreous.  Thorax 
and  abdomen  ochreous-brown.  Legs  whitish-ochreous.  Fore- 
wings  elongate-triangular,  costa  nearl}^  straight  ;  apex  round- 
pointed,  termen  slightly  rounded,  oblique;  reddish-brown;  a 
narrow  inwardly  oblique  white  fascia  from  mid-costa  to  dorsum 
at  f,  followed  by  a  broader  grey  suffusion:  cilia  white,  mixed 
with  fuscous-brown.  Hindwings  with  termen  rounded;  veins  3 
and  4  stalked;  grey;  cilia  grey. 

Townsville,  Q.;  in  May,  July,  and  August;  three  specimens 
received  from  Mr.  F.  P.  Dodd. 

Z  0  p  H  0  c  H  R  o  A,   n.g. 

Face  rounded,  with  appressed  scales.  Palpi  moderate,  porrect; 
second  joint  thickened  with  appressed  scales;  terminal  joint  short, 
obtuse.  Antennae  in  male  with  long  double  pectinations,  apical 
fifth  simple.  Thorax  not  crested.  Abdomen  with  a  small  basal 
crest.  Legs  normal.  Forewings  with  vein  2  from  §,  3  from  well 
before  angle,  7,  8,  9  stalked,  no  areole.  Hindwings  with  5  some- 
what approximated  to  4  at  base,  S  anastomosing  with  cell  near 
base. 

Allied  to  Eublemma,  Hb.  Differing  in  the  abdominal  crest, 
and  pectinated  antenna?  of  male. 

ZOPHOCHROA    LEUCOCHORPA,    n.Sp. 

^.  19-20  mm.  Head,  thorax,  and  palpi  black.  Antenme  grey 
in  male  with  long  pectinations  (5).      Abdomen  dark  grey,  basal 


BY    A.    JEFFERIS    TURNER.  119 

crest  black.  Legs  dark  fuscous.  Fore  wings  elongate-triangular, 
costa  straight,  apex  rounded,  termen  rounded,  slightly  oblique; 
black;  a  white  dot  on  fold  above  |  dorsum;  a  finely  waved  con- 
spicuous white  line  from  costa  beyond  middle  to  dorsum  at  |; 
three  minute  white  dots  on  apical  ^  of  costa:  cilia  black.  Hind- 
wings  with  termen  rounded;  grey;  cilia  grey. 

Townsville,  Q  ;  in  March,  two  specimens  received  from  Mr.  F. 
P.  Dodd. 

ZOPHOCHROA    AXTIIRACITIS,  n.sp. 

^.  15  mm.  Head,  palpi,  and  thorax  black.  Antennae  grey, 
towards  base  blackish;  pectinations  in  male  moderate  (2).  Abdo- 
men grey,  basal  crest  black.  Legs  dark  fuscous;  posterior  pair 
grey,  Forewings  elongate-triangular,  costa  gently  arched,  apex 
rounded,  termen  rounded,  oblique;  blackish-fuscous;  an  ochreous- 
whitish  spot  with  some  black  scales  on  fold  at  4,  representing  a 
basal  line ;  a  transversely  elongate  black  discal  dot  tending  to 
become  double ;  a  fine  dentate  blackish  line  followed  by  an 
ochreous-whitish  line,  obsolete  near  costa,  from  mid-costa  obliquel}' 
outwards,  bent  abruptly  near  costa,  thence  transverse,  bent 
inwards  below  mid-disc,  and  again  to  dorsum  atf ;  an  interrupted 
black  terminal  line:  cilia  blackish-fuscous.  Hindwings  with 
termen  rounded;  dark  grey;  cilia  dark  grey. 

Townsville,  Q.;  in  March;  one  specimen  received  from  Mr.  F. 
P.  Dodd. 

ZoPHOCHROA    MELANICA,   n.sp. 

(J.  15  mm.  Head,  palpi,  and  thorax  black.  Antennae  grey. 
Abdomen  dark  fuscous.  Legs  dark  fuscous.  Forewings  triangu- 
lar, costa  straight,  except  close  to  base  and  aj)ex,  apex  round- 
pointed,  termen  scarcely  rounded,  oblique;  black;  an  ochreous- 
whitish  spot  in  disc  at  |^,  second  on  dorsum  at  ^,  and  a  third 
above  second:  cilia  black.  Hindwings  with  termen  rounded; 
fuscous-grey;  cilia  fuscous-grey. 

Townsville,  Q.;  in  January;  one  specimen  received  from  Mr. 
F.  P.  Dodd. 

The  terminal  joints  of  palpi  are  less  developed  in  this  than  in 
the  other  two  species  of  the  genus. 


120  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

Zagira  polycesta,  n.-sp. 

(J.  26  mm.  Head  purple-reddish,  mixed  with  whitish,  face 
whitish.  Palpi  ochreous- whitish,  base  of  second  joint  externally 
reddish;  second  joint  rather  loosely  scaled  towards  apex,  terminal 
joint  short,  obtuse.  Antennae  ochreous-whitish ;  in  male  with 
short  ciliations  (h)  and  longer  bristles  (IJ).  Thorax  purple-grey, 
anterior  third  whitish.  Abdomen  purple-grey,  mixed  with 
whitish.  Legs  whitish  ;  anterior  pair  fuscous.  Forewings 
elongate-triangular,  costa  straight,  apex  acute,  termen  crenate, 
rounded,  oblique ;  purple-grey,  suffused  with  pale  brownish- 
ochreous;  a  broad  whitish  streak  along  costa  nearly  to  apex;  two 
or  three  fuscous  dots  on  costal  edge  in  terminal  part  of  streak; 
lines  slender,  white,  not  crossing  streak;  first  transverse,  slightly 
angled  at  i,  followed  by  a  white,  dark-edged  discal  dot  nearer 
costa  than  dorsum;  a  small  fuscous  suffusion  above  centre  of  disc; 
second  line  dentate  from  beneath  costa  at  #,  first  outwardl}'  then 
inwardly  curved  to  dorsum  at  |;  beyond  it  is  a  whitish  blotch 
before  tornus;  a  similar  subterminal  line  parallel  to  termen;  a 
series  of  dark  fuscous  terminal  dots:  cilia  whitish,  mixed  with 
purple-grey.  Hindwings  with  termen  crenate,  rounded;  colour 
and  markings  as  forewings,  but  basal  line  and  dot  absent. 

Brisbane,  Q.;  one  specimen  taken  by  Mr.  R.  Illidge. 

A  X  I  0  R  a  T  a,  n.g. 

Face  flat.  Palpi  recurved,  ascending,  not  reaching  or  exceeding 
vertex.  Tongue  well  developed.  AntenniB  in  male  minutely 
ciliated.  Thorax  and  abdomen  without  crests.  Forewings  with 
2  from  |,  3  from  angle,  G  from  well  below  upper  angle  of  cell, 
7,  8,  9,  10  stalked,  no  areole.  Hindwings  with  veins  3  and  4 
stalked,  o  from  middle  of  discocellulars,  6  and  7  stalked. 

Besides  the  tj^pe  described  below  this  genus  contains  Acontht 
tripartita,  Butl.,  which  is  closely  allied  though  with  larger  palpi. 

Sir  George  Hampson  referred  these  species  to  Hyela,  Steph., 
but  I  do  not  think  this  can  be  maintained. 


BY    A.    JEFFERIS    TURNER.  121 

AXIORATA    LEUCOZONA,   11. Sp. 

f^^.  15-17  mm.  Head  brovvnish-ochreous,  mixed  with  whitish. 
Palpi  brownish-ochreous;  external  surface  of  second  joint  fuscous. 
Thorax  dark  fuscous;  a  posterior  spot  and  apices  of  patagia'white. 
Abdomen  whitish-ochreous.  Legs  whitish-ochreous;  anterior  and 
middle  pairs  mixed  with  fuscous.  Fore  wings  elongate,  posteriorly 
dilated,  costa  gently  arched,  apex  rounded,  termen  scarcely 
rounded,  slightly  oblique;  white,  faintly  ochreous-tinged,  markings 
dark  fuscous;  a  triangular  spot  at  base  of  costa;  a  broad  central 
transverse  fascia,  anterior  edge  nearly  straight  or  slightly  concave, 
posterior  edge  straight,  wavy;  a  broad  terminal  fascia,  broadest 
in  middle  of  disc,  where  it  forms  an  angular  projection:  cilia 
dark  fuscous.  Hindwings  with  termen  gently  rounded,  slightly 
indented  beneath  apex;  grey;  cilia  ochreous-whitish,  with  a  grey 
median  line. 

Townsville,  Rockhampton,  and  Brisbane,  Q.;  from  Januarj''  to 
March;  several  specimens. 

H  I  M  E  R  0  1  s,   n.g. 

Face  flat.  Palpi  moderate,  recurved,  not  reaching  vertex,  with 
appressed  scales  ;  terminal  joint  well  developed,  stout,  obtuse. 
Antenna?  in  male  simple,  minutely  ciliated  (j).  Thorax  and 
abdomen  not  crested.  Posterior  tibiae  with  two  pairs  of  spurs. 
Forewings  with  7,  8,  9,  10  stalked,  no  areole.  Hindwings  with 
3,  4  connate,  5  from  middle  of  discocellulars  not  curved  towards 
4  at  base,  6,  7  connate,  8  connected  with  cell  near  base. 

In  structure  this  approaches  Pachylepis,  Feld.,  as  described  in 
Hamson's  '  Moths  of  India,'  to  which  the  type  does  not  appear 
related.  Probably  the  structure  of  vein  5  of  hindwing  will  prove 
a  sufficient  distinction. 

HlMEROIS   THIOCHROA,  n.sp. 

,^.  14  mm.  Head  and  thorax  pale  yellow.  Palpi  pale  ochreous, 
second  joint  dark  fuscous  on  external  surface,  terminal  joint  with 
a  few  dark  fuscous  scales  on  external  surface.  Antennae  fuscous. 
Abdomen    grey,   dorsum    near   base   yellowish.       Legs   fuscous ; 


•122  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

posterior  pair,  except  tarsi,  oclireous-whitish.  Eorewings  narrow 
elongate-triangular,  costa  gently  arched,  apex  round -pointed, 
termen  scarcely  rounded,  oblique;  pale  yellow;  costal  edge  at 
base  blackish;  a  black  dot  in  centre  of  disc;  an  interrupted  black 
terminal  line :  cilia  dark  fuscous,  at  apex  and  tornus  yellowish. 
Hind  wings  with  termen  rounded;  grey,  towards  base  paler;  cilia 
grey. 

Townsville,  Q.;  in  February;  one  specimen  received  from  Mr. 
F.  P.  Dodd. 

Anachrostis  aphanta,  n.sp. 

^Q.  13-14  mm.  Head,  antennae,  thorax,  and  abdomen  brown- 
whitish.  Palpi  brown-whitish,  irrorated  with  fuscous;  second 
joint  exceeding  vertex,  with  long  spreading  hairs  towards  apex. 
Legs  ochreous-whitish.  Forewings  elongate-triangular,  costa 
gently  arched,  apex  round-pointed,  termen  slight^  rounded, 
oblique;  brown-whitish,  with  fuscous  markings;  a  line  from  costa 
at  J  angulated  beneath  costa  obliquely  inwards  to  dorsum  at  -J;  a 
large  dot  in  mid-disc;  a  posterior  line  from  costa  at  f,  crenulate, 
bent  inwards  in  disc  and  continued  to  dorsum  at  |;  a  suffused 
subterminal  line;  a  row  of  terminal  dots:  cilia  brown-whitish, 
irrorated  wdth  fuscous.  Hind  wings  with  termen  rounded;  grey- 
whitish;  in  male  whitish-ochreous  towards  inner  margin;  cih'a 
whitish,  in  male  whitish-ochreous  on  inner  margin. 

Brisbane,  Q.:  two  specimens. 

Hypeninae. 
Panilla  cautophanes,  n.sp. 

9.  20  mm.  Head  brown.  Palpi  brown- whitish,  base  of  second 
joint  dark  fuscous  ;  second  joint  long,  considerably  exceeding- 
vertex,  with  long  appressed  hairs,  looser  towards  apex,  terminal 
joint  well  developed.  Antennae  fuscous.  Thorax  brown,  mixed 
with  fuscous.  Abdomen  brown,  mixed  with  fuscous  and  whitish, 
Legs  dark  fuscous,  annulated  with  ochreous-whitish.  Forewings 
elongate-triangular,  costa  nearly  straight,  apex  round-pointed, 
termen  rounded,  oblique,  crenate;  brown;  a  broad  dark  fuscous 


BY    A.    JEFFERIS    TURNER.  123^^ 

basal  fascia;  two  deeper  brown  outwardly  ciun  ed  wav}'  parallel 
lines  from  costa  before  middle  to  mid-dorsum,  enclosing  a  fuscous 
dot  near  costa;  a  fine  brown  dentate  line  from  costa  at  J  obliqueh^ 
outwards  forming  a  prominent  acute  projection  in  disc,  thence 
obliquely  inwards  to  dorsum  at  |;  a  large  triangular  dark  fuscous 
blotch  on  costa  from  |^  to  apex;  a  paler  fuscous  suffusion  on  mid- 
termen;  a  finely  dentate  pale  subterminal  line  crossing  costal 
blotch,  less  distinct  in  disc;  a  fuscous  terminal  line:  cilia  pale 
brown,  bases  barred  with  fuscous.  Hind  wings  with  termen 
rounded,  crenate;  colour  and  markings  as  forewings,  but  without 
basal,  costal,  and  terminal  fuscous  blotches;  a  fuscous  spot  above 
tornus  from  which  a  dark  subterminal  line  runs  to  costa;  a  large 
crest  of  scales  at  end  of  cell. 
Brisbane,  Q.;  one  specimen. 

E  P  I  T  R  I  PT  A,    n.g. 

Face  flat.  Palpi  long,  obliquely  ascending;  second  joint  twice 
breadth  of  eye,  smooth-scaled;  terminal  joint  more  than  one-half 
second,  slender,  smooth,  acute.  Thorax  and  abdomen  not  crested. 
Forewings  with  7  from  cell,  8,  9,  10  stalked,  no  areole.  Hind- 
wings  with  vein  3,  4  connate,  5  from  near  4.- 

Allied  to  Falcimala,  Hmps.,  but  veins  3  and  4  of  hind  wings 
are  not  stalked,  and  5  is  approximated  to  them  at  base,  and  there- 
is  no  abdominal  crest.  In  neuration  it  agrees  with  Alelimma, 
Hmps.,  but  lacks  the  peculiar  palpi. 

Epitripta  acosmopis,  n.sp. 

9.  23  mm.  Head,  thorax,  and  abdomen  gre\^  Palpi  fuscous.. 
Antennae  grey.  Legs  pale  grey ;  posterior  pair,  except  tarsi, 
whitish.  Forewings  narrow  elongate-triangular,  costa  gentl}^ 
arched,  apex  rectangular,  termen  rounded,  slightly  oblique;  grey, 
markings  dark  fuscous;  a  line  from  costa  near  base  to  fold,  pro- 
duced to  base  along  fold;  an  irregularl}'  dentate  line  from  i  costa 
to  ^  dorsum;  a  fuscous  median  shade  l^roadly  interrupted  in  disc, 
followed  by  a  whitish  discal  dot;  a  posterior  line  from  ;|  costa 
obliquely  outwards,  very  line  in  disc,  forming  a  rounded  projec- 


124  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

tion,  very  distinct  near  dorsum,  ending  at  f ;  a  triangular  l^lotcli 
on  costa  before  apex,  succeeded  by  a  whitish  subterminal  line;  an 
interrupted  fuscous  terminal  line:  cilia  whitish,  with  a  broad 
fuscous  median  line.  Hindwings  with  termen  rounded,  wavy; 
pale  grey;  a  darker  terminal  line;  cilia  pale  grey. 

P  A  D  R  O  P  H  Y  L  L  A,    n.g. 

Face  flat.  Palpi  long,  ascending,  with  closely  appressed  scales; 
second  joint  reaching  or  exceeding  vertex;  terminal  joint  nearly 
as  long  as  second,  stout,  acute.  Tongue  present.  Thorax  and 
abdomen  without  crests.  Forewings  with  vein  7  from  cell,  8,  9 
stalked,  10  absent.  Hindwings  with  veins  3,  4  stalked,  5  from 
rather  below  centre  of  discocellulars,  6,  7  stalked,  8  anastomosing 
with  cell  to  middle.  This  genus  probably  corresponds  to  Section 
ii.,  Division  A,  of  the  genus  Catada,  Wlk.,  in  Hampson's  'Moths 
of  India,'  and  if  so  includes  Catada  prominent,  Hmps. 

Paurophylla  aleuropasta,  n.sp. 

9.  14  mm.  Head,  thorax,  palpi,  and  antenna?  dark  fuscous. 
Abdomen  whitish-grey.  Legs  fuscous-grey.  Forewings  narrow 
elongate,  posteriori}^  dilated,  costa  gently  arched,  apex  rounded, 
termen  rounded,  scarcely  oblique ;  dark  fuscous,  sparsely  and 
irregularly  irrorated  with  wdiite;  a  transversel}'^  elongate  white 
mark  in  disc  beyond  middle:  cilia  dark  fuscous.  Hindwings 
with  termen  rounded;  whitish-grey;  cilia  whitish-gre}^ 

Brisbane,  Q.;  in  March,  one  specimen. 

Nodaria  ctenucha,  n.sp. 

(J.  19  mm.  Head  and  thorax  pale  grey;  forehead  with  a  tuft 
of  hairs  directed  forwards.  Palpi  grey;  very  long;  loosely  scaled, 
second  joint  four  times  breadth  of  eye,  upper  edge  with  a  ridge 
of  loose  hairs,  terminal  joint  well  developed,  acute.  Antennae 
grey;  in  male  with  long  fine  pectinations  (8),  apical  fifth  simple. 
Abdomen  grey,  apices  of  segments  whitish.  Legs  grey.  Fore- 
wings elongate-triangular,  apex  round-pointed,  termen  rounded, 
oblique;  whitish-grey,  irrorated  with  fuscous:  a  fuscous  line  from 


BY    A.    JEFFERIS    TURNER.  125 

^  costa  to  ^  dorsum,  outwardly  curved,  a  blackish  discal  dot 
beneath  mid-costa,  transversely  elongate  and  constricted  in  centre; 
a  wavy  fuscous  line  from  costa  at  4  obliquely  outwards,  sharply 
bent  in  disc,  and  thence  inwardly  oblique  to  dorsum  at  |;  a 
whitish  oblique  line  preceded  by  a  dark  shade  from  apex  to  before 
tornus:  cilia  whitish-grey,  with  two  fine  grey  lines.  Hindwings^ 
with  termen  rounded;  whitish-grey;  a  fuscous  dot  or  short  line  in 
disc  opposite  i  inner  margin;  a  short  white  line  preceded  by  a 
darker  shade  from  tornus,  lost  in  disc;  cilia  as  forewings. 

Differs  markedly  from  other  members  of  the  genus  in  the 
structure  of  the  male  antennae. 

Kuranda,  Q.;  in  October,  one  specimen. 

NODARIA    ACKOSEMA,   n.sp. 

J$.  21-25  mm.  Head,  thorax,  and  abdomen  pale  grey;  fore- 
head with  a  tuft  of  hairs  directed  forwards.  Palj^i  very  long, 
loosely  scaled;  second  joint  five  times  breadth  of  eye,  with  a  ridge 
of  loose  hairs  on  discal  |  of  upper  edge;  terminal  joint  well 
developed  ;  grey.  Antennae  grey ;  in  male  slightly  laminate, 
evenly  ciliated  (1),  with  longer  bristles  (3).  Legs  ochreous- 
whitish,  irrorated  with  grey.  Forewings  elongate-triangular, 
costa  straight  to  near  apex,  apex  acute,  termen  slightly  rounded,, 
oblique;  grey;  traces  of  an  anterior  line;  a  faintly  indicated  discal 
dot;  a  faint  posterior  line  from  costa  at  |  obliquely  outwards^ 
forming  a  rounded  projection  in  disc,  and  ending  in  dorsum  at  |; 
a  dark  fuscous  spot  at  apex,  preceded  by  a  very  short  oblique 
whitish  mark,  and  that  again  by  a  fuscous  dot  on  costa;  a  fine 
dark  fuscous  terminal  line:  cilia  grey,  with  a  whitish  basal  line. 
Hind  wings  with  termen  rounded;  grey;  a  faintly  darker  line  from 
inner  margin  at  |,  and  a  faint  pale  line  from  tornus,  both  lost  in 
disc;  terminal  line  and  cilia  as  forewings. 

Kuranda  and  Geraldton,  Q. ;  in  October  and  November,  three 
specimens. 

SiMPLICIA    PHRICOZOXA,  n.Sp. 

(J 9.  34  mm.  Head,  palpi,  antennae,  and  thorax  dark  brown. 
Palpi  in  male  with  terminal  joint  enlarged  and  thickened,  extend- 


126  NEW    CEN'KRA    AND    SPECIES    OF    LEPIDOPTKRA, 

ing  backwards  to  edge  of  collar,  and  bearing  an  expansive  tuft  of 
hairs  internall}''.  Antennae  of  male  with  rather  long  even  cilia- 
tions  (2),  and  longer  bristles  (4).  Abdomen  brownish.  Legs 
brownish,  mixed  with  dark  fnscous.  Forewings  triangular,  costa 
slightly  arched,  apex  rounded,  termen'  rounded,  slightly  oblique; 
brow^n,  with  dark  fuscous  lines;  a  line  near  base  not  reaching 
dorsum;  a  dot  on  costa  at  J;;  a  thick  wavy  line  from  costa  at  ^  to 
dorsum  at  4;  a  tine  acutely  dentate  line  from  costa  before  ^  to 
mid-dorsum,  edged  posteriorly  with  brown-whitish;  an  irregularly 
dentate  submarginal  line,  thickened  towards  costa  and  dorsum, 
edged  posteriorly  with  brown-whitish;  some  dark  fuscous  scales 
on  termen:  cilia  brown-whitish.  Hindwings  with  termen  rounded ; 
brownish-fuscous,  suffused  more  or  less  with  w^hitish  ;  cilia 
w^hitish. 

Brisbane,  Q. ;  one  specimen  taken  by  Mr.  R.  Illidge  ;  there  is 
a  second  in  the  Museum  of  the  Agricultural  Department. 

TiPASA    MACROBELA,   n.sp. 

9.  18  mm.  Head,  thorax,  antenna?,  and  abdomen  ochreous- 
whitish.  Palpi  ochreous- whitish,,  sparsely  irrorated  with  fuscous; 
porrect,  nearly  as  long  as  aV)domen,  second  joint  excessively  long, 
twelve  times  breadth  of  eye,  rough-haired  above  and  beneath, 
terminal  joint  slender,  acute.  Thorax  and  abdomen  without 
■crests.  Legs  ochreous-whitish.  Forewings  elongate-triangular, 
costa  moderately  arched,  apex  acute,  termen  rounded,  oblique; 
ochreous-whitish,  sparsely  irrorated  with  fuscous;  a  whitish  spot 
containing  two  fuscous  dots  arranged  transversely  in  mid-disc;  a 
short  oblique  fuscous  streak  at  apex  giving  origin  to  a  line  of 
fuscous  dots  to  dorsum  at  t;  an  interrupted  fuscous  terminal  line: 
cilia  ochreous-whitish.  Hindwings  rather  broad,  termen  sinuate; 
whitish:  cilia  whitish. 

Brisbane,  Q.;  one  specimen. 

1  have  also  received  from  Mr.  R.  Illidge  what  is  doul)tless  the 
male  of  this  species.  The  antennae  have  very  long,  fine,  slender 
l)ipectinations.      The  forewings  differ  from  the   female  type  in 


BY    A.    JEFFERIS    TURNER.  127 

having  a  circular  dark  fuscous  spot  beyond  middle,  and  the  other 
dark  markings  rather  better  developed. 

Prolophota  camptoloma,  n.sp. 

9.  20  mm.  Head,  thorax,  palpi,  and  abdomen  ochreous 
whitish,  sparsely  irrorated  with  fuscous.  Palpi  porrect,  long; 
second  joint  five  times  breadth  of  eye,  rough ty  haired  above  and 
beneath  ;  terminal  joint  slender,  acute.  Antennie  ochreous- 
whitish.  Legs  ochreous-whitish,  sparsely  irrorated  with  fuscous. 
Forewings  elongate-triangular,  costa  nearl}^  straight,  but  strongh^ 
bent  before  apex,  apex  acute,  termen  oblique,  sigmoid,  being 
slightly  concave  beneath  apex,  and  strongly  bowed  on  vein  4; 
ochreous-whitish,  irrorated  with  fuscous;  a  brownish  dot  on  costa 
at  :j,  a  fuscous  dot  at  middle,  and  two  or  three  minute  fuscous 
dots  before  apex;  a  brownish  discal  dot  at  ^,  and  a  dark  fuscous 
dot  above  middle;  a  brownish  line,  first  sigmoid,  then  wav}^  from 
dorsum  at  J,  obliquely  outwards,  ending  beneath  costa  at  -i:  an 
obscure  dentate  fuscous  subterminal  line;  a  row  of  dark  fuscous 
terminal  dots:  cilia  ochreous-whitish,  mixed  with  fuscous.  Hind- 
wings  with  termen  wavy,  slightly  sigmoid  ;  svhitish,  towards 
tsrmen  whitish-grey;  a  waved  fuscous  line  from  inner  margin  at 
1^,  not  reaching  costa;  cilia  whitish. 

Brisbane,  Q  ;  in  August,  one  specimen. 

S  Y  M  M  0  L  P  I  S,    n.g 

Forehead  with  a  tuft  of  scales  directed  forwards.  Palpi  long, 
second  joint  three  times  breadth  of  eye,  terminal  joint  about  half 
second,  acute;  both  second  and  terminal  joints  clothed  with  long 
rough  hairs  above  and  beneath.  Tongue  present.  Thorax  and 
abdomen  not  crested.  Legs  normal.  Forewings  with  inner 
margin  strongly  arched;  vein  3  from  before  angle,  7,  8,  9  stalked, 
10  connected  by  a  bar  with  8,  9  just  behind  7.  Hindwings  with 
3  from  well  before  angle,  well  separated  from  4,  5  approximated 
to  4  at  ba?e,  but  separate,  6  and  7  connate,  8  anastomosing  with 
cell  to  ^. 


128        NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

In  neuration  the  type  nearly  approaches  Orthozona,  Hmps.  (to 
which  Sir  George  Hampson  referred  it),  but  in  the  hairy  third 
joint  of  palpi  and  strongly  arched  dorsum  of  forewins:  it  resembles 
Corcohaea,  Moore. 

Symmolpis  hyporrhoda,  n.sp. 

^.  28  mm.  Head  and  thorax  pinkish-grey.  Palpi  pinkish- 
grey,  with  a  few  dark  fuscous  scales.  Antennae  ochreous-whitish. 
Abdomen  ochreous-whitish.  Legs  ochreous-whitish;  anterior  and 
middle  pairs  tinged  with  pink.  Forewings  elongate-oval,  costa 
strongly  arched,  apex  acute,  termen  rounded,  strongly  oblique, 
dorsum  strongly  arched  at  base,  pinkish-grey;  a  suffused  dark 
fuscous  median  streak  from  base  to  4:  cilia  pinkish-grey.  Hind- 
wings  with  termen  rounded ;  ochreous-whitish,  irrorated  w4th 
pinkish  scales;  cilia  whitish.  Underside  of  fore  wing  dull  pink; 
of  hindwing  as  upper  surface. 

Stradbroke  Island,  Q.;  in  October,  one  specimen. 

Marapana  nycteropis,  n.sp. 

9.  28  mm.  Head  and  thorax  fuscous-grey.  Palpi  fuscous- 
gre}^  apex  of  terminal  joint  ochreous-whitish  ;  long,  porrect, 
second  joint  three  times  breadth  of  eye,  clothed  with  loose  hairs 
above,  terminal  joint  J  second,  stout,  tolerably  pointed.  Antennae 
fuscous.  Abdomen  dark  grey.  Legs  dark  fuscous;  tarsi  annu- 
lated  with  white.  Forewings  elonojate-triangular,  costa  modera- 
tely arched,  apex  rounded,  termen  bowed,  oblique;  dark  fuscous, 
glossy  in  oblique  light;  a  whitish  dot  beneath  J  costa,  and  an 
outwardly  curved  dark  median  shade  are  just  perceptible:  cilia 
dark  fuscous.  Hind  wings  with  termen  angled  on  vein  4,  thence 
straight  to  tornus;  colour  as  forewings;  traces  of  a  fine  dark 
median  line;  cilia  as  forewings. 

Brisbane,  Q.;  in  January,  one  specimen. 

L  o  p  H  o  T  o  M  A,  n.g. 

Face  with  projecting  cone  of  scales.  Palpi  moderate,  ascending, 
but  slightly  exceeding  vertex;  second  joint  with  appressed  scales: 


BY    A.    JEFFERIS    TURNER.  129 

terminal  joint  moderate,  stout,  tolerably  pointed.  Tongue  well 
developed.  Antennae  in  male  simple  or  laminate,  ciliated,  with 
a  pair  of  bristles  on  each  joint.  Thorax  with  a  prominent 
anterior  and  a  prominent  posterior  crest.  Abdomen  not  crested. 
Legs  normal.  Forewings  with  vein  2  from  -|,  3  from  near  angle, 
6  from  well  below  upper  angle,  7,  8,  9  stalked,  10  connected  by 
a  bar  with  8,  9  opposite  7.  Hindwings  with  3  and  4  connate,  5 
approximated  to  4  at  base,  6  and  7  connate,  8  anastomosing  with 
cell  to  1-. 

Distinguished  by  the  doubly  crested  thorax. 

LOPHOTOMA    DIAGRAPHA,   n.sp. 

Head  and  thorax  fuscous.  Palpi  ochreous-whitish,  mixed  with 
fuscous.  Antennye  fuscous  ;  in  male  shortly  laminate,  except 
near  apex,  shortly  ciliated  (1)  with  longer  bristles  (2).  Abdomen 
whitish-grey.  Legs  ochreous-whitish ;  anterior  pair  fuscous. 
Forewings  elongate-triangular,  costa  rather  strongly  arched,  apex 
pointed,  termen  bowed  on  vein  3,  oblique  ;  fuscous,  markings 
whitish;  a  broad  streak  from  base  along  fold,  becoming  lost  in 
disc;  several  streaks  along  veins  from  end  of  cell;  a  narrow 
oblique  blackish  discal  dot  beneath  mid-costa;  three  whitish  dots 
on  terminal  third  of  costa;  a  well  marked  oblique  line  from  costa 
just  before  apex  nearly  to  dorsum  at  |;  an  inwardly  oblique 
whitish  apical  shade;  a  row  of  dark  fuscous  dots  succeeded  by  a 
whitish  line  immediately  before  termen;  terminal  edge  fuscous  : 
cilia  fuscous,  with  a  whitish  basal  line.  Hindwings  with  termen 
rounded;  whitish,  suffused  with  fuscous:  a  fine  fuscous  terminal 
line  preceded  by  an  indistinct  subterminal  line;  cilia  whitish. 

Brisbane,  Q.;  in  May,  one  specimen. 

LOPHOTOMA    METABULA,  n.sp. 

(^2-  2S-37  mm.  Head,  thorax,  palpi,  antennae,  and  abdomen 
ochreous-grey.  Antennae  in  male  ciliated  (1)  with  longer  bristles 
(2).  Legs  ochreous-grey;  anterior  pair  mixed  with  dark  fuscous. 
Forewings  elongate-triangular,  costa  gently  arched,  apex  pointed, 
termen  nearly  straight,  slightly  oblique ;  ochreous-grey ;  with 
9 


130  NKW    GENERA    AND    SPECIES    OF    LEPIDOPTERA, 

fuscous  or  brownish  lines,  often  almost  obsolete;  a  transverse 
twice-dentate  line  from  |  costa  to  J  dorsum;  two  discal  dots,  one 
in  mid-disc,  the  other  between  this  and  costa;  these  are  sometimes 
combined  in  one  large  dot;  an  oblique  line  from  costa  before  apex 
to  mid-dorsum,  sometimes  followed  by  an  interrupted  parallel 
line;  a  series  of  dark  fuscous  terminal  dots:  cilia  ochreous-  or 
brownish-grey.  Hind  wings  with  termen  rounded ;  grey  or 
ochreous-grey,  towards  base  paler:  cilia  ochreous-whitish. 

Kuranda  (in  October),  Gympie,  and  Brisbane,  Q.;  four  speci- 
mens. 

Very  variable  in  intensity  of  markings. 

ESTHLODORA,    n.g. 

Face  flat,  forehead  with  loose  projecting  scales.  Palpi  long, 
obliquely  ascending;  second  joint  exceeding  apex,  with  appressed 
scales,  except  on  posterior  edge  which  is  clothed  with  long  hairs; 
terminal  joint  half  second,  stout,  clothed  with  appressed  scales, 
except  apex,  which  is  naked  and  tolerably  pointed.  Antennae  in 
male  shortly  ciliated  (J).  Thorax  and  abdomen  not  crested. 
Leo"s  normal.  Forewings  with  vein  2  from  y,  3  from  well  before 
angle  widely  separate  from  3,  3  from  angle,  7,  8,  9  stalked,  10 
connected  by  a  bar  with  8,  9  just  beyond  7  to  form  an  areole. 
Hindwings  deeply  excavated  at  tornus;  with  3,  4  connate.  5  from 
rather  below  middle  of  cell,  6,  7  connate,  8  anastomosing  with 
cell  near  base. 

ESTHLODORA    VERSICOLOR,   n.Sp. 

(J^.  17-18  mm.  Head,  thorax,  and  antennas  whitish-grey. 
Palpi  ochreous-grey.  Abdomen  ochreous-grey,  towards  base 
whitish-grey.  Leg^  purple-grey ;  anterior  pair  irrorated  with 
dark  fuscous;  tarsi  dark  fuscous,  annulated  with  white.  Fore- 
wings  elongate-triangular,  costa  slightly  concave,  apex  round - 
pointed,  termen  sinuate,  oblique  ;  ochreous-grey,  basal  third 
whitish-grey;  black  dots  on  costa  near  base  and  at  1;  from  second 
dot  a  faint  ontwardly  curved  line  proceeds  to  near  base  of  dorsum, 
a  fine  line  separating  ground-colours  from  costa  at  |,  outwardly 


BY    A.    JEFFRRIS    TURNKR.  131 

oblique,  angled  near  costa  and  continued  to  dorsum  at  4;  two 
minute  black  dots  placed  transversely  just  beyond  angle  of  this 
line;  a  tine  angulated  fuscous  line  followed  by  a  whitish  line  from 
§  costa  to  f  dorsum;  a  similar  sul)terminal  line  preceded  by  a 
whitish  line;  an  interrupted  fuscous  terminal  line :  cilia  ochreous- 
grey.  Hindwings  with  termen  rounded,  deeply  excavated  at 
tornus;  colour  as  fore  wings,  but  whitish-gre}'-  towards  costa; 
markings  as  forewings,  but  median  line  not  angulated,  discal  dot 
single,  posterior  lines  partly  obsolete;  cilia  as  forewings,  but  on 
excavation  purple-red.  with  apices  whifcish-ochreous.  at  each  angle 
of  excavation  apices  dark  fuscous. 

Var.  A.  Forewings  with  posterior  area  pinkish-ochreous,  median 
line  broadl}^  dark  fuscous,  discal  dots  conspicuous,  posterior  lines 
obsolete.  Hindwings  pinkish-ochreous,  towards  base  on  costa 
whitish,  a  dark  fuscous  suffusion  near  tornus. 

Brisbane,  Q. ;  in  February ;  three  specimens,  a  pair  of  the 
normal  form  and  one  female  of  the  variety. 

Probably  this  species  will  have  many  colour-varieties. 

T  H  E  L  X  I  N  0  A,    n.g. 

Face  Hat.  Palpi  moderate,  recurved,  not  reaching  vertex; 
second  joint  rather  loosely  scaled ;  terminal  joint  truncate. 
Tongue  present.  Antennae  in  male  slightly  serrate  towards  apex. 
Thorax  and  abdomen  not  crested.  Legs  normal.  Forewings  with 
vein  2  from  |,  3,  4  stalked  from  angle,  7,  8,  10,  11  stalked,  9 
absent.  Hindwings  with  3,  1  stalked,  5  from  near  middle  of  cell, 
6,  7  stalked,  8  anastomosing  with  cell  near  base. 

Allied  to  Arcteopteron,  Hmps.,  which,  however,  has  all  veins 
present  in  forewing,  and  vein  7  arising  from  cell.  Type  T. 
ejnphracta. 

Besides  the  three  species  described  below  there  is  a  fourth, 
which  Sir  G.  Hampson  has  identified  as  canescens,  Wlk.  ( ^Jlavil- 
lalis,  Wlk.). 


132        NEW  GENERA  AND  SPECIES  OF  LEPIDOPTERA, 

1.  Fore  and  hindwitigs  white 2. 

Fore  and  hindwings  grey 3. 

2.  Abdomen  with  a  reddish-brown  band;  face  white calliscia 

Abdomen  without  a  reddish-brown  band;  face  fuscous epiphracta 

3.  Forewings   with    ochreous    markings    on    costa.    hindwings 

moderately  broad lyleurotypa 

Forewings  without  ochreous  markings  on  costa,  hindwings 

narrow canescem 

THELXINOA    EPIPHRACTA,  11. sp. 

(J.  11-1  J:  mm.  Head  and  thorax  white;  face  fuscous.  Palpi 
dark  fuscous;  internal  surface  of  second  joint  whitish.  Antennae 
whitish-grey,  towards  base  white.  Abdomen  whitish,  towards 
apex  irrorated  with  dark  fuscous;  a  pale  ochreous  spot  on  each 
side  of  first  segment.  Legs  whitish;  anterior  pair  dark  fuscous 
on  dorsal  sui-face.  Forewings  narrow  elongate-triangular,  costa 
gently  arched,  apex  rounded,  termen  ronnded,  oblique;  white, 
markings  dark  fuscous  and  ochreous-fuscons;  a  dot  on  costa  near 
base  and  another  at  i,  from  which  ochreous-fuscous  lines  proceed 
to  dorsum  near  base  and  at  4;  a  dot  on  costa  before  middle 
separated  b}^  an  ochreous  spot  from  a  large  dark  fuscous  spot  in 
disc,  containing  an  incomplete  whitish  circle:  a  slightly  waved 
interrupted  line  from  'i  costa  forming  an  oblong  projection  in 
disc,  and  bent  inwards  to  dorsum  at  |;  between  bend  and  tornus 
is  a  fuscous  suffusion;  a  faintly  marked  acutely  dentate  line  from 
costa  at  4  to  tornus;  a  subapical  fuscous  suffusion;  a  broad  dark 
fuscous  terminal  line  interrupted  by  ochreous  on  veins:  cilia 
whitish,  mixed  with  fuscous.  Hindwings  with  termen  slightly 
rounded;  white:  a  few  fuscous  scales  near  base;  a  conspicuous 
dark  fuscous  dot  in  disc  at  4;  connected  by  a  suffused  line  with 
inner  margin;  wav}^  fuscous  posterior  and  subterminal  lines; 
subapical  blotch,  terminal  line,  and  cilia  as  forewings. 

Brisbane,  Q.;  four  specimens. 

The  species  of  this  and  the  following  genus  have  the  habit  of 
resting  with  the  wings  widely  outspread  on  tree-trunks  and  fences, 
resembling  small  species  of  Geometridje. 


BY    A.    JEFFERIS    TURNER.  133 

Thelxinoa  calliscia,  n.sp. 

(J.  1 1  mm.  Head,  thorax,  and  antennae  white.  Palpi  white, 
base  of  second  joint  dark  fuscous.  Abdomen  white,  irrorated 
with  dark  fuscous;  basal  joint  white;  second  joint  reddish-brown; 
third  joint  whitish,  mixed  with  reddish-brown ;  tuft  whitish. 
Legs  whitish;  dorsal  surface  of  anterior  pair  dark  fuscous.  Fore- 
wings  narrow  elongate-triangular,  costa  gently  arched,  apex 
round-pointed;  termen  rounded,  very  oblique;  white,  markings 
pale  fuscous,  mixed  with  darker  fuscous;  a  dot  on  costa  at  i,  from 
which  a  pale  waved  line  proceeds  to  dorsum  at  -J;  a  black  dot  on 
costa  at  ^  also  giving  rise  to  a  pale  waved  transverse  line;  a  line 
closely  approximated  to  this  posteriorly,  tirst  bent  outwards  to 
form  an  obtuse  projection  in  disc,  then  inwards  to  dorsum  at  ^; 
a  pale  suffused  line  from  costa  at  f  to  near  tornus;  a  fuscous 
suffusion  at  tornus,  and  another  above  mid-termen  ;  a  row  of 
fuscous  dots  on  termen:  cilia  grey,  at  apex  white.  Hindwings 
with  termen  nearly  straight;  white  ;  a  suffused  basal  fuscous 
fascia;  a  similar  fascia  before  middle,  succeeded  by  a  narrow 
waved  black  line;  a  subterminal  fuscous  sufi'usion;  cilia  whitish- 
grey. 

Brisbane,  Q.;  one  specimen  received  from  Mr.  F.  P.  Dodd. 

Thelxinoa  pleurotypa,  n.sp. 

9.  10-11  mm.  Head  white;  face  dark  grey.  Palpi  dark 
fuscous,  internal  surface  whitish-grey.  Antennae  whitish-grey, 
towards  base  white.  Abdomen  whitish-grey,  towards  apex  dark 
grey.  Legs  whitish,  mixed  with  fuscous.  Forewings  narrow 
elongate-oval,  costa  moderately  arched,  apex  round-pointed,  termen 
rounded,  oblique;  whitish-grey;  costal  edge  ochreous  in  basal 
half;  a  dark  fuscous  dot  on  costa  near  base,  and  another  at  J; 
from  the  latter  a  faint  ochreous  line  proceeds  to  dorsum  at  ^;  a 
short  dark  fuscous  streak  on  costa  before  middle,  bordered  beneath 
with  ochreous;  beneath  this  is  a  dark  fuscous  discal  dot;  a  fine 
grey  median  transverse  line,  sometimes  obsolete ;  a  grey  sub- 
terminal  suffusion;  a  series  of  dark  fuscous  terminal  dots:  cilia 


134  NEW    GENE K A    AND    SPECIES    OF    LEPIDOPTERA, 

grey,  with  a  few  dark  fuscous  scales.  Hind  wings  with  termen 
sHghtly  sigmoid;  whitish-grey;  an  obHque  grey  line  from  beyond 
middle  of  inner  margin  towards  costa  at  J;  terminal  dots  and 
cilia  as  forewings. 

Cairns  and  Townsville,  Q.;  in  June  and  February,  two  speci- 
mens. 

B  R  Y  o  M  I  3M  A,   n.g. 

Face  flat.  Palpi  moderate,  recurved,  rather  short,  not  reaching 
vertex;  second  joint  rather  loosely  scaled;  terminal  joint  short, 
truncate.  Tongue  present.  Antenna?  in  male  simple,  minutely 
ciliated  (i).  Thorax  and  abdomen  not  crested.  Legs  normal. 
Forewings  with  vein  2  from  4,  3  from  near  angle  separate  from  4, 
7,  8,  10  stalked,  9  absent,  11  from  cell.  Hindw^ngs  with  3,  4 
connate,  5  from  below  middle  of  cell,  6,  7  connate,  8  anastomo&ing 
with  cell  near  base. 

Certainly  allied  to  the  preceding,  though  the  neuration  is 
different. 

Bryomima  el.eodes,  n.sp. 

^.  12-13  mm.  Head  and  thorax  whitish,  irrorated  with 
greenish.  Palpi  and  antennse  fuscous.  Abdomen  dark  grey  ; 
first  two  segments  whitish,  irrorated  with  greenish;  third  segment 
reddish-ochreous.  Legs  whitish;  anterior  pair  fuscous.  Fore- 
wings narrow  elongate- triangular,  costa  scarcel}"  arched,  apex 
rounded,  termen  scarcely  rounded,  oblique;  whitish,  mixed  with 
ofreenish,  with  a  few  scattered  black  scales ;  five  or  six  short 
blackish  linear  marks  on  costal  edge,  bounded  beneath  b}^  ochreous; 
two  inwardly  oblique  greenish  lines  near  base;  a  squarish  suffused 
black  discal  spot,  from  which  two  greenish  lines,  irrorated  with 
black,  proceed  to  dorsum;  three  suffused  and  blotched  posterior 
greenish  lines:  cilia  grey.  Hindwings  with  termen  slightly 
rounded,  colour  and  markings  as  forewings,  but  without  discal 
spot. 

Brisbane,  Q.;  four  specimens  receixed  from  Mr.  F.  P.  Dodd. 

Best  distinguished  from  the  following  by  the  absence  of  black 
lines  across  hindwings.      It  is  also  of  larger  size,  and  the  reddish- 


BY    A.    JEFFEUIS    TURNKR.  135 

ochreoiis  band  on  abdomen  appears  constant,  as  also  do  the 
ochreous  markings  near  costa  of  forewings.  Both  are  absent  in 
a  series  of  B.  ochrochlora. 

BUYOMIMA    OCHROCHLORA,   n.sp. 

(J9.  10-11  mm.  Head,  thorax,  and  abdomen  whitish,  more  or 
less  mixed  with  greenish.  Palpi  blackish-fuscous,  apices  ochreous- 
whitish.  Antenucie  grey,  bases  whitish.  Legs  whitish  ;  dorsal 
surface  of  anterior  and  middle  pairs  dark  fuscous.  Forewings 
narrow  elongate-triangular,  costa  straight,  except  towards  apex, 
apex  rounded,  termen  slightly  rounded,  oblique;  whitish,  more  or 
less  mixed  with  greenish,  lines  black;  a  short  dentate  line  from 
costa  at  J  to  dorsum  near  base,  often  obsolete  towards  dorsum,  a 
suffused  blackish  discal  spot;  a  line  from  mid-costa  very  obliquely 
outwards,  bent  at  right  angles  beneath  costa  so  as  to  form  a  long- 
oblong  projection  with  rounded  angles,  and  then  continued  to  mid- 
dorsum;  a  strongly  angled  whitish  subterminal  line,  often  obsolete: 
cilia  grey,  mixed  with  whitish.  Hindwings  with  termen  rounded; 
colour  as  forewings;  two  parallel  approximated  transverse  black 
lines  before  middle,  slightl}'-  angled  in  disc;  cilia  as  forewings. 

Var.  A.  With  a  broad  dark  fuscous  fascia  before  middle  of 
forewings,  including  first  line  and  discal  spot. 

Brisbane,  Q.;  from  December  to  February;  ten  specimens. 

Hyblaeinse. 

HYBLiEA   IBIDIAS,  n.Sp. 

(J.  34  mm.  Head  and  thorax  ochreous-brown.  Palpi  purple- 
fuscous,  basal  half  of  lower  surface  whitish.  AntenntB  ochreous- 
brown.  Abdomen  dull  ochreous,  beneath  reddish.  Legs  pale 
red  ;  anterior  coxye  and  femora  whitish,  with  ochreous  hairs  ; 
tarsi  dark  fuscous,  irrorated  with  whitish.  Forewinsfs  elonaate- 
triangular,  costa  slighly  arched,  apex  rounded-rectangular,  termen 
angled  on  vein  3,  above  angle  straight,  beneath  slightly  excavated 
to  tornus,  dorsum  strongly  curved  near  base;  ochreous-brown,  in 
some  lights  purplish-tinged;  a  few  black  scales  on  veins  near  apex: 
cilia  brownish-ochreous.      Hindwings  with  termen  rounded;  bright 


136  NEW    GENERA    AND    SPECIES    OF    LEPIDOPTERA. 

crimson;  termen  near  apex  ochreous-tinged;  a  few  fuscous  scales 
at  end  of  cell;  three  or  four  subterminal  fuscous  dots  opposite 
mid-termen;  cilia  brownish-ochreous,  a  fuscous  dot  at  apex,  at 
tornus  crimson,  on  inner  margin  whitish. 

Brisbane,  Q.;  in  March;  one  specimen  at  light  (Central  Rail- 
way Station). 

The  bright  crimson  hindwings  make  this  an  exceptionally 
distinct  and  brilliant  species. 

HyBL^A    SYN.EMA,  n.Sp. 

Differs  from  Hyhlcea  puera,  Cram.,  as  follows  : — Size  smaller, 
26-30  mm.  (as  against  33-40  mm.  in  ^:>we?Yt).  Antennae  of  male 
slightly  thickened,  and  with  slightly  projecting  scales  on  upper 
surface  (in  puern  they  are  quite  smooth).  Forewings  narrowei-; 
markings  brighter  and  clearer;  usually  with  a  conspicuous  black 
discal  dot ;  an  outwardly  curved  oblique  whitish  fascia  from 
dorsum  at  ^,  narrowing  to  a  point  at  mid-disc;  the  oblique  sub- 
apical  line  preceded  by  a  whitish  suffusion.  Hindwings  with 
spots  yellowish,  not  or  only  slightly  tinged  with  red. 

I  owe  the  discrimination  of  this  species  to  Mr.  F.  P.  Dodd, 
who  has  bred  both  it  and  puera  from  the  larvae.  H.  synoima  is 
extremely  constant  in  the  markings  of  the  forewings;  in  puera 
these  vary  much,  but  never  have  the  whitish  markings  so  well 
developed.  On  comparing  a  long  series  of  both  species  their 
distinctness  is  evident,  and  if  any  doubt  remained  the  structural 
difference  in  the  male  antennae  (though  slight)  w^ould  be  sufficient 
to  remove  it.  H.  puera  very  frequently  goes  greasy;  synoima  does 
not 

Townsville,  Q. ;  in  January  and  February;  a  series  received 
from  Mr.  F.  P.  Dodd. 

The  larvae  of  syncenia  were  found  on  Tecoma  australis,  and 
were,  I  am  informed,  much  darker  on  the  dorsal  surface  than 
those  of  puera  which  were  found  on  Vitex  trifolia. 


137 


AN  ASCOBACTERIUM  FROM  THE  SUGAR-CANE, 

WITH  NOTES  UPON  THE  NATURE 

OF  THE  SLIME 

(Bacterium  sacchari,  n.sp.) 

By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

Society. 

(Plate   vi.) 

During  an  investigation  into  the  gummosis  of  the  sugar-cane 
an  organism  was  met  with  great  frequency ;  in  fact,  so  often  was 
it  found  that  I  cannot  but  regard  it  as  a  normal  inhabitant  of 
the  cane.  In  nine  cases  out  of  ten  it  was  found  in  tubes  of  media 
inoculated  in  the  fields  from  the  gum  of  diseased  plants,  and  also 
from  the  juice  of  healthy  stalks.  It  was  also  separated  in  the 
laboratory  from  the  tissue  of  gummed  Rappoe,  and  of  healthy 
Malabar  and  Tanna  canes.  The  organism  was  peculiar,  inasmuch 
as  under  certain  conditions  it  formed  asci,  which  are  well  defined 
masses  of  capsulated  bacteria.  The  conditions  under  which  these 
were  produced  included  growth  upon  a  solid  medium  and  the 
presence  of  a  sugar. 

The  asci  were  occasionally  formed  in  fluid  media,  but  only  when 
the  layer  of  fluid  was  shallow,  and  the  bacteria  grew  as  a  ring- 
around  the  junction  of  the  surface  with  the  glass.  The  solid 
medium  which  was  most  suited  to  the  growth  of  the  bacterium, 
and  for  the  formation  of  asci  was  cane  gelatine,  which  is  prepared 
by  dissolving  10  per  cent,  of  gelatine  in  cane  juice,  or  in  strong- 
diffusion  liquor,  and  neutralising  the  medium  to  phenolphthalein 
with  dilute  potassium  hydrate. 


138  AN    ASCOBACTERIUM    FROM    THE    SUGAR-CANE, 

The  bacterium  was  grown  in  the  presence  of  various  sugars, 
and  it  was  found  that  either  dextrose,  levulose  maltose,  or  saccha- 
rose would  serve  equally  well  as  a  source  from  which  the  asci 
could  be  formed.  Starch  and  lactose  were  useless  for  this  purpose. 
In  the  presence  of  a  suitable  sugar,  salts  such  as  calcium  chloride, 
magnesium  sulphate,  did  not  show  any  advantage  over  potassium 
phosphate  in  accelerating  the  growth  in  gelatine  media. 

There  are  many  races  of  the  bacterium,  and  these  may  be 
classified  into  two  groups.  The  bacteria  of  one  of  the  groups 
form  a  pale  yellow  growth  on  gelatine  and  agar  media;  they 
liquefy  gelatine  slowly,  and  produce  many  asci.  The  organisms 
of  the  other  group  are  deep  yellow  on  agar  and  gelatine;  they 
liquefy  gelatine  quickl}^,  and  produce  few  asci.  The  gummy 
substance  of  the  asci  from  both  groups  is  identical.  On  continued 
cultivation  in  the  laborator}^,  the  ^''ellow  rapidly  liquefying  races 
become  paler  yellow  or  cream-coloured,  and,  losing  the  greater 
part  of  their  liquefying  power,  they  become  identical  with  the  first 
or  normal  type  whose  specific  characters  are  given  at  the  end  of 
this  paper. 

When  the  growth  was  scraped  from  the  surface  of  saccharose- 
agar  and  heated  with  water,  a  slimy  emulsion,  like  unbeaten 
white  of  egg,  was  obtained;  and  as  I  was  at  that  time  searching 
for  the  gummosis  bacterium,  this  organism  seemed  to  be  very 
promising.  But  as  the  slime  had  to  be  tested  chemically,  and  its 
relation  to  the  gum  of  the  sugar-cane  investigated,  a  considerable 
quantity  of  the  culture  with  the  accompanying  asci  was  necessary. 
To  obtain  a  sufficiency  of  material  the  bacteria  were  sown  upon 
the  surface  of  a  neutral  medium  contained  in  large  covered 
vessels.  The  medium  contained  peptone  10,  saccharose  100, 
sodium  phosphate  2,  potassium  chloride  5,  agar  20,  and  tap-water 
1,000  grms.  In  about  a  week  at  30°  the  growth  seemed  to  have 
reached  a  maximum,  and  after  soaking  in  water  for  about  a 
quarter  of  an  hour  the  culture,  which  had  become  considerably 
swollen,  was  easily  separated  from  the  agar. 

The  swollen  emulsion  was  of  a  deep  yellow  colour,  and  had  the 
consistency  of  unbeaten  white  of  egg.     Numerous  attempts  were 


BY    R.    GREIG    SMITH.  139 

made  to  break  up  the  asci  and  obtain  a  solution  of  the  slimj'^ 
material.  The  most  successful  method  was  found  to  consist  in 
heating  the  emulsion  in  the  autoclave  up  to  three  atmospheres 
pressure.  With  this  treatment  the  emulsion  separated  into  a 
viscous  solution  and  a  yellow  sediment.  The  slightly  opalescent 
viscous  solution  was  clarified  by  using  small  quantities  of  alu- 
minium hydrate,  care  being  taken  to  avoid  an  excess  of  the 
hydrate,  which  coagulated  the  gummy  material  of  the  solution. 

The  clear  solution  was  viscous,  but  readily  passed  through  filter 
paper.  On  the  addition  of  alcohol  a  voluminous  white  curd}^ 
precipitate  was  thrown  down  This  readily  swelled  up  in  water 
to  form  a  clear  gelatinous  unfilterable  emulsion.  On  standing  for 
some  weeks  the  sterilised  and  clear  emulsion  became  slighth^ 
turbid,  a  light  flocculent  portion  separating  out,  leaving  a  filterable 
solution.  From  this  behaviour  it  would  appear  that  treatment 
with  alcohol  alters  the  outside  surface  of  the  lloccules  to  an 
insoluble  modification,  so  that  on  subsequent  treatment  with 
water  each  particle  of  dissolved  slime  is  enclosed  in  a  thin  mem- 
brane. The  addition  of  alcohol  to  the  gelatinous  solution  again 
gave  a  curdy  precipitate,  but  on  repeating  the  precipitation 
several  times  a  stage  was  reached  when  the  addition  of  alcohol 
produced  no  precipitate,  and  formed  an  opalescent  solution. 
However,  on  adding  traces  of  salts — such  as  common  salt — the 
curdy  precipitate  was  again  formed.  The  aqueous  solution  was 
tested  with  a  number  of  reagents,  and  the  reactions  obtained  are 
as  follows  : — 

EkACTIONS    of    the    mucilaginous   slime    of   the    ASCI. 

Lead  acetate...  ...         ...         ...  opalescence;  opaque  solution  on  heating 

Basic  lead  acetate    ...          ..  ...  precipitate 

Ammoniacal  lead  acetate  ...         ...  precipitate 

Barium  hydrate       ...  ..  .  .  no  precipitate 

Calcium  hydrate      ...          ...          ...  precipitate 

Copper  sulphate       no  precipitate 

Ferric  chloride          ...         ...         ...  no  ppt.  followed  by  ammonia  gave  no 

ppt.  of  Fe  (OH). 

Hydrochloric  acid,  dil opalescence 

Alcohol  ...         ...         ...         ...  curdy  precipitate  in  presence  of  salts 


l-tO  AN    ASCOBACTERIUM    FROM    THE    SUGAK-CANE, 

Aluminium  hydrate  .  coagulation 

Acetic  acid,  dil.        ...  ...  ...   no  precipitate 

,,         ,,      glacial...          ...         ...   precipitate 

Acid  mercuric  nitrate         ...         ...  ppt.  soluble  in  excess;  ppt,  reddens  on 

heating 
Xanthoproteic  reaction       ...          ...  positive 

Sulphuric  &  phosphotungstic  acids   precipitate 
Acetic  and  tannic  acids      ..  ...  opalescence 

Acetic  acid  and  pot.  ferrocyanide...  strong  opalescence 
Hydrochloric  acid  and  pot.  mercuric 

iodide opalescence 

Sodium  hydrate  &  copper  sulphate     purplish  colour  with  slight  precipitate 
Strong  mineral  acids  ...  ..  reddish-brown  colour  on  heating 

No  reaction  with  KI3,  AgNOs,  KOH,  NaOH,  BaCl*.  picric  acid. 

These  reactions  indicate  that  the  gummy  matter  which  forms 
the  substance  of  the  asci  has  relations  with  the  carbohydrates  as 
well  as  the  proteids,  and  at  first  sight  might  be  taken  as  being  a 
mucin  body.  The  slimy  capsule  of  some  bacteria  has  been  said 
to  consist  of  a  substance  *' related  to  mucin,  or  probably  identical 
therewith."* 

Lepierref  claims  to  have  obtained  a  true  mucin  from  cultures 
of  a  fluorescent  bacterium.  Charrin  and  Desgrez;  obtained  a 
mucinous  body  from  bouillon  cultures  of  Bact.  pyocyaneum.  This 
body  swelled  up  with  water,  and  was  filterable.  It  was  precipi- 
tated by  alcohol,  acetic  acid,  the  mineral  acids,  common  salt  and 
magnesium  sulphate.  The  acetic  acid  precipitate  was  insoluble 
in  excess,  but  was  dissolved  by  dilute  alkalies.  According  to  the 
authors,  these  properties  showed  it  to  be  a  compound  mucinoid, 
but  as  it  contained  phosphoric  acid  it  was  probably  accompanied 
by  a  nucleoproteid.  A  nitrogen  determination  was  apparently 
not  made. 

Although  the  gummy  substance  of  the  asci  forms  a  viscous 
solution,  and  behaves  like  mucin  in  some  of  its  reactions,  still 
<lifferences  can  be  found.     The  mucins  are  soluble  in  lime  and 


*  Lafar.  Technical  Mycology,  189S,  40. 

t  Lepierre,  Comptes  Kendus,  1898  (126),  761. 

+  Charrin  and  Desgrez,  ibid,  596. 


BY    R.    GREIG    SMITH.  141 

baryta  waters;  this  is  insoluble  in  lime  water.  The  colour  re- 
actions of  the  albumens,  which  are  generally  decided  in  the  case 
of  the  included  mucins,  are  not  so  with  this  substance.  Finally, 
the  nitrogen  percentage  shows  that  it  is  neither  a  mucin  nor  an 
allied  substance. 

In  the  estimation  of  the  nitrogen  the  clear  aqueous  solution 
was  precipitated  with  alcohol  and  redissolved  in  water  several 
times  until  an  opalescent  alcoholic  solution  was  obtained.  This 
was  divided  into  two  parts,  one  of  which  was  precipitated  with 
common  salt,  the  other  with  barium  chloride.  The  precipitates 
were  then  washed  with  alcohol  and  dried  until  they  ceased  to 
lose  weight.  Portions  were  taken  for  the  ash  determination  and 
for  the  nitrogen,  which  was  estimated  by  the  Kjeldahl  process,, 
due  allowance  being  made  for  the  nitrogen  in  the  sulphuric  acid,, 
etc.,  by  check  tests  with  pure  saccharose. 

Portion  precipitated  by  common  salt : — 

0-1542  grm.  gave  0-0128  grm.  ash  =  8-30%. 

The  ash  consisted  of  carbonate,  phosphate,  sulphate, 

and  chloride  of  soda. 
0-2164  grm.   required  0-5  c.c.  ^/o   acid  to  neutralise  the 

ammonia  .-1-76%  nitrogen  in  the  ash-free  substance. 
Portion  precipitated  b}'  barium  chloride  : — 

0-1054  grm.  gave  0*0184  grm.  ash=  17*47%. 

The  ash  consisted  of  carbonate,  phosphate,  sulphate, 

and  chloride  of  barium. 
0-1820  grm.  required  0-25  c.c.  ^/.^   acid   to  neutralise  the 

ammonia  =  1"16%  nitrogen  in  ash-free  substance. 

The  amount  of  dry  gum  in  the  cultures  is  very  small — a  ver}^ 
viscous  solution  contains  very  little  solid  gum,  and  on  this  account 
only  a  small  quantity  was  available  for  the  determination.  But 
although  these  results  differ  from  one  another,  still  they  are 
sufficient  to  show  that  the  nitrogen  in  the  dry  and  ash-free  slime 
is  less  than  2%.  This  indicates  that  it  is  far  removed  from  mucin, 
which  contains  about  12%,  from  pseudomucin  with  about  10%, 
and  from  colloid  with  about  7%.     The  presence  of  phosphoric  acid 


142  AN    ASCOBACTERIUM    FROM    THE    SUGAR-CANE, 

in  the  ash  has  no  signiticance  because  the  culture  media  contained 
0*2%  of  sodium  phosphate.  On  the  whole  the  albuminoid  re- 
actions are  rather  undecided;  the  two  which  were  undoubtedly 
obtained  were  the  xanthoproteic  and  the  phosphotungstic;  but 
these  tests  alone  do  not  give  an}^  definite  information  regarding 
the  nature  of  the  slime. 

If  the  slime  were  originally  pectose,  the  method  of  separating 
it  from  bacteria  by  heating  under  pressure  would  have  converted 
it  to  pectin,  which  gives  a  precipitate  on  the  addition  of  barium 
hydrate;  and  as  the  purified  slime  does  not  precipitate  with 
barium  hydrate,  it  is  evident  that  it  cannot  be  pectose.  Nor 
can  it  be  any  of  the  pectin  bodies.  That  the  slime  contains 
a  pentosan  is  shown  by  the  production  of  furfural  on  dis- 
tillation with  hydrochloric  acid,  sp.g.  1*16,  and  also  on  careful 
heating  with  strong  sulphuric  acid.  On  hydrolysis,  with 
dilute  sulphuric  acid  at  three  atmospheres  pressure,  a  reducing 
substance  is  formed.  This,  upon  prolonged  heating  on  the  water 
bath  with  phenylh3^drazine,  gives  an  osazone  consisting  of  balls  of 
short  needle-shaped  crystals,  which,  when  recrystallised,  have  a 
melting  point  of  153°  C.  The  product  of  acid  hydrolysis  is  pro- 
bably a  pentose,  and  appears  to  be  similar  to  that  obtained  b}^ 
Bendix*  on  hydrolysing  tubercle  and  other  bacteria.  The  pentose 
from  tubercle  bacteria  reduced  Fehling's  solution,  and  formed  an 
osazone  which  melted  at  153°- 155°. 

The  formation  of  a  pentose  shows  that  the  bacterial  slime  is 
related  to  the  vegetable  gums. 

Bacterium  sacchari,  n.sp. 

Shape,  etc.—Ps.  short  rod,  co^i-like,  with  rounded  ends  measures 
0*6:l-2/M,  and  occurs  singly  in  pairs  and  threads.  On  solid  media 
•containing  sugar,  and  sometimes  on  the  surface  of  fiuid  sugary 
media,  it  forms  asci.  The  rods  are  motile,  and  the  flagella  when 
stained  by  the  night-blue  method  were  seen  to  vary  from  one 
(terminal)  to  nine  (peritricial).      With  the  blues  the  cells  stain 

*  Bendix,  Jour.  Chem.  Soc.  1901,  Abs.  ii.  'J06. 


BY    R.    GREIG    SMITH.  143 

feebly;  fuchsin,  and  especially  violet,  stain  well.  Staining  l)y 
Gram's  method  is  negative.     No  spores  are  formed. 

Kelation  to  oxygen,  etc. — The  bacterium  does  not  grow  anaero- 
bically,  and  the  optimum  temperature  is  28°  C. 

;^%  glucose-gelatine  ■plate.- — The  colonies  are  white  glistening, 
slightly  raised  and  irregular.  When  magnified  they  are  seen  to 
have  a  dark  clouded,  convoluted  or  areolate  centre,  and  a  finely 
granular  margin,  w^ith  a  crenate  structure  near  the  smooth  edge. 

Nutrient  gelatine  plate. — Whitish  or  cream-coloured,  round, 
raised,  glistening  colonies.  When  magnified  the  surface  colonies 
are  finely  granular,  the  subsurface  colonies  are  coarsely  granular, 
and  the  deep  colonies  are  rounded  and  opaque. 

Nutrient  agar  plate. — The  colonies  vary  in  colour  from  dirty 
white  to  buff-white;  they  are  rounded,  flat  and  glistening;  the 
centre  becomes  depressed.  When  magnified  the  rounded  colonies 
are  fineh''  granular,  with  large  granules  distributed  chiefly 
round  tlie  centre.  The  older  surface  colonies  are  coarsely  granular, 
with  a  puckered  or  wrinkled  centre.  In  crowded  plates  the  small, 
round,  coarsely  granular  colonies  maytiave  a  marginal  ring  (halo). 
The  deep  colonies  are  rounded  or  lenticular  with  large  coarse 
granules. 

Nutrient  agar  stroke. — The  growth  is  cream-coloured  at  37°, 
yellowish-white  at  22°,  slightly  raised,  glistening,  and  has  a 
lobular  margin.     The  condensed  water  carries  a  slight  film. 

Neutral  cane-agar  stroke. — A  luxuriant,  almost  transparent, 
yellowish-white,  raised,  spreading  growth  is  formed.  It  has  a 
gelatinous  consistency,  and  the  condensed  water  is  slightly  viscous. 

Neutral  cane-gelatine  stroke. —  An  ivory-white  or  yellowish-white 
glistening,  irregularly  raised  growth,  with  an  irrregular  serrated 
margin,  and  gummy  or  gelatinous  consistency  is  produced.  The 
culture  slowly  gravitates,  and  forms  a  thick  gummy  mass  at  the 
base  of  the  slope.  Sometimes  there  is  no  apparent  liquefaction 
of  the  medium  until  the  gravitated  gummy  mass  is  removed  and 
a  pit  revealed ;  at  other  times  the  site  of  the  stroke  l^ecomes 
furrowed  by  a  partial  liquefaction  of  the  medium. 


144  AN    ASCOBACTERIUM    FROM    THE    SUGAH-CANE, 

Nculral  cane-gelatine  stab. — The  needle  track  shows  a  filiform 
i^rowth,  with  a  hemispherical  yellowish  nail-head.  Neither  in 
this  nor  in  a  glucose-gelatine  shake  were  gas  bubbles  produced. 

Nutrient  gelatine  stab. — The  growth  is  filiform,  with  a  thin, 
flat,  white,  glistening  and  spreading  nail-head.  The  nail-head  is 
depressed,  but  there  is  no  sign  of  liquefaction  within  a  week  with 
the  normal  races.  After  the  seventh  day  the  gelatine  beneath 
the  nail  head  either  appears  consumed  or  shows  a  slight  crateri- 
form  liquefaction. 

Potato. — The  growth  is  thin,  dry,  deep  yellow,  flat  and  glisten- 
ing. 

Carrot  and  turnip. — A  slimy,  translucent,  whitish  growth 
rapidly  spreads  over  the  surface. 

Sugar-cane.  —A  yellow,  glistening,  gummy  growth  spreads  over 
the  surface,  and  growing  down  the  vessels  exudes  at  the  lower 
ends  as  raised  globules  similar  in  appearance  to  the  exuded  gum 
of  gummy  cane. 

Siveet  ivort. — The  growth  is  luxuriant. 

Bouillon. — The  medium  becomes  turbid,  and  forms  a  film  and 
sediment.      A  faint  indol  reaction  is  sometimes  obtained. 

Nitrate  bouillon.  —Nitrate  is  reduced  to  nitrite. 

Milk.  —  The  medium  is  coagulated  about  the  tenth  da}'  at 
28^  C,  and  the  reaction  is  faintly  acid. 

The  ascobacteria  which  have  been  described  are  few  in  number. 
Bact.  luteum  (List.),  Adametz,*  is  non-motile,  and  produces  asci 
in  the  absence  of  sugar.  The  colour  of  the  gelatine  cultures  is 
decidedly  yellow,  and  it  does  not  appear  to  liquefy  gelatine. 
Bac  citreus,  Xlnna,"^  gro\\'s  ver}^  slowl}'-  in  gelatine,  and  produces 
asci  in  sugar-free  media.  Ascobacillus  aqitatilis,  Moreno,!  is 
stained  b}-  Gram's  method,  and  grows  quickly  at  37°.  Like  the 
others  the  asci  are  formed  in  sugar-free  media.  Since  this 
organism  forms  asci  only  in  the  presence  of  sugar,  and  otherwise 
differs  from  the  bacteria  hitherto  described,  I  have  named  it 
Bacte^num  sacchari. 

*  Migula,  System  der  Bakt.  1900. 
t  Moreno,  Cent.  f.  Bakt.  1  Abt.  80,  111. 


P.L.S.N.S.W.,    1  902 


^  /^dc;^fCi^ 


AUSTRALIAN     MOLLUSCA. 


P.L.S.N.S.W.,    1902. 


C./%<:;i^^  a^ 


AUSTRALIAN     MOLLUSCA. 


^  /¥c<^^  a^ 


AUSTRALIAN     MOLLUSCA. 


P.L.S.N.S.W..    1902. 


1     .. 


t. 


!-d 

.i.lr'^-^i 

i 

1 

^ 

^ 

L 

«:    'i 

1%  „  1 

BACTERIUM    VASCULARUM. 


P.L.S.N.S.W.,    1902. 


Fig.  1. 


Fig.  2. 

.BACTERIUM    VASCULARUM,    Cohh    (x'/| 


P  L.S.N. SW.,   1902 


UCTERI'JM    A3C0FCRMANS    SACCMARI. 


PLSN.SW.   1902 


PI  VII. 


IV  AH.   del 


GYROCOTVLE. 


BY    R.    GREIG    SMITH. 


145 


DESCRIPTION  OF  PLATE. 


-Balsam  film  from  growth  upon  cane-gelatine.  The  asci  are  shrunken 
by  the  dehydration.  The  cellular  structure  of  some  of  the  asci  can 
be  made  out  from  the  ascus,  which  is  slightly  out  of  focus,  at  the 
bottom  left-hand  corner.  Stained  with  dilute  carbol  fuchsin,  and 
washed  with  alcohol,      x  1000. 

-Flagella  stained  by  the  night-blue  method,      x  1000. 


10 


146 


PRELIMINARY  NOTE  ON  THE  GEOLOGY  OF  THE 
QUEENSLAND  COAST  WITH  REFERENCES  TO 
THE  GEOGRAPHY  OF  THE  QUEENSLAND  AND 
N.S.   WALES  PLATEAU. 

By  E.  C.  Andrews,  B.A. 


SYNOPSIS  OF  CONTENTS. 

A. — Introduction     ...         ...         ...         ...         ...         ...         ...         ...  p.  14(i 

B. — Physiography p.  148 

1.  General  Appearance  of  Coast  from  Tweed  Heads  to  Cairns. 

2.  Coastal  Plains. 

3.  The  Continental  Shelf. 

4.  Islands. 

General  appearance  and  structure. — Stradbroke,  Moreton,  Frazer, 
Curtis,  Gloucester,  The  Palms,  Hinchinbrook,  Goold,  Dunk, 
The  Barnards,  Fitzroy,  and  Green  Island. 

5.  Marine  Erosion. 

C. — Elevation  p.  164 

D. — Petrological  Notes  on  some  Queensl.\nd  Rocks p.  167 

E. — Summary  AND  Conclusion       p.  173 


Introduction. 

Rock  composition,  geologic  structure,  coastal  movements,  and 
the  agents  of  denudation  are  the  codepcndent  criteria  of  any 
theory  as  to  the  origin  of  present  land  forms. 

Of  late  years  the  study  of  the  topographical  features  of  a 
country  as  illustrating  its  geological  histor}'  has  received  much 
attention. 

The  evolution  of  mature  river  systems  from  plateaux  of  accu- 
mulation and  erosion  ;  the  appearances  of  land  forms  in   gently 


BY    E.    C.     ANDREWS.  '  147 

sloping  and  steeply  inclined  strata  and  the  influences  of  subsi- 
dence, elevation  and  marine  erosion  in  the  formation  of  coast 
lines  are  clearly  brought  out  by  this  branch  of  geological  research. 

The  advantage  of  grasping  the  meaning  of  the  more  important 
scenic  features  of  a  locality  is  that  a  cursory  visit  to  the  place 
will  often  suffice  to  make  clear  the  leading  points  in  its  history. 

For  instance,  a  mere  cruise  along  the  coast  is  sufficient  to  note 
"  the  regular  and  rhythmical  curves  "*  which  mark  the  effect  of 
long -continued  marine  erosion;  the  multitude  of  islands  surround- 
ing an  irregular  coast  which  speak  of  recent  subsidence ;  or 
pyramidal  and  flat-topped  mountains,  skirted  by  coastal  plains, 
which  tell  the  passer-by  of  plains  of  accumulation  elevated  by 
crustal  movements  to  plateaux  of  accumulation,  the  subsequent 
loss  suffered  through  the  agents  of  deundation;  of  the  subsidence 
which  followed  the  formation  of  river  systems  in  the  plateaux  of 
accumulation,  and  the  elevation  which  transformed  the  marine 
beds  (formed  during  subsidence)  to  coastal  plains. 

The  following  brief  notes  on  the  geology  of  Queensland  are 
based  mostly  on  observations  during  a  short  trip  taken  by  Mr. 
C.  Hedley  and  myself  along  the  Queensland  coast,  and  are 
intended  merely  as  a  preliminary  note  to  a  more  extended  paper 
on  the  Tertiary  history  of  the  Cordillera,  the  evolution  of  the 
present  river  systems,  and  the  various  stages  of  development  of 
the  Queensland  and  New  South  Wales  coasts. 

To  these  notes  on  Queensland  coast  topography  are  added 
references  to  N.S.  Wales  topography  based  on  personal  observa- 
tions along  the  plateau  and  coast. 

I  desire  here  to  express  my  sincere  thanks  to  Mr.  Hedley  foi~ 
much  valuable  information  concerning  the  Queensland  coast,  and 
at  whose  suggestion  the  trip  was  undertaken;  to  Mr.  J.  M. 
Newman,  B.E.;  Captain  Almond,  of  Brisbane;  Mr.  Patience,  of 
Townsville ;  Captain  Reader,  of  Lucinda  Pt.  ;  Mr.  Brooks,  of 
Tully  River;  Messrs.  Cutten  Bros.,  of  Clump  Pt.;  Mr.  George 
Butler,  of  Towmsville ;  and  many  others  for  information  and 
hospitality. 

*  Reclus. 


148  GEOLOGY  OF  QUEENSLAND  COAST, 


Physiography. 


1 .  General  apj^earance,  (he.  — From  the  Tweed  Heads  to  Thursday 
Island  the  coast  is  of  the  highly  indented  type,  bordered  with 
broad  coastal  plains.  A  wide  continental  shelf  exists  from  north 
to  south,  dotted  over  with  innumerable  islands,  many  of  them 
rugged  and  mountainous  in  character. 

South  of  the  Great  Barrier  Reef  we  have  the  large  islands 
known  as  Stradbroke  and  Moreton,  from  which  to  the  north  and 
west  extend  wide  coastal  plains,  especially  wide  in  the  neighbour- 
hood of  the  Glass  House  Mountains,  which  consist  of  trachytic 
lavas  weathered  into  most  fantastic  shapes. 

Further  north  Frazer  (80  miles  in  length)  and  Curtis  Islands 
occur,  both  separated  from  the  mainland  by  long,  narrow,  and 
shallow  channels,  also  accompanied  by  great  sandy  flats  and  small 
islands.  Lady  Elliot  Island  and  The  Bunkers  form  the  southern 
outposts  of  the  Great  Barrier  Reef,  and  occur  a  little  north  of 
Frazer  Island. 

From  ]Mackay  to  Cairns  the  coast  is  very  nigged.  Great 
mountains  advance  into  the  sea;  numerous  bold  promontories  and 
islands  such  as  Cape  Cleveland,  Cape  Upstart,  Castle  Hill,  Cape 
Grafton,  Lion  Island,  Gloucester,  Hinchinbrook,  AYhitsunday 
and  Palm  Islands  are  separated  from  the  mainland  by  long  and 
narrow  channels  or  broad  flats  a  few  feet  only  above  high- water 
mark. 

The  flats  which  gather  at  the  bases  of  these  headlands  and 
coastal  ranges  are  ver}''  conspicuous. 

In  many  cases  these  headlands  consist  of  barren  rocks,  showing 
a  great  wealth  of  enormous  precipices  and  escarpments  covered 
with  rounded  boulders,  illustrating  the  granitic  type  of  weathering. 

2.  Coastal  plains. — These  are  very  extensive  along  the  Queens- 
land coast,  and  represent  the  redistribution  b)'  tidal  action  of 
fluviatile  material,  and  to  a  less  extent  that  derived  from  the 
headlands  by  marine  erosion. 

Around  Brisbane  and  the  Glass  House  Mountains  these  flats 
are  as  much  as  20  miles  in  width.     To  the  west  the  mountains 


I 


BY    E.    C.    ANDREWS.  149 

rise  precipitously  from  them.  As  seen  during  a  journey  up  the 
Brisbane  River  they,  together  with  St.  Helena,  Mud  and  Peel 
Islands,  appear  due  to  a  slight  elevation.  The  Glass  House 
Mountains  stand  out  like  islands  in  the  centre  of  this  plain.  In 
this  connection  Mr.  J.  M.  Newman,  B.E.,  writes: — "I  think  they 
are  undoubtedly  the  result  of  recent  elevation.  For  fully  10-20 
miles  inland  the  country  is  more  or  less  flat  and  sand3\  For  a 
few  miles  in  from  the  coast  one  finds  even  now  bits  of  recent 
shells;  also  there  are  large  swamps  and  lagoons,  the  remnants 
apparently  of  rivers  whose  mouths  have  been  silted  up  by  Ijars. 
The  water  runs  in  them  now  only  in  flood  time.  What  ridges 
there  are  are  Ioav  and  sandy,  and  the  swamps  and  paddymelon- 
liole  gullies  run  between  them.  As  you  go  inland  rhe  ridges 
become  higher  and  less  sandy,  forming  gradually  the  spurs  of  a 
low  range  of  mountains  at  the  back  of  the  Glass  House  Mountains. 
These  latter  rise  sheer  out  of  the  coastal  flats,  and  seem  to  have 
been  submarine  in  origin.  ...  In  the  coastal  regions  one 
finds  a  few  feet  beneath  the  sand  a  sort  of  pipeclay,  with  ironstone 
nodules  extending  to  a  great  depth." 

Dense  mangrove  swamps  have  fastened  on  to  the  seaward  edges 
of  these  plains. 

Another  great  "  coastal  plain "  exists  off  the  mouth  of  the 
Burnett,  and  others  around  Maryborough. 

Well  marked  also  is  the  recent  gain  to  the  coast  by  these  flats 
in  the  neighbourhood  of  Rockhampton,  and  due  to  the  action  of 
the  Fitzroy  River. 

At  Keppel  Bay  great  dismal  mangrove  swamps  extend  for 
miles.     These  pass  into  the  flats  just  mentioned. 

Again  at  Mackay  similar  broad  flats  exist. 

When  nearing  Townsville  the  enormous  headlands  and  atten- 
dant flats  are  marked  features  in  the  scenery.  In  almost  every 
case  the  mountains  of  the  coast  line  rise  sheer  out  of  the  low 
ground  until  far  out  at  sea  they  appear  as  islands. 

Especially  emphasised  is  this  association  of  steep  mountain 
and  lowly  flats  in  the  neighbourhood  of  Hinchinbrook  Island  and 
the  Herbert  River  (Fig.  1). 


150 


GEOLOGY  OF  QUEENSLAND  COAST, 


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KY    E.    C.    ANDREWS.  151 

Here  great  mangrove  swamps,  miles  in  width,  pass  into  flats 
as  much  as  20  miles  across  in  places,  and  from  which  the  great 
coast  range  rises.  From  these  extensive  flats  monadnock-like"^ 
forms  arise,  for  example.  Glass  House  Mountains.  North  of 
Card  well  the  great  extent  of  coastal  plain  around  Rockingham 
Bay,  traversed  by  the  Tully,  Murray  and  Hull  Rivers,  forms  a 
conspicuous  object  in  the  topography. 

At  Cairns  other  great  coastal  flats  exist.  One  interesting 
plain  between  Cape  Grafton  and  False  Cape  may  be  noted  here. 
It  is  about  a  mile  across  and  two  miles  long  ;  and  but  a  few  feet 
above  high  tides. 

We  have  thus  found  these  large  flats  or  "  coastal  plains  "  raised 
but  a  few  feet  above  high  tides  to  exist  right  along  the  eastern 
coast  of  Queensland. 

They  appear  almost  undoubtedly  to  be  matter  derived  from  the 
large  coastal  rivers  and  from  the  forces  of  marine  erosion  redistri- 
buted by  the  various  agents  of  along  shore  action. 

"They  are  due  to  fluviatile  action  being  invariably  absent  from 
coasts  possessing  no  considerable  rivers."! 

A  slight  elevation  would  account  for  their  present  position. 

Similar  flats  will  be  described  under  the  head  of  islands. 

Associated  with  these  plains  are  numerous  lagoons,  lagoon 
marsh  meadows,  and  shallow  creeks.  Especially  well  are  these 
seen  along  the  coast  between  Rockhampton  and  the  Tweed  Heads. 

3.  The  Continental  Shelf. — This,  in  Queensland,  maintains 
generally  a  great  width.  To  the  south  of  the  Great  Barrier  Reef 
it  is  represented  by  reefs  of  old  stratified  rocks,  enormous  shoals, 
and  soft  deposits  uniformly  distributed  over  the  surface,  and 
having  a  gradual  dip  to  the  east,  and  which  rise  40  or  50  miles 
away  to  the  east  from  enormous  depths. 


""  An  isolated  hummock  or  hill  rising  from  a  surface  developed  by  subaerial 
Agencies  to  extreme  old  age. 

t  Prof.  Pencke,  Morphologie  der  Erdoberfliiche. 


152  GEOLOGY  OF  QUEENSLAND  COAST, 

On  the  northern  submarine  extension  of  Breaksea  ^Spit, 
which  is  composed  almost  entirely  of  "  siliceous  material,' "^  are 
situated  Lady  Elliot  Island  and  the  Bunker  Groups. 

Northwards  again  the  outer  edge  of  the  shelf  is  represented  by 
the  Great  Barrier  Reef  itself,  occurring  at  distances  varying  from 
15  to  100  miles  from  the  coast.  Long  lanes  of  coral  sand  and 
wide  openings  l)reak  into  the  integrity  of  the  ocean  front  of  the 
Reef. 

Deep  gutters  of  valley  form  occur  in  a  few  instances  on  the 
continental  shelf,  and  in  the  vicinity  of  the  outer  barrier,  t 

A  few  miles  from  the  shore  line  great  numbers  of  precipitous 
islands  exist,  rising  from  shallow  water,  rarel}'  if  ever  exceeding 
ten  fathoms  in  depth.  These  islands  are  often  accompanied  by 
small  fringing  reefs,  and  in  many  cases  also  by  plains,  miniatures 
of  the  large  occurrences  on  the  mainland,  and  described  under 
"Coastal  Plains."  In  man}'-  cases  the  axes  of  the  mountain 
ridges  composing  the  islands  are  parallel  to  the  main  coast  line, 
and  are  separated  therefrom  by  narrow  channels  {e.g.,  Hincliin- 
l:)rook,  MoUe,  and  Albany  Passages). 

4.  Islands — Stradbroke,  Moreton  and  Frazer  Islands  are  huge 
sand  piles  lying  to  the  south  of  the  Great  Barrier  Reef.  They 
are  respectively  35,  20  and  SO  miles  in  length,  and  appear  due  to 
the  redistribution  of  tluviatile  and  other  material  by  marine  and 
ieolian  agencies.  In  the  cases  of  8tradbroke  and  Moreton  Islands 
the  northern  extremities  consist  of  older  stratified  rocks,  and 
these  uncovered  masses  of  continental  rocks  doubtless  belong  to 
a  series  of  small  insular  patches  on  which  the  threefold  action  of 
rivers,  wind  and  currents  have  worked,  first  to  tie  the  islands 
together,  and  afterwards  to  fashion  the  large  sand}^  masses  nito 
their  present  appearance.  As  a  result  of  long-continued  residence 
and  observation  on  those  islands,  Mr.  Hedley  is  of  ojiinion  that 


*  Jukes,  Voya^^e  of   "FI3',"  Vol.  i.,  p.  318.     See  also  A.  Agassiz,  '•  Great 
Barrier  Reef  of  Australia,"  pp.  64,  105. 

t  Ibid.,  Yol  i.,  p.  845. 


BY    E.    C.    ANDREWS.  .  153 

the  only  continental  rocks  now  visible  are  those  to  be  found  at 
tlie  nortliern  island  extremities. 

Frazer  Island  likewise  appears  to  be  composed  of  sand.  In  all 
cases  these  islands  are  associated  with  large  rivers  and  coastal 
plains.  They  have  straight  eastern  coast  lines  50  or  60  miles 
long,  while  straight  or  swinging  sandy  coasts  with  enormous  coast 
plains,  lagoons,  creeks,  monadnocks,  lagoon-marsh  meadow^s,  and 
w  ith  ,2;ently  sloping  off-shore  deposits  constitute  the  chief  features 
of  the  neighbouring  mainland. 

Near  the  mouth  of  the  Fitzroy  River  numerous  insular  patches 
occur,  the  longest  of  which  is  Curtis  Island,  running  parallel  to 
the  mainland,  and  separated  from  it  by  a  long  narrow  channel,  in 
wliich  the  variation  in  the  tides  amounts  to  as  much  as  27  feet. 
This  island  is  composed  in  great  part  of  sfind;  it  contains  signs  of 
elevated  beaches,*  and  is  associated  with  extensive  coastal  plains. 

The  Beverley  and  Northumberland  Groups  comprise  numerous 
small  rocky  and  pine-clothed  islands,  usually  presenting  mural 
fronts  to  the  sea — the  result  of  marine  erosion. 

In  the  Whitsunday  Group  are  islands  of  considerable  size, 
separated  from  each  other  and  the  main  coast  by  narrow  channels 
of  great  length. 


*  As  no  notice  of  this  has  appeared  in  literature,  Iledley  supplies  the 
following  note  :  — 

"  A  striking  instance  of  apparent  elevation  was  shown  to  me  by  Mr,  J.  B. 
Paterson,  the  lessee  of  Curtis  Island,  Q.  A  few  miles  south-west  of  the  Cape 
Capricorn  Lighthouse  occur  what  are  locally  called  the  Marine  Plains. 
These  are  an  extensive  series  of  swampy  Hats,  some  miles  inland,  occupying 
bays  in  the  hilly  country  and  margined  by  a  continuous  beach-bank,  com- 
posed of  sand  and  recent  sea  shells.  Within  the  bays  are  small  rocky 
'  islands '  with  wave -eroded  flanks,  and  also  encircled  with  beach-banks. 

"  When  I  saw' it  fifteen  years  ago  this  scene  impressed  me  as  indicating  a 
recent  and  slight  upheaval.  But  further  study  of  beach-formation  convinces 
me  that  elevation  is  not  the  only  possible  explanation  of  the  phenomena 
described.  Should  the  entrance  of  a  shallow  bay  become  blocked  by  beach- 
banks  or  dunes,  the  consequent  shutting  out  of  the  sea  might  leave  the  head 
of  the  bay  in  such  a  condition  as  the  Marine  Plains  of  Curtis  Island,  without 
the  intervention  of  elevation." 


154  GEOLOGY    OF    QUEENSLAND    COAST, 

Gloucester  Island  rises  abruptly  from  the  sea  to  a  height  of 
2,000  feet,  and  from  a  short  distance  much  resembles  a  giant 
headland,  so  narrow  is  the  channel  between  it  and  the  Queensland 
shore. 

Magnetic  Island,  1,600  feet  in  height,  is  roughly  rectangular 
in  plan,  and  composed  of  a  dark  granite  admirably  adapted  for 
building  purposes.  Unlike  the  granite  of  the  Towns ville  foreshores 
(five  miles  distant)  and  Castle  Hill,  it  is  homogeneous  in  character, 
and  iDossesses  cuboidal  jointing.  This  island  is  further  interesting 
by  reason  of  certain  rock-markings  which  point  to  recent  eleva- 
tion. Mr.  G.  Butler,  of  Towns  ville,  mentions  the  presence  of 
masses  of  oyster  shells  at  a  height  of  from  10  to  15  feet  above 
H  W.M.,  and  removed  some  50  feet  from  the  sea.  These  shells, 
he  says,  are  attached  to  the  solid  granite  composing  the  island, 
and  not  to  loose  boulders.  Maitland,  also,  from  a  discover}'  of 
pumice  fragments  some  distance  from  the  present  sea  level,  claims 
recent  elevation  for  this  island."^ 

The  Palm  Islands. — This  cluster  was  examined  in  some  slight 
detail,  and  was  found  to  consist  of  granites  of  varying  acidity 
and  quartz-porphyry,  traversed  by  regular  networks  of  basalt 
dykes. 

Generally  speaking,  they  are  rugged  and  barren  in  character, 
and,  with  the  exception  of  a  few  sandy  bays,  possess  precipitous 
and  rocky  coasts.  Large  black  rounded  granite  tors  cover  the 
hill  sides. 

South  Palm  is  7  miles  in  length  and  about  3  in  width.  It 
consists  of  two  peaks  1,400  and  1,820  feet  respectively  in  height. 
The  rock  of  the  island  is  an  ordinary  ternary  granite,  with  a 
tendency,  however,  to  pass  into  acid  types  by  reason  of  a  diminu- 
tion in  the  quantity  of  biotite.  Large  segregations  of  quartz  and 
pink  orthoclase  may  be  seen  in  the  granite  outcrops.  Masses  of 
orthoclase  occur  as  much  as  1 2  to  1 5  inches  in  diameter,  as  also 
quartz.      On  the  north-east  of  the  island  a  remarkable  floor  or 

*  Maitland,  Kept.  Magnetic  Island.     Brisbane,  bv  authority,  1892. 


BY    E.    C.    ANDREWS.  155 

table  of  aplite  occurs,  traversing  the  ternary  granite  of  the  locality. 
It  has  the  appearance  of  a  fine-grained  white  sandstone.  Its  dip 
is  about  10°,  and  it  may  be  traced  over  an  area  of  at  least  five 
acres. 

At  Challenger  Bay  an  interesting  Hat  occurs  joining  two  high 
points  of  the  island,  which,  but  for  its  presence,  would  exist  as 
separate  islands. 

The  flat  is  about  one  and  a  half  miles  in  length  and  500  yards  in 
width,  with  an  average  height  above  H.W.M.  of  at  least  15  feet. 

A  gutter  some  six  feet  wide  and  ten  feet  deep  has  been 
excavated  through  the  centre  of  this  flat  by  the  forces  of  erosion. 
The  structure  of  the  plain  as  revealed  by  this  miniature  valley  is 
a  sandy  clay,  coarse  sand,  and  a  clay  containing  angular  frag- 
ments of  granite.  No  shells  or  calcareous  material  were  seen, 
although  careful  search  was  made. 


Fig.  2. — Sketch  Section  across  Palm  Island  "  Katsed  Plain." 

a.  Mt.    Bentley,   1800  feet  high.  />6.    Granite  of    Palm  Island. 

cc.  Raised  Plain,  H  miles  long,  600  yards  wide,  and  from  10  to  15  feet 
above  H.W.M.         dd.  Probable  Contour  of  underlying  Granite. 

On  the  southern  side  of  this  flat  large  quantities  of  pumice 
occur,  high  above  high  water  mark,  and  stretching  inland  for 
some  ten  chains.  The  flat  is  covered  at  present  with  growths  of 
eucalypts,  pandanus,  and  coarse  grass  about  four  feet  in  height. 

A  similar  though  smaller  occurrence  was  observed  b}'^  Mr. 
Hedley  near  Bentley's  Creek. 

These  flats,  in  my  opinion,  point  undoubtedly  to  the  redistri- 
bution by  tidal  action  of  matter  lost  to  the  mainland  and  adjacent 
islands,  but  principally  by  loss  to  the  South  Palm  itself  by  the 
forces  of  marine  erosion  acting  on  the  windward  side. 


156 


GEOLOGY  OF  QUEENSLAND  COAST, 


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BY    E.    C.    ANDREWS. 


157 


JSigiis  of  elevation  were  also  present  on  this  island  in  the  shape 
of  beach  rock  apparently  in  situ,  and  some  eight  feet  above  high 
tides. 

The  reefs  around  the  >S.  Palm  Island  are  of  the  small  fringing 
type,  and  in  common  with  those  existing  on  the  other  islands  on 
the  Queensland  continental  platform  lack  the  luxuriance  and 
beauty  of  those  seen  on  the  outer  edge  of  the  Great  Barrier  Reef 
or  those  in  Fiji  waters  (Fig.  3a). 


%i||!i|||IP^ 


Fig.  3a. — Sketch  Plan  of  Frinuin(4  Reef  at  South  Palm  Island. 

1.  Coast  Mountains.  2.  Beach.  3.  Fringing  Keef.  4.  Gutter  in 
Coral  Reef  (5  chains  long).         5.  Sea  (10  fathoms  deep;. 

A  curious  and  instructive  feature  was  observed  in  the  frinsin^- 
reef  to  windward.  Here  the  reef  is  about  10  or  12  chains  in 
width,  and  rises  from  10  fathoms  water.  A  long  lane  of  water, 
a  chain  in  width,  is  cut  clean  as  with  a  knife  across  the  reef, 


/Vjf^^ 


Fig.  36.— Sketch  Section  across  Gutter  in  S.  Palm  Island  Fringinu 

Reef. 
1.  Coral  Keef.         2.  Gutter  (60  feet  deep,  1  chain  wide), 

reaching  almost  to  the  shore.  No  corals  whatever  were  observed 
growing  in  this  lane;  the  sides  are  precipitous,  and  10  feet  below 
the  surface  appear  to  be  slightly  overhanging  in  character  (Fig.  36). 


158  GEOLOGY  OF  QUEENSLAND  COAST, 

Orpheus  Island,  also  composed  of  granite  and  quartz  porphyry, 
crossed  with  basalt  dykes,  contains  on  its  sheltered  side  an 
extensive  flat  some  12  feet  above  H.W.M.,  and  composed  of 
black  coral  fragments;  some  as  much  as  three  feet  in  diameter. 

Fantome  Island,  also  of  granite  and  quartz  porphyry,  possesses 
two  flats — one  on  the  windward,  the  other  on  the  leeward  of  the 
island.  The  more  important  one  lies  between  two  neighbouring 
o-ranite  hills.  The  seaward  edge  consists  of  a  long  sloping  white 
beach,  skirting  a  fringing  reef,  which  is  exposed  in  great  measure 
during  low  tides.  Beyond  the  present  line  of  beach  another  rises 
in  similar  form  to  a  height  of  12  to  15  feet  above  spring  tides, 
forming  a  wide  and  distinct  terrace  as  much  as  five  chains  wide 
in  places. 

Laroe  fragments  of  "  beach  rock ''  exist  on  this  upper  l^each 
apparently  in  situ,  the  pieces  having  a  gentle  slope  seawards.  A 
forest  growth  has  seized  upon  this  higher  beach,  and  almost 
concealed  it  from  view. 

A  similar  though  smaller  flat  occurs  on  the  leeward  side.  In 
the  cases  of  Orpheus  and  Fantome  beaches  the  arms  of  the  sea  in 
a  couple  of  instances  are  less  than  half-a-mile  in  width  in  the 
locality,  and  in  these  waves  could  not  have  such  play  as  on  exjDosed 
ocean  beaches. 

Curious  umbrella-like  coral  growths  occur  in  Juno  Bay,  Fantome 
Island.  The  accompanying  diagram  is  a  sketch  section  of  two  of 
these  growths  near  one  of  our  anchorages  (Fig.  5). 

These  interesting  growths  rise  from  as  much  as  30  and  40  feet 
depth  of  water,  and  form  shelters  for  fish.  It  is  very  probable 
that  in  time  to  come  some  of  the  upjDer  portions  may  coalesce  by 
fusion  of  the  growing  walls,  leaving  a  species  of  cavern  below.* 

Curaroa  Island  is  small  and  conical  in  shape,  comjDOsed  of 
granite  and  porphyry.  It  is  920  feet  in  height.  On  its  western 
side  a  flat  of  some  50  acres  exists.  Mr.  G.  Butler,  of  Town8^  ille, 
states  that  this  flat  is  composed  almost  entirely  of  large  black 
coral  fragments,  at  least  12  feet  above  high  water  mark. 

*  Sawyer  and  Andrews,  "  Caves  of  Fiji."     Proc.  Linn.  Soe.  N.S.  W.  1901 
xxvi.,  Pt.  1,  pp.  91-106. 


BY    K.    C.    ANDREWS. 


159 


Hinchiiibrook  Island  consists  of  huge  rock  escarpments,  sur- 
mounted by  aiguilles— a  mountain  range  20  miles  in  length,  whose 


Fig.  5.— Sketch  Section  illustrating  interesting  Coral  Growths  at 
Juno  Bay,  Fantome  Island. 

1.  Flat  growing  coral  tables,  from  6  to  10  feet  in  diameter,  10  feet  below 
H.W.M.         2.  Rockshelters  under  coral  tables.         3.  Sand  lanes. 
Height  of  pillars  about  30  feet. 

axis  is  approximately  parallel  to  the  neighbouring  mainland  coast. 
Immense  mangrove  flats,  miles  in  width,  fringe  its  western  edge, 
and  from  these  rise  precipitously  the  great  granite  peaks  known 
as  Straloch,  Diamantina,  Pitt  and  Bowen,  the  latter  being  nearly 
4,000  feet  in  height.  The  southern  portion  contains  many  barren 
peaks,  and  exhibits  the  granite  type  of  weathering.  The  northern 
end  of  the  island  is  devoid  of  the  rugged  peaks  of  the  south,  and 
is  composed  principally  of  a  dark  quartz  porphyry. 

A  long  narrow  channel  (in  places  nine  fathoms  deep)  separates 
the  island  and  the  mainland.  If,  as  seems  undoubtedly  the  case 
to  me,  the  Hinchinbrook  flats  argue  recent  elevation,  then  this 
channel  must  previously  ha^■e  been  much  wider,  yet  of  the  nature 
of  a  shoal  at  low  tides  over  which  no  heavy  waves  could  beat,  the 
present  channel  having  been  determined  by  subaerial  denudation 
ante-dating  the  period  of  subsidence.     Consequent  drainage  is  a 


160  GEOLOGY  OF  QUEENSLAND  COAST, 

striking  feature  of  these  elevated  flats,  and  islands  connected  by 
sand  ridges  are  common  on  the  north-eastern  side. 

Goold  Island  \^  1,400  feet  in  height,  and  composed  of  granite 
of  acid  type.  Here,  as  at  the  Palms  and  Hinchinbrook,  quartz 
porphyry  is  a  common  rock-type,  weathering  generally  into  rect- 
angular blocks,  at  times  simulating  the  structures  observable  in 
stratified  rocks,  thus  differing  from  the  ordinary  spheroidal  and 
dome-shaped  masses  characteristic  of  granitic  weathering. 

Dunk  Island  is  of  small  extent,  and  covered  with  a  dense 
jungle  or  "brush."  It  consists  of  Silurian  slates,  quartzites,  tuffs 
and  schists,  and  lies  some  five  miles  distant  from  the  mainland, 
which  at  this  point  also  is  composed  of  similar  rocks,  and  supports 
exceedingly  dense  forest  and  jungle  growths  (Fig.  6). 


Fig.  6. — Section  across  Dunk  Island. 
a.  Present  reef.        />6,  Coastal  plain  20  feet  above  sea-level.        c.  Canon, 
20  feet  deep,  exposing  section  of  coastal  plain.       d.  Contorted  rocks  (slates, 
schists  and  quartz  rocks). 

Horizontal  Scale,  1  inch  =250  yards. 

The  island  proper  consists  of  high  rough  land  meridionally 
disposed.  On  the  side  facing  the  mainLand  (leeward)  a  most 
interesting  flat  occurs.  It  is  between  one  and  two  miles  in 
length,  and  500  or  600  yards  in  width.  A  low  bench,  two  or 
three  feet  above  high  water  mark  and  several  chains  in  width, 
accompanies  it  for  a  considerable  distance  on  its  seaward  edge, 
and  to  this  the  flat  presents  a  mural  front  20  feet  in  height.  Its 
structure,  as  revealed  by  an  examination  of  the  seaward  edge, 
and  that  exposed  in  a  20  feet  section  made  by  a  small  canon 
cutting  across  its  breadth  is  that  of  a  stiff  clay,  or  clay  and  sand 
admixtures,  containing  numerous  angular  and  subangular  frag- 


BY    E.    C.    ANDREWS.  161 

meiits  of  the  local  rocks.  Shells  and  other  calcareous  matter  are 
conspicuous  by  their  absence.  At  present  its  surface  is  covered 
with  thick  growths  of  eucalypts,  jungle,  pandanus  and  grass. 
This  flat  is  doubtless  due  to  redistribution  by  along  shore  action 
of  matter  lost  to  the  island  by  marine  erosion. 

South  Barnard  is  a  very  small  island  composed  of  basic  tuffs, 
strengthened  by  several  great  vertical  basaltic  dykes,  whose 
powers  of  resistance  have  prevented  the  little  island  from  being- 
reduced  to  a  terrace  of  erosion. 

A  small  sandy  flat  exists  on  one  side  of  the  island,  and  at  a 
height  of  20  feet  above  high  water  mark,  on  the  southern  side, 
indistinct  traces  of  a  line  of  former  beach  erosion  were  seen, 
which  reminded  the  writer  somewhat  of  the  elevated  lines  of 
marine  erosion  seen  in  many  of  the  Fiji  Islands.* 

The  Xorth  Barnards  are  small,  tuifaceons  in  character,  densely 
brushed,  and  present  mural  fronts  at  times  to  the  sea. 

Fitzroy  Island,  nearly  1,000  feet  in  height,  is  rugged  in 
character,  and,  in  common  with  the  other  islands  of  granitic 
composition  on  this  coast,  is  surrounded,  as  to  its  shore  line,  with 
great  spheroidal  blocks. 

A  small  beach  exists  on  the  north-west  side  of  this  island, 
exceedingly  interesting  by  reason  of  : — 

{a)  Its  steep  slopes. 

(6)  Its  terraced  appearance. 

(c)  The  occurrence,  in  the  bed  of  a  small  creek  cutting  across 
it,  of  dense  hard  "  beach  rock." 

A  section  determined  by  Mr.  Hedley  and  myself  is  supplied 
illustrating  its  excessive  steepness  (27°),  and  the  great  height 
(18  feet)  of  the  second  terrace  above  high  tide  mark. 

The  lower  beach  consists  of  white  coral  and  shell  fragments, 
while  the  upper  one  is  composed  of  similar  material,  but  quite 
black  in  colour.  The  section  was  taken  near  the  south-western 
extremity  of  the  beach.  ^ 

"  Andrews,  "  Notes  on  Geology  of  Fiji."       Bull.  Mus.  Comp.  Zool.  Harv. 
Coll.,  1900,  xxxviii. 
11 


162 


GEOLOGY  OF  QUEENSLAND  COAST, 


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BY    E.    C.    ANDREWS.  163 

King*  ascribes  the  origin  of  the  higher  beach  to  gale  action, 
and  Prof.  Agassiz  refers  it  also  to  similar  agencies,  f  G.  Elphin- 
stone  Dalyrniple  also  refers  it  to  elevation.  |  In  this  connection 
Brazier§  writes  : — "  The  beach  (at  the  Fitzroy  landing  place)  was 
composed  of  nothing  but  coral  and  shells,  about  30  yards  long, 
six  to  seven  feet  in  thickness,  and  the  same  in  breadth.  During 
my  second  visit  (1875)  the  whole  of  the  bank  had  been  washed 
away  by  some  heavy  gale."  As,  however,  the  present  beach  is 
500  or  600  yards  long  and  several  chains  in  breadth  the  excision 
of  such  an  insignificant  fragment  as  mentioned  by  Brazier  would 
not  affect  the  general  appearance  of  the  beach. 

The  occurrence  of  "  beach  rock  "  in  the  bed  of  a  stream  of 
fresh  water,  compact  and  hard  almost  as  building  limestone,  is 
very  interesting,  pointing  to  its  origin  (as  suggested  b}^  many 
geologists)  being  referable  to  the  action  of  fresh  water. 

From  the  summit  of  Fitzroy  the  sunken  coral  patches  of  the 
outer  Barrier  could  be  seen  for  miles. 

Green  Island  is  small,  and  consists  of  sand  scattered  over  a 
base  of  coral.  It  is  merely  a  portion  of  the  Great  Barrier  Reef 
itself,  determined  partly  by  elevatory  forces,  and  partly  by  the 
combined  efforts  of  seolian  and  marine  agencies,  and  populated  by 
animal  and  vegetable  waifs  from  the  neighbouring  mainland, 
which  at  this  point  is  but  a  few  miles  distant. 

On  the  reef  surrounding  this  island  coral  and  other  growths 
are  more  luxuriant  than  in  the  turbid  waters  around  the  small 
islands  dottins:  the  continental  shelf. 


*  Narrative  of  a  Survey  of  the  Intertropical  and  Western  Coasts  of  Aus- 
tralia, i.,  1827,  p.  206. 

t  "  A  Visit  to  the  Great  Barrier  Reef  of  AustraUa,  &c."     Bull.  Mus.  Couip. 
Zool.  Harv.  Coll.  xxviii.,  1898,  p.  110. 

X  Brisbane.     By  authority,  1874. 

§  Journal  of  Conchology,  ii.,  pt.  6,  June,  1S7&. 


164  geology  of  queensland  coast, 

The  Elevation  of  Eastern  Australia  compared  with  that 
OF  Melanesia  and  Polynesia. 

In  recent  times  a  slight  elevation  appears  to  have  obtained 
over  the  whole  eastern  coast  of  Australia,  traces  of  the  elevatory 
movement  being  recorded  at  Raine's  Islet,*  Cooktown,t  the 
islands  between  Thursday  Island  and  Cairns,;  Cape  Grafton, 
Dunk  Island,  Hinchinbrook,  the  Palm  Islands,  Magnetic  Island,^ 
Curtis  Island,|l  Moreton  Bay, ^  the  various  "coastal  plains"  of 
Queensland,  Ballina  beaches,"*^*  Hunter  River,tt  Port  Stephens, 
Hawkesbur}^  River,  Botany  Bay,  WoUongong;!:!  and  other  places. 

This  elevation  is,  as  before  remarked,  so  recent  that  corals, 
"  beach  rock,"  and  plains  around  small  islands  have  suffered 
scarcely  any  disintegration. 

A  similarly  slight  and  recent  movement  of  elevation  occurred 
throughout  such  island  groups  as  the  Fijis,  Tonga,  and  the 
Solomons  so  recently  that  raised  "lines  of  marine  erosion"  left 
on  the  clifts  in  places  exist  almost  in  their  original  integrity, 
though  sometimes  30  feet  above  H.W.M.  So  also  in  certain 
islands  in  Fijiv^§  and  the   Solomons,  lilj   raised   beaches  and  coral 


*  Dr.  Alex.  Rattray,  "Notes  on  the  Geology  of  Cape  York  Peninsula," 
Quart.  Journ.  Geol.  Soc.  xxv.,  1869. 

+  J.  E.  T.  Woods,  Proc.  Linn.  Soc.  N.S.  Wales,  v.,  1880,  pp.  187-189. 

I  Jukes,  Voyage  of  H.M.S.  "Fly,"  Vol.  i. ;  A.  Agassiz,  Bull.  Mus.  Comp. 
Zool.  Harv.  Coll.  xxviii. 

§  Maitland,  "Report  on  Magnetic  Island."  Brisbane,  1892;  G.  Butler, 
Esq.,  Townsville. 

li  Fidt  C.  Hedley, 

IT  Stutchbury,  Twelfth,  Thirteenth,  and  Sixteenth  Tri-monthly  Reports, 
N.S.  Wales  Leg.  Council  Papers,  1854;  R.  L.  Jack,  Geology  of  Queensland, 
p.  617. 

•  •    J.  E.  Carne,  Ann.  Report  Dept.  Mines  N.S.  Wales,  1896,  p.  151. 

+t  Prof.  David  and  R.  Etheridge,  Junr.,  Rec.  Geol.  Survey  N.S.  Wales,  ii., 
1890. 

tt  Jukes,  Voyage  of  H.M.S.  "Fly,"  Vol.  i.,  p.  336. 

§§  Andrews,  "Notes  on  the  Limestones  and  General  Geology  of  the  Fiji 
Islands/'  &c.,  Bull.  Mus.  Comp.  Zool.  Harv.  Coll.  xxxviii.,  1900. 
1111  Guppy,  The  Solomon  Islands  :  their  Geology,  &c. 


BY    E.    C.    ANDREWS.  165 

platforms  exist  with  numerous  loose  and  blackened  corals  scattered 
over  them. 

In  these  islands,  however,  this  recent  uplift  represents  the 
final  stage  in  a  series  of  repeated  slight  elevatory  movements, 
alternating  with  periods  of  stable  equilibrium,  evidenced  by 
**  terraces  "  of  coral  growths  and  successive  lines  of  marine  erosion. 
Australia  appears  to  have  participated  in  the  most  recent  one 
only.  Had  elevation  proceeded  in  Eastern  Australia  along  similar 
lines  to  those  pursued  in  the  mid-Pacific,  we  should  have  elevated 
coral  islands  along  the  site  of  the  Great  Barrier  Reef  in  much  the 
same  relation  to  the  Queensland  coast  as  the  Loyalties  (raised 
coral)  are  to  New  Caledonia,  the  Lau  Group  to  main  Fiji  (Viti  and 
Vanua  Levu),  or  the  smaller  coralline  islands  immediately  east  of 
the  largest  Solomon  and  New  Hebridean  Islands  to  these  same 
land  masses. 

In  certain  groups,  as  in  Fiji,  coral  reefs  and  atolls  were  formed 
during  these  movements  of  elevation,  which  followed  a  period  of 
great  subsidence  in  Tertiary  times,  and  during  which  period 
immense  beds  of  limestone,  volcanic  conglomerates  and  Fiji 
"  soapstones "  had  been  laid  down  to  constitute,  during  the 
succeeding  elevations,  a  base  for  the  later  (recent)  "raised  reefs." 
In  the  New  Hebrides,  the  Solomons,"^  Fiji,t  Tonga, |  New  Guineai^ 
and  elsewhere  traces  of  elevated  coral  reefs  are  found  from  high 
water  mark  to  a  height  of  2,000  feet  above  the  same  datum  line. 

The  participation  by  Australia  (as  regards  the  Queensland  and 
New  South  Wales  coasts)  in  the  most  recent  only  of  these  uplifts 
is  very  interesting.      Hedley|[  has  conclusively  shown  frofn  biolo- 

*  Guppy,  The  Solomon  Islands  :  their  Geology,  &c.,  pp.  102,  113. 
t  Agassiz,  Bull.    Mus.    Comp.    Zool.    Harv.    Coll.    xxviii.,  pp.  132-133; 
Andrews,  ihid.,  xxxviii. ,  p.  27. 

It:  Lister,  "Notes  on  the  Geology  of  the  Tonga  Islands,"  Quart.  Journ 
Geol.  Soc,  Vol.  47,  1891,  pp.  590-616. 

§  Maitland,  Geological  Observations  in  New  Guinea,  1891,  p.  10. 
11  C.  Hedley,  "  A  Zoogeographic  Scheme  for  the  Mid  Pacific,"  Proc.  Linn. 
Soc.  N.S.  Wales,  1899,  pp.  391-417.     See  also  references  in  same  paper. 


166  GEOLOGY  OF  QUEENSLAND  COAST, 

gical  data  that  the  main  islands  of  Fiji,  New  Caledonia,  the  New 
Hebrides  and  the  Solomons  Avere  previously  coextensive,  and 
these,  in  turn  with  New  Guinea,  continental  in  origin.  Since 
then  geologists*  have  proved  a  continental  origin  for  Fiji. 
Similarly  for  the  New  Hebrides,  New  Caledonia,  and  the  Solomon 
Islands.  If,  then,  the  deep  ocean  separating  these  groups  repre- 
sents faulted  or  warped  areas,  it  is  very  probable  that  some 
relation  exists  between  the  subsidences  in  the  ocean  area  between 
the  island  clusters  and  the  repeated  uplifts  in  the  groups  them- 
selves. The  elevation  of  main  Fiji  probably  proceeded  along 
much  more  rapid  lines  than  those  pursued  in  the  Lau  Group, 
inasmuch  as  the  Tertiary  "  soapstones,"  limestones  and  volcanic 
conglomerates  of  Viti  Levu  appear  to  have  no  capping  of  recent 
"raised  reef,"  whereas  each  pause  in  the  elevation  of  Lau  was 
attended  b}^  the  formation  of  "raised  reefs"  of  recent  age,  having 
as  a  base  Tertiary  rocks  indistinguishable  from  the  Viti  Lcau 
strata  of  similar  age.  (It  must  be  remembered,  however,  that  a 
very  recent  uplift  seems  to  have  affected  main  Fiji,  Lau,  the 
Solomons  and  North  Australia;!  and  that,  too,  after  a  period  of 
quiescence,  as  is  shown  by  the  raised  "coral  reef  platforms.'') 
The  elevations  of  main  Fiji  and  Lau  probably  w^ere  not  synchro- 
nous, the  main  islands  of  continental  origin  being  rapidly  elevated 
to  a  great  height  in  one  movem.ent,  while  it  is  more  than  jDossible 
that  the  intermittent  elevations  of  Lau  were  due  in  great  measure 
to  the  existence  of  the  large  island  masses  (Viti  and  Vanua  Levu) 
lying  to  the  West.  New  Caledonia  and  the  Loyalties  form  a 
similar  group  to  Fiji  and  Lau,  so  also  possibly  do  New  Guinea 
and  the  small  "  terraced  "  coral  islands  lying  to  the  east.|  Thus, 
whereas  the  island  groups  of  the  Fijis,  Solomons,  Hebrides, 
Tonga  and  New  Caledonia  have  undergone  a  cycle  of  elevation, 
Eastern  Australia,  as  will  be  shown  later,  is  in  a  cycle  of  develop- 
ment attendant  on  subsidence. 

*W.  G.  Woolnough,  B.Sc,  Adelaide  University,  and  Dr.  H.  B.  Guppy, 
in  lift. 

t  This  elevation  most  probably  occurred  well  into  the  historic  period, 
although  the  individual  island  upUfts  may  not  have  been  contemporaneous. 
J  Maitland,  Geological  Observations  in  New  Guinea,  1891,  pp.  9,  10. 


BY    E.    C.    ANDREWS.  161 


Petrological  Notes. 


The  igneous  rocks  consist  mostly  of  granites  of  acid  types,  quartz 
and  quartz-felspar  porphyries,  both  Palaeozoic  in  age,  and  intruded 
alike  by  regular  networks  of  basaltic  dykes. 

Granites. — These  have  a  marvellous  development,  and  are 
principally  of  acid  types.  Many  of  the  islands,  as  also  the  high 
coastal  ranges,  are  composed  entirely  of  these  plutonic  rocks. 
They  are  probably  Carboniferous  and  Permo-Carboniferous  in  age. 

Quartz  and  felspar  are  the  chief  constituents,  although  horn 
blende  and  biotite  are  frequently  present.  In  some  of  the  island 
groups,  such  as  the  Palms,  the  rocks  may  consist  of  quartz  and  fel- 
spar, with  idiomorphic  hornblende  and  biotite  flakes.  The  horn- 
blende, which  is  strongly  pleochroic,  occurs  in  hexagonal,  rhombic 
and  rectangular  forms  in  thin  sections,  and  post-dates  the  formation 
of  the  biotite.  Again,  hornblende  may  be  absent,  while  in  other 
cases  flat  veins  of  aplite  with  ill-defined  boundaries  occur;  they 
are  composed  mainly  of  micrographic  growths  of  quartz  and 
felspar.  Large  segregations  of  orthoclase  and  quartz  may  be 
seen  in  the  rock  exposures,  and  epidote  at  times  replaces  the 
ferromagnesian  constituents.  Zircons  are  associated  with  the 
hornblendic  varieties.  Here,  also,  crushing  is  a  marked  feature. 
Under  the  microscope  the  quartzes  and  felspars  are  seen  to  have 
undergone  great  peripheral  crushing,  besides  wliolesale  fracturing 
of  the  different  minerals  constituting  the  rock.  The  grains  of 
quartz  and  felspar  appear  as  if  cemented  in  a  base  of  the  same 
material  (mortar  structure).  Splendid  examples  of  undulose 
extinction  are  shown  by  both  quartz  and  felspar  crystals;  long 
lines  of  liquid  inclusions  occur  in  the  quartz,  while  the  felspars 
(orthoclase  and  acid  plagioclase)  show  zonary  structure,  and  the 
development  of  fresh  minerals  along  solution  planes.  Biotite  also 
occurs  in  bent  flakes,  and  secondary  quartz  is  present  in  great 
abundance. 

Other  granites,  collected  from  Fitzroy  and  Goold  Islands 
although  altered,  do  not  show  the  effects  of  metamorphism  in  so 
marked  a  desfree  as  the  Palm  Islands  varieties. 


168  GEOLOGY  OF  QUEENSLAND  COAST, 

Porphyries. — These  consist  of  quartz  and  felspar  idiomorpbs 
scattered  plentifully  through  a  fine  granular  base  of  the  same 
minerals.  The  quartz  and  felspar  may  be  porph3^ritic  towards 
the  rest  of  the  rock.      Mica  plates  are  common. 

Felspar  occurs  as  stout  orthoclase  and  plagioclase  crj'stals,  and 
much  altered.  The  quartz  contains  abundant  liquid  inclusions. 
Some  of  the  crystals  are  much  corroded,  the  magma  having  almost 
penetrated  to  the  crystal  centres  from  every  direction. 

The  base  is  rarely  felsophyric  with  abundant  magnetite  in 
grains. 

Dolerites  and  Basalts. — Throughout  the  acid  rocks  just  men- 
tioned basic  dykes  occur  in  great  numbers,  so  as  to  appear  at 
times  in  the  cliff  faces  as  regular  networks  of  veins. 

The  rocks  are  both  holocrystalline  and  hypocrystalline,  and 
consist  principally  of  plagioclase  laths  and  augite  grains.  Olivine 
is  frequently  present.  Hornblende  also  occurs  in  certain  types, 
but  much  decomposed.     The  olivine  is  principally  decomposed. 

Ophitic  structure  is  noticeable  in  certain  sections. 

These  basic  intrusions  belong  probably  to  the  Tertiary  period. 

Mr.  Jukes  and  Professor  Agassiz  spent  some  considerable  time 
on  the  Great  Barrier  Reef.  Saville  Kent  also  studied  Barrier  Reef 
problems  on  the  spot  for  a  period  of  twelve  months.  Jukes 
furnishes  a  section  (reproduced  by  Agassiz,  I.e.,  p.  137)  across  the 
Barrier  Reef  and  to  the  mainland  which  sums  up  his  ideas  of  its 
origin,  viz.,  that  (even  allowing  for  his  exaggerated  vertical  scale), 
generally  speaking,  the  continental  shelf  from  the  coastal  plains 
of  the  mainland  to  the  outer  Barrier  is  composed  of  reef  and  reef 
debris.  Notwithstanding  this,  his  description  is  at  variance  with 
his  section  (postea,  pp.  170-172).  Jukes  also  sa3^s*  that  the  Cxreat 
Barrier  Reef,  longitudinally  considered,  "  would  be  found  to ha\e 
a  considerable  resemblance  to  a  gigantic  and  irregular  fortification, 
a  steep  glacis  crowned  with  a  broken  parapet  wall,  and  carried 
from  one  rising  ground  [Sir  C.  Hardy's  Islands]  to  another" 
[Breaksea  Spit]. 

*  Jukes,  Voyage  of  H.M.S.  "Fly,"  Vol.  i.,  p.  332-333. 


BY    E.    C.    ANDREWS.  169 

Saville  Kent*  also  concurs  in  Jukes's  conclusions  as  to  the  origin 
of  the  continental  shelf  and  the  Barrier  Reef  as  supplied  by  his 
(Mr.  Jukes's)  section. 

Prof.  Agassiz,  however,  claims  that  the  forces  of  subsidence 
are  unnecessarily  called  in  by  these  writers,  and  that  the  coast 
(formed  of  granites  and  allied  rocks)  was  planed  down  by  the 
forces  of  marine  erosion,  and  that  coral  and  other  reef-building 
organisms  seized  upon  "  the  terraces  of  erosion  "  to  which  the 
coastal  area  had  been  reduced,  and  in  this  way  by  the  coalescence 
of  once  isolated  masses  of  coral,  the  Great  Barrier  Reef  was 
determined  in  position. 

The  following  extracts!  illustrate  the  opinions  held  by  Pro- 
fessor Agassiz  and  Mr.  Jukes  as  to  the  origin  of  the  Reef : — 

"While  it  is  undoubtedly  true,  as  mentioned  by  Kent,  that 
Jukes  considered  Darwin's  hypothesis  as  '  perfectly  satisfactory  to 
my  [his]  mind,'  yet  I  cannot  help  analysing  Jukes's  summary  to 
show  how  correctly  he  had  analysed  the  main  features  of  the 
Great  Barrier  Reef,  and  of  its  relations  to  the  mainland  and 
intervening  islands,  and  was  led  to  what  seem  to  me  erroneous 
conclusions,  from  the  inferences  he  drew  from  the  diagram  he 
gives  of  an  imaginary  section  of  the  Great  Barrier  Reef,  and 
which  I  here  reproduce,  i. 

"  It  seems  strange  that  Jukes  should  have  given  a  section  across 
the  Great  Barrier  Reef,  and  have  left  out  the  islands  which  crop 
up  nearly  all  along  the  coast  of  Queensland  between  the  mainland 
and  the  outer  or  inner  line  of  reefs.  This  would  have  given  his 
section  an  entirely  different  aspect,  as  he  would  have  had, 
cropping  up  and  connected  with  the  line  of  the  mainland,  a  series 
of  peaks  rising  from  ten  to  thirty  fathoms,  round  which  alone,  or 
round  the  flat  bases  of  islands  and  peaks  which  had  disappeared 
from  erosion  or  other  atmospheric  causes,  corals  had  grown 
Such  a  section  is  not  an  imaginary  one,  for  the  channels  between 

*  Saville  Kent,  The  Great  Barrier  Reef  of  Australia;  London,  1893. 

+  A.  Agassiz,  "Great  Barrier  Reef  of  Australia,"  pp.  136-139. 

X  Voyage  of  H.M.S.  "Fly,"  Vol.  i.,  p.  333. 


170  GEOLOG\  OF  QUEENSLAND  COAST, 

the  outcropping  peaks,  islands,  reefs,  or  reef  flats  are  covered 
with  the  telluric  detritus  derived  from  the  decomposition  of  the 
rocks  forming  those  islands,  or  obtained  from  the  slopes  of  the 
mainland.  Such  a  section  would  have  shown  the  layer  of  corals 
to  be  comparativel}^  thin,  of  not  more  than  twelve  to  fifteen 
fathoms,  and  it  would  have  shown  the  great  probability  that  the 
outer  line  of  reefs,  even  built  upon  similar  bases,  once  connected 
with  the  mainland,  had  not  attained  a  much  greater  thickness. 
8ee  sections  across  the  Great  Barrier  Reef  (Plates  xxxvii.  toxli.). 
That  Jukes  himself  felt  his  imaginary  section  and  his  explanation 
of  it  not  to  be  of  universal  application  can  be  proved  from  his 
own  words."^     He  says  : — 

"  '  The  most  remarkable^deviations  from  this  condition  are  in 
the  spaces  between  Cape  Melville  and  Lizard  Island,  and  at  the 
back  of  Wreck  Bay  and  Raine's  Islet.  Now  in  each  of  these 
cases  there  are  islands  of  granite  or  other  rocks  advanced  from 
the  mainland,  and  thus  causing  an  original  irregularity  in  the 
depth  of  water,  as  it  would  be  independent  of  the  coral  reef. 
This  is  very  remarkable  in  the  space  between  12°  20'  and  11°  30', 
where  we  have  Cape  Grenville,  Cockburn  Islands,  and  8ir  C. 
Hardy's  Islands,  projecting  towards  Raine's  Islet  opening,  and 
Fair  Cape  and  Cape  Weymouth,  Avith  Forbes  Island  and  Quoin 
Island  projecting  towards  Wreck  Bay.  Near  Sir  C.  Hardy's 
Islands  there  is  also  a  remarkable  narrow  channel  of  deep  water, 
between  them  and  the  large  Cockburn  Reef,  in  which  there  is  a 
depth  of  thirty  fathoms,  while  on  each  side  of  it  is  either  a  reef 
nearly  dry  at  low  water,  or  a  depth  not  exceeding  ten  fathoms. 
This  channel  is  about  twenty  miles  long,  rarely  more  than  two 
miles  broad,  and  it  runs  in  the  same  direction  as  the  islands  lie 
off  Cape  Grenville,  or  about  east-north-east,  and  points  in  a 
straight  line  for  Raine's  Islet  opening.' 

"  It  seems  tome  tljat  Jukes  has  here  struck  the  correct  explana- 
tion of  the  structure  of  the  Great  Barrier  Reef.  But  having 
examined  only  the  two  extremes,  he  did  not  perhaps  realise  that  the 


Jukes,  Voyage  of  H.M.S.  "  Fly,"  Vol.  i.,  p.  883. 


BY    E.    C.    ANDREWS.  171 

same  condition  of  things  existed  off  any  line  in  which  such  islands 
were  found.  He  allowed  his  admiration  for  the  simplicity  of  the 
explanation  of  the  theory  of  coral  reefs  b}^  Darwin  to  blind  him  to 
his  own  still  simpler  explanation,  which  I  will  here  quote.* 

'' '  In  the  lirst  place,  speaking  generally,  the  outline  of  the 
Great  Barrier  Reef  is  parallel  to  the  outline  of  the  north-east 
coast.  The  one  follows  the  other  in  all  its  curves  and  flexures  with 
quite  sufficient  conformabilit}^  to  show  that  the  two  are  connected. 
This  is  perceptible  even  in  the  small  chart  attached  to  this  work, 
but  still  more  remarkably  so  when  the  large  Admiralty  Charts 
are  examined.  It  is  evident  that  the  circumstances  that  modified 
the  outline  of  the  coast  likewise  determined  the  general  outline 
of  the  reefs.  This  is  nothing  else  than  to  sa}',  that  the  outline  of 
the  reefs  depends  upon  the  depth  of  the  water.  Just  as  in  a 
large  and  accurate  chart  of  an}'  line  of  coast  we  should  find  the 
boundary  of  any  certain  line  of  soundings,  such  as  20,  50,  or  100 
fathoms,  conforming  generally  to  the  outline  of  the  coast,  following 
its  larger  flexures  and  more  important  features;  so  we  find  the 
outline  of  the  Barrier  Reefs  conforming  to  the  north-east  coast 
of  Australia.  Granting  that  the  mean  slope  of  the  rocks,  forming 
the  original  sea-bottom  of  this  coast,  was  tolerably  regular  and 
conformable  to  the  slope  of  the  land,  it  is  evident  that  if  we  took 
away  the  coral  reefs  and  raised  the  land  to  any  given  height  as, 
for  instance,  100  fathoms,  we  should  not  greatly  alter  the  outline 
of  the  coast,  but  only  shift  its  situation.  It  would  be  thrown  so 
much  further  forward,  or  towards  the  east.  Now,  sujDpose  the 
coast  cleared  of  coral  reef,  and  raised  so  much  that  it  emerged 
from  the  sea  just  widiin  the  line  of  the  present  Barrier  Reef. 
Then  let  the  reef  commence  in  the  shallow  water  along  that  shore, 
and  a  very  slow  and  gradual  depression  take  place,  giving  time 
for  the  polyps  to  build  up  so  as  to  keep  near  the  surface  of  the 
water.  The  result  of  this  action  would  be  the  present  Barrier 
with  its  steep  outer  slope,  and  its  gradual  extension  over  the 
sinking  rocks  that  were  once  dry  land  within  it.      Portions  that 

*■  Voyage  of  the  "  Fly,"  Vol.  i.,  p.  345. 


172  GEOLOGY    OF    QUEENS1.AND    COAST, 

were  once  hills  oa  the  dry  land  would  now  be  islands  between  the 
Barrier  and  the  main,  such  as  Sir  C.  Hardy's  Island  and  those 
about  it.  Islands  that  once  existed  in  front  of  the  mainland 
would  now  be  altogether  submerged,  and  their  places  only  marked 
by  detached  reefs  outside  the  Barrier,  such  as  those  north  and 
south  of  Wreck  Bay.  According  to  the  old  rule  of  high  land  and 
deep  water  going  together  (in  other  words,  the  slope  of  the  ground 
below  water  being  only  a  continuation  of  that  above),  we  should 
have  the  Barrier  much  closer  to  the  present  land  in  its  more 
abrupt  and  lofty  portions  than  in  those  which  were  lower  and 
less  highly  inclined.  We  see  accordingly  the  reefs  approach  the 
present  land  about  Cape  Melville,  where  the  land  is  steep  and 
lofty,  and  recede  from  it  as  we  go  further  north  in  proportion  as 
the  land  becomes  flatter  and  more  gentle  in  its  inclination.  Deep 
holes  and  ravines,  full  perhaps  of  fresh  water,  may  have  existed 
on  the  old  land,  so  that  when  the  surface  of  these  lakes  and 
hollows  first  sank  to  the  surface  of  the  sea,  and  admitted  its 
waters,  the  bottom  may  have  been  too  deep  for  the  coral  animals 
to  live  on.  This  would  explain  such  a  phenomenon  as  the  deep 
narrow  channel  just  north  of  Sir  C.  Hardy's  Islands,  with  reefs 
running  along  each  side  of  it.  In  short,  every  modification  in  the 
form  and  structure  of  the  reefs  is  explicable  by  this  h3"pothesis, 
and  many  difficulties  solved,  which  admit  of  no  other  explanation.' 

"  He  assumes,  as  we  do,  that  the  Australian  coast  at  one  time 
was  just  within  the  line  of  the  present  Barrier  Reef;  but  it  seems 
to  me  that  the  causes  given  by  Jukes  for  the  formation  of  the 
Barrier  Reef  are  equally  well  explained  by  erosion  and  denuda- 
tion. He  assumes  a  great  thickness  for  the  corals  on  the  outer 
edge  of  the  Barrier  Reef, — a  thickness  to  have  grown  b}'  the 
synchronism  of  the  subsidence  and  the  growth  of  the  corals, — a 
thickness  the  extent  of  which  no  one  can  even  guess  at.  We 
assume  for  the  corals  a  thickness  that  can  be  determined  fairly 
accurately  as  only  a  veneer  of  at  most  20  fathoms  upon  the  faces 
of  the  denuded  platforms  of  the  islands  which  once  formed  the 
outer  line  of  the  Australian  continent.     .     .     ."' 


by  e.  c.  andrews.  173 

Deductions  from  foregoing  Observations  with  References 
TO  Geograpiiv  of  the  Cordillera. 

It  is  well  known  that  coast  lines  of  the  regular  or  smooth  type 
indicate  (except  in  cases  of  riverless  foreshores)  a  verj^  advanced 
stage  of  marine  erosion  either  in  the  present  cycle  or  in  one 
immediately  preceding  recent  elevation,  the  tendency  of  marine 
erosion  being  to  reduce  irregularities,  whereas  the  recession  of  the 
coast  line  from  the  edge  of  the  continental  shelf,  combined  with 
highly  indented  coast  lines  and  numerous  outlying  islands,  evi- 
dences the  effect  of  recent  subsidence. 

The  general  trend  of  a  coast  line  is  due  to  earth  movements,  a 
very  subsidiary  part  being  plaj^ed  by  the  agencies  of  marine 
erosion,  no  inlets*  or  fiords  arising  as  the  result  of  such  action, 
these  being  attributable  to  the  subsidence  of  deeply  eroded 
plateaux  of  accumulation  or  erosion. 

Sedimentation  also  plays  an  important  j^art  in  the  genesis  of 
coastal  topograph}'-,  the  w^aste  resulting  from  subaerial  denudation 
being  arranged  to  form  deltas,  coastal  plains,  shoals  and  spits. 
Marine  erosion  also  helps  in  a  lesser  degree,  but  always  in  the 
way  of  forming  a  regular  coast,  viz.,  by  attacking  the  headlands, 
the  waste  thus  derived  being  formed  into  bars,  or  being  swept 
into  sheltered  spots. 

In  forming  an}'  theory  as  to  the  origin  of  the  present  aspect  of 
the  Queensland  and  New  South  Wales  coastal  areas  we  may 
rememl^er  that  :  — 

(1)  The  outer  edge  of  the  Great  Barrier  Reef  appears  to  follow 
the  general  trend  of  the  coast. 

(2)  The  seaward  edge  of  the  Great  Barrier  Reef  is  not  the  only 
portion  of  the  continental  shelf-margin  removed  to  a  considerable 
distance  from  the  coast  line,  since  that  part  lying  between  Lady 
Elliot  Isle  (where  temperature  places  a  limit  on  the  southward 


*  Under  "  inlets"  are  not  included  those  small  irregularities  in  a  coast  such 
as  exist  between  Port  Jackson  and  Botany  Bay  Heads,  but  indentations  like 
the  above-mentioned  ports  themselves. 


174  GEOLOGY  OF  QUEENSLAND  COAST, 

extension  of  the  Reef)  and  the  Tweed  Heads  is  also  of  consider- 
able width. 

(3)  The  Barrier  Reef  rises  suddenly  from  great  depths,  but  in 
a  less  marked  degree  than  does  the  shelf  eastwards  of  Moreton 
Bay  and  Sydney  Heads.* 

(4)  The  great  shoals  of  the  shelf  in  the  neighbourhood  of 
Stradbroke,  Moreton  and  Frazer  Islands  are  arenaceous  and 
argillaceous  in  character,  and  small  calcareous  patches,  as  Lady 
Elliot  Isle,  have  grown  on  the  northern  extension  of  one  of  these 
banks,  t 

(5)  Between  the  outer  Barrier  and  the  mainland  the  average 
depth  of  the  water  is  from  10  to  20  fathoms.  A  few  dejDths 
exceeeding  30  fathoms  have,  however,  been  recorded  in  the  vicinity 
of  Wreck  Ba3^l 

(6)  The  Great  Barrier  Reef  itself  is  of  variable  width,  and  is 
broken  up  by  long  passages  arranged  parallel  and  transversely  to 
its  length.  The  depth  of  the  water  in  these  '*  lanes  "  or  passages 
varies  from  about  20  to  40  fathoms. 

(7)  A  maze  of  islands  exists  on  the  shelf,  occupying  a  zone  of 
from  10  to  15  miles  from  the  coast,  close  together  inshore  and 
scattered  in  an  easterly  direction;  20  miles  off  shore  no  trace  of 
continental  island  occurs  above  H.W.M.  In  size  these  insular 
areas  var}-^  from  several  acres  to  rugged  mountainous  masses,  like 
Hinchinbrook  Island  20  miles  in  length  and  3,600  feet  in  height, 
or  huge  masses  of  sand  dunes  as  Moreton,  Stradbroke  and  Frazer 
Islands.  Frazer  Island  is  80  miles  in  length,  has  a  great  spit 
continuing  from  its  northern  extremity,  with  a  fairl}^  meridional 
disposition,  and  the  eastern  coast  forms  a  long  uninterrujDted 
curve.  Moreton  and  Stradbroke  Islands  have  lines  of  sand 
dunes  as  much  as  800  feet  in  height,  and  show  masses  of  conti- 
nental rocks  at  their  northern  extremities.  All  of  these  three 
large  sandy  islands  are  associated  with  large  sand  shoals,  island 

*  See  also  A.  Agassiz,  Bull.  Mus.  Comp.  Zool.  Harv.  Coll.  xxviii.,  pp.  129, 

140,  141. 

t  Jukes,  Voyage  of  H.M.S.  "Fly,"  Vol.  i.,  p.  318. 
+  Jukes,  Ibid.,  p.  333. 


BY    E.    C.    ANDREWS.  175 

tying  in  various  stages,  and  extensive  coastal  plains.      They  occur 
also  in  proximity  to  large  rivers. 

(8)  On  the  mainland  and  islands  of  Queensland  the  occasional 
evidence  of  small  cliffs  is  present,  although  typically  conspicuous 
by  their  absence.  (Clifls  are  noticeable  on  the  JST.S.  Wales  coast, 
and  are  very  pronounced  on  the  Tasmanian  seaboard.)  Innnelise 
rugged  escarpments  exist  on  both  mainland  and  insular  areas. 
These  abut  in  many  cases  on  to  wide  coastal  plains.  Large 
beaches  and  coast  plains  exist  also  on  the  leeward  sides  of  the 
islands  sometimes  as  much  as  20  feet  above  H.W. M.,  while 
examples  of  tied  islands  are  frequent,  as  at  the  Palms,  Hinchin- 
brook  and  Dunk  Islands. 

Fringing  coral  reefs  are  associated  with  most  of  the  islands. 

(9)  Lagoons  and  lagoon-marsh  meadows  are  of  frequent  occur- 
rence along  the  coast,  especially  so  between  Townsville  and  Sydney 
Heads.  At  Ballina"^  (just  south  of  the  Queensland  area)  there 
exists  a  great  extent  of  low  land  encircled  by  an  amphitheatre  of 
hills.  Long  lines  of  sand  dunes  on  this  area  curve  sympathetically 
with  the  coast  line.  Lagoons  have  been  formed  between  successive 
dunes  to  be  changed  subsequentl}^  to  lagoon-marsh  meadows.  A 
still  later  phase  here  is  elevation  and  encroachment  by  the  sea,  as 
shown  by  the  peaty  products  of  the  marshes  being  exposed  in  the 
present  coast  nips. 

(10)  Bay  bars  and  spits  have  a  great  distribution. 

(11)  South  of  the  Great  Barrier  Reef  numerous  rivers,  of  which 
the  Brisbane,  Tweed,  Richmond,  Clarence  and  Macleay  may  be 
taken  as  types,  pursue  general  easterly  directions  to  within  a  few 
miles  of  the  coast,  when  they  flow  to  sea  along  northerly  channels 
sometimes  as  much  as  50  miles  in  length. 

The  country  east  of  these  deviations  is  typicall}^  of  flat,  low, 
sandy  or  swampy  nature.  River  bars  exist  at  the  entrances. 
Although  the  rivers  have  a  general  easterly  flow,  the  head  waters 
frequently  follow  meridionally  disposed  channels. 


*  J.  E.  Carne,  Ann.  Eeport  Dept.  Mines  N.S.  Wales,  1896,  p.  151. 


176  GEOLOGY  OF  QUEENSLAND  COAST, 

(12)  Along  the  present  seaboard  the  coastal  plains  maintain  a 
fairly  constant  height  above  sea  level.  A  few  miles  inland  from 
Townsville,  however,  along  the  flanks  of  the  ranges,  recent 
deposits  occur  at  elevations'^  of  as  much  as  300  feet  above  sea 
level.  Variable  heights  at  which  Post-Tertiary  deposits  occur 
may  be  seen  at  the  Hunter  River  mouth  and  Raymond  Terrace 
(E.  C.  Andrews),  Maitland  district!  (50  feet;  David  and  Etheridge). 

(13)  Splendid  examples  of  consequent  drainage  occur  on  the 
coastal  plains  of  Hinchinbrook  Island,  Townsville  flats,  Palm, 
Dunk  Island  and  other  places. 

(11)  The  coast  is  attended  by  a  wide  upland,  rising  gradually 
towards  the  west  until  an  elevation  of  about  4,000  feet  is  attained. 
Considered  broadly,  this  mountainous  belt  is  represented  at  the 
coast  merely  by  undulating  country,  rising  occasionall}^  into 
isolated  mountains.  Still  further  west  the  monadnocks  crowd 
together  until  the  country  has  the  aspect  of  a  maze  of  wild 
ravines  from  2,000  to  4,000  feet  in  depth,  separated  by  long  razor- 
backs,  the  summits  of  which  would  lie  on  the  surface  of  a  flattened 
hemi-cylinder,  the  decrease  in  height  taking  place  in  an  easterly 
direction.  The  central  and  highest  portion  of  the  mountain  mass 
consists  of  undulating  tableland.  Flat  basaltic  hills  occupy  the 
highest  points,  and  overlie  numerous  extensive  and  deep  masses 
of  auriferous  river  drift.  These  old  water  courses  far  exceed  in 
size  those  of  the  present  streams.  The  central  portion  of  the 
Cordillera  consists  principally  of  various  granitic  rocks,  while  to 
east  and  west  lie  the  variously  inclined  rocks  belonging  to  the 
Palaeozoic  era.  On  the  upturned  edges  of  these  the  old  rivers  ran. 
A  most  interesting  feature  in  the  present  topography  is  the 
generally  wide  and  shallow  series  of  basins  (with  basaltic  outliers), 
in  which  the  present  streams  run,  and  the  numerous  long  and 
narrow  canons,  2,000'  to  4,000'  in  depth,  which  are  entrenched 
along  the  recent  broad  basins  of  the  present  streams. 

*  Jack  and  Etheiidge,  Geology  of  Queensland,  p.  617. 

+  David  and  Etheridge,  Rec.  Geol.  Survey  N.S.  Wales,  1S90,  ii.,  Pt.  2, 
pp.  37-52,  pi.  3. 


>> 


BY    E.    C.    ANDREWS.  177 

111  a  mere  advance-note  like  the  present  the  proper  discussion  of 
the  above  observations  would  occupy  too  much  space,  but  briefly 
their  philosophy  appears  to  me  to  be  much  as  follows  : — 

The  coincidence  of  Barrier  Reef  and  present  coastal  contours 
points  to  an  occupation  by  coral  growths  of  a  former  coast  line, 
but  the  continuance  in  width  of  the  shelf  southwards  of  the  limits 
of  reefs  (coralline),  and  the  great  shoals  thereon,  points  to  a  minor 
part  only  of  the  shelf  being  formed  of  coral  growths. 

It  would  appear  that  the  almost  uniform  and  smooth  bottom 
of  the  outer  centre  and  eastern  portions  of  the  continental  plateau, 
combined  with  the  great  depths  from  which  the  Barrier  rises, 
argues  a  long  period  of  marine  erosion  preceding  the  present 
cycle*  during  which  a  uniform  coast  and  smooth  off-shore  bottom 
had  been  formed.  The  sinking  of  this  uniform  area  allowed  the 
sea  to  trespass  far  over  the  old  coast  sands  into  the  ranges,  and 
the  corals  —  formerly  prevented  from  forming  barrier  reefs,  b}'' 
reason  of  the  practical  coincidence  of  continental  shelf  margin  and 
shore  line,  and  the  excessively  turbid  character  of  the  water  on  the 
narrow  fringe  of  the  continental  shelf — proceeded  in  the  clear 
waters  of  the  shelf  margin,  now  removed  far  seaward,  to  invest  the 
whole  width  of  the  smooth  ofF-shore  deposits  with  their  masses, 
and  establish  themselves  as  the  Barrier  Reef. 

Some  connection  probably  exists  between  the  present  Reef 
passages  (and  parallel  channels)  and  the  old  watercourses  of  the 
coastal  area. 

The  association  of  numerous  rugged  mountainous  islands  on  the 
inner  centre  of  the  shelf  (and  close  in  shore)  and  smooth  shallow 
interinsular  seabottom,  combined  with  traces  only  of  coastal  nips, 
is  suggestive  of  gradual  subsidence  with  concomitant  sedimenta- 
tion rather  than  a  period  of  equilibrium  succeeding  subsiden(^e 
which   would   result  in   the   formation  of   cliffs. f      Magnificent 

*  Cycle — the  time  involved  in  a  movement  of  considerable  extent  or  in  the 
development  of  a  submarine  plain  or  peneplain. 

t  The  existence  of  bar-bound  rivers  has,  however,  to  be  taken  into  con- 
sideration. ^-^   -J  ■>^ 

12  ^<^^C><2 


178  GEOLOGY  OF  QUEENSLAND  COAST, 

examples  of  drowned  topography  occur  in  the  neighbourhood  of 
Hinchinbrook  Island,  also  inaST.S.  Wales  in  the  vicinity  of  Port  Jack- 
son and  Hawkesbury  River.  To  the  south  of  Queensland  longer 
continued  marine  erosion  during  periods  of  more  stable  equilibrium 
is  shown  by  the  cliffs  of  Sydney  district,  Victoria  and  Tasmania. 
(This  after  pronounced  movement  or  movements  of  depression.) 

The  existence  of  islands  like  Frazer,  Moreton  and  Stradbroke, 
and  widespread  extensive  coastal  plains  of  the  Queensland  area, 
the  tombolos*  of  the  shelf  islands,  the  lagoons  and  lagoon-marsh 
meadows,  the  bay  bars,  etc  ,  all  point  to  the  adolescent  and  late 
adolescent  stage  of  development  attained  by  the  Queensland  coast. 
These  features  are  necessarily  more  accentuated  south  of  the 
Barrier  owing  to  the  increased  wave-action  in  Extra-Barrier 
areas.  When  the  harbours  of  the  coast  become  silted  uj^,  the 
headlands  cut  back,  and  the  sea  able  to  carry  out  its  intention  of 
establishing  a  straight  or  gently  swinging  shore  line,  then  will  the 
coast  have  advanced  to  maturit}''.  That  period  is  far  away  at 
present,  although  even  this  stage  has  been  attained  in  certain 
small  areas,  seeing  that  elevation  has  accelerated  the  movement. 
The  straight  eastern  shore-line  of  Frazer  Island  is  suggestive  of 
maturity  for  that  particular  spot.  Here,  however,  the  sea  has 
had  a  sand  mass  only  to  work  on. 

The  situation  of  Frazer  Island,  the  distribution  of  the  great 
coastal  flats,  the  deflection  northwards  of  such  streams  as  the 
Brisbane,  Tweed,  Clarence,  Richmond  and  Macleay,  points  to  a 
dominant  along-shore  current  from  the  south,  and  a  weak  fluvia- 
tile  action.  The  streams,  in  their  attempts  to  push  their  loads  off 
shore,  are  opposed  by  a  stronger  wave  action  which  piles  up  river 
bars;  a  dominant  current  redistributes  the  debris  along  its  own 
path,  offsets  and  overlaps  occur,  and  finally  the  stream  is  deflected 
for  miles  out  of  its  normal  course. 

The  occurrence  and  present  aspect  of  the  aoastal  plains  and 
allied  features  at  heights  above  sea  level  argues  a  general  elevation 
in  historic  times  for  the  coast  line  itself. 

*  A  sand  bar  connecting  two  islands,  or  islands  with  mainland.  Gulliver 
"  Shoreline  Topography."     Proc.  Am.  Acad.  Arts  &  Sci.,  1899,  xxxiv. 


BY    E.    C.    ANDREWS.  179 

The  appearance  of  the  coasfcal  area  is  explained  by  the  forces 
of  subaerial  denudation  acting  on  a  plateau  till  a  period  of 
maturity  and  old  age,  when  the  general  outline  of  the  upland 
would  have  vanished,  with  only  a  few  mountains  left  amidst  the 
undulating  country  to  show  its  former  extent.  The  extreme  old 
age  of  this  area  will  be  reached  when  it  is  reduced  to  a  peneplain. 
It  must  be  remembered  that  mature  dissection  of  an  upland  is 
not  necessarily  associated  with  a  mature  neighbouring  coast,  since 
the  ranges  may  have  been  developed  to  this  stage  in  a  cycle  pre- 
ceding a  crustal  movement  which  would  place  the  coast  line  in  its 
initial  stages  when  the  other  was  alread}'-  mature. 

The  frequent  occurrence  of  wide  and  very  thick  masses  of  river 
gravel  underlying  the  various  Tertiary  basaltic  flows  of  the  table- 
land, and  now  associated  with  the  present  insignificant  strearas 
only,  points  to  a  cycle  of  erosion  antedating  the  present  one. 
In  this  first  cycle  several  basaltic  outbursts  occurred,  and 
the  present  3,000  feet  level  of  the  New  England  plateau  was 
developed  near  sea-level,  since  which  period  a  3,000  feet  eleva- 
tion has  taken  place  and  the  present  coastal  forms  evolved.  The 
upland  elevation  and  coastal  subsidence"*  indicate  a  differential 
movement  from  east  to  west,  while  the  various  phases  of  shore 
development  show  a  variable  movement  from  north  to  south. 
The  subsidence  was  accentuated  seawards,  therefore  the  Barrier 
Reef  is  probably  of  some  considerable  thickness,  though  forming 
but  a  mere  fraction  of  the  mass  of  the  continental  shelf. 

One  of  the  most  instructive  and  interesting  features  in  connec- 
tion with  this  subject  is  the  still  more  recent  differential  movement 
from  east  to  west.  On  the  most  easterly  limit  of  the  Great 
Barrier  no  records  exist  of  any  trace  of  elevatory  movement; 
closer  in  shore  Mr.  Jukes  and  Professor  Agassiz  found  abundant 
signs  of  a  slight  movement  varying  from  5  to  20  feet  above 
H.  W.M.  Large  terraces  and  coastal  plains  exist  on  the  mainland, 
up  to  as  much  as  300  feet  in  one  instance. 

On  the  eastern  uplands  themselves  the  recent  streams  which 
ha^•e  flowed  over  the  plateau  in  broad  and  shallow  basins  now 

*  The  coastal  subsidence,  however,  postdated  by  a  long  period  of  time  the 
last  considerable  elevation  of  the  plateau. 


180  GEOLOGY  OF  QUEEN'SLAND  COAST, 

precipitate  themselves  over  precipices  in  places  1,000  feet  in 
height. 

Lower  down"^  the  canons  attain  a  depth  of  3,000  feet,  and 
are  entrenched  upon  their  former  basins,  while  the  general 
level  of  the  plateau  persists  for  as  much  as  30  miles 
beyond,  only  broken  up  by  these  inaccessible  falls,  often 
but  a  few  hundred  yards  in  width,  and  not  noticeable 
a  quarter  of  a  mile  away.  The  traveller  may  wish  to  reach 
a  hill  four  or  five  miles  distant  across  the  gently  undulating 
plateau,  but  a  detour  of  many  milesf  must  be  made  in  order  to 
cover  the  short  intervening  space,  the  gullies  trenching  the  upland 
being  practically  impassable.  This  points  to  a  late  rejuvenescence 
of  the  plateau — the  hydrographic  system  was  disturbed,  and 
*'  valley  in  valley  "  structures  induced,  terminating  in  gigantic 
waterfalls.  Withal,  however,  there  appears  to  have  been  no 
deflection  of  the  streams  by  the  formation  of  the  anticline,  since 
the  canons  open  along  the  recent  stream  basins  into  the  old  valley 
and  plain  tracks.     This  then  points  to  stream  revival. 

The  general  east  and  west  trend  of  the  coastal  streams  shows 
consequent  drainage  attendant  on  the  seaward  slope  of  the  upland 
during  previous  elevations,  while  the  long  north  and  south  courses 
of  the  head  waters  of  such  rivers  as  the  Fitzroy,  Brisbane, 
Clarence,  Hawkesbur}'-  and  Shoalhaven  tell  a  tale  of  subsequent 
drainage  determined  by  the  dominant  meridional  strike  of 
Palaeozoic  rocks  varying  in  hardness.  These  "  strike  "  streams 
point  to  a  youthful  stage  in  stream  development.  Such  rivers  as 
the  Hastings,  now  completely  encircled  by  the  "strike"  streams 
of  the  Manning  and  Macleay,  may  in  time  capture  the  latter, 
owing  to  their  steeper  grade  allowing  them  to  eat  back  more 
quickly  into  the  plateau  (unless  checked  by  very  resistant 
structures  the  while  the  individual  streams  cut  their  way  down 
approximately  to  base  level).     This  will  mark  the  graded  stage 

*  5  to  15  miles, 
t  Special  reference  is  here  made  to  the  canons  of  the  Upper  Macleay 
waters  (New  England). 


BY    E.    C.    ANDREWS.  181 

for  the  upper  courses  of  our  coastal  streams,  since  waterfalls  and 
rapids  will  then  be  no  more. 

Instances  of   these    interesting  phases   of    river  development 
might  be  multiplied  from  the  cordillera. 

It  appears,  then,  that  probably  at  some  period  in  Tertiary  time, 
immediately  preceding  the  present  cycle,  the  Queensland  coast  (as 
also  that  of  N.S.  Wales)  ran  parallel  to,  although  at  a  consider- 
able distance  from,  the  present  shoreline,  and  that  shore  develop- 
ment had  advanced  to  the  advanced  old  age  stage  as  a  result 
of  long-continued  marine  erosion  and  sedimentation.  An  old 
upland*  curved  sympathetically  with  the  coast,  and  was  coeval 
with  it.  The  evidence  goes  to  show  that  immediately  prior  to 
the  great  movement  obtaining  at  present,  this  old  land  was 
dissected  by  subaerial  agencies  as  to  its  eastern  and  western 
portions  to  a  stage  of  old  age.  A  differential  movement 
was  set  up,  involving  such  forms  of  stress  as  tilting,  folding, 
faulting  and  warping,  the  motion  varying  both  from  east  to 
west,  and  from  north  to  south.  The  pivotal  axis  was  probably 
an  inconstant  quantity,  although  confined  in  the  main  to  the 
neighbourhood  of  the  centre  of  the  cordillera.  It  may  be  stated 
here,  although  the  discussion  is  reserved  for  a  future  paper,  that  at 
the  close  of  the  Cretaceous  period  a  long  protracted  cjx'le  of 
erosion  resulted  in  the  formation  of  a  plain  almost  at  sea-level  in 
the  area  occupied  now  by  the  tableland.  An  elevation  of  1,000- 
1,500  feet  took  place  in  the  early  Tertiary  period  ;  while  the 
new  cycle  of  development  was  in  a  fairly  advanced  stage,  several 
distinct  basaltic  outbursts  occurred,  each  separated  by  long- 
intervals  of  time.  The  plateau  was  then  worn  down  to  the  old 
age  stage,  when  another  cycle  of  elevation  intervened,  forcing 
the  Cretaceous  and  Tertiar}^  peneplains  an  additional  3,000 
feet  above  sea-level.  Reduction  of  the  plateau  was  then 
started  afresh  by  the  swiftly  falling  streams,  and  after  an 
anastomosing  series  of  canons  had  been  imposed  upon  the 
plateau,  accompanied  by  mature  or  old  age  coastal  develop- 
ment, a  minor  cycle  of  subsidence  ensued  for  the  coastal  area,  of 

*  Plateau  of  erosion. 


182  GEOLOGY  OF  QUEENSLAND  COAST, 

Pleistocene  a<]je.  This  subsidence  determined  the  present  Barrier 
Reef.  What  the  immediately  antecedent  stage  to  the  early- 
Tertiary  uplift  of  the  Cretaceous  peneplain  may  have  been  is  not 
known  at  present.  Secular  movement  is,  however,  doubtless  the 
key  to  the  initial  stages,  while  subsidence  with  concomitant  sedi- 
mentation was  merely  an  after  factor  in  the  evolution  of  the  main 
features  of  the  present  shore  line  topography.  The  presence  of 
small  cliffs  only  on  the  markedl}^  irregular  Queensland  coast, 
combined  with  the  even  seabottom  of  interinsular  areas,  and  the 
great  extent  of  coastal  plains,  argues  a  gentle  subsidence  combined 
with  filling  of  sunken  coast  valleys,  while  marked  depression 
succeeded  by  more  stable  conditions  during  which  marine  erosion 
could  cut  cliffs  is  hinted  at  b}"  a  study  of  N.S.  Wales,  Victoria, 
and  Tasmanian  coasts.  Shore-grade"*  was  attained  in  places,  and 
island-tying  was  effected  by  the  complicated  processes  of  along- 
shore action.  Splendid  examples  of  this  action  are  illustrated  at 
Challenger  Bay  in  the  Palms  (marine  erosion),  at  Hinchinbrook 
Island  (complication  of  tide,  along  shore  current,  marine  erosion, 
fluviatile  action,  and  elevation),  Stradbroke,  Moreton,  Frazer, 
Curtis  and  other  islands.  Gain  to  the  coast  was  accomplished 
by  the  development  of  coastal  plains.  Bay  bars  were  produced 
by  wave  and  current  action,  while  similar  processes  resulted  in 
the  foi'mation  of  lagoons,  lagoon-marsh  meadows,  and  successive 
aggradation  lines  curving  sympathetically  with  the  shore  line,t 
while  bays  were  becoming  silted  uj^  b}'  fluviatile  and  {eolian 
agencies  after  the  formation  of  bay  bars.|      During  this  period 

*  An  adolescent  feature — attained  when  transportation  occurs  along  shore, 
with  island  tying  and  formation  of  bay  bars. 

t  Line  of  intersection  of  the  sea  with  the  land.  Gulliver,  "  Shoreline 
Topography." 

X  Port  Jackson  is  an  example  of  a  ria  or  submerged  subaerially  curved 
valley,  which  owes  its  commercial  importance  to  its  steep  off- shores,  the 
absence  of  large  streams  discharging  either  into  it  or  into  the  sea  immediately 
south  of  the  Heads,  thus  preventing  silting  up  by  river  action,  or  the  redis- 
tribution of  river  loads  by  wave  and  dominant  along  shore  current  (south) 
action  to  form  spits  and  bay  bars. 

Botany  Bay  is  an  instructive  lesson  in  the  filling  up  of  a  submerged  valley 
by  river  and  ieolian  action  aided  by  recent  elevation.  Aggradation  lines  are 
seen  on  the  large  flat  near  Lady  llobinson's  Beach. 


BY    E.    C.    ANDREWS.  183 

the  dissection  of  the  eastern  uplands  was  carried  on  from  the 
maturity  of  the  previous  cycle  to  a  very  slightly  advanced  stage 
during  the  cycle  of  subsidence.  The  subsidence  of  the  coast 
gave  diminished  stream  action  along  the  lower  portions  of  the 
watercourses.  River  grade  was  also  reached,  no  falls  interrupting 
the  courses  of  the  streams. 

The  sinking  of  the  maturely  (or  adolescently)  developed  coast 
and  shore  of  the  previous  cycle  was  the  occasion  of  its  investiture 
by  coral  growths  to  form  the  Great  Barrier  Reef.  These  growths 
kept  pace  with  the  gradual  subsidence,  and,  as  in  other  coral  reef 
regions,  numerous  deep  water  lanes  became  coexistent  with  the 
growth  of  the  Great  Barrier,  being  determined  probably  in  part 
by  tlie  old  watercourses  of  this  area  of  drowned  topography. 
The  evidence  points  to  the  limitation  of  luxuriantly  growing 
coral  masses  to  the  outer  centre  and  edge  of  the  shelf,  owing  to 
the  turbid  water  west  of  the  centre  of  the  shelf,  the  recession  of 
the  shore  line  from  the  edge  of  the  continental  plateau  permitting 
of  their  existence  in  the  clear  water  to  seaward,  while  the  turbid 
water  of  the  closer  inshore  area  militates  against  its  western 
expansion.  Exceptions  occur  in  the  deeper  off-shore  areas  of 
mountainous  coasts  possessing  inconsiderable  streams  (coast 
north  of  Cairns).  The  inner  and  central  portions  of  the  fairly 
flat  continental  shelf  seem  determined  by  the  conjoint  influences 
of  tides  and  dominant  currents  in  the  way  of  redistributing  the 
loads  brought  down  by  the  rivers,  and  the  material  lost  to  the 
islands  and  mainland  by  the  agencies  of  marine  erosion.  Gain 
to  this  area  is  also  represented  by  various  animal  and  plant 
growths  containing  calcareous  tissues,  coral  growths  being  con- 
fined to  the  formation  of  island  fringing  reefs. 

This  cycle  of  coastal  depression  was  interrupted  in  modern 
times,  as  regards  the  shore,*  by  an  epicycle  of  elevation,  which 
may  have  originated  ni  a  shifting  of  the  pivotal  axis,  inasmuch 
as  that,  while  apparent  elevations  along  the  coast  line  are  con 


*  The  anticline  to  the  west  across  which  the  canons  of  the  coastal  rivers 
have  sawn  their  way  appears  to  have  been  in  process  of  formation  for  a  con- 
siderable period. 


18-1:  GEOLOGY  OF  QUEENSLAND  COAST, 

fined  between  vertical  limits  of  5  and  20  feet,  traces  of  contem- 
poraneous elevation  exist  inland  from  Townsville  to  the  extent 
of  300  feet;  terraces  also  occur  at  Raymond  Terrace  about  20 
feet  above  sea  level,  while  some  10  miles  to  the  west  traces  of 
the  upward  movement  occur  as  much  as  50  feet  above  present 
high  water  mark.  From  observations  made  in  North  Queensland 
and  along  the  New  England  plateau  the  topographical  features 
of  the  central  and  eastern  portions  of  the  cordillera  record  a 
double  cycle  of  elevation.  The  hydrographic  system  was  dis- 
turbed ;  the  broad  shallow  basins  which  had  been  developed 
in  the  plateau  after  the  Tertiary  basalt  outbursts  were  forced 
upwards  again  in  much  more  recent  times,  the  criteria  of  later 
elevation  consisting  in  the  ''valley  in  valle}- "  form  of  the  river 
systems.  The  canons  of  such  rivers  as  the  Macleay,  2,000  to 
4,000  feet  deep,  occupy  the  centres  of  their  former  basins,  and 
end  in  gigantic  waterfalls  sometimes  1,000  feet  in  height.  Other 
e.xamples  of  recently  disturbed  drainage  systems  are  : — 

(1)  The  Barron,  Tull}^,  and  Johnson  falls  in  N.  Queensland. 

(2)  The  various  Clarence  and  Macleay  falls  in  New  England. 

(3)  The  Nepean  and  Shoalhaven  falls  in  southern  N.S.  Wales. 
A  youthful  stage  in  stream  development  is  also  hinted  at  by 

the  "strike"  streams  of  the  upper  waters  of  the  coastal  rivers. 
More  steeply  graded  rivers  travelling  westwards  from  the  coast 
may  hereafter  catch  the  long  meridionally  disposed  headwaters  of 
streams  like  the  Clarence,  Shoalhaven  and  Nepean. 

The  recent  epicycle  of  elevatory  coastal  movement  accentuates 
such  features  as  island  tying,  the  formation  of  lagoons,  coastal 
plains,  bay  bars  and  silting  up  of  bar-bound  harbours. 

Such  rivers  as  the  Brisbane,  Richmond,  Clarence  and  Macleay 
furnish  magnificent  examples  of  the  dominance  of  a  southern 
along-shore  current,  and  the  evident  intention  of  the  sea  to 
establish  a  straight  shore  line  in  the  enormous  deflection  north- 
wards of  these  streams  and  the  presence  of  liuge  bars  at  their 
mouths.  Other  evidence  of  the  influence  of  sea  action  is  manifest 
from  the  position  of  Frazer  Island,  the  uninterrupted  sweep  of 
its   eastern   coast,  its  extensive   northerly  spit,  and   the  gently 


BY    E.    C.    ANDREWS.  IS") 

swinging  coast  south  of  the  island  backed  up  by  lagoons  and 
lagoon-marsh  meadows. 

Criteria  of  the  adolescent  stage  in  coast  formation  appear  to 
be  frequent,  while  maturity  is  still  far  away  in  point  of  time,  as 
is  shown  b}''  the  presence  of  numerous  irregularly  shaped  islands 
and  headlands  with  attendant  unfilled  lagoons,  bays  and  harbours. 

From  a  consideration  of  the  plateau  and  coast  features,  there 
appears  to  be  no  reason  for  assigning  an  age  later  than  Pliocene, 
possibly  late  Pliocene,  for  the  movement  of  elevation  which 
carried  the  Tertiary  and  Cretaceous  peneplains  3,000  feet  above 
their  former  positions  and  resulted  in  the  formation  of  the  canons 
and  upper  valleys  of  the  coastal  area.  The  subsidence  which 
determined  the  present  Barrier  Reef  may  be  referred  to  Pleisto- 
cene times  on  topograjDhical  grounds,  while  the  latest  epicycle  or 
vibration  of  coastal  elevation  may  be  referred  to  the  historical 
period. 


186 


NOTES  ON  THE  BOTANY  OF  THE  INTERIOR   OF 
NEW  SOUTH  WALES. 

By  R.  H.  Cambage. 

Pakt  VI. — From  Marsden  to  Narrandera. 

(Plates  viii.-x.) 

The  distance  from  Marsden  to  W3'along  is  about  20  miles 
westerly.  The  country  is  generally  level,  except  for  a  ridge  of 
sedimentary  formation  at  about  half-way.  Various  trees  and 
shrubs  noticed  were  : — Ereinofhila  Mitchelli  (Budtha  or  Sandal- 
wood), Geijera  parviflora  (Wilga,  getting  scarce),  Heterodendron 
olecefolium  (Rosewood),  Casuarina  Luehmanni  (Bull  Oak),  C. 
Cainhagei  (Belah),  Callitris  robusta  (White  or  Cyj^ress  Pine), 
Sterculia  diversifoLia  (Currajong),  and  Myoporum  deserti  (Dog- 
wood). The  above  were  all  passed  between  Marsden  and  the 
10-mile  post  from  Wyalong.  On  the  ridge  betAveen  the  10-  and 
<S-mile  posts  from  Wyalong  were  Grevillen  floribunda,  Calythrix 
tetragona,  Callitris  calcarata  (Mountain  Pine),  Casuarina  quadri- 
valvis  (She  Oak  or  Mountain  Oak),  Exocarpiis  cuj^ressiformis 
(Native  Cherry),  Caladenia  cceridea  (small  blue  Orchid),  Dodoiicea 
viscosa  (Hopbush),  and  Prostanthera  nvalifolia,  R.Br,  a  shrub 
about  fi^'e  feet  high,  co\ered  with  blue  flowers  in  September. 
Between  the  8-  and  4-mile  posts  were  Hakea  leucoptera  (Needle- 
wood),  Cassia  eremophila,  Casuarina  Cambagei,  C.  Lue1i7nanni ^ 
Callitris  I'obusta,  Birtya  Gunninghamii  (Broom  Bush),  Heteroden- 
dron olecHfolium,  Melaleuca  parviflora^  Lindl.,  and  Eriostemon 
difformis.  Between  the  4-mile  post  and  Wyalong  the  following 
were  noticed : — Calythrix  tetragona,  Melaleuca  uncinata  ( Youngie 
Bush),  Fusanus  acuminatus  (Quandong).  Myoporum  deserti, 
Bursaria  spi^wsa,  Cav.  (Black  Thorn),  Sterculia  diversi/olia,  and 


BY    R.    H.    CAMBAGE.  187 

a  few  trees  of  Eremophila  longifolia  (Emu  Bush).  The  plants  of 
Prostanthera  ovalifolia  and  Bursaria  spinosa  were  the  first  seen 
in  coming  from  Bourke. 

In  the  drought  of  1895,  when  all  grass  and  herbage  had  dis- 
appeared, the  rabbits  in  the  Wyalong  district  turned  their 
attention  to  the  bark  of  various  shrubs  and  small  trees,  but  the 
species  which  suB'ered  most  from  these  rodents  was  Myoporum 
deserti  (Dogwood).  The  bark  was  eaten  off  right  round  the  stem, 
and  as  far  up  as  Bunny  could  reach  when  standing  on  two  legs. 
In  this  way  the  lower  parts  of  these  trees  were  whitened  as  if 
covered  with  white  stockings,  and  presented  a  striking  appearance 
when  growing  in  clusters  in  the  forest.  Scarcely  a  tree  of  this 
species  escaped,  and  the  great  quantity  of  whitened  stems  bore 
testimony  both  to  the  numbers  and  energy  of  their  assailants, 
while  a  great  many  of  the  trees  died  from  this  novel  process  of 
ringbarking.  From  various  sources  I  gathered  that  the  rabbits 
did  not  thrive  on  this  bark,  but  simply  prolonged  existence. 

Throughout  various  localities  in  my  western  travels  I  have  seen 
instances  of  rabbits  eating  the  bark  oti'  the  trunks  and  exposed 
roots  of  very  many  trees  aud  shrubs,  but  have  never  noticed  their 
showing  such  a  decided  preference  for  any  other  species  as  was 
shown  for  this  one  in  the  Wyalong  district.  Usually  their 
favourite  bark  can  be  detected  in  any  forest  by  noticing  which 
trees  are  most  constantly  attacked.  The  observations  of  others 
on  this  point  would  be  of  interest.  In  the  Cobar  district  Capparis 
Mitchelli  (Wild  Orange)  was  one  of  the  shrubs  most  constantl}'- 
attacked.  The  various  Eucalypts,  including  the  Mallees,  are 
evidently  not  in  favour  with  them,  as  these  are  left  untouched 
while  anything  else  can  be  obtained. 

Mijopori.ini  de<erti  has  been  noticed  nearly  the  whole  way  from 
Bourke.  In  the  Nymagee  district  it  is  often  found  growing  near 
the  various  Mallees,  but  is  also  quite  common  in  places  to  the 
eastward  where  these  Eucalypts  do  not  grow.  It  is  usually 
known  as  Dogwood,  though  that  name  is  given  to  other  trees  in 
different  localities.  In  the  Lachlan  district  the  chief  flowering 
time  for  this  species  is  October,  and  in  November  the  ground  may 


188  BOTANY  OF  THE  INTERIOR  OF    NEW  SOUTH  WALES, 

be  seen  in  places  whitened  with  the  fallen  petals.  More  than 
once  I  have  met  with  Western  men  who  have  confused  these  trees 
with  the  Budtha  or  Sandalwood,  Eremo2)hila  Mitchelli:  but  by- 
exercising  a  little  care  the  two  species  are  easily  separated  with- 
out reference  to  flowers  or  fruits.  The  general  appearance  of 
Budtha  is  paler  than  that  of  Dogwood  both  as  regards  bark  and 
foliage.  The  former  flowers  a  little  earlier  than  the  latter,  though 
of  course  occasional  flowers  of  either  may  be  found  during  a 
period  extending  over  several  months. 

Myoporum  deserti  contains  some  kind  of  resinous  substance 
which  may  at  times  be  seen  exuding  from  the  bark.  I  knew  of 
an  instance  in  the  Forbes  district  where  a  flock  of  domestic 
turke3^s  were  in  the  habit  of  making  periodical  visits  to  a  number 
of  these  little  trees  in  order  to  feed  on  this  exudation  while  it 
was  fresh.  Possibly  the  same  substance  was  also  the  attraction 
for  the  rabbits."^ 

In  the  following  list  of  Acacias  passed  between  Marsden  and 
Wyalong  their  positions  are  given  in  regard  to  the  mile-posts; 
those  Acacias  whose  names  precede  any  given  mile-post  were  seen 
before  that  post  was  reached.  The  various  species  were : — A. 
stenophyV a  (on  the  Bland  Creek  only),  A.  pendula  (Boree  or 
Myall);  13  m.  A.  homalophylla  (Yarran);  11m.  A.  dealbata  (green 
variety),  A.  hakeoides,  A.  doratoxylon  (Currawong)  ;  10  m.  A. 
cardiophylla^  A.  Cunn.;  8  m.  A,  homalophylla ;  7  m.  A.  Osivaldi 
(Miljee);  5  m.  A.  rigeyis,  A.  ixiophylla,  Benth.  (?);  4  m  A.difformis, 
Baker  (Black  Wattle);  2  m.  and  A.  decora. 

A.  pendida  ceases  on  this  road  with  the  river  soil  at  about  ISA- 
miles  from  Wyalong,  but  occurs  again  to  the  south  and  south- 
west. As  previously  stated  (Partii.,  p.  719,  and  Part  v.,  p.  690), 
this  species  is  more  often  called  Boree  south  of  the  Lachlan  than 
Myall,  W'hile  A.  homalophylla  is  here  known  both  as  Yarran  and 
Myall. 


*  For  some  notes  on  exudates  from  ^1/.  platycarpiiiii,  by  the  late  Mr.  K.  H. 
Bennett,  see  these  Proceedings,  vii.,  1882,  pp.  350-351. 


BY    R.    H.    CAMBAGE.  189 

A.  cardiophylla  was  found  on  the  sedimentary  ridge  previously 
mentioned,  and  was  not  noticed  in  any  other  part  of  the  countrj^ 
described  in  these  papers.  It  was  flowering  earl}'-  in  September, 
and  in  general  appearance,  as  well  as  size,  much  resembles  A. 
pubesceiis,  R.Br.,  of  the  Parramatta  district. 

A.  rigens  was  found  growing  as  trees  up  to  12  feet  high  with  a 
peculiar  rough  crinkled  bark,  and  needle-shaped  leaves  generally 
having  straight  points.  The  trees  were  covered  with  flowers  earl}' 
in  September. 

A.  ixiophylla  (*?)  was  also  flowering  at  the  same  time,  but  is  a 
smaller  tree,  with  fairly  smooth  grey  bark  and  ^Dale-coloured 
leaves  with  numerous  parallel  veins.  It  was  not  seen  previously, 
and  there  is  some  doubt,  in  the  absence  of  pods,  as  to  its  identity. 

A.  diff'ormis  was  also  first  noticed  here.  This  species,  which  is 
common  in  the  Wyalong  district,  somewhat  resembles  A.  hakeoideSy 
but  has  decidedly  darker  bark  and  foliage.  The  latter  was 
flowering  in  August  and  September,  but  young  pods  were  found 
on  A.  diff'ormis,  showing  that  it  must  have  flowered  some  time 
previously.  These  trees,  which  average  about  10  or  12  feet  high, 
often  form  scrubs  extending  over  many  acres,  and  sufticiently 
thick  to  afford  shelter  for  the  nests  of  the  Mallee  Hen  [Leipoa 
ocellata). 

The  Eucalypts  noticed  between  Marsden  and  Wyalong  were : — 
E.  rostraia  (River  Red  Gum),  E.  Woollsiana  (Black  or  N^arrow- 
leaved  Box) ;  10  m.  E.  sideroxylon  (Ironbark),  E.  tereticoriiis, 
v&v.  dealbata  (Gum);  8  m.  E.  co7iica  (Apple  IBox);  4  m.  E.  diimosa 
(White  Mallee),  E.  sideroxylon  (Ironbark),  E.  jjopulifolia  (Box, 
scarce),  E.  Behriana,  F.v.M.,  and  a  few  trees  of  E.  melliodora 
(Yellow  Box). 

The  district  around  Wyalong  is  interesting  to  lovers  of  either 
geology  or  botany.  The  formation  in  which  the  rich  gold  reefs 
occur  is  granite,  whereas  many,  though  not  all,  of  the  reefing  fields 
in  New  South  Wales  are  in  sedimentary  formations,  ver}'  often 
Silurian  slate,  or,  if  in  Plutonic,  the  rock  is  often  other  than 
granite.  The  depth  to  which  the  rock  has  decomposed,  and  the 
level  at  which  water  too  salt  for  use  is  found,  are  among  the 


190  BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

points  of  geological  interest.  Plants  are  very  plentiful,  and  many 
are  met  with  here  which  do  not  occur  to  the  eastward,  but  extend 
across  into  South  Australia.  The  localit}-  may  be  regarded  as 
being  on  the  eastern  boundary  of  the  true  interior  flora. 

Various  trees  and  shrubs  noticed  around  Wyalong  and  tov.  ards 
Barmedman  were  : — Bertya  Cunninghamii,  Sterculia  diversifoJia, 
Myoporum  deserti,  Melaleuca  parvijlora,  J/,  uncinata,  Olearia 
phneloides,  A.  Cunn.,  0.  lepidopht/lla,  Benth.,  Eriostemon  difformis, 
Philotheca  ciliata,  Hook.,  Bursaria  spinosa,  Pittosporum  phiHy- 
rceoidesy  Fxocarpus  aphylla,  Phehalium  glandidosum,  P.  ohcorda- 
tum,  A.  Cunn.,  Callit7'is  robusta,  C.  calcarata,  Halea  leucopdera, 
Cassia  eremophila,  Calythrix  tetragona^  Casuarina  Luehmaiini, 
C.  Cambagei,  Cassytha  sp.,  Dodonce.a  sp.,  Heterodendron  olecefolium, 
Templetonia  egena,  Benth.,  Fusanus  acuminatus,  Thryptomene 
Tninutiflora,  F.v.M.,  Lissanthe  strigosa,  R.Br.,  Prostanthera 
coccinea,  F.v.^l.,  Solanuin  simile,  F.v.M.,  Helichrysum  obcordatum, 
F.v.M. ,  and  Leptospermum  myrsinoides,  Schl.  This  last-named 
species  was  also  found  north  of  the  Lachlan,  and  was  previously 
mentioned  as  Tea-tree  (Part  iv.,  p.  325),  but  not  then  definitely 
identified. 

The  species  of  Cassytha  growing  at  AVyalong  is  found  as  a 
parasite  clinging  to  and  often  almost  smothering  the  top  of  the 
Mallee  or  other  bushes,  the  leafless  vines  being  so  numerous  as  in 
many  cases  to  form  quite  a  canopy.  They  are  often  known  as 
Mallee-vines,  and  are  useful  as  a  fodder.  The  fruits  are  con- 
sidered edible  by  children. 

Eriostemon  difformis  flowers  profusely  around  Wyalong  ■  in 
September,  and  contributes  much  to  the  beauty  of  the  flora. 

Pliebalhtm  obcordatum  was  found  at  Wyalong  only,  and, 
judging  by  its  absence  from  herbaria,  is  evidently  a  plant  not 
often  met  with  by  the  collector.  It  is  somewhat  difl'use  in  its 
habit,  and  when  covered  w^ith  yellow  flowers  is  an  ornamental 
little  shrub.  Usually  the  colour  of  a  flower  is  decided  by  the 
colour  of  its  petals,  but  in  this  species  it  is  from  the  anthers  that 
the  yellow  appearance  emanates,  a  feature,  of  course,  not  confined 
to  this  genus.     I  have  noticed  these  flowers  apparentl}^  change 


BY    R.    H.    CAMBAGE.  191 

colour  after  a  shower  of  rain,  but  an  inspection  showed  that  it  was 
simply  owing  to  the  removal  of  the  pollen  b}^  the  rain  drops. 

Melaleuca  parviflora  is  fairly  common  around  Wyalong,  and  in 
general  appearance,  as  well  as  habit,  is  exactly  similar  to  this 
species  growing  in  the  Ninety-Mile  Desert,  South  Australia.  On 
the  coast  of  New  South  Wales  it  is  generally  a  darker  green,  and 
the  leaves  are  often  more  flexuose. 

The  Acacias  around  Wyalong  and  towards  Barmedman  are 
numerous,  and  include  several  which  had  not  been  seen  along  the 
country  described  to  the  northward,  as  well  as  one  not  previously 
recorded  for  New  South  Wales.  Those  noticed  were  : — A.  mon- 
tana,  Benth.,  A.  microcar^Kt,  F.v.M.,  A.  rigens,  A.  aspera,  Lindl., 
A.  conferta,  A.  acinacea,  Lindl.,  A.  t^ernicifiua^  A.  hakeoides,  A. 
decora,  A.  cidtri/ormis,  A.  Cunn.  (west  of  Wyalong ',  A.  homalo- 
jihylla,  A.  pendula  (towards  Barmedman),  .1.  Osivaldi,  A. 
ixiophylla  (?),  A.  difformis,  A.  dealbata  {green  \ariety),  A.  ohliqiia, 
A.  Cunn.,  and  A.  rhigiophylla,  F.v.M. 

A .  microcarpa  grows  as  a  shrub  3  or  4  feet  high,  spreading  across 
the  top  to  4  and  5  feet.  In  general  appearance  it  somewhat 
resembles  A.  montana,  but  the  latter  is  slightly  taller,  not  so 
spreading,  and  very  viscid,  while  its  leaves  have  several  nerves, 
those  of  A.  micTocar'pa  having  but  the  central  one,  and  that  not 
very  prominent. 

A.  rhigiophylla  occurs  a  few  hundred  yards  north-westerly  of 
West  Wyalong,  and  has  not  previously  been  recorded  for  New 
South  Wales.  It  was  originally  described  from  specimens 
collected  in  South  Australia;  but  the  country  to  the  westward  of 
Wyalong  is  of  much  the  same  nature  as  some  of  the  eastern  parts 
of  South  Australia,  so  the  plant  does  not  appear  to  be  out  of 
place  in  this  part  of  New  South  Wales,  and  will  probably  be 
found  to  extend  over  a  considerable  area  in  the  south-west.  It 
grows  as  a  somewhat  spreading  shrub,  a  few  feet  high,  with  rigid 
pointed  leaves. 

Between  the  two  towns  of  Wyalong  and  West  Wyalong,  which 
are  situated  about  two  miles  apart,  the  country  is  almost  level  and 
was   formerly  covered  with  Mallee,  though   some    of    it  is  now 


192  BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

cleared.  There  appear  to  be  four  species,  but  the  scrub  is  made 
up  chiefly  of  two,  viz..  Eucalyptus  Behriana  and  E.  polyhractea  ^ 
Baker.  The  other  two,  which  are  not  so  plentiful,  are  E.  dumosa 
and  E.  viridis,  with  an  occasional  Box  (E.  Woollsiana)  and  Iron- 
bark  [E.  sideroxylon). 

E.  Behriana  usually  grows  as  a  Mallee  about  8  to  15  feet  high, 
though  in  rare  instances  I  found  trees  up  to  about  35  feet  high, 
with  a  diameter  of  9  or  10  inches,  the  bark  being  smooth  and 
white,  though  often  toning  off  to  greenish  and  grey.  Perhaps 
the  most  striking  feature  of  this  Mallee  is  that  it  has  fairly  broad, 
smooth,  and  generally  shining  leaves,  though  this  latter  is  not  so 
marked  as  in  the  case  of  E.  popidifolia.  Yet  to  the  westward  of 
Wyalong  I  have  collected  a  few  specimens  of  this  species  having 
silvery-white  powdered  leaves,  though  on  no  other  occasion  was 
this  glaucous  form  met  with.  The  flowers  of  E.  Behriana  are 
small  and  the  fruits  almost  sessile.  Its  timber,  which  is  of  a  pale 
red  colour,  is  usually  too  small  to  be  of  use,  but  from  the  larger 
trees  it  is  sometimes  brought  in  and  used  for  timbering  the  mining 
shafts,  though  that  growing  near  the  mines  on  the  goldfields  is 
too  small  for  the  purpose. 

E.  pohjhractea  has  never  been  seen  by  me  except  around 
Wyalong,  though  it  probably  occurs  to  the  westward.  So  far  as 
observed  it  has  alwaj's  a  glaucous  appearance,  and  is  in  conse- 
quence known  as  Blue  Mallee.  If  it  were  not  for  this  white 
powder  on  the  leaves  there  would  be  some  difiiculty,  without 
examination,  in  separating  it  from  E.  viridis^  and  even  in  the 
herbarium  specimens  the  similarit}'  in  buds  and  fruits  is  notice- 
able, but  the  identification  is  here  also  assisted  by  the  glaucous 
appearance  of  the  leaves.  When  I  first  found  this  species  I 
adopted  the  method  of  crushing  a  few  leaves  to  ascertain  vvhether 
it  could  be  separated  from  E.  viridis,  and  soon  found  this  could 
readily  be  done,  as  the  Blue  Mallee  seems  exceedingly  rich  in  oil. 
In  no  case  was  this  species  found  growing  into  a  tree,  but  usually 
from  6  to  10  feet  high  and  forming  dense  scrubs. 

The  Eucalypts  noticed  on  the  roadside  between  Wyalong  and 
Barmedman    were:  -E.   dinnosa,    K.     Behriana,    E.    poJyhractea 


BY    K.    H.    CAMBAGE.  193 

(only  near  Wj^along),  E.  melliodora  (scarce),  E.  tereticornis 
(Forest  Red  Gum),  E.  conica  (scarce),  E.  WooUsiana,  E.  sider- 
oxi//o)i,  and  E.  tereticornis  var.  dealbata. 

The  first  three  Eucalypts  mentioned  are  Mallees,  but  a  few 
trees  of  E.  dumosa  were  noticed  about  50  feet  high  and  a  foot  in 
diameter.  After  passing  Barmedman  no  more  Mallee  was  seen, 
but  it  is  to  be  found  extending  away  to  the  south-west.  Though 
it  is  very  plentiful  just  at  Wj'-along,  it  may  be  said  that  between 
the  Lachlan  and  Murrumbidgee  Rivers  it  is  not  common  east  of  a 
line  joining  Condobolin,  Wyalong,  Temora  and  Wagga,  and  does 
not  come  up  to  this  line  except  in  places. 

E.  conica  was  last  seen  at  a  few  miles  south  of  Wyalong,  and 
this  is  probably  one  of  its  most  southern  points  in  this  locality. 

E.  populifolia  was  not  noticed  south  of  the  Wyalong  district, 
but  from  this  point  northwards  it  is  one  of  the  commonest  trees 
in  the  interior.  Although  usually  known  as  Bimble  Box,  I  was 
informed  that  away  to  the  westward  of  Wyalong  it  bears  the 
name  of  Mint}^  Box,  but  as  I  did  not  visit  the  locality  indicated 
I  cannot  be  certain  of  the  species.  E.  populifolia  is  the  Box  tree 
which  usually  produces  such  a  number  of  seedlings  after  the  land 
has  been  ringbarked,  and  in  this  respect  is  a  considerable  source 
of  annoyance  to  the  landowner. 

From  Barmedman  to  Temora  is  south-easterly  about  20  miles, 
and  various  trees  and  shrubs  noticed  along  the  roadside  were  : — 
Heterodendron  olecf^folium,  Myoporum  deserti,  CaHitris  calcarata, 
C.  robusta,  Casuarina  Luehmanni,  Cpaludosa,  Sieb.  (?),  Dodoncea 
viscosa,  Exocarpus  cupres  si/or  mis,  Cassinia  Theodori,  F.v.M., 
(Sifting  Bush),  Eusamcs  acwminatus,  Cassia  ereinop>hila,  Caly- 
thrix  tetragona,  Hihbertia  stricta,  R.Br.,  Brachyloma  daphnoides, 
Pimelea  linifolia,  Sm.,  Eremophila  longifolia  (very  scarce), 
Sterculia  diversi/olia,  and  Indigofera  australis,  Wild.  On  a  ridge 
just  west  of  Barmedman  a  species  of  Olearia  was  found,  but  so 
far  not  identified. 

The  tree  mentioned  as   Casuarina  paludosa  (1)    is    found    in 
various  parts  of  Central   New  South  Wales,  but  chiefly  to  the 
eastward  of  the  route  followed  in  these  papers.     It  is  doubtful 
13 


194      BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

whether  this  is  really  C.  paludosa  at  all,  or  a  distinct  species.  I 
have  collected  it  on  the  Harvey  Range  near  Peak  Hill,  near 
Manildra,  and  on  the  hills  around  Cow  Flat  in  the  Bathurst  dis- 
trict. It  grows  rather  more  as  a  little  tree  than  a  shrub,  and 
ranges  from  about  6  to  12  and  sometimes  15  feet  high.  The 
medullary  rays  in  the  wood,  though  fine,  are  quite  distinct,  while 
the  bark  is  generally  smooth.  A  feature  of  this  species  is  that  in 
all  the  localities  mentioned  it  usually  grows  on  dry  ridges,  and  not 
in  damp  places,  as  its  botanical  name  would  suggest.  Its  affinities 
appear  to  be  with  C.  suherofia  on  the  one  hand,  and  C.  distyla 
on  the  other;  but  I  have  never  met  with  either  of  these  sjDecies 
between  the  Macquarie  and  Murrumbidgee  Rivers.  C.  distyla  is 
mentioned,  however  (B.Fl.,  vi.,  198),  as  having  been  collected  on 
the  Lachlan  by  Allan  Cunningham,  though  imperfect  specimens  of 
C.  pahidosa  C?)  might  possibly  be  confused  w^ith  those  of  C.  distyla. 

Cassinia  Theodori  is  very  plentiful  in  the  Temora  district, 
where  it  is  known  by  the  name  of  Sifting  Bush.  This  name  is 
suggested  by  the  similarity  of  the  numerous  fallen  florets  to 
the  "  sif tings  "  which  are  blown  away  from  grain  by  a  winnowing 
machine.  Although  only  a  few  feet  high,  it  forms  a  dense  under- 
growth in  places  where  the  soil  is  slightly  inferior,  often  covering 
many  acres;  and  in  addition  to  affording  a  shelter  for  vermin,  it 
sometimes  encroaches  on  wheattields  which  are  adjacent  to  these 
areas. 

At  intervals  along  the  whole  of  the  route  followed  from 
Bourke,  patches  of  a  species  of  Cucumis,  usually  known  as  melons, 
were  seen.  They  are  found  on  good  soil,  and  the  vines  from  each 
plant  radiate  several  feet,  being  covered  with  small,  round  melons 
nearly  an  inch  in  diameter,  and  exceedingly  bitter.  In  the 
autumn  when  the  vines  are  dead,  and  especially  in  times  of 
drought  when  the  grass  has  disappeared  as  well,  these  green 
marbles  are  very  conspicuous,  and  sometimes  are  scattered  over 
several  acres  where  the  vines  have  been  numerous.  So  far  as  I  could 
judge  they  seem  to  be  untouched  by  stock  of  any  kind,  though 
recently  a  report  came  from  Lake  Cowal  to  a  Sydney  daily  jDaper 


BY    R.    H.    CAMBAGE.  195 

that  150  sheep  had  died  presumably  from  eating  these  wild  melon 
seeds. 

The  Acacias  noticed  between  Barmedman  and  Temora  were  : — 
A.  honialophylla  (only  near  Barmedman),  A.  hakeoides,  A.  dealhata 
(green  variety),  A.  verniciflua,  A.  flexifolia^  A.  Cunn.,  (which  had 
just  ceased  flowering  in  September),  A.  difformis^  A.  decora,  and 
A.  ixiophylla  (?). 

The  Eucalypts  were  represented  by  ^.  Woollsiana,  E.  sideroxylon, 
E.  melliodora  (increasing  in  quantity),  E,  tereticornis  and  var. 
decdbata,  one  tree  of  E.  ajfinis  and  three  of  Ironbark  Box  similar 
to  the  questionable  hybrid  of  ]Sr3^magee. 

E.  Woollsiana,  which  is  known  locally  as  Black  Box,  was  being- 
cut  for  railway  sleepers.  Finding  it  on  a  ridge  near  Barmedman 
with  E.  sideroxylo7i,  I  searched  a  considerable  area  for  the 
questionable  hybrid,  and  eventually  found  three  trees  fairly  close 
together,  with  the  bark,  timber  and  fruits  about  midway  between 
those  of  the  other  two,  and  corresponding  with  trees  previously 
found  in  similar  compsmy  (vide  Part  ii.,  p.  716,  and  Part  iii.,  p.  324). 
I  am  still  unable  to  offer  any  definite  opinion  as  to  what  these 
trees  really  are.  In  a  general  way  they  seem  to  more  nearly 
approach  E.  sideroxylon  than  any  other  local  species,  but  the 
fruits  are  smaller,  the  bark  less  rough  and  inclined  to  be  like  that 
of  the  Box,  and  the  wood  yellowish,  while  they  seem  too  scarce 
to  represent  a  distinct  species.  In  this  instance,  as  previously, 
they  were  only  found  as  the  result  of  special  search.  Although 
these  trees  appear  to  be  very  rare,  it  may  be  mentioned  that  in 
nearly  every  case  they  have  been  found  in  twos  or  threes. 

E.  Woollsiayia,  between  Barmedman  and  Temora,  showed  a  dis- 
tinctly glaucous  appearance  in  September,  a  feature  never  noticed 
by  me  at  an}?-  other  place. 

In  going  southerly  from  Temora  towards  Sebastopol  for  about 
10  miles,  the  following  plants  were  noticed : — Pittosporum 
phillyrceoides,  Daviesia  corymhosa,  Sm.,  var.,  Myoporum  deserti, 
Fusa7ius  acumi^iatios,  Cassinia  Theodori,  Callitris  rohusta,  C. 
calcarata,  Dodoncea  viscosa,  Exocaryus  cupressi/ormis,  Bursaria 


196  BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

n'pinosa^  Pterostylis  mutica,  R.Br.,  CaJadenia  claviyera,  A.  Cunn., 
and  Glossodia  major,  R.Br. 

The  Acacias  passed  were  A.  hakeoides,  A.  difformis,  A.  conferta, 
A.Jlexifolia,  A,  dealbata,  and  var.  (green  variet}^). 

This  is  the  second  instance  in  coming  from  Bourke  that  the 
typical  A.  dealbata  has  been  found.  Easterly  and  southerly  from 
this  point  it  becomes  more  plentiful. 

The  Eucalypts  noticed  southerl}^  from  Temora  were: — E. 
Woollsiana  (still  known  as  Black  Box),  E.  melliodora,  E.  tereti- 
cornis,  E.  tereticornis  var.  dealbata,  E.  sideroxylon  and  E.  macro- 
rhyncha,  F.v.M.  (Stringybark). 

Over  the  area  described  in  these  papers  this  is  the  first  record 
of  Stringybark.  This  species  prefers  a  colder  climate  than  is 
found  in  the  interior,  and  the  fact  of  its  being  so  far  west  as 
Temora  is  possibly  owing  to  the  influence  of  southern  latitudes. 
North  of  the  Murrumbidgee  this  is  the  most  western  point  at  which 
I  have  ever  found  the  species;  but  a  settler  informed  me  that 
there  are  similar  Stringybark  trees  on  Scrubby  Mountain,  near 
Rankin's  Springs,  about  70  miles  north-west  of  Temora.  If  this 
is  the  case,  the  species  is  probably  E.  macrorhyncha,  it  being  by 
far  the  most  westerly  of  our  Stringybarks.  The  statement  is 
probably  correct,  but  requires  investigation.  The  most  western 
point  in  New  South  Wales  at  w^hich  I  have  collected  this  species 
is  Albury.  E.  rtiacrorhyyicha  has  an  extensive  range  in  this  State, 
and  is  fairly  plentiful  over  the  area  which  extends  north  and 
south  along  the  Great  Dividing  Range,  spreading  sometimes  to 
the  westward  and  again  to  the  eastward  of  the  range.  South  of 
Mudgee  its  western  boundary  coincides  very  nearly  with  the 
eastern  boundary  of  E.  hemiphloia  var.  albens,  except  in  isolated 
cases  as  at  Temora,  Grenfell  and  Manildra,  svhere  it  occupies 
ridges  to  the  westward.  So  far  as  my  observations  have  gone  I 
find  that  it  undoubtedly  prefers  a  geological  formation  of  sedi- 
mentary origin,  and  may,  therefore,  often  be  found  on  goldfields 
in  Silurian  slate  areas.  To  the  miner  and  settler  it  is  a  most 
useful  tree.  The  bark  is  used  freely  for  roofing  huts  and  out- 
buildings, while  its  timber  is  greatly  in  request  in  all  kinds  of 


BY    R.    H.    CAMBAGE.  197 

mining  work,  building  and  fencing.  In  the  highlands  around 
Bathurst  it  is  one  of  the  most  generally  useful  trees  the  inhabi- 
tants possess,  its  timber  being  superior,  both  in  or  out  of  the 
ground,  to  that  of  many  of  the  others  growing  in  the  same 
elevated  localities,  besides  being  easy  to  work.  On  the  western 
slopes  it  is  known  as  Stringy  bark,  but  often  in  places  to  the  east- 
ward it  is  called  Red  Stringybark  from  the  colour  of  the  wood, 
and  to  separate  it  from  other  String^'^barks  which  grow  in  the 
same  localities  and  ha\e  paler  timber. 

FromTemora  toCootamundra,  via  Sprnigdale  and  Stockinbingal, 
is  about  36  miles  south-easterly,  and  a  careful  study  of  plants 
along  this  route  will  show  that  the  western  flora  is  being  gradually 
left  behind  and  replaced  by  some  of  the  forms  which  are  usually 
found  on  slightly  higher  levels  to  the  eastward.  Thus  many  trees 
wdiich  have  been  noticed  along  the  greater  part  of  the  road  from 
Bourke  are  not  seen  east  of  Temora,  except  perhaps  in  very  rare 
instances,  among  others  being  Hakea  leucoptera^  Heterodendron 
olecefoliutn,  E^'emophila  Mitchelli,  Geijer'a  parvijiora,  Casuarina 
Cambagei,  Acacia  pendida  and  A.  homalophylla. 

Various  trees  and  shrubs  noted  before  reaching  Springdale  at 
11  miles  were: — Myoporuiin  deserti,  Casua7'ina  Luehmanni,  a 
very  little  of  Eremophila  longijolia,  Callitris  rohusta,  C.  calcarata 
(generally  on  ridges  with  E.  sideroxylon),  Dodonce,a  viscosa, 
Cassinia  Theodori,  Cassia  eremophila^  Bursaria  spinosa,  Fusanus 
acunmiatus,  and  Exocarjnis  cupressiformis. 

Between  Springdale  and  Stockinbingal,  a  distance  of  about  1 1 
miles,  the  following  were  passed  :—  Callitris  calcarata,  C.  robusta, 
Cassinia  Theodori,  Fusanus  acuininatus,  Grevillea  jioribunda, 
Dodoncea  viscosa,  Bursaria  spinosa,  Casuarina  Luehmanni,  C. 
quadrivalvis,  Drosera  sp.,  and  Eiitaxia  empetrifolia,  Schl, 

Between  Stockinbingal  and  Cootamundra,  a  distance  of  14 
miles,  most  of  the  land  is  under  cultivation,  but  on  one  hill 
Exocarpus  cupressiformis  and  Casuay'ina  quadrivalvis  were 
noticed. 

The  Acacias  passed  between  Temora  and  Springdale  were  :  — 
A.  difformis,  A.  montana,  A.  hakeoides  and  A.  dealbaia  (green 
variety). 


198  130TAXY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

Between  Springdale  and  Stockinbingal  there  were :  —  A. 
hakeoides,  A.  conferta,  A.  dealhata,  A.  difformis,  and  A.  ai^mataj 
R.Br.,  the  first  noticed. 

Between  Stockinbingal  and  Cootamundra  A.  diffusa,  Edw., 
was  seen  for  the  first  time. 

Acacia  Bailey  ana,  F.v.M.,  (Cootamundra  Wattle),  is  fairly 
plentiful  a  few  miles  to  the  westward  of  the  road  travelled,  being 
common  along  parts  of  the  main  coach  road  from  Cootamundra 
to  Temora;  and  is  to  botanists  a  most  interesting  species.  It  is 
now  a  well  known  ornamental  tree  in  many  gardens  in  various 
States,  but  there  apparently  was  a  time  when  it  grew  nowhere 
except  near  Cootamundra.  From  considerable  enquiries  made,  I 
conclude  that,  prior  to  its  cultivation,  a  circle  with  a  radius  of 
30  miles,  or  perhaps  less,  described  around  a  centre  somewhere 
between  Cootamundra  and  Temora,  would  have  included  every 
tree  of  Cootamundra  Wattle  in  the  known  world.  The  question 
naturally  arises  whether  it  is  a  new  Acacia  which  originated  here, 
or  a  remnant  once  more  plentiful,  but  now  gradually  disappearing 
altogether.  Without  a  lengthy  investigation,  the  evidence  avail- 
able on  which  to  solve  the  matter  is  very  slight,  so  that  I  am 
unable  to  express  any  opinion  concerning  it. 

The  Eucalypts  passed  between  Temora  and  Stockinbingal 
were  :  — E.  Woollsiana,  E.  melliodora,  E.  liemiphloia  var.  alhens  (a 
few  trees  were  flowering  in  September,  which  is  very  late),  E. 
sideroxyhn,  E.  affnis,  E.  tereticornis  var.  dealhata,  and  E.  macro- 
rhyncha. 

Between  Stockinbingal  and  Cootamundra  were: — E.  meUiodora, 
E.  liemiphloia  var.  albens,  E.  Woolhiana  (becoming  scarce),  E. 
tereticornis,  E.  macrorhyncha,  E.  Cambagei,  Deane  and  Maiden, 
and  E.  Bridgesiana,  Baker. 

E.  Woollsiana  may  be  said  to  cease  at  Cootamundra.  South  of 
the  Macquarie  River  the  eastern  boundary  of  this  species  may  be 
roughly  described  by  lines  joining  Wellington,  Molong,  Cudal, 
Canowindra,  Mount  McDonald,  Koorawatha,  Cootamundra,  and 
Albury,  passing  thence  into  Victoria. 


BY    R.    H.    CAMBAGE.  199 

E.  sideroxyJon  is  also  rare  to  the  east  of  Cootamundra  except 
in  some  isolated  coast  spots,  the  elevation  being  too  great  for  it. 
Its  eastern  boundary,  south  of  the  Macquarie,  corresponds  very 
nearly  with  that  of  E.  Woollsiana.  It  also  extends  into  Victoria  at 
least  as  far  as  Chiltern,  showing  little  or  no  variation.  The  other 
Victorian  tree,  known  as  E.  leucoxylon,  F.v.M.,  though  having  a 
smooth  white  bark  and  yellowish  wood,  is  thought  by  some  to  be 
the  same  species  as  the  New  South  Wales  E.  sideroxylon,  but  the 
distinct  difference  in  both  bark  and  timber  seems  to  me  sufficient 
to  show  that  they  are  separable,  notwithstanding  a  considerable 
similarity  in  the  fruits.  So  far  I  have  not  met  with  any  trees 
showing  a  decided  gradation  from  one  species  to  the  other,  those 
which  have  come  under  my  notice  in  the  forests  being  either 
typical  Ironbarks  or  Gums. 

E.  Cambagei  is  a  species  extending  over  a  considerable  area  on 
the  highlands  of  New  South  Wales  and  in  Victoria,  though  in 
the  latter  State  it  is  also  found  almost  down  to  sea-level,  which  is 
probably  another  instance  of  the  effect  of  southern  latitudes.  In 
the  Mudgee,  and  Bathurst  to  Goulburn  districts  it  is  seldom  found 
below  an  altitude  of  2,000  feet  above  sea-level,  though  near 
Cootamundra  it  is  growing  at  about  1,200  feet,  and  at  about  550 
near  Albury.  It  is  known  under  the  names  of  Apple,  Mountain 
Apple,  Bastard  Box,  and  Bundy,  the  latter  being  the  local  name 
south  of  Bathurst  around  Rockley  and  Burraga,  where  it  is  in  con- 
siderable request  as  a  fuel  in  the  copper  smelting  furnaces.  South 
of  the  Macquarie  River  E.  Cambagei  is  seldom  found  west  of  a  line 
joining  Wellington,  Molong,  Cargo,  Mount  McDonald,  Gundagai 
and  Albury.  There  are,  however,  a  few  patches  of  it  to  be  found 
west  of  this  line,  one  being  near  Bumberry  between  Molong  and 
Parkes,  and  that  now  under  discussion  near  Cootamundra,  while 
there  are  probably  other  small  areas  of  it  in  isolated  spots.  In 
these  extreme  western  localities  it  is  usually  found  occupying  the 
tops  of  hills,  and  is  undoubtedly  more  in  its  regular  home  on  the 
higher  lands  to  the  eastward.  The  wood  of  this  tree  is  hard  for 
a  mountain  species,  but  as  the  trees  are  seldom  straight  or  tall 
the  timber  is  not  considered  to  be  of  much  value.     In  the  Bathurst 


200  BOTANi:  OF  THE  INTERIOR  OF  NBW  SOUTH  WALES, 

and  Orange  districts  it  may  generally  be  found  growing  on  ridges 
of  Silurian  slate;  and  although  it  evidently  prefers  a  sedimentary 
formation,  it  is  occasionally  to  be  found  on  hills  of  igneous  origin, 
though  in  such  cases  the  wood  often  appears  to  be  softer,  probably 
owing  to  a  more  rapid  growth  through  being  in  a  better  soil.  In 
no  case  does  it  appear  to  grow  on  an  alluvial  flat.  The  bark  of 
this  tree  is  a  dark  grey,  somewhat  resembling  a  Box  bark,  but  of 
a  more  woolly  nature,  toning  off  to  smooth  on  the  ultimate 
branches.  It  is,  however,  only  in  rare  instances  that  it  is  found 
in  company  with  typical  Box  trees  such  as  E.  hemiphloia  var. 
albens,  or  F.  Woollsiana,  preferring  a  colder  climate. 

For  many  years  E.  Camhagei  was  confused  in  various  herbaria 
with  E.  goniocalyx,  F.v.M.,  but  in  the  forest  the  two  species  are 
scarcely  ever,  or  it  might  almost  be  said  never,  confounded.  The 
latter  is  usually  a  large  straight  Gum  tree,  found  along  the  south 
coast  and  in  Victoria,  having  smooth  bark  and  moderatel}^  hard 
fissile  timber;  while  the  former  has  more  of  a  Box  or  Apple  tree 
appearance,  with  hard,  interlocked  timber.  In  some  cases,  how- 
ever, E.  goniocalyx  has  a  rough  bark  somewhat  similar  to  that 
at  times  seen  on  E.  saliyna,  Sm.,  the  Sydney  Blue  Gum,  covering 
the  trunk  almost  up  to  the  branches.  I  have  seen  this  at  Fern 
Tree  Gully,  near  Melbourne,  where  my  attention  was  drawn  to  it 
by  Mr.  J.  G.  Luehmann,  F.L.S.  The  sucker-foliage,  however, 
separates  these  two  species  very  clearly. 

The  tree  with  which  E.  Camhagei  is  often  confused  in  the 
forest  is  that  named  E.  Bridyesiana  by  Mr.  R.  T.  Baker  (Proc. 
Linn.  Soc.  N.S.W.  1898,  Part  2)  and  formerly  recognised  as^. 
Stuartiana,  F.v.M.,  there  being  a  fancy  with  many  bushmen  that 
the  latter,  often  known  as  Apple,  and  which  has  a  soft  wood  and 
grows  on  flats,  graduates  into  the  former  species,  which  grows 
on  the  hills  and  has  a  hard  wood.  Botanically  these  two  trees 
are  very  distinct,  but  the  similarity  in  the  bark  causes  the  con- 
fusion among  casual  observers. 

E.  Bridgesiana  was  not  met  with  in  coming  from  Bourke  until 
Cootamundra  was  reached.  It  extends,  north  and  south,  almost, 
if  not  quite,  through  the  entire  length   of  New  South  Wales, 


BY    R.    H.    CAMBAGE  201 

following  in  a  general  way  the  course  of  the  Great  Dividing- 
Range,  being  found  perhaps  more  on  its  western  than  eastern 
slopes  and  seldom,  if  ever,  coming  down  to  the  sea-level.  It  is 
usually  known  as  either  Apple  or  Woolly-butt,  the  former  name 
being  in  general  use  to  the  south  of  Bathurst,  while  it  is  known 
by  the  latter  name  to  the  northward,  though  these  names  are  not 
constantly  allotted  according  to  the  above  divisions.  In  some 
localities  it  is  called  Peppermint,  \vhile  in  others  it  bears  the  name 
of  Box  from  a  similarity  in  the  appearance  of  the  bark  to  some 
of  the  Box  trees.  It  takes  its  name  of  Apple  from  a  general  out- 
ward similarity  which  it  has  to  Angophora  intermedia,  DC,  that 
species  being  known  as  Apple  almost  wherever  it  is  found  in  New 
South  Wales;  and  when  these  trees  grow  together  the  Eucalypt 
is  often  designated  Woolly-butt  by  way  of  distinction.  Although 
Angophora  intermedia  is  plentiful  all  along  the  coastal  districts, 
and  crosses  to  the  western  slopes  in  many  places  to  the  northward, 
it  is  rare  within  that  large  area  lying  to  the  south  of  the  Great 
Western  Railwa}^  Line,  and  to  the  west  of  the  Great  Dividing- 
Range,  though  it  touches  the  boundary  of  this  area  near  Welling- 
ton, and  may  possibly  be  found  somewhere  in  the  Yass  and 
Tumut  districts.  Within  this  area  £.  Bridgesiana  is  usually 
known  as  Apple.  But  the  name  changes  to  Woolly-butt  north 
of  Bathurst  and  in  the  Rylstone  district,  as  A.  intermedia  begins 
at  about  7  miles  from  Bathurst  towards  Sofala  and  continues 
northwards. 

E.  Bridgesiana  is  generally  found  growing  on  flats,  but  some- 
times on  elevated  land  in  basaltic  country,  and  in  the  colder  parts 
often  in  company  with  a  white  drooping  Gum,  E.  vlminalis, 
Labill.  In  habit  these  trees  somewhat  resemble  each  other,  but 
the  white  smooth  bark  of  the  latter  is  very  distinct  from  that  of 
the  former,  which  has  a  grey  Box  bark,  usually  covering  all  the 
trunk  and  part  of  the  branches.  Although  these  two  species 
grow  in  company  in  the  cold  parts,  it  is  found  in  going  to  the 
lower  western  country  that  the  Gum  ceases  considerably  before 
the  Apple.  The  timber  of  this  Apple-tree  is  considered  useless, 
beinsc  even  valueless  as  a  fuel. 


202      BOTAXY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

Near  its  western  limit  it  is  often  found  on  the  river  flats  in 
company  with  E.  melliodora  (Yellow  Box),  and  reaches  its  most 
western  points  along  the  rivers  to  the  south.  From  the  Mac- 
quarie  River  southwards  its  western  boundar}^  may  be  roughly 
defined  by  lines  joining  Wellington,  Eugowra,  Grenfell,  Coota- 
mundra  and  Albiuy,  although  it  apparently  continues  down  the 
Murray  at  least  to  Mulwala,  having  been  recorded  from  that 
locality  by  Mr.  Baker  (Proc.  Linn.  Soc.  N.S.W.  xxv.,  667).  It 
is  common  on  the  Murrumbidgee  around  Gundagai.  It  may  also 
be  seen  in  Victoria  from  the  railway  line  at  various  points 
between  Albury  and  Seymour. 

Baron  von  Mueller  always  placed  this  tree  under  his  E. 
Stuartiana,  but  it  differs  from  the  tree  growing  at  Ringwood  and 
other  places  near  Melbourne,  also  included  by  the  Baron  under 
E.  Stv^artiana,  in  timber,  bark  and  foliage,  the  Melbourne  tree 
showing  strong  aflinities  to  the  Argyle  Apple  of  New  South  Wales, 
E.  pulveridenta,  Sims,  to  which  E.  Brldgesiana  shows  much  less, 
excepting  in  the  sucker-foliage.  Attention  was  first  drawn  to  this 
difference  by  Mr.  A.  W.  Howitt,  F.G.S.,  in  1898,  before  a  meeting 
of  the  Australasian  Association  for  the  Advancement  of  Science, 
w^iich  resulted  in  the  New  South  Wales  tree  being  described  by 
Mr.  Baker  under  the  name  of  E.  Bridgesiana. 

Messrs.  Deane  k  Maiden  in  their  "  Observations  on  the 
Eucalypts  of  New  South  Wales  "  have  retained  the  name  of  E. 
Stuartiana  for  the  N.  S.  Wales  tree,  and  Mr.  Maiden  has  since 
included  the  Victorian  tree  under  the  lanceolar-leaved  form  of  E. 
pulveridenta  (Proc.  Linn.  Soc.  N.S.W.,  xxvi.,  547). 

Between  Cootamundra  and  Junee  the  following  trees  and 
shrubs  were  noticed  from  the  train  : — Callitris  calcarata,  C. 
robusta,  Sterculia  diversi/oUa,  Cassinia  Theodori,  Casiiarina 
quadrivalvis,  C.  Luehmanni,  and  Bicrsaria  spinosa. 

The  Acacias  noticed  were  : — A.  doratoxylon,  A.  diffusa,  A. 
montana,  A.  difformis,  and  A.  Bailey  ana. 

The  Eucalypts  passed  were  : — E.  melliodora,  E.  Bridgesiana, 
E.   tereticornis,   E.  macrorhyncJia,   E.   heiniphloia  var.  albens,   E. 


BY    R.    H.    CAMBAGE.  203 

sideroxylon^  E.  afflnis,  E.  Woollsiana,  E.  Camhagei  (?),  and  E.  tere- 
ticornis  var.  dealhata. 

E.  sideroxylon,  E.  ajfinis,  E.  macrorhyncha,  and  trees  which 
appeared  to  be  E.  Camhagei  were  on  a  ridge  between  Frampton 
and  Bethun^ra. 

Between  Junee  and  Narrandera  the  following  trees  and  shrubs 
were  seen  from  the  train  : — Sterculia  divei^sifolia,  Cassinia  Theo- 
do7'i,  Exocarpus  cuiyressiformis,  Casuarina  guadrivalvis,  C.  Lueh- 
manni,  C.  C unninghamiana  (only  seen  at  Narrandera  on  the  banks 
of  the  Miirrumbidgee),  Bui^saria  spinosa,  Gallitris  rohusta,  Dodo- 
iicea  sp.,  Heterodendron  olece folium  (trees  up  to  18  inches  in 
diameter),  Myoporum  deserti,  Hakea  leit^Goptera,  Pimelea  linifolia 
and  Calythrix  tetragona,  the  last  two  being  collected  at  Narran- 
dera. 

The  Acacias  passed  were  : — A.  homalopliylla,  A.  difformis,  A. 
decora,  A.  peiidida,  A.  montana  (?)  and  A.  doratoxylon,  the  latter 
seen  only  at  Narrandera. 

A.  pendida  was  only  noticed  in  two  localities,  near  Grong  Grong^ 
where  it  is  locally  known  as  Boree.  In  going  from  Sydney  on 
the  Southern  line,  this  is  the  first  place  at  which  this  species  is 
met  with,  while  on  the  western  line  it  is  first  seen  from  the  train 
near  Narromine. 

The  Eucalypts  identified  between  Junee  and  Narrandera  were  : 
—  E.  meUiodora,  E.  Woollsiana  (both  continuing  all  the  way),  E. 
tereticornis  and  v^ar.  dealhata,  E.  hemiphloia  var.  alhens  (not 
noticed  past  Coolamon)  and  E.  rostrata. 

The  complete  list  of  Eucalypts  passed  between  Marsden  and 
Narrandera  is : — E.  rostrata,  E.  Woollsiana,  E.  sideroxylon,  E. 
tereticornis  and  var.  dealhata,  E.  conica  (only  in  the  northern 
part),  E.  populifolia  (not  plentiful),  E.  dumosa,  E.  melliodora,  E. 
Behriana,  E.  polyhractea,  E.  viridis,  E.  afflnis,  E.  macrorhyncha, 
E.  hemiphloia  var.  alhens,  E.  Camhagei,  E.  Bridgesiana  and  a  few 
trees  of  Ironbark-Box  not  definitely  identified. 

The  Acacias  were  numerous  and  included  A.  stenophylla,  A. 
pendida,  A.  homalophylla,  A.  dealhata  and  the  green  variety,  A. 
hakeoides,  A.  doratoxylon,  A.  cardiophylla,  A.  Osivaldi,  A.  rigens^ 


204      BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES. 

A,  ixiophylla  (1),  A,  diffusa,  A.  difforinis,  A.  decora,  A.  moiitana, 
A.  microcarjm,  A.  aspera,  A.  conferta  A.  acinacea,  A.  verniciffua, 
A.  cidtriformis,  A.  obliqiia,  A.  rhigiophylla,  A.  ffexifolia,  A.  armata 
and  A.  BaiJeyana. 

The  Casuarinas  noted  were  : — C.  Litehmanni^   C.  pahidosa  (1) 
(scarce),  C.  quadrivalvis,  C.  Cunning hamiana  and  C.  Camhagei. 


EXPLANATION  OF  PLATES  VIIL-X. 

Plate  viii. 
Casnarina  Camhagei,  Baker  (Belah),  Gilgandra,  N.S.W. 

Plate  ix. 
Geijera  2iarvifIora,'Lmd\.,  (Wilga),  Gilgandra,  N.S.W. 

Plate  X. 

Fig.  1.— Eucalyptus    Woollsiana,  Baker   (Black  or  Narrow-leaved   Box), 

Forbes,  N.S.W. 
Fig.  2. — Eucalyptus  conica,  Deane  &  Maiden  (Apple  Box),  Forbes,  N.S.W. 


205 


NOTES    AND    EXHIBITS. 


Ml-.  D.  G.  Stead  exhibited  two  specimens  of  the  rather 
uiicoinmon  "Doctor''-  or  ^^Snrgeon^^-Fifih,  Prionurus  microle2ndotus, 
Lacep,,  (so-called  on  account  of  the  keeled,  sharp,  bony  jDlates 
running  along  each  side  of  the  tail,  which  are  used  for  purposes 
of  offence  and  defence).  These  were  both  procured  from  Port 
Jackson  by  means  of  the  trammel  net.  This  means  of  capture  is 
worth}"  of  note,  as  there  is  only  one  man  at  present  using  this 
kind  of  net  in  Port  Jackson.  It  will  be  of  interest  to  record 
that  the  contents  of  the  intestinal  tracts  of  both  fishes  consisted 
of  the  remains  of  two  or  three  species  of  Fucus-like  seaweeds, 
for  the  plucking  of  which  the  curiously  denticulated  teeth  are 
splendidly  adapted.  He  also  showed  a  specimen  of  the  "  Flute- 
mouth  "  (Fistularia  serrata,  Cuv.),  also  from  Port  Jackson. 
Although  the  mouth  of  this  curious  fish  is  only  J  inch  in  length 
as  compared  with  a  snout  SJ  inches  long,  the  stomach  contained 
a  small  fish  2  inches  long,  which  could  not  be  determined,  as  all 
the  external  portion  had  been  digested. 

Mr.  Froggatt  exhibited  specimens  and  photographs  of  Aus- 
tralian fleas,  in  illustration  of  the  well  known  fact  that  members 
of  the  Siphonaptera  often  li\e  upon  animals  of  different  species, 
and  of  diverse  habits.  The  largest  known  flea,  Hystrichopsylla 
talpcSy  described  from  the  mole,  is  common  upon  the  field-mouse, 
and  in  the  nest  of  humble  bees  whither  field-mice  go  in  order  to 
feed  upon  the  bees'  nests.  One  of  the  most  peculiar  Australian 
species,  Echidnojohaga  ambulans,  was  described  from  an  Echidna 
by  Mr.  Olliff  (Proceedings  of  this  Society,  1886).  The  specimens 
of  this  species  now  exhibited  were  recently  received  from  Mr. 
Steel,  and  were  caught  upon  a  native  cat  {Dasyurus).  Mr. 
Turner,  of  Mackay,  had  lately  sent  him  the  exhibited  photographs 
of  a  flea  taken  upon  a  bandicoot  [Perameles),  which  agreed  with 
Mr.  Skuse's  description  of  Ste2:)hanocircus  dasyuri,  taken  upon  a 
native  cat. 


206  NOTES    AND    EXHIBITS. 

Mr.  H.  L.  Kesteven  exhibited  specimens,  and  contributed  the 
subjoined  list,  of  sixteen  species  of  Mollusca  of  the  Family 
Rissoidce  from  Sandgate,  Moreton  Bay,  Queensland,  none  of 
which  had  previously  been  recorded  from  that  State  : — 

Rissoa  scrohiculata^  Watson. 

,,       dissimilis,  Watson. 

,,      flammea,  Frauenfeld. 

,,       salehrosa,  Frauenfeld. 

,,        incidata,  Frauenfeld. 

,,       olivacea,  Frauenfeld. 

,,        contahiilata^  Frauenfeld. 

,,        australice^  Frauenf.,  var.  ochroleuca,  Brazier. 

5,       atropurpurea^  Frauenfeld. 

,,       nitens^  Frauenfeld. 

,,       fraiienfeldi,  Dunker. 

,,       jacksoni,  Brazier. 

,5       devecta,  Tate. 

,,        cheilostoma,  Ten. -Woods. 
Rissoiiia  lianleyi,  Schwartz. 
,,         fasciata,  A.  Adams. 
All  the  foregoing  species  occur  in  New  South  Wales  and,  with 
the  exception  of  the  first  three,  in  South  Australia  likewise,  so 
that  the  extension  of  range  so  far  north  is  therefore  interesting. 
The  BissoidcE  enumerated  by  Tate  from  Queensland  comprised 
three  species  of  Rissoa  and  three  of  Bissoina.     The  specimens 
exhibited  were,  it  was  said,  to  be  presented  to  the  Australian 
Museum. 

Mr.  R.  H.  Cambage  exhibited  specimens  of  EiicalypUhs  eximia 
from  Picton  Lakes,  Hawkesbury  River,  and  Maitland  (Sawyer's 
Gully,  where  it  is  known  as  Rock  Apple);  and  he  pointed  out 
that  the  species  did  not  appear  to  have  flowered  last  spring  at 
any  of  these  places,  though  since  January  last  the  trees  had  been 
covered  with  buds,  showing  they  will  flower  profusely  next  spring. 
It  would  seem,  therefore,  that  the  drought  was  not  responsible  for 
the  failure  of  last  spring. 


NOTES    AND    EXHIBITS.  207 

Mr.  Trebeck  showed  the  foliage  of  a  seedling  peach  much 
spotted  with  Uromyces  amygdali,  Cooke. 

Mr.  Cheel  exhibited  growing  seedlings  of  Juncus  2:>rismatocar- 
pus,  R.Br.,  and  J.  hohschcemis,  R.Br.,  in  illustration  of  a  paper 
showing  that  Mr.  Bentham  was  in  error  in  uniting  these  two 
species,  to  be  read  at  next  Meeting. 

Mr.  R.  Greig  Smith  exhibited  cultures  of  the  Ascobacterium 
from  the  sugar-cane,  described  in  his  paper.  Also  a  culture  of 
Penicillium  glaucum  obtained  from  the  fresh  kino  of  Eucalyptus 
piperita,  which  was  remarkable  for  the  development  of  a  crimson 
colour  in  the  deeper  portions  of  the  medium,  the  colour  being 
probably  due  to  catalase,  the  oxidising  enzyme  secreted  by  the 
Penicillium,  acting  on  fragments  of  kino. 

Mr.  A.  J.  North  exhibited  skins  and  eggs  of  Sisura  nana,  and 
Rhijndura  dryas  from  the  Northern  Territory  of  South  Australia. 
An  egg  of  Sisura  nana,  taken  near  the  Daly  River,  in  January, 
1902,  is  oval  in  form,  the  shell  being  close-grained,  and  lustreless. 
It  is  of  a  dull  buffy-white  groundcolour  irregularly  spotted  and 
blotched  with  umber-brown,  and  similar  underlying  markings  of 
greyish-lilac  which  form  an  irregular  band  around  the  larger  end. 
Length  0*71  x  0*5  inch.  An  egg  of  Rhipidura  dryas,  taken 
during  January,  in  the  same  locality,  is  a  short  oval  in  form,  the 
shell  being  close-grained  and  slightly  lustrous.  It  is  of  a  dull 
yellowish-white  groundcolour  with  an  indistinct  zone  of  confluent 
spots  and  blotches  of  dark  yellowish-brown,  and  bluish-grey 
around  the  thicker  end.  Length  0*65  x  0*52  inch.  The  specimens 
described  were  from  the  collection  of  Mr.  Charles  French,  Junr. 

He  also  exhibited  a  skin  of  a  Grass  Pinch  with  a  pale  wax- 
yellow  bill  from  Wyndham,  N.W.  Australia,  in  illustration  of  the 
following 

Note  on  some  Northern  and  North-western  Austrcdian  Grass 
Finches. 

Among  a  large  number  of  live  birds  brought  to  Sydney  a  few 
years  ago  by  M.  Octave  Le  Bon,  who  had  trapped  them  in  North- 


208  NOTES    AND    EXHIBITS. 

western  Australia,  my  attention  was  arrested  by  numerous 
examples  of  Long-tailed  Grass  Finches.  The  greater  number, 
captured  at  Derby,  were  of  the  well  known  type  of  Poephila 
acuticauda  with  pale  wax-yellow  bills,  while  those  caught  at 
Wyndham  were  almost  similar  in  colour,  but  Avere  distinguished 
by  having  their  bills  orange-scarlet,  forming  a  marked  contrast, 
especiall}^  when  seen  together  in  the  same  cage.  This  distinction 
was  also  pointed  out  to  me  some  time  ago  by  Mr.  G.  A.  Keartland, 
of  Melbourne,  who  had  both  the  pale  yellow  and  orange-billed 
birds  in  confinement;  and  again  more  recently  by  sending  me  a 
skin  of  one  of  the  latter  that  had  died  the  previous  daj'-  in  his 
aviary.  Mr.  Keartland,  who  had  many  opportunities  of  observing 
Poephila  acuticauda  while  at  Derb}^,  informs  me  that  the  bills  of 
all  the  specimens  he  collected,  and  of  sixteen  birds  he  brought 
back  with  him  alive,  were  all  pale  wax-yellow.  Since  his  return 
others  that  were  caught  at  Wyndham  and  Port  Darwin,  and 
which  he  had  in  confinement,  were  all  distinguished  by  their 
orange-scarlet  bills.  Except  that  the  birds  from  these  localities 
are  a  slightly  darker  shade  of  plumage  than  those  obtained  at 
Derb}",  there  is  nothing  except  the  colour  of  the  bill  to  separate 
them.  Age  or  sex  has  nothing  to  do  with  this  distinction,  for  I 
have  seen  many  hundreds  of  Poephila  acuticauda^  and  have  had 
them  under  observation  from  the  nestling  to  the  adult,  while 
breeding  in  confinement.  The  type  of  this  species  was  obtained 
b}^  Mr.  Bynoe  on  the  north-Avest  coast  of  Australia;  and  the  bill 
was  described  by  the  late  Mr.  Gould  as  yellow.  The  e3'es  and  feet 
of  all  I  have  examined  are  coral-red.  If  the  colour  of  the  bills  of 
the  Wyndham  and  Port  Darwin  birds  is  not  a  specific  character 
it  certainly  constitutes  a  very  distinct  variety  of  P.  acuticauda^ 
which  I  propose  to  distinguish  under  the  name  of  Poephila 
aurantiirostrns.  Vernacularly  it  ma}^  be  known  as  the  Orange- 
billed  Grass  Finch. 


209 


WEDNESDAY,  JUNE  25th,  1902. 

The  Ordiiiaiy  Monthly  Meeting  of  the  Society  was  held  in 
the  Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening,  June  25th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S.,  &c..  President,  in  the  Chair. 

Dr.  F.  A.  GoDiNG,  U.S.  Consulate,  Newcastle,  N.S.W.;  and 
Mr.  A.  J.  Walkom,  Sydney,  were  elected  Ordinary  Members  of 
the  Society. 

A  letter  from  Mr.  J.  R.  Garland,  M.A.,  thanking  the  Members 
of  the  Society  for  their  expression  of  sympathy,  was  read  to  the 
Meeting. 

By  the  wish  of  the  Council,  the  President  explained  to  the 
Meeting  that,  in  consequence  of  the  retrenchment  policy  which 
untoward  circumstances  had  forced  upon  the  neighbouring 
State  of  Queensland,  it  was  to  be  feared  that  Mr.  F.  M.  Bailey's 
"  Queensland  Flora,"  now  in  course  of  publication,  would  be 
brought  to  an  abrupt  termination  in  the  middle  of  the  sixth  or 
concluding  Part.  An  expression  of  the  views  of  scientific  men 
in  other  States  would  perhaps  help  to  justify  the  Queensland 
Government  in  making  some  special  effort  to  utilise  to  the  full 
Mr.  Bailey's  experience  and  unrivalled  knowledge  of  the  flora  of 
Queensland  in  completing  the  important  publication  in  question. 

On  the  motion  of  the  President,  it  was  unanimously 
resolved  :  — 

That  this  Meeting  desires  respectfully  to  give  expression  to  the 
hope  that,  in  the  interests  of  science,  the  Queensland  Government 
may  see  its  way  to  allow^  Mr.  F.  M.  Bailey  to  take  the  steps 
necessary  to  complete  the  "  Queensland  Flora." 

The  Donations  and  Exchanges  for  the  month,  amounting  to  7 
Vols.,  32  Parts  or  Nos.,  3  Reports,  1  Separate,  and  3  Miscellanea, 
received  from  35  Societies,  &c.,  and  1  Individual,  were  laid  upon 
the  table. 

14 


210 


NOTES  ON  JUNCUS  HOLOSCFL^NU%  R.Br.,  AND  J. 
PBISMATOCARPUS,  R.Br.;  AND  ON  CERTAIN 
OTHER  NEW  SOUTH  WALES  PLANTS. 

By  Edwin  Ciieel. 

In  his  '  Prodromus '  (p.  259)  R.  Brown  described  two  species 
of  Juncus  under  the  names  of  J.  holoschcenus  and  J.  prismato- 
carpus,  which  he  distinguished  chiefly  by  the  following  characters  : 

J.  holoschcenus.  J.  pvUimatocarpus. 
Stems  terete,  leaves  nearly  iS^e^^is  compressed,  Zea-uesalter- 
terete,  both  articulate.  Panicle  nate,  upright  [verticalibus]. 
terminal,  with  one  leafy  bract  Panicle  terminal,  spreading, 
at  its  base  articulated  like  the  Flowers  numerous,  in  globular 
leaves.  Flowers  numerous,  in  clusters;  perianth-segments  sub- 
globular  clusters;  perianth-seg-  ulate.  Stamens  3.  Capsules 
ments  acute.  Stamens  6.  Cap-  prismatic,  longer  than  the  peri- 
sules  prismatic,  as  long  as  the  anth-segments.  Seeds  oh^cuvelj 
perianth-segments.  Seeds  striate, 
striate. 

Robert  Brown  had  seen  both  species  under  natural  conditions. 
From  the  study  of  herbarium  material  Bentham  came  to  the  con- 
clusion that  R.  Brown's  distinctions  could  not  be  upheld  (B.Fl. 
vii.,  131);  he  therefore  united  the  species  and  adopted  the  name 
J.  prismatocarpus.  It  appears  to  me,  however,  that  these  plants 
are  quite  distinct;  for  they  may  readily  be  distinguished  in  the 
field  by  their  different  habit,  as  well  as  by  some  very  important 
structural  differences. 

J.  holoschtfinus  is  a  more  dwarf  plant  than  J.  prismatocarpus, 
producing  erect,  nearly  terete  stems  from  a  creeping  perennial 
rhizome.  The  leaves  are  slender  and  semiterete,  of  a  dark  green 
colour  and  quite  hollow  inside  except  for  transverse  partitions  of 


BY    EDWIN    CHEEL.  211 

pith  which  give  them  a  jointed  appearance  readily  seen  in  her- 
barium specimens.  These,  I  presume,  are  the  "  articulations " 
mentioned  by  R.  Brown.  I  have  examined  numbers  of  plants  of 
this  species  throughout  the  Port  Jackson  district  in  the  living 
state,  and  have  never  found  less  than  six  stamens  present.  These 
are  best  seen  on  a  dull  damp  morning  when  the  flowers  may  be 
found  to  be  fully  expanded.  The  seeds  are  of  a  brownish  colour, 
and  prominently  striate,  as  may  be  easily  seen  under  an  ordinary 
pocket  lens. 

./.  prismatocaiyus  is  a  much  taller  plant  than  the  former  species, 
and  usually  grows  in  tufts,  hardly  ever  producing  a  creeping 
rhizome.  The  leaves  are  much  broader  and  more  compressed, 
appearing  almost  flat;  they  are  of  a  yellowish-green  colour  and 
are  not  hollow  inside  like  those  of  J.  holoschcenics,  but  have  three 
longitudinal  partitions  of  pith  extending  from  base  to  apex;  these 
are  again  divided  by  transverse  partitions,  but  are  not  so  pro- 
minent as  are  those  of  J.holoschcenus.  The  stamens  are  only  three 
in  number.  The  seeds  are  much  lighter  in  colour,  appearing 
almost  transparent,  and  are  obscurely  striate. 

The  internal  structure  of  the  leaves  appears  to  me  to  be  a  very 
important  character  to  be  observed  in  the  determination  of  these 
two  species;  and  may  easily  be  detected  by  splitting  open  the 
leaves.  By  kind  permission  of  Mr.  J.  H.  Maiden,  F.L.S.,  Director 
of  the  Sydney  Botanic  Gardens,  I  have  examined  all  the  specimens 
labelled  ./.  prismatocarpus  in  the  National  Herbarium,  which  are 
from  various  parts  of  Australia  and  Tasmania.  Some  of  the  Tas- 
manian  specimens  closely  resemble  in  outward  appearance  the 
plants  of  J.  prismatocarpus  grown  in  dry  situations  in  New 
South  Wales.  I  find  that  plants  of  both  species,  if  grown  in  dry 
situations,  so  closely  resemble  each  other  as  to  appear  almost 
identical,  "  their  natural  habitat  being  in  swampy  ground."  But 
by  applying  the  test  of  splitting  open  the  leaves  it  will  be  found 
that  the  hollow-leaved  plants  (J.  holoschcenus)  have  six  stamens, 
whilst  the  longitudinally  partitioned  ones  have  only  three.  A 
further  test  was  made  by  me  by  sowing  some  carefully  selected 
seeds  of  each  of  these  species  in  separate  pots  which  received  the 


212  JUXCrS  HOLOSCH.EXUS  AND  J.  PRISM ATOCARPCS,  &c., 

same  treatment.  At  a  very  early  stage  a  dififerent  habit  of 
growth  could  readily  be  distinguished,  as  well  as  the  difference  in 
leaf -structure,  which  is  identical  with  that  of  the  respective  adult 
plants. 

The  specimens  in  the  National  Herbarium  examined  and 
separated  by  me  are  from  the  following  localities  : — 

J.    HOLOSCH^XUS. 

New  South  Wales : —Bondi,  near  Sydney  (E.  Betche;  May, 
1883);  Bowral  (A.  H.  S.  Lucas;  January,  1894);  Snow}^  Moun- 
tains (W.  Bauerlen;  1890);  Walcha  District,  New  England  (E. 
Betche;  December,  1898);  Centennial  Park,  Sydney  (E.  Cheel; 
January,  1898). 

Victoria : —Wimmera  and  Oakleigh(C.  Walter;  October,  1900). 

Tasmania: — Cascades  (L.  Rodney;  1898). 

J.    PRISMATOCARPUS. 

New  South  Wales : — (Dr.  Leichhardt);  Kogarah  (J.  H.  Cam- 
field;  No\ember,  1893,  with  note  as  follows  : — "  Tall  variety  with 
flat  leaves  and  with  ^'ery  indistinct  cross-partitions;  apparently 
always  aquatic  ");  Tia  River,  New  England  (E.  Betche;  December, 
1898,  with  note  as  follows  :  — "  Identical  with  an  almost  aquatic 
form  from  Kogarah;  cross-partitions  of  leaves  very  indistinct "); 
Conjola  (W.  Heron;  February,  1899);  Menangle  (Mr.  Harper); 
Centennial  Park,  Sydney  (E.  Cheel;  Januarj^,  1898;  with  note 
drawing  attention  to  the  indistinct  cross-partitions  of  the  leaves, 
and  the  differences  in  the  number  of  stamens  and  in  the  seed  as 
compared  with  the  preceding  species). 

Caryophylle.^. 

Silene  inflata,  Sm.  (Fl.  Brit.  467  =  Cucuhahis  Behen,  L.,  Eng. 
Bot.,  p.  164).  Centennial  Park  (E.  Cheel;  December,  1901).  An 
introduced  weed  not  previously  recorded  except  from  the  Wagga 
district  (these  Proceedings,  2nd  Ser.,  iv.,  p.  1055). 


BY    EDWIN    CHEEL.  213 

LEGUMINOSiE. 

Ornithopus  perpusillus,  Linn. — Centennial  Park,  Sydney  (E. 
Cheel;  November.  1899).  An  introduced  naturalised  weed  not 
previously  recorded  from  New  South  Wales. 

EPACRIDEiE. 

Leucopogoii  exolasius,  F.v.M. — Woronora  River  (E.  Cheel; 
October,  1901).  Previously  recorded  only  from  near  Camden 
(Bentham's  'Fl.  Aust.' ;  and  the  'Handbook  of  the  Flora  of 
N.S.  Wales'). 

Monotoca  ledifolia,  A.  Cunn. — Woronora  River  CE.  Cheel; 
October,  1901).  Previously  recorded  only  from  the  Blue  Moun- 
tains (Benth.  '  Fl.  Aust.';  and  'Handbook  of  the  Flora  of  N.S. 
Wales').  There  are  also  specimens  in  the  National  Herbarium 
from  Woronora  River  (collected  by  Mr.  E.  Betche  in  January, 
1S94,  but  not  recorded). 

OnCHIDEiE. 

Thelymitra  pauciflora,  R.Br. — Woronora  River  (E.  Cheel; 
October,  1901).  Previously  recorded  from  Hunter's  Hill,  Sydney, 
and  Mount  Wilson  (Benth.  '  Fl.  Aust.';  and  '  Handbook  of  the 
Flora  of  N.S.  Wales'). 


214 


4. —ON  EUCALYPTUS  BAUERIANA,  8chauer. 

By  J.  H.  Maiden. 

Ramulis  patentib.  subfastigiatis  teretib. ;  foil,  coriaceis  ovatis  v.  subrhombeo- 
ovatis,  basi  obliqua  in  petiolum  longum  contractis  attenuatisve  tenuiter 
acuminatis,  margine  cartilagineo  subrevolutis  unclulatisq.  subglancescentib. 
impunctatis  reticulatis;  umbellis  5-7  floris  axillarib.  terminalibq.  subpani- 
culatis;  pedunculis  subteretib.  petiolo  triple  breviorib.;  pedicellis  subangu- 
latis  cupulae  turbinatae  continuis  eamq.  subaequantibus  ;  operculo  subconico- 
hemisphaerico  apiculato  cupula  triente  breviore.— Foliorum  lamina  2-2^ 
poll,  longa,  1-2  poll,  lata,  petiolus  pollicaris,  pedunculus  3-4  lin.  raetiens; 
cupula  cum  pedicello  3^  lineam  longa.  — In  Nova  Hollandia  legit  F.  Bauer. 
(Walp.  Repert.  ii.  924;  Suppl.  i.,  1845.) 

The  type  is  in  the  Vienna  herbarium  (Herb.  Musei  Caesarei 
Palatini  Vindobonensis).  It  is  in  plump  bud  and  expanded 
flower.  It  bears  a  label,  in  Schauer's  handwriting,  "  Eucalyptus 
Baueriana,  Schauer,  in  Walp.  Repert.,"  and  also  the  words  "  E. 
Baueriana,  Schauer,"  written  over  a  label  "Eucalyptus  rhombi- 
folia,  Tausch." 

Access  to  the  type  has  enabled  me  to  settle  the  identity  of 
Schauer's  plant.  Collected  by  Ferdinand  Bauer,  it  was  doubtless 
obtained  in  the  Port  Jackson  district;  and  it  is  the  Lignum-vitae 
or  Poplar-leaved  Box  which  was  always  looked  upon  by  Mueller 
as  a  form  of  E.  jyolyantheinos,  Schauer,  and  w^iich,  later  on,  was, 
as  will  be  indicated  presently,  recognised  as  a  distinct  species. 

Bentham  (B.Fl.  iii.  214,  under  E.  polyanlhemos)  says: — "The 
tropical  specimens  to  which,  from  the  character  given,  belongs  E, 
Baueriana^  Schau  ,  in  Walp.  Pvep.  ii.  924,  have  generally  smaller 
flowers  and  fruits  than  the  southern  ones,  but  do  not  otherwise 
diff-er." 

I  am  not  clear  as  to  the  allusion  to  "  tropical  specimens." 
There  is  no  evidence  that  Bauer  obtained  the  specimens,  described 
by  Schauer,  in  the  tropics,  or  was,  in  fact,  in  troj^ical  Australia 


BY    J.    H.    MAIDEN.  215 

at  all;  but  Bentham,  who,  I  believe,  never  saw  the  type  of  E. 
Baueriaiia,  is  probably  referring  to  certain  tropical  specimens  he 
thinks  are  referable  to  U.  Baueriana. 

Mueller  (Eucalytographia,  under  E.  'polyanihema)  says : — 
"  Bentham  unites  with  this  \j)olyantheina\  E.  Baueriana^  Schauer, 
the  diagnosis  of  which  agrees  sufficiently  "  [with  i)olyanthema\.   .   . 

Bauer  most  probably  obtained  his  specimens  from  the 

vicinit}-  of  Sydney,  and  not  from  the  tropical  regions  of  Australia." 
I  understand  that  Mueller  never  saw  a  type  specimen  of  E. 
Baueriana,  Schauer. 

E.  Baueriana,  Miquel  (Ned.  Kruidk.  Arch.  iv.  137),  collected 
by  Charles  Stuart  in  Tasmania  is  E.  Gur\nii,  Hook,  f.,  var. 
acervula. 

Synonyms. 

1.  E.  suhrotunda,  R.Br. — I  do  not  know  whether  Brown 
described  this  species.  Even  if  it  is  a  nomen  nudum,  I  think  it 
right  to  draw  attention  to  it,  for  the  case  of  Eucalyptus  is  a 
special  one.  The  nomenclature  of  the  genus  is  so  complex,  and 
the  same  species  has  been  distributed  amongst  first-class  herbaria 
under  so  many  names,  that  it  seems  desirable  to  draw  attention 
to  some  of  them  as  a  matter  of  practical  convenience. 

2.  E.  rhomhifolia,  Tausch. — A  specimen  in  Herb.  Vindob. 
bears  the  label: — "Eucalyptus  rhombifolia,  Tausch,  Hb.  Bauer; 
Ferd.  Bauer  del.  No.  859."  On  this  label  Schauer  has  endorsed 
the  words  "  E.  Baueriana,  Schauer."  I  cannot  trace  that  E. 
rhomhifolia  is  more  than  a  name. 

3.  E.  ohtusifolia,  Tausch. — A  specimen  in  Herb.  Vindob.  bears 
the  label : — "  Eucalyptus  obtusifolia,  Tausch,  Hb.  Bauer;  Ferd. 
Bauer."  It  has  leaves  rather  narrower  than  those  of  the  preced- 
ing specimen.     It  is  E.  Baueriana,  Schauer. 

4.  E.  polyanthemos,  Schauer. — As  already  indicated,  E,  Baueri- 
ana was  both  by  Bentham  and  Mueller  looked  upon  as  a  form  of 
E.  })olyanthemos.  It  will  be  observed  that  in  Exhibition  litera- 
ture and  other  publications  our  "  Lignum-vitae  "  is  referred  to^. 
'polyanthemos  by  other  authors  as  well. 


216  ON  EUCALYPTUS  BAUERIANA,  SCHAUER, 

5.  E.  conica,  Deane  and  Maiden  (these  Proceedings,  1899,  612). 
— Although  different  enough  at  first  sight,  this  is  indubitably, 
in  my  opinion,  the  western  or  narrow-leaved  form  of  B.  Baueriana^ 
but  it  insensibly  connects  with  the  typical  species  found  on  the 
coast.  The  type-species  has  also  more  glaucous  leaves  than  the 
interior  form.  This  interior  form  might  be  known  under  the 
name  of  var.  conica. 

6.  E.  Fletcheri,  R.  T.  Baker  (these  Proceedings,  1900,  682).— 
This  is  absolutely  typical  for  E.  Baueriana^  Schauer. 

Affinities. 

{a).  With  E.  polyanthemos. — That  it  is  liable  to  be  confused 
with  this  species  is  evident  when  it  is  borne  in  mind  that  eminent 
botanists  have  confounded  them.  Following  are  some  points  in 
which  they  differ  : — 

(1)  The  bark  of  E.  Baueriana  is  fibrous,  "  fuzzy,"  or  woolly; 
that  of  E.  polyanthemos  being  ribbony  rather  than  box-like. 

(2)  The  wood  of  E.  Baueriana  is  pale  brown,  and  that  of  E. 
polyanthemos  red. 

(3)  The  leaves  of  E.  Baueriana  are  thinner,  and  the  rim  of  the 
fruit  likewise  thinner  than  that  of  E.  polyanthemos. 

(b).  With  E.  hemiphloia. — The  fruits  of  this  species  are  sub- 
cylindrical,  not  conical  as  is  the  case  with  E.  Baueriana.  The 
western  form  of  E.  Baueriana  is  more  likely  to  be  confused  with 
the  western  form  of  E.  hemiphloia  (var.  microcarpa,  Maiden), 
than  are  the  coast  forms.  E.  hemiphloia  has  the  true  box-bark. 
The  timber  of  E.  hemiphloia  is  paler  than  that  of  E.  Baueriana. 

(c).  With  E.  largijlorens,  F.v.M. — The  two  species  are  con- 
fused in  some  herbaria.  E.  largijlorens  is  a  western  species,  and 
can  only  be  confused  wdth  the  western  form  of  E.  Baueriana. 
But  their  fruits  will  readily  separate  them;  the  timber  of  E. 
largijlorens  is  red.     Both  species  have  subfibrous  ('"box")  bark. 

Range  of  Typical  form. 

New  South  Wales.— George's  River  (collected  by  Robert  Brown, 
1802-5;  and  named  ])y  him  E.  subrotiuida.     Distributed  from  the 


BY    J.    H.    MAIDEN.  217 

British  Museum  at  least  as  early  as  1876  under  the  number  4734, 
under  the  above  name  and  also  that  of  E.  jiolyanthetnos,  Schauer). 

"  Blue  Box,"  Bankstown  and  Cabramatta  (J.  H.  Maiden  and 
J.  L.  Boorman). 

Liverpool  (H.  Deane). 

Thirlmere  (W.  Cuneo).  Type  of  E.  Fletcheri.  Also  collected 
by  R.  H.  Cambage.  Milton  (R.  H.  Cambage,  who  furnishes  the 
note  :  "  Wood  good,  posts  of  building  good  in  ground  for  upwards 
of  40  years.     Bark  rough,  up  to  the  ultimate  branchlets  "). 

Mr.  J.  S.  Allan,  Inspecting  Forester  of  the  district,  says  that 
it  occurs  from  Milton  to  Eden. 

I  collected  it  about  Pambula.  At  the  saw-mill  there  it  is 
known  as  "  Black  Box,"  because  of  the  dark  foliage  of  the  tree. 
The  timber  is  much  valued  locally,  though  in  this  district  it  is 
rare  to  get  a  log  large  enough  for  milling  purposes.  The  rough 
bark  is  up  to  the  ultimate  branchlets.  It  is  hard  to  grub  out, 
and  it  suckers  badly.  It  seems  to  be  usually  found  in  good  land. 
It  also  goes  by  the  names  of  '*  Round-leaf  Box  "  and  "  Brown 
Box." 

Richmond  (W.  Woolls).  Dr.  Woolls  always  called  it  Lignum- 
vitae  or  Poplar-leaved  Box. 

St.  Mary's,  South  Creek  (R.  T.  Baker).     Type  of  E.  Fletcheri. 

Penrith  (J.  H.  Maiden  and  J.  L.  Boorman). 

Victoria. — This  is  a  tree  known  in  Victoria  as  E.  polyaiithemos, 
being  included,  with  the  Red  Box,  under  that  name  by  Mueller. 

Mr.  A.  W.  Howitt  deserves  the  credit  of  working  out  the  range 
of  E.  Bauer iana  in  Gippsland.  In  that  portion  of  Victoria  it  is 
known  as  "Cabbage  Box."  It  occurs  about  Metung,  Heyfield, 
and  Bairnsdale;  on  the  littoral  strip  about  the  Lakes'  entrance, 
also  on  river  flats  only.  It  does  not  occur  in  South  Gippsland. 
"The  timber  is  brown,  and  much  softer  than  that  of  the  Red 
Box  "  {E.  polyanthemos). 

Var.  CONICA,  var.nov. 

New  South  Wales. -West  of  Wyalong,  "  Apple  Box  "  (R.  H. 
Cambage). 


218  ON  EUCALYPTUS  HAUERIANA,  SCHAUER, 

Grenfell.  Received  under  the  name  of  "  Bimble  Box."  The 
true  Bimble  Box  is  E.  populifoHa. 

Weddin  Forest  'Reserve  (J.  H.  Mcaiden). 

Young  (W.  W.  Froggatt). 

Cowra,  a  "Box"  with  persistent  bark  on  small  branches  (H. 
Deane). 

Banks  of  the  Lachlan,  six  miles  south-east  of  Cowra;  also  two 
miles  north-east  of  Cowra,  a  "Grey  Box"  or  "White  Box"  (R. 
H.  Cambage). 

Murga  (H.  Deane). 

Forbes  district  (H.  Deane;  also  R.  H.  Cambage). 

Parkes  (H.  Deane). 

"  Fuzzy  box,"  Wellington.  "  Formerly  looked  upon  as  E. 
largiflorens!''  (Received  from  Rev.  Dr.  Woolls  with  this  informa- 
tion). 

Dubbo  (R.  H.  Cambage,  also  J.  V.  de  Coque).  It  is  the  "  Grey 
Box,  No.  2  "  of  Mr.  de  Coque's  notes. 

"  Blue  Box,"  Minore  (J.  L.  Boorman). 

Toraingley,  Peak  Hill,  Narromine  and  Dubbo  (J.  H.  Maiden). 

Gulgong  (J.  H.  Maiden  and  J.  L.  Boorman).  In  swampy  or 
low-lying  country,  rarely  on  hills;  sometimes  known  as  "  Bastard 
Box."  With  broader  leaves  than  those  of  the  Dubbo  trees;  the 
sucker-leaves  identical  with  those  of  the  coast  form,  even  if  not 
quite  so  broad.  The  stems  of  the  suckers  are  yellow,  turning  red 
later.  The  tree  has  a  glaucous  cast  of  foliage  like  E.  polyantheinos: 
the  surrounding  trees  of  E.  heniiph/oia,  F.v.M.,  var.  microcarpa, 
Maiden, are  glabrous.  Known  locally  as  "Fuzzy Box"  or  "Bastard 
Box,"  with  rough  bark  up  to  the  branchlets.  The  fuzziness  or 
woolliness  of  the  bark  is  a  useful  diagnostic  character  in  this 
species.  The  timber  is  hard  to  cut,  but  more  chippy  and  short 
grained  (brittle)  than  E.  hemiphloia.  It  is  locally  esteemed  as  a 
durable  timber  and  a  valuable  firewood.  The  bark  and  timber 
appear  to  be  in  no  way  different  from  the  coast  or  typical  form. 

Merriwa  (J.  H.  Maiden  and  J.  L.  Boorman),  with  much 
Loranthus  on  it.  On  the  river  fiats  and  taluses  of  the  ridges — a 
usual  situation. 


BY    J.    H.    MAIDEN.  219 

Mr.  J.  L.  Boorman  has  collected  it  around  Wallaiigarra,  on  the 
New  South  Wales-Queensland  border.  He  states  that  it  is 
abundant  in  the  district,  both  on  the  New  South  Wales  and 
Queensland  sides,  so  that  another  species  is  added  to  the  Queens- 
land flora. 

The  species  thus  has  extensive  range,  and  should  be  looked  for 
north  of  Gulgong  and  Merriwa,  and  east  of  Merriwa  and  Tenter- 
field,  wliile  further  localities  should  be  found  in  Queensland. 


5.   ON  EUCALYPTUS  CALYCOGONA,  Turcz. 

By  J.  H.  Maiden. 

The  description  of  this  species  is  as  follows.  I  am  not  aware 
that  it  has  previously  been  seen  in  Australia,  and  I  am  indebted 
to  Kew  for  a  copy  of  it : — 

"  49.  Eacah/ptus  calycogona  (Dri>m.  5,  n.l84).  E.  glabra;  ramis  teretibus; 
foliis  alternis  lineari-lanceolatis  utrinque  attenuatis  acuminato-mucronatis : 
mucrone  interdum  uncinato,  marginatis  pellucido-punctatis;  umbellis  later- 
alibus  3-6-floris;  pedunculis  angulatis  petiolo  paulo  brevioribus ;  cupulis 
obpyramidatis  tetragonis,  nigro-punctatis  subsessilibus  vel  cum  pedicello 
confluentibus,  pedunculo  longioribus  ;  operculo  conico  laevi,  cupula  plus 
quam  duplo  breviore. 

E.  foecunchr,  Schauer,  cujus  operculum  ignotum,  stirps  nostra  affinis  est, 
sed  folia  minora,  pellucida  et  cupula  angulis  4  acutis  marginata.  Filamenta 
alba.  Capsula  4-locularis,  cupula  duplo  brevior.  Folia  bipollicaria,  2^  lin. 
lata.     (Turcz.,  Bull.  Phys,-Math.  Acad.  P^tersb.  10,  1852,  p.  338.) 

The  type  is,  as  Turczaninow  states,  No.  1S4  of  Drummond's 
5th  collection. 

Range. 

Western  Australia  —The  original  specimens  came  from  "  Swan 
River  to  Cape  Riche,"  Western  Australia.  I  have  examined  co- 
types  from  many  herbaria. 

South  Australia. — I  have  seen  a  specimen  from  Murray  Desert 
(S.A  ),  and  labelled  E.  gracilis  by  Mueller. 

Victoria.—"  The  Mallee  Country  "  (a  very  angled  coarse  form). 
Lake  Albacutya  (also  a  ver}'^  angled  coarse  form.  Both  from  C. 
Walter). 

Swan  Hill  (J.  G.  Luehmann). 

Lake  Hindmarsh  (C.  AValter). 

The  Wimmera  (F.  Reader).      Very  coarse  form. 


BY    J.    H.    MAIDEN.  221 

Kamerooka  "  No.  1  Mallee  "  (A.  W.  Howitt).  "  Tall,  up  to 
15ft.,  bark  smooth."  Broad,  shiny  thick  leaves,  with  angled  buds 
and  fruits.     The  coarsest  form  of  the  species  I  have  seen. 

"  Kamerooka  is  near  Bendigo,  being  on  the  fringe  of  the 
country  where  Mallee  is  found,  not  in  large  tracts,  but  in  patches" 
(A.  W.  H.  in  litt.). 

Synonyms. 

B.  calycogona,  Turcz.,  and  U.  celastroides,  Turcz.,  were  omitted 
byBentham  from  the  "Flora  Australiensis"  by  accident,  together 
with  seventy -five  other  species  of  Myrtacese  described  by 
Turczaninow  (Bull.  Phys.  Math.  Acad.  Sc.  St.  Petersb.  x.  p.  321, 
1852). 

Mueller  (Eucalyptographia,  also  Fragm.  viii.  184)  simply  gives 
F.  calycogona,  Turcz.,  and  E.  celastroides,  Turcz.,  as  synonyms  of 
E.  gracilis,  F.v.M.,  but  makes  no  reference  in  the  text  to  them, 
the  date  of  publication  of  Turczaninow's  species  being  presumably 
unknown  to  him. 

1.   E.  celastroides,  Turcz. 

"50.  EJucalyptus  celastroides  (Drum.  5,  n.34).  E.  glabra;  ramis  tereti- 
bus  superne  subangulatis;  foliis  alternis  lineari-Ianceolatis  utrinque  attenu- 
atis  abrupte  et  breviter  acuminatis  subinaequilateris,  marginatis,  obscure 
trinerviis  venosisque;  umbellis  axillaribus  3-6-floris;  pedunculis  angulatis 
petiolum  subaequantibus,  pedicellos  triple,  cupulam  paulo  superantibus  ; 
cupula  obconica  4-costata,  operculum  depresso-hemisphaericum  muticum 
quadruple  excedente.  Folia  bipollicaria  aut  parum  longiora,  3-8^  lin.  lata, 
punctis  aliis  opacis,  interdum  nigricantibus,  aliis  paucioribus  pellucidis  con- 
spersa,  petiolus  fere  trilinealis.  Filamenta  alba.  Cupula  fructus  parum 
aucta,  prope  orificium  leviter  constricta.  Capsula  inclusa,  vertice  plana  4- 
locularis.  Ad  descriptionem  E.  amygdaliiw,  Labill.  in  multis  accedit,  sed 
nullam  reticulationem  in  foliis  video,  folia  breviora,  operculum  depressum 
nee  subconicum,  forsan  etiam  operculi  forma  prae  caeteris  dignoscitur.  E. 
cneorifolia  et  E.  stricta  floribus  sessilibus  recedunt,  E.  pallens  pedunculis 
compressis  et  foliis  5-pollicaribus,  E.  obtusijlora  calycibus  ecostatis  (Turcz. 
in  Bull.  Phys.  Math.  Acad.  Petersb.  10,  1852,  p.  338). 

The  type  is,  as  Turczaninow  states,  No.  34  of  Drummond's  5th 
collection. 


•222  ON  EUCALYPTUS  CALYCOGOXA,  TURCZ., 

This  is   a  glaucous   form   with   fruits    slightly  urceolate,  and 

slightly  rimmed.     I   think   it  is   a  well   marked   variety  of   E. 

calycogonch^  Turcz.,  and  therefore  propose  the  name  of  var.  celas- 

tr aides  for  it. 

Range. 

It  appears  to  be  confined  to  Western  Australia.  Following 
are  some  specimens  in  the  National  Herbarium  of  New  South 
Wales  :— Elder  Exploring  Expedition.  Camp.  63,  W.  A.,  27.9.91, 
and  40  miles  N.W.  of  Eraser's  Range  (R.  Helms;  4/xi/91).  These 
specimens  were  labelled  E.  fmcunda  by  Prof.  Tate.  They  have 
leaves  rather  broader  than  the  Coolgardie  specimens. 

''Goldfields"  (Conservator  of  Forests,  Perth). 

Coolgardie  (W.  L.  Webster). 

2.  E.  gracilis,  F.v.M. :  Fruticose;  leaves  coriaceous,  alternate,  shining, 
narrow-lanceolate,  hooked-acuminate,  a  little  oblique,  thinly  veined,  dotted; 
umbels  axillary  and  terminal  pedunculate:  flowers  small,  short-stalked;  lid 
blunt,  depressed-hemispherical;  tube  of  the  calyx  obconical,  bell-shaped,  a 
little  broader  and  three  times  longer  than  the  lid;  fruit  nearly  hemispherical; 
not  contracted  at  the  top;  valves  of  the  capsule  almost  enclosed. 

In  the  desert  on  the  Murray  River,  where  it  forms  the  Mallee  Scrub 
together  with  E.  dumosa,  santalij'olia  and  other  species.  (Trans.  Vict.  Inst, 
i.  35,  1855.) 

Miquel's  description  is  in  the  following  words.: — 

3.  E.  gracilis,  Ferd.  Milll.  E,  perforata  Behr.  Herb,  partim  :  arbuscula 
gracilis,  ramulis  teretibus  apice  angulatis  lanceolato  linearibus  vulgo  subfal- 
catis  in  acumen  vel  apiculum  uncinatum  excurrentibus  glabris  coriaceis 
crebro  pellucido-punctatis,  umbellis  axillaribus  et  lateralibus  3-6  floris,  calycis 
tubo  turbinato  operculum  depresso-hemisphaericum  apiculatum  triplo  ex- 
cedente. 

Ab  E.  amygdalina  proxime  afiine  differt  foliis  non  venosis,  ab  E.  ambigaa 
operculo  vix  apiculato,  petiolis  longioribus,  umbellis  plerumque  5-fioris  ab 
E.  cneorifolia  floribus  breviter  pedicellatis  (Miiller). 

Frutex  vel  saepe  arbuscula  gracilis  5-8  pedum  altitudinis,  partem  magnam 
fruticetorum  extensorum  aliquot  miliaria  a  fl.  Murray  remotorum  sistens, 
aestate  florens,  ramuli  junioresrubri  (F.  Miiller  Herb,  et  observ.  manuscript.); 
Stuart  Herb.  Tasman.  n.  3).  Folia  2  poll,  longa,  H  lin.  lata.  Calycis 
tubus  pallidis  ^-2  lin.  lata  (Ned.  Kruidk.  Arch,  iv,  1856). 

C.  Stuart's  Tasmanian  specimens  No.  3  are  E.  amygdalina^ 
Labill.     T  have  seen  them.      Micjuel's  statement  that  E.  gracilis, 


BY    J.    H.    MAIDEN.  223 

F.V.M.,  is  near  to  E.  arnygdaluia,  Labill.,  applies  with  a  good 
deal  of  force  to  Tasmanian  specimens,  the  superficial  resemblances 
of  herbarium  specimens  being  frequently  very  considerable. 

The  type  of  E.  gracilis,  F.v.M.,  is  a  South  Australian  specimen, 
and  was  collected  by  Dr.  H.  Behr.  It  was  labelled  by  Mueller  in 
Herb.  Melb.  as  "Eucalyptus  gracilis,  Ferd.  Muell.  Murray 
Scrub.     Dec.  1848.     Behr,  Nov.  Holl.  Austr.,  Dr.  Ferd.  Miiller." 

It  will  be  observed  that  neither  in  Mueller's  nor  Miquel's 
description  is  there  any  mention  of  an  angular  calj^x  which  is  so 
obvious  a  feature  of  the  typical  E.  calycogona,  Turcz. 

It,  however,  passes  by  insensible  gradations  into  the  angular 
form. 

3,  E.  gracilis,  F.v.M.,  var.  hreciflora,  Benth.  Calyx-tube  scarcely  angled, 
1^  to  nearly  2  lines  long.  Fruit  about  2  lines  only,  but  the  deeply  sunk 
capsule  and  the  stamens  entirely  as  in  the  ordinary  form — Darling  and 
Murray  Desert,  also  F.  Mueller's  Spencer's  Gulf  specimens,  which  being  in 
fruit  only  are  somewhat  doubtful  (B.Fl.  iii.  211). 

Bentham's  specimens  cannot  be  traced  in  Herb.  Melb.,  but  I 
am  indebted  to  Kew  for  a  fine  drawing  of  the  original  specimens, 
and  for  fragments  of  the  specimen  which  place  its  identity  beyond 
doubt. 

I  am  of  opinion  that  E.  gracilis,  F.v.M.,  and  E.  gracilis,  F.v.M., 
var.  hreviflora,  Benth.,  are  so  closely  allied  that  it  is  impossible 
to  separate  them  even  as  two  varieties.  I  think  that  they  should 
form  one  variety  readily  noted  by  its  hemispherical  operculum 
and  almost  entire  absence  of  angularity  in  calyx  or  operculum. 
I  propose  the  name  gracilis  for  this  variety,  i.e.,  E.  calycogona, 
Turcz.,  var.  gracilis. 

Range. 

Western  Australia. -Coolgardie  (Nos.  100  and  101;  1899;  K. 
Helms).  Sap-green  leaves,  very  shiny;  fruits  small,  pear-shaped, 
constricted  at  the  mouth,  but  not  ripe ;  operculum  a  little 
pointed. 

Specimens  from  South  Australia  [no  locality]  (W.  Gill,  1896 
and  1900)  are  very  close  to  this  form. 


224  ON  EUCALYPTUS  CALYCOGOXA,  TURCZ. 

Fifty  miles  west  of  Golden  Valley,  W.  A.  (E.  Merrall;  1888,  in 
Herb.  Melb.). 

Both  these  forms  show  transit  to  E.  odorata. 

South  Australia.— York  Peninsula  (J.  G.  O.  Tepper,  1880  ; 
No.  938).  "Middle-sized  trees  10-20  ft.  x  3-8  in.  Coast  plain." 
Herb.  Melb.,  labelled  E.  gracilis  by  Mueller. 

"  White  Mallee,"  Flinders'  Range,  Foot-hills  of  Mt.  Brown 
(Port  Augusta:  W.  Gill,  Conservator  of  Forests,  with  the  note, 
"as  figured  in  Brown's  'Forest  Flora  of  South  Australia'"). 

Ninety  Mile  Desert  (R.  H.  Cambage  ;  March,  1901).  The 
Ninet}^  Mile  Desert  is  the  modern  name  for  the  Murray  Desert 
where  the  type  was  collected. 

Victoria— Mildura  (A.  W.  Howitt's  No.  130);  Wimmera  (C. 
Walter);  8wan  Hill  (Dr.  Griffiths). 

New  South  Wales— Gol  Gol,  near  Went  worth  ("No.  3  Mallee," 
A.  W.  Howitt).  With  slender  rather  tapering  fruits;  the  leaves 
and  buds  precisely  those  of  Tepper's  938  (South  Australia). 

Mt.  Hope  Road  to  Euabalong,  Condobolin  district  (August, 
1899;  R.  H.  Cambage).  The  Mallee  referred  to  by  Mr.  Cambage 
in  these  Proceedings  (1901,  p.  209). 


225 


LI  ©R  A  R^ 


ON    EUCALYPTUS  MELANOPHLOIA,  F.v.M.,   AND    ITS 
COGNATE  SPECIES. 

By  R.  T.  Baker,  F.L.S.,  Curator  and  Economic  Botanist, 
Technological  Museum,  Sydney. 

(Plate  xi.) 

This  species  was  originally  described  by  Mueller  in  1858  (Journ. 
Linn.  Soc.  iii.  93);  and  Bentham  in  the  '  Flora  Australiensis  ' 
(iii.  220)  reproduced  this  description. 

In  the  above  and  all  later  references  to  this  tree  in  scientific 
literature,  it  is  always  described  or  mentioned  as  having  uniformly 
cordate,  sessile  leaves  throughout  its  life  ;  and  such  was  my 
experience  of  it  until  the  research  on  Eucalypts  recently  com- 
pleted at  the  Technological  Museum  proved  that  such  was  not 
the  case. 

It  appears  now  that  the  original  description  applies  only  to  one 
form  of  this  tree's  life-history,  i.e  ,  the  opposite,  sessile,  cordate 
stage;  and  this  being  the  only  described  form  up  till  now  it  can 
be  easily  understood  how,  as  regards  its  foliage,  it  was  taken  to 
belong  to  that  class  of  Eucalypts  having  similar  leaves.  This 
includes  such  Eucal3^pts  as  E.  cordata,  Labill.,  and  E.  indveru- 
lentci,  Sims,  for  Eastern  Australia ;  and,  as  far  as  at  present 
known,  no  lanceolate  leaves  have  ever  been  found  to  occur  on 
them. 

It  can  now  be  shown  that  E.  melanophloia,  F.v.M.,  on  a 
morphological  classification  of  foliage,  must  be  grouped  with  E. 
dives,  Schau.,  E.  Risdoni,  Hook,  f.,  E.  cinerea,  F.v.M.,  and  others, 
all  of  which  in  the  sessile,  cordate-leaved  stage,  bear  both  buds, 
flowers  and  fruits;  and  it  is  a  coincidence  that  all  these,  as  well 
as  this  species,  should  have  been  described  originally  from  this 
particular  leaf-stage.  It  is  now  also  known  that  all  and  each  of 
15 


226  ox  EUCALYPTUS  MELANOPHLOIA,  F.v  M., 

these  develop  lanceolate,  alternate  leaves  as  the  trees  attain   a 
maximum  height. 

As  stated  above,  E.  cor  data,  LabilL,  and  E.  puh-erulenta,  Sims, 
are  the  only  two  Eucalypts  in  Eastern  Australia  and  Tasmania 
which  preserve  the  sessile,  cordate  form  or  shape  of  leaf  through- 
out their  life-history.  The  other  species  also  recorded  as  posses- 
sing this  character  are  : — E.  setosa,  Schau.,  Gulf  of  Carpentaria; 
E.  gamophylla,  F.v.M.,  Northern  Territory,  S.A.;  E.  macrocarpa, 
Hook,  f.,  West  Australia ;  and  E.  pricinosa,  Schau.,  Gulf  of 
Carpentaria.  I  know  nothing  of  the  life-history  of  these  species; 
but  possibly  future  research  may  also  show  that  some  at  least 
develop  lanceolate  leaves  in  their  mature  stages  of  growth. 

The  lanceolate  form  of  leaf  of  E.  melanophloia,  F.^■.M.,  first 
came  under  my  notice  in  January,  1900,  in  the  shape  of  material 
collected  at  Coolabah,  and  also  between  Girilambone  and  Cobar, 
by  Mr.  Bauerlen,  the  Museum  Collector,  and  forwarded  under  the 
name  of  "  Ironbark."  At  that  time  it  was  not  even  suspected  of 
being  E.  melanophloia;  and  as  the  specimens  dia  not  agree  with 
any  species  known  to  me,  its  naming  and  investigation  were  held 
over. 

Later,  having  occasion  to  collect  material  of  E.  melanophloia  at 
Narrabri,  the  "Silver-leaf  Ironbark"  of  that  district  and  a  species 
well  known  there  bj''  its  cordate,  sessile,  opposite  leaves,  it  was 
found  that  E.  microtheca,  F.v.M.,  occurred  there  also,  and  was 
known  vernacularly  as  "  Swamp  Box."  Amongst  some  of  the 
material  of  this  latter  species  forwarded  for  oil-investigation  were 
found  leaves  identical  with  the  "  Ironbark  "  of  the  West  above 
mentioned.  The  ver}'-  greatest  care  had  then  to  be  exercised  that 
none  but  true  leaves  of  E.  microtheca  were  distilled,  and  several 
distillations  were  undertaken  in  order  to  verify  the  results. 

The  lanceolate  form  of  leaves  corresponding  to  the  Western 
"Ironbark"  were  next  traced  to  their  botanical  source,  which  turned 
out  to  be  none  other  than  E.  melanophloia,  and  which  species 
Avas  found  to  have  a  complete  gradation  of  leaves  from  the 
narrow-lanceolate  to  sessile-cordate,  and  also  that  the  far  Western 
"  Ironbark  "  is  the  same  species,  only  differing  in  having  a  lanceo- 


BY    R.    T.    BAKER.  227 

late  form  of  leaf  throughout  its  life.  At  Narrabri  the  trees  of  E. 
mclanophloia  having  all  lanceolate  leaves,  are  not  easily  distin- 
guished from  those  of  E.  microtheca,  as  the  barks  are  also  identical; 
in  herbarium  material,  however,  the  leaves  of  the  latter  species 
can  easily  be  separated  from  the  former,  as  they  always  dry  a 
light  slate  colour  in  contradistinction  to  the  brownish  colour  of 
those  of  the  "  Ironbark." 

The  fruits  of  each  are  characteristically  distinct,  as  mentioned 
by  all  writers  on  the  species.  Although  E.  melanojMoia  is  now 
shown  to  possess  such  a  variation  in  foliage,  no  such  variability 
is  known  or  recorded  of  E.  microtheca,  which  is  probably  one  of, 
if  not  the  most  widely  distributed  species  in  Australia,  occurring 
as  it  does^  in  the  northern  interior  half  of  New  South  Wales, 
Western  Queensland,  Northern  Territory  of  South  Australia  into 
the  northern  parts  of  Western  Australia. 

Bentham  {loc.  ciL),  under  E.  melanophloia,  states  "the  species 
is  very  nearly  allied  to  E.  crehra,  and  may  prove  to  be  an 
opposite-leaved  state  of  the  form  described  as  the  '  Mackenzie 
River  Box-tree.'  It  sometimes  resembles  E.  cine7^ea,  F.v.M.,  but 
differs  in  the  bark,  the  stamens,  and  the  fruit." 

Under  E.  crehra,  F.v.M.,  Bentham  also  states  : — "  '  Box-tree  ' 
of  the  Mackenzie  River,  Leichhardt,  also  on  the  Suttor  River, 
Bowman,  described  by  both  as  having  the  bark  persistent  and 
fissured.  The  specimens  are  somewhat  glaucous.  .  .  .  Fruit 
not  seen.  This  is  very  probably  an  alternate-leaved  state  of  E- 
melanophloia,  F.v.M." 

In  my  opinion  I  think  there  can  be  little  doubt  but  that 
Bentham's  surmises  are  correct,  and  that  the  Mackenzie  River 
^'Box"  is  "an  alternate-leaved  state  of  E.  melanophloia,'^  and  is 
identical  with  that  at  Narrabri,  Nyngan,  Dubbo,  &c.  His  refer- 
ence to  E.  cinerea  resembling  the  opposite,  sessile,  cordate-leaved 
state  of  E.  melanophloia  proves  conclusively  that  it  was  the 
"  Argyle  Apple"  he  recognised  as  that  species,  and  not  E.  pulveru- 
lenta,  Sims,  as  has  been  conjectured. 

Although  the  gradation  of  leaf-form  of  E.  melanophloia  is 
towards  E.  microtheca,  yet  there  is  still  wanting  the  connecting 
link  between  them. 


228  ON  EUCALYPTUS  MELAXOPHLOIA,  F.v.M., 

That  the  two  are  closely  allied  there  appears  to  be  little  doubt, 
but  still  the  hiatus  exists,  and  the  connection,  as  far  as  my 
researches  go,  is  not  extant  to-day. 

The  timbers  are  much  alike  in  colour  and  figure,  but  differ  in 
specific  gravity  and  hardness.  That  of  F.  microtheca  is  more 
durable,  harder,  and  not  so  easily  worked  as  that  of  E.  melano- 
phloia. 

The  barks  are  identical  in  colour  and  texture. 

The  chemical  constituents  of  the  oils  of  these  two  species  are 
also  almost  identical,  so  that  the  differences  are  to  be  found  in 
the  shape  of  the  fruits,  in  the  timber,  and  in  the  leaves,  particu- 
larly in  the  dried  state.  In  the  field  E.  microtheca  is  always 
known  vernacularly  as  "  Swamp  Box  "  or  "  Coolabah  ";^  whilst  E. 
melanophloia  appears  to  be  invariably  known  as  "  Ironbark  "  or 
"Silver-leaved  Ironbark,"  although  when  found  growing  along 
with  the  former,  and  with  all  its  leaves  of  the  lanceolate  form,  it 
is  regarded  by  bushmen  as  "  Swamp  Box."  As  it  is  this  latter 
state  that  extends  westward,  it  is  very  possible  that  the  sessile, 
cordate  leaved-form  is  the  parent  tree,  and  a  survivor  of  the  Miocene 
times  when  the  main  coast  range  remained  above  water.  As  the 
western  plains  were  raised  above  the  sea-level  the  lanceolate  state 
probably  developed. 

The  life-history  of  the  foliage  of  E.  melanophloia  is  thus 
identical  with  the  Tasmanian  species  E.  Risdoni,  Hook,  f.,  and 
E.  dives,  Schau.,  of  the  mainland. 

The  description  of  E.  melanophloia  should  now  read  as  follows  : 

A  medium-sized  forest  tree  attaining  a  height  of  30  to  50  feet, 
with  a  very  rough,  hard,  deeply  furrowed,  dark-coloured  bark, 
extending  nearl}^  right  out  to  the  branchlets.  Foliage  glaucous  or 
Vjrownish-green,  variable  in  form.  Some  trees  preserve  the  sessile 
cordate-shaped  form  of  leaf  entirely,  whilst  others  show  a  grada- 
tion into  lanceolate  ones.  The  w^estern  trees  have  entirely 
lanceolate  leaves  even  on  the  "  suckers."  The  former  leaves  are 
glaucous,  whilst  the  latter  are  only  rarely  so,  being  generally  of 
a  pale  yellowish  or  dull  olive-green  or  pale  brownish  colour, 
slightly  coriaceous,  the  margins  sometimes  recurved  ;    venation 


HY    R.    T.    BAKER.  229 

distinct,  the  lateral  veins  oblique,  spreading,  the  marginal  one 
removed  from  the  edge.  Branches  axillary,  or  se.veral  together 
forming  a  terminal  funicle,  terete,  or  flattened,  from  6  to  12  lines 
long,  bearing  from  7  to  8  flowers.  Buds  on  a  short,  almost  filiform 
pedicel.  Calyx  pyriform,  IJ  lines  in  diameter,  slightly  angular 
at  the  base.  Operculum  domed,  shortly  acuminate.  Stamens 
very  short;  anthers  small,  parallel,  opening  by  longitudinal  slits. 
Pistil  thick,  clavate;  ovary  flat-topped.  Fruit  slightly  angular 
at  the  base,  truncate,  globular  or  pyriform,  about  3  lines  in 
diameter,  but  less  in  the  elongated  form  of  the  narrow-leaved 
variety,  constricted  at  the  orifice,  giving  it  a  kind  of  thin  rim 
which  dries  red;  valves  occasionally  exserted. 

flab. — The  range  of  this  species  is  now  extended  \evy  much 
further  west  : — Nyngan,  Girilambone,  Cobar,  Coolabah  (W. 
Bciuerlen);  Dubbo,  Narrabri,  Angledool  (A.  Paddison). 


EXPLANATION  OF  PLATE  XI. 

Eucalyptus  melanophloia,  F.v.M. 

Fig.  L — Specimen  of  the  cordate,  sessile-leaved  state  in  flower. 

Fig.  2. — Specimen  of  the  cordate,  sessile-leaved  state  in  fruit. 

Fig.  3. — Specimen  of  the  cordate-acuminate  leaved  state  in  fruit. 

Fig.  4. — Specimen  of  the  lanceolate-leaved  state  in  fruit. 

Fig.  5. — Specimen  of  the  narrow-lanceolate  leaves. 

Fig.  6- — "  Sucker  "  leaves  of  the  lanceolate  state. 

Fig.  7. —Buds  (enlarged). 

Fig.  8. — Section  of  bud  (enlarged). 

Fiff.  9. — Anther,  back  and  front  views. 


230 


A  GUM  (LEVAN)  BACTERIUM  FROM  A  SACCHARINE 
EXUDATE   OF  EUCALYPTUS  STUART  IAN  A. 

[Bacterium  eucalyjHi,  n.sp.) 

By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

Society. 

(Plate  xii.) 

A  sweet  exudate  from  a  species  of  Eucalyptus  was  upon  exami- 
nation found  to  contain  a  quantity  of  gum  precipitable  by  alcohol, 
and,  as  several  gum-forming  bacteria  had  been  under  investigation 
in  the  Society's  laboratory,  the  specimen  was  tested  to  see  if  the 
gum  could  possibly  have  a  microbic  origin.     Plates  of  saccharose- 
gelatine*  were  infected  in  the  usual  manner,  and  upon  these  there 
developed   the  dome-shaped  colonies   so  characteristic  of  many 
gum  bacteria. 

The  exudate  was  a  pale  straw-coloured  syrup,  very  similar  in 
appearance  and  consistency  to  honey  or  golden  syrup,  and  had 
fragments  of  bark,  Eucalyptus  capsules,  etc.,  scattered  throughout 
the  mass.      When  dissolved  in  water  and  separated  from  woody 
debris,  a  portion  contained  : — 

Non-reducing  but  hydrolysable  sugarf  calcu- 
lated to  saccharose  ...  ...  ...      1*1  grm. 

Reducing  sugars  ...  ...  ...  ...     2-5     ,, 

Crude  gum       ...  ...  ...  ...  ...     0*8     ,, 

*  Saccharose  10,  peptone  0*25,  potassium  chloride  0"5,  sodium  phosphate 
0*2,  gelatine  10,  water  to  100.  Acidity  to  phenolphthalein  10  c.c.  =0'1  c.c. 
tenth  normal  acid. 

t  This  is  probably  raftinose,  the  sugar  of  Eucalyptus  manna.  The  reducing 
sugars  probably  consist  of  a  mixture  of  levulose  and  melibiose. 


BY    K.    GREIG    SMITH.  231 

I  obtained  the  specimen  from  Mr.  J.  H.  Maiden,  Government 
Botanist.  It  had  been  taken  from  the  bark  of  a  Eitcalyptus 
Stuarfiana,  F.v.M.,  by  Mr.  A.  M.  N.  Rose  at  Dalgety,  Southern 
Monaro.  Mr.  Maiden  obtained  for  me  two  more  samples  from 
the  same  tree.  The  second  specimen  consisted  of  the  exudate  in 
situ  adhering  to  the  bark,  and  containing  fragments  of  a  ruby- 
coloured  kino.  The  third  specimen  consisted  of  a  mixture  of  the 
same  exudate  with  Eucalyptus  manna  of  various  colours  ranging 
from  white  to  reddish-brown.  In  portions  of  the  white  manna  I 
found  small  quantities  of  the  same  gum  that  was  obtained  from 
the  first  exudate,  and  after  separating  the  gum  spherical  masses 
of  prismatic  crystals  of  raffinose  were  readily  obtained. 

In  all  three  specimens  the  same  bacterium  was  obtained  in 
practically  pure  culture. 

A  quantity  of  the  gum  was  prepared  by  growing  the  bacterium 
in  saccharose-peptone  fluid,  and  after  a  sufficient  amount  had 
been  formed,  as  indicated  b}^  the  medium  being  very  opalescent, 
the  gum  was  precipitated  with  alcohol,  and  purified  by  repeated 
solution  in  water  and  precipitation  with  alcohol.  When  free 
from  reducing  sugars  the  gum  was  tested  with  the  following- 
results.  Fehling's  solution  was  not  reduced,  and  the  gum  readily 
hydrolysed  with  dilute  acids  producing  a  reducing  sugar  which 
yielded  glucosazone.  Basic  lead  acetate  gave  a  strong  opalescence, 
and  the  solution  passed  through  filter  paper  unaltered.  Ammonia- 
cal  lead  acetate,  barium  hydrate,  strontium  hydrate  and  lime 
water  in  excess,  each  gave  a  white  precipitate.  Lead  acetate, 
tannic  acid,  ferric  chloride,  copper  sulphate,  aluminium  hydrate, 
iodine,  sodium  hj^drate  and  ammoniacal  silver  nitrate  gave  no 
reaction.  The  melting  point  of  the  dry  and  powdered  gum  was 
199°  C.  Mr.  T.  U.  Walton,  B.Sc,  of  the  Colonial  Sugar  Refining 
Co.,  found  the  sugar  to  be  laevorotatory,  and  to  hydrolyse  com- 
pletely to  levulose. 

From  these  results  it  is  evident  that  the  gum  is  levan,  which  I 
first  obtained  on  cultivating  Bac.  /evaniformans  in  saccharose 
media. 


232 


A    GUM    BACTERIUM    FROM    A    KUCALYPT, 


The  bacterium  differs  absolutely  morphologically  from  Bac. 
levaniformans,  the  organism  which  occurs  in  cane-juice,  and  in 
raw  and  refined  sugars;  and  it  is  interesting  that  the  same  gum 
should  be  formed  by  two  widely  differing  species  of  bacteria. 
The  specific  characters  of  the  bacterium,  which  I  have  named 
Bacterium  eucalypti^  are  given  at  the  end  of  this  paper. 

In  testing  the  various  points  connected  with  the  growth  of  the 
organism,  it  was  found  that  growth  occurred  at  22°,  30°,  and  37°  C. 
The  bacterium  did  not  appear  to  have  a  preference  for  either  of 
the  higher  temperatures;  the  growths  appeared  equally  copious, 
although  at  37°  it  was  dry  and  stiff,  while  at  30°  it  was  moist 
and  flowing.  A  faintly  acid  medium  (acidity  =  0-075%  tartaric 
acid)  enables  the  organism  to  grow  better,  and  to  produce  more 
gum  than  neutral  or  slightly  alkaline  media.  Saccharose  and 
raffinose  («.p'.,  Eucalyptus  manna)  are  the  only  carbohydrates 
from  which  the  bacterium  appears  to  form  gum.  No  levan  was 
produced,  and  the  growth  was  always  scanty  when  dextrin, 
starch,  levulose,  dextrose,  lactose  or  maltose  was  substituted  for 
saccharose  in  the  medium. 

The  composition  of  the  fluid  saccharose  culture*  as  regards 
sugars  and  gum  was  tested  in  the  manner  described  in  a  former 
paper t  at  the  end  of  one,  nine  and  nineteen  days.  The  results 
are  calculated  upon  100  parts  of  saccharose  which  the  medium 
contained  in  the  litre,  and  due  allowance  has  been  made  for  the 
evaporation  of  the  culture  fluid  daring  its  incubation  at  30°. 

Production  of  Levan  and  Reducing  Sugars  from  Saccharose. 


Temperature  =  30^  C. 

At  Start. 

1  day. 

1 
9  days.         19  days. 

Saccharose 

Reducing  sugars      

Levan           

100 

95-2 
1-5 

2-8 

11-4                3-8 
54  1              63-2 
33-4              31-8 

*  Saccharose  100,  peptone  10,  potassium  chloride  ,'),  sodium  phosphate  2, 
tap  water  to  1000. 

t  These  Proceedings,  1901,  Pt.  iv.,  o93. 


BY    R.    GREIG    SMITH.  233 

These  results  show  that  the  action  of  this  bacterium  upon 
saccharose  is  precisely  similar  to  that  of  Bac.  levaniformans.  The 
relative  amounts  of  levan  and  reducing  sugars  are  about  the  same, 
and  there  is  also  a  similar  hydrolysis  of  the  levan  by  the  secreted 
acids  in  the  old  culture.  On  the  19th  day  the  culture  medium 
contained  an  acidity  equal  to  0-135%  of  lactic  acid. 

For  the  same  reason  that  levaniformans  was  shown  to  secrete 
invertase,  so  is  it  with  this  bacterium  :  the  amount  of  acids 
secreted  in  both  cases  is  similar,  and  too  small  to  account  for 
the  heavy  inversion,  which  must,  therefore,  be  ascribed  to  the 
action  of  an  enzyme. 

During  the  bacterial  fermentation,  carbon  dioxide  is  evolved. 
This  was  made  manifest  by  connecting  the  cultivation  flask  with 
a  bottle  containing  baryta  water  and  aspirating  air  which  had 
passed  over  soda-lime,  through  the  apparatus.  A  copious  forma- 
tion of  barium  carbonate  occurred. 

The  acids  secreted  by  the  bacterium  were  tested  in  the  manner 
already  described  for  Bac.  levaniformans*  The  chief  acid  formed 
is  lactic;  capric,  formic  and  acetic  acids  were  detected.  The 
presence  of  butyric  acid  could  not  be  definitely  proved,  which 
may  be  accounted  for  by  the  fact  that  a  young  (6  days')  culture 
was  used  for  the  separation,  and  it  is  admitted  in  some  cases  of 
butyric  fermentation  that  the  butyric  acid  is  formed  from  the 
calcium  salt  of  lactic  acid,  which  means  that  it  is  formed  at  a 
later  stage  of  the  fermentation. 

It  would  appear  that  in  the  presence  of  calcium  carbonate  the 
reducing  sugars  are  used  for  the  formation  of  acid.  A  chalk 
culture  which  had  been  incubated  for  12  days  at  30"  contained 
the  following  ccmstituents  per  litre  : — 

Saccharose       ...  ...  ...  ...  ..       12  £frms. 

o 

Mixed  reducing  sugars  ...  ...  ...      18     ,, 

Levan  .,  ...  ...  ...  ...     32     ,, 

The  saccharose  and  levan  are  in  the  proportions  and  amount 
which  were  found  under  ordinary  conditions  of  cultivation  both 


*  These  Proceedings,  1901,  Pt.  iv.,  p.  605  et  seq. 


234  A    GUM    BACTERIUM    FROM    A    EUCALYPT. 

with  this  bacterium  and  with  Bac.  fevaniforvians,  but  instead  of 
the  calculated  55  grms.  of  reducing  sugars  which  should  have 
been  present  there  are  only  18  grms.;  the  difference  (37  grms.) 
has  disappeared,  i.e.,  it  has  been  converted  into  acids. 

Although  levan  can  be  formed  from  saccharose,  it  must  not  be 
forgotten  that  the  gum  found  naturally  in  the  exudate  had  in  all 
probability  been  formed  from  raffinose,  the  sugar  of  Eucalyptus 
manna.  This  is  indicated  by  the  presence  of  manna  in  one  of 
the  samples.  That  levan  could  be  produced  by  the  organism 
from  raffinose  is  to  be  expected  from  the  fact  found  in  the  study 
of  Bac.  levaniformans,  viz.,  that  the  gum  was  formed  chiefly  from 
nascent  levulose,  and  from  the  fact  that  raffinose  under  the 
influence  of  invertase  splits  up  into  levulose  and  melibiose. 

Bacterium  eucalypti,  n.sp. 

Shape,  etc. — An  actively  motile,  short  coJi-\ike  bacterium, 
measuring  generally  in  the  stained  and  imbedded  condition 
0*5  : 1  /x.  It  stains  well  with  violet  and  fuchsin,  but  feebly  with 
blue;  it  is  decolourised  by  the  Gram  method.  The  flagella  are 
long,  and  vary  in  number  from  one  to  nine,  and  are  studded  over 
the  surface  of  the  cell.     No  spores  are  formed. 

Relation  to  temperature,  etc. — The  bacterium  is  aerobic,  and 
appears  to  grow  equally  well  at  28°  and  at  37°. 

Nutrient  gelatine  plate. — Small  punctiform  colonies  are  visible 
in  seven  days,  and  by  the  eleventh  day  they  have  become  rounded, 
translucent  white,  and  3  mm.  in  diameter.  When  magnified  they 
appear  round  or  rounded,  and  finely  granular,  sometimes  with 
central  granules.  The  edge  is  smooth  and  slightly  waved.  The 
deep  colonies  are  irregular  and  finely  granular. 

Glucose-gelatine  plate. — The  colonies  are  glistening,  translucent 
white  and  rounded.  When  magnified  they  appear  rounc^ed  and 
erose  with  co/^-like  striations.  The  deep  colonies  are  rounded  to 
elliptical,  and  have  a  striated  margin. 


BY    R.    GREIG    SMITH.  235 

Wort^'-gelatine. — The  colonies  are  raised  and  like  drops  of  whey. 
When  magnified  they  appear  round  and  uniformly  granular. 
Sometimes  the  margin  is  striated  as  if  from  the  growth  flowing 
down  the  dome-shaped  colony.  The  deep  and  subsurface  colonies 
are  small  and  coarsely  granular. 

Saccharose-gelatine  plate. — The  colonies  are  transparent  and 
hemispherical,  like  exuded  drops  of  glycerine. 

Nutrient  agar  plate. — The  colonies  are  round,  slightly  raised, 
translucent  white  and  moist  glistening.  When  magnified  the}'' 
appear  round  with  a  smooth  edge.  There  are  granules  around  the 
centre,  but  otherwise  the  structure  appears  homogeneous.  The 
deep  colonies  are  rounded  to  elliptical,  and  contain  large  granules. 

Saccharose-2jepto7ie-agar  plate. — The  colonies  are  hemispherical 
and  whitish,  like  drops  of  starch  paste.  When  magnified  they 
appear  rounded,  and  have  a  blistered  surface;  the  margin  is 
apparently  smooth. 

Nutrient  gelatine  stab. — The  stab  is  filiform  and  white;  the 
nail-head  is  round,  flat,  white  and  glistening.  The  medium  is 
slowly  liquefied;  in  14  days  at  22^  the  liquefied  area  is  slightly 
funicular,  and  at  the  top  of  the  stab  the  medium  has  been  con- 
sumed, leaving  an  air-bubble. 

Glucose-gelatine  stab. — As  with  nutrient  gelatine. 

Saccharose-gelatine  stab. — A  filiform  stab  with  an  hemispherical 
drop  of  transparent  fluid  as  a  nail-head.  The  fluid  increases  and 
flows  over  the  surface  of  the  gelatine.  As  this  occurs,  the  stab 
develops  lenticular  and  spherical  pockets  of  almost  transparent 
gum.  A  liquefaction  of  the  medium  below  the  surface  was  noted 
after  a  month. 

Glucose-gelatine  stroke.— The  growth  is  rough  and  narrow,  with 
an  expanded  lower  portion  like  the  nail-head  on  nutrient  gelatine. 
The  stroke  becomes  rough  and  ribbed,  dry  glistening  and  whitish. 

Nutrient  agar  stroke. — A  thin  translucent  white,  almost  trans- 
parent, layer  is  formed.  It  is  always  scanty,  and  the  condensed 
water  has  a  white  sediment  and  no  film. 


The  wort  contained  saccharose. 


^c>V' 


^Cq 


^  ^. 


236  A  GUM  BACTERIUM  FROM  A  EUCALYPT. 

Saccharose-peptone  agar. — The  stroke  becomes  broad,  raised  or 
hemispherical  in  section,  sometimes  undulating,  and  slowly  gravi- 
tates. The  culture  is  of  the  appearance  and  consistency  of  thin 
starch  paste.  The  luxuriance  of  the  growth  is  in  striking  contrast 
to  the  growth  on  nutrient  agar. 

Potato. — The  growth  is  dry  and  glistening,  whitish  or  slightly 
yellow  and  constricted.  The  colour  deej)ens  to  cream  or  pale 
buff,  and  the  grov/th  becomes  raised.  It  is  always  meagre,  and 
does  not  spread  over  the  surface  of  the  medium. 

Bouillon. — The  medium  Ijecomes  turbid,  and  a  fine  white,  loose 
sediment  is  deposited,  while  a  slight  film  forms  on  the  surface. 
A  faint  indol  reaction  is  obtained  In  nitrate-bouillon  the 
nitrate  is  not  reduced. 

Milk. — The  medium  is  unaltered. 

Saccharose-peptone  fluid. — The  medium  becomes  milk-white,  and 
at  a  later  stage  becomes  brownish-yellow.  The  gum  can  be 
readily  precipitated  by  alcohol,  and  Fehling's  solution  is  strongly 
reduced. 

The  nearest  allied  bacterium  capable  of  forming  slime  from 
saccharose  appears  to  be  Bact.  gelatinosum  hetoi,  Glaser,  which 
forms  dextran  and  alcohol,  but  no  lactic  acid.  As  the  gum, 
moreover,  has  only  been  obtained  previously  from  Bac.  levani- 
fovinans  the  bacterium  is  evidently  a  new  species,  and  therefore 
I  have  named  it,  on  account  of  its  origin.  Bacterium  eucalypti. 


EXPLANATION  OF  PLATE  XIL 

Film   of    culture   on   saccharose-peptone-agar,   stained   with    carbol-violet. 

X  1000. 
Flagella   stained  by  the   night-blue  method  ;   bacteria  grown  on  ordinary 

nutrient  agar,     x  1000. 
Dome-shaped  colonies  on  saccharose-gelatine  plate,      x  \, 


I 


237 


NOTES    AND    EXHIBITS. 

As  instances  of  the  severity  of  the  prevailing  drought,  Mr. 
North  exhibited,  with  the  permission  of  the  Curator,  specimens 
of  Chlamydodera  maculata  and  Philemon  citreogvlaris,  which  had 
been  received  in  the  flesh  by  the  Trustees  of  the  Australian 
Museum.  The  former  was  shot  in  an  apple  tree  on  the  19th  May 
in  a  garden  at  Smithfield,  about  twenty  miles  from  Sydney,  b}^  a 
son  of  Mr.  James  Stein,  the  donor.  The  spotted  Bower-bird, 
Chlamydodera  maculata,  is  an  inhabitant  of  the  scrubs  on  the 
inland  plains  in  the  western  and  north-western  portions  of  the 
State,  and  seldom  occurs  east  of  Byrock.  Philemon  citreogularis, 
another  inland  species,  was  procured  on  the  31st  May  by  Mr.  H. 
Newcombe  at  Kurnell,  or,  as  it  is  now  called.  Cook's  Landing 
Place,  on  the  shores  of  Botan}^  Bay.  Previously  neither  of  these 
species  had  been  recorded  from  the  County  of  Cumberland.  On 
the  16th  May  Mr.  North  saw  a  flock  of  Pied  Crow-shrikes 
{Strepera  graculina)  in  the  Botanic  Gardens,  Sydney,  several 
individuals  of  which  have  been  since  trapped,  and  are  now  in  one 
of  the  aviaries.  Although  this  species  frequents  during  the 
autumn  months  the  northern  and  western  suburbs,  he  had  never 
observed  it  in  a  wild  state  in  the  city  before.  What  might  be 
regarded  as  an  irruption  of  Blood-birds  (Myzomela  sanguinolenta) 
has  taken  place  in  the  Sydney  coastal  districts.  In  ordinary 
seasons  during  winter  this  species  occurs  near  the  coast  onl}^  in 
limited  numbers.  At  present  they  are  in  hundreds  at  Middle 
Harbour,  Long  Reef,  Bondi,  Randwick,  Kurnell  and  intermediate 
localities  where  previously  they  had  been  unobserved.  Mr.  H. 
Newcombe,  who  presented  eight  specimens  to  the  Trustees  of  the 
Museum,  supplied  the  information  that  at  Kurnell  on  the  31st 
ult.,  some  boys  had  about  thirty  specimens  that  they  had  killed 
with  sticks  and  stones. 


23S  NOTES    AND    EXHIBITS. 

Mr.  Cheel  exhibited  a  series  of  botanical  specimens  in  illustra- 
tion of  his  paper. 

Mr.  H.  G.  Smith  exhibited  a  fine  sample  of  crude  eudesmol 
from  Eucalytus  camphora,  Baker.  This  sample  of  the  Stearoptene 
of  Eucalyptus  oil  was  separated  commercially  from  9  lbs.  of  the 
crude  oil  by  redistillation.  It  had  crystallised  into  a  solid  hard 
mass,  and,  as  shown,  represented  about  20  per  cent,  of  the  crude 
oil,  distilled  from  trees  growing  at  Kareela,  Paddy's  River,  thus 
bringing  the  species  within  about  90  miles  of  83'dne3\  The 
species  was  named  E.  camphora  b}'^  Mr.  R.  T.  Baker,  F.L.S.,  on 
account  of  the  large  amount  of  eudesmol  occurring  in  the  oil. 
The  commercial  possibilities  of  eudesmol  have  yet  to  be  investi- 
gated. 

Mr.  Baker  exhibited  flowering  and  fruiting  specimens  of 
Eucalyptus  vimincdis,  LabilL,  from  Johannesburg,  Transvaal, 
South  Africa.  In  morphological  characters  the  specimens  show 
no  discernible  differences  from  typical  Australian  examples  ;  so 
that  in  this  instance  environment  had  produced  no  ol^vious 
alteration  of  characters. 

Mr.  Greig  Smith  exhibited  cultures  of  Bact.  eucalypti  described 
in  his  paper,  and  also  a  specimen  of  gum  levan  produced  by  the 
bacterium.  He  also  communicated  the  substance  of  a  report 
received  from  Mr.  T.  U.  Walton,  B.Sc,  of  the  Colonial  Sugar 
Refining  Co.,  giving  the  opinions  of  several  mill  managers  upon 
the  prevalence  of  gummosis  in  crops  of  sugar-cane  grown  on  land 
impregnated  with  salt  through  being  subjected  to  the  influence 
of  tidal  waters.  A  summar}^  of  the  opinions  of  most  of  the 
officers  consulted  is  that  cane  grown  on  salt  lands  is  quite  as 
liable  to  develop  gummosis  as  that  grown  elsewhere;  but  the  main 
reasons  for  the  presence  of  the  disease  are  probably  the  use  of 
gummed  seed  and  bad  drainage,  and  the  latter  is  as  a  rule  found 
in  low  coastal  lands  within  the  influence  of  tides.  One  officer 
instanced  the  case  of  a  farmer  who  grew  Mauritius  Ribbon  cane 
on  salt  land   long  after    this  variety   had    been    abandoned    as 


NOTES    AND    EXHIBITS.  239 

gummed,  but  the  apparent  protective  influence  of  the  salt  only 
extended  to  that  variety;  crops  of  Rappoe  grown  at  the  same 
time  were  badly  gummed. 

Mr.  Maiden  exhibited  (1)  specimens  of  Eucalypts  to  illustrate 
his  papers.  (2)  Fruiting  and  flowering  specimens  of  Cocos 
Yatayi,  Mart.,  cultivated  in  the  Sydney  Botanic  Gardens  under 
the  names  of  Cocos  capitata  C?),  Mart.,  and  Dlplothemium  mariti- 
mum,  Hort.  Syd.,  the  Brazilian  Wine  Palm.  It  has  been  pre- 
viously exhibited  before  this  Society  (it  is  largely  grown  in  New 
South  Wales)  as  a  Diplothemiuiii^  and  recorded  in  the  Agricultu- 
ral Gazette  of  June,  1891,  p.  356,  under  the  .same  name.  The 
correct  identification  of  the  palm  is  due  to  Dr.  Udo  Dammer  of 
Berlin.  (3)  A  water-colour  drawing  of  the  fruits  (receptacles) 
of  Ficus  Bennettii,  Seem.,  a  Fijian  species  described  by  Seemann 
from  specimens  in  the  Sydney  Botanic  Gardens.  Seemann  des- 
cribed the  fruits  as  very  much  less  than  shown  in  the  drawing,  he 
probably  having  only  windfalls.  This  year,  however,  they  have 
attained  a  size  (  3  by  1  j  inches)  which  is  greater  than  previously 
observed,  and  their  colouring,  rich  dark  orange-red,  is  more 
brilliant  than  has  hitherto  been  noted.  Mr.  Maiden  also 
reported  the  occurrence  of  Manna  from  Dalgety,  near  Bombala, 
on  Eucalyptus  Gunnii,  Hook,  f.,  var.  rubicla,  Maiden,  and  on  U. 
Stuartiana,  F.v.M.,  it  occurring  annually  on  the  former  tree  and 
about  every  ten  years  on  the  latter. 


240 


WEDNESDAY,  JULY  30th,  190: 


The  Ordinary  Monthly  Meeting  of  the  Society  was  held  in 
the  Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening,  July  30th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S..  cfec.  President,  in  the  Chair. 

The  President  announced  that  the  Council  had  elected  Mr.  E. 
Meyrick,  B.A.,  F.Z.S.,  Marlborough,  England;  Dr.  R.  Broom, 
B.Sc,  Pearston,  Cape  Colony;  and  Mr.  D.  Mc  Alpine,  Melbourne, 
Vic  ,  to  be  Corresponding  Members  of  the  Society. 

Also  that  the  Members  of  the  Society  would  be  very  glad  to 
hear  that  official  notification  of  the  intention  of  the  Queensland 
Government  to  continue  Mr.  F.  M.  Bailey's  services  until  the 
end  of  the  year  in  order  that  he  may  be  able  to  complete  the 
"Queensland  Flora,"  had  been  received. 

The  Donations  and  Exchanges  for  the  month,  amounting  to  21 
Vols.,  80  Parts  or  Nos,,  4  Bulletins,  6  Scientific  Reports,  2 
Administrative  Reports,  2  Pamphlets,  and  2  Miscellanea,  received 
from  55  Societies,  itc,  were  laid  upon  the  table. 


241 


NOTES  ON  SOME  HITHERTO  UNRECORDED  SPECIES 

OF  PLANTS  INDIGENOUS  IN  THE  STATE  OF 

WESTERN  AUSTRALIA. 

By  W.  V.  Fitzgerald,  F.S.Sc,  Lond.,  F.R.H.S.,  Eng. 
(Communicated  by  J.  H.  Maiden,  F.L.S.) 

C  0  M  P  O  S  I  T  JE. 
MiNURIA    INTEGERRIMA,   Bentll. 

Lawless  (W.  V.  Fitzgerald;  Jul}^,  1899).  Ray  purple  or  bluish. 
This  is  the  first  recorded  locality  for  this  species  in  extratropical 
Western  Australia. 

LOGANIACEiE. 
MiTRASACME     PALUSTRIS,   Sp.nOV. 

A  minute  glabrous,  often  almost  stemless  herb,  with  a  com- 
paratively thick  spongy  white  radix.  Leaves  linear-spathulate, 
connate,  rather  thick  and  succulent,  not  numerous,  and  in  many 
instances  apparently  but  not  actually  radical.  Flowers  pedicel- 
late, numerous,  in  terminal  umbel-like  clusters;  pedicels  filiform, 
ascending,  ebracteate.  Calyx  about  f  line  long  when  in  flower, 
little  longer  in  fruit,  broad,  with  two  short  herbaceous,  deltoid, 
often  divergent  acute  lobes,  not  above  ^  the  length  of  the  tube. 
Corolla  nearly  as  long  as  the  calyx,  white,  campanulate,  glabrous, 
with  short  acute  lobes.  Stamens  inserted  below  the  middle  of 
the  tube ;  anthers  not  exserted,  small.  Styles  distinct  and 
parallel,  with  the  stigmatic  ends  adjacent  but  not  cohering. 
Capsule  much  shorter  than  the  calyx,  ovoid  and  slightly  flattened, 
membranous;  seeds  numerous,  almost  black,  with  a  reticulate 
testa. 

Zoc  — Midland  Junction;  in  mud  (W.  V.  Fitzgerald;  October, 
1901). 

16 


242  PLANTS    FROM    WESTERN    AUSTRALIA, 

Stems  when  produced  comparatively  stout,  and  not  exceeding 
J  inch  in  height.  Leaves  from  J  to  1  inch  long.  Pedicels  from 
J-J  inch  in  length. 

In  floral  characteristics  this  species  closely  approximates  M. 
distylis,  F.v.M.,  but  differs  widely  in  the  stouter  although  dwarfed 
habit,  in  the  peculiar  spongy,  white  taproot,  the  foliage,  and  in 
the  terminal  umbellate  inflorescence. 

M  Y  0  P  0  R  I  N  E  ^. 

Eremophila  virgata,  sp.nov. 

An  erect  compact  shrub  with  virgate  branches,  the  slender 
branchlets  and  leaves  viscid  and  scented.  Leaves  alternate,  erect, 
semicylindrical,  with  acute  hooked  points,  thick,  channelled  above. 
Flowers  white,  axillary,  on  slender  recurved  viscid  pedicels, 
usually  solitary  or  occasionally  2  from  the  same  axil,  ebracteate. 
Calyx  divided  nearl}'  to  the  base,  segments  hardl}'  overlapping, 
ovate,  acute,  thin,  viscid,  ciliate  with  white  viscid  hairs,  veined 
after  flowering  but  hardly  enlarged.  Corolla  broad,  the  base  very 
short,  the  upper  part  campanulate  and  much  longer  than  the 
base;  sprinkled  outside  with  a  few  hairs,  throat  denseh^  hirsute; 
lobes  very  short,  nearly  equal,  upper  ones  reflexed,  lower  spread- 
ing. Stamens  didynamous,  included.  Style  sparingh'  hirsute, 
prominently  hooked  at  the  end ;  stigma  prominent :  ovulary 
slightly  hairy  at  the  tapering  apex. 

Loc. — Paddington;  in  ferruginous  gravelly  or  sandy  soil  (W. 
Y.  Fitzgerald;  September,  1898). 

Height  of  plant  10-12  feet;  leaves  not  exceeding  ^  inch  in 
length;  pedicels  1^-2  lines;  calyx  IJ  lines,  segments  1  line  long; 
corolla  6  lines  with  lobes  not  exceeding  1  line  long.  Ovulary 
2-celled  with  apparently  only  1  pair  of  ovules  at  the  summit  of 
each  cell.     Fruit  not  seen. 

The  species  differs  from  E.  Paideyi,  F.v.M.,  chiefly  in  the 
foliage  and  in  the  not  enlarging  cal3^x-lobes;  and  rightly  belongs 
to  the  section  including  R.  Brown's  Pholidia. 


BY    W.    V.    FITZGERALD.  243 

PROTEACE.E. 

Grevillea  Jamesoniana,  sp.nov. 

An  erect  rigid  shrub;  branclilets  stout,  ferruginous  or  greyish- 
tomentellous;  young  leaves  and  shoots  with  a  closely  appressed 
silky- white  vestiture,  the  older  leaves  glabrous.  Leaves  crowded 
at  the  ends  of  the  branclilets,  entire,  linear-terete,  on  short 
petioles,  erect,  slender  but  rigid,  pungent-pointed,  closely  refracted 
along  the  margins,  doubly  grooved  beneath.  Racemes  very  short, 
almost  umbel-like,  axillary  or  terminal;  rhachis  densely  invested 
with  appressed  white  silky  hairlets.  Flowers  comparativel}'  large, 
not  numerous,  scarlet,  on  conspicuous,  stout  glabrous  pedicels; 
perianth  quite  glabrous,  striate,  tube  broad,  limb  short,  globular, 
closely  revolute.  Pistil  glabrous.  Hypogynous  gland  semiannular, 
truncate.  Torus  very  oblique.  Ovulary  glabrous,  gibbous,  on 
a  prominent  free  stipes;  stigmatic-disk  lateral,  orbicular. 

Log. — Lakeside ;  in  wet  soil  (W.  V.  Fitzgerald  ;  September, 
1898). 

Total  height  of  plant  8-10  feet;  leaves  3-5  inches  long  and 
about  1  line  broad;  rhachis  1-2  lines  long;  pedicels  2-3  lines  long; 
perianth  nearly  or  quite  1  inch,  with  a  limb  2  lines  long;  ovulary 
stipes  \  inch  and  style  f  inch  in  length.  Stigmatic-disk  very 
thick.      Ripe  fruit  not  known. 

This  species  differs  from  G.  acuaria,  F.v.M.,  in  the  greater 
length  of  the  leaves,  and  in  the  perianth  and  other  minor 
characters.  The  foliage  is  not  unlike  that  of  some  forms  of  G 
nematojihyUa,  F.v.M. 

This  species  is  named  in  honour  of  the  Hon.  Dr.  Adam  Jameson, 
M.L.C.,  Minister  for  Lands  for  the  State  of  Western  Australia, 
a  gentleman  who  has  displaj^ed  more  than  ordinary  interest  in 
scientific  matters. 

Cyperace.e. 

SCHOENUS  RODWAYANUS,   sp.nOV. 

Caespitose ;  stems  slender,  slightly  compressed,  prominently 
striate,  more  or  less  scabrous,  simple,  leafless.     Leaves  reduced 


244  PLANTS  FROM  WESTERN  AUSTRALIA, 

to  several  basal,  sheathing,  shining  brown  or  dark  brown  bracts, 
with  slightl}^  membranous  margins;  lamina  abbreviated,  obtuse, 
usually  involute,  with  finely  scabrous  margins;  floral  bract  similar. 
Inflorescence  consisting  of  a  solitary,  terminal,  sessile  spikelet,  or 
from  2-4:  pedicellate  terminal  spikelets  on  filiform  pedicels  of 
unequal  length.  Spikelets  not  very  flat,  lanceolate-acuminate,  9 
lines  long,  containing  2  flow^ers,  the  lowest  maturing  fruit. 
Glumes  rather  blunt,  with  a  prominent  black  keel;  margins  pale, 
membranous,  minutely  woolly-ciliate  at  the  apex;  five  outer  empty 
ones  gradually  shorter.  Stamens  3.  Stigmas  almost  plumose, 
much  shorter  than  the  hirsute  style.  Hypogynous  bristles  6, 
equal,  shorter  than  the  fruit,  ovate,  acute,  not  fringed.  Nut 
ovoid,  smooth,  obtusely  3-angled,  very  blunt,  1  line  long,  pale 
with  black  blotches,  on  a  thick  stipes  of  J  line. 

Loc. — Bayswater,  in  dry  sandy  soil;  Leederville,  in  swampy 
spots  (\V.  V.  Fitzgerald;  November,  1901). 

Stems  1-2  feet  high;  sheath-lamina  not  exceeding  1  line  long; 
longest  pedicel  about  1^  inches  long. 

In  the  Bayswater  form  the  spikelets  are  apparentl}^  all  pedicel- 
late, the  inflorescence  appearing  more  or  less  umbellate.  In  the 
Leederville  plant  the  inflorescence  varies  from  the  above  to 
spikelet  solitary  and  sessile,  and  the  stems  more  compressed. 
There  are  no  other  diff'erences. 

In  habit  and  general  appearance  the  new  plant  bears  a  close 
resemblance  to  S.  pedicellaiiis,  Poiret,  but  diff'ers  in  the  mem- 
branous and  not  bearded  margins  to  the  leaf -sheaths,  the  larger 
size  of  the  spikelets,  the  prominently  keeled  glumes,  only  2 
flowers  in  the  spikelet,  equal  short  hypognous  bristles,  and  in  the 
stipitate  nut. 

In  the  latter  characteristic  it  bears  some  aflinity  to  S.  minu- 
tulus,  F.V.M.,  and  S.  tr  achy  carpus,  F.v.M. 

The  species  is  named  in  honour  of  Mr.  L.  Rod  way,  the  well- 
know^n  investigator  of  the  Tasmanian  flora,  as  a  recognition  of 
much  service  rendered  to  the  writer. 


BY    W.    V.    FITZGERALD.  245 

SCHOENUS    JaMESOXIANUS,  sp.llOV. 

Csespitose;  stems  slender,  prominently  striate,  simple.  Leaves 
numerous  at  the  base  of  the  stem,  involute,  linear,  flexuose,  blunt, 
much  dilated  at  the  base  into  open  brown  sheaths,  hyaline  on  the 
margin.  Sheathing  bracts  with  closed  black  sheaths  and  hyaline 
margins,  lower  one  produced  into  a  leaf-like  lamina.  Spikelets 
usually  2  together,  rarely  solitary,  on  unequal  pedicels  within 
the  sheath,  occasionally  the  longer  pedicel  bears  a  second  pedicel- 
late spikelet;  spikelets  much  compressed,  ovate-lanceolate,  4-5 
lines  long,  containing  from  3-4  flowers,  apparently  all  fertile. 
Glumes  dull  black,  truncate  or  bifid,  with  a  prominent  keel, 
which  in  the  outer  glumes  is  green;  margins  slightly  ciliate  near 
the  apex;  3-4  outer  empty  ones  gradually  shorter.  Stamens  3. 
Stigmas  much  shorter  than  the  style.  Hypogynous  bristles 
none.  Nut  obovoid,  obtusely  3-angled,  J  line  long,  tubercular- 
rugose. 

Loc. — Midland  Junction,  in  wet  spots  (W.  V.  Fitzgerald; 
September,  1900). 

Stems  usually  about  9  inches  high;  leaves  from  4-6  inches  long; 
lamina  of  lower  bract  from  li-2J  inches  long. 

This  species  is  very  close  to  S.  bijidus,  Bceckel.,  from  which  it 
differs  chiefly  in  the  more  numerous  flowers  within  the  spikelet, 
and  in  the  entire  absence  of  h3'pogynous  bristles. 

The  species  is  dedicated  to  the  Hon.  Dr.  Adam  Jameson, 
M.L C,  Minister  for  Lands  for  Western  Australia. 


246 


NEW  SPECIES  OF  QUEENSLAND  LEPIDOPTERA. 

By  Thomas  P.  Lucas,  L.R.C.P.,  Ed.,  L.S.A.,  Lond.,  M.R.C.S., 

Eng. 

Family  COSSID^. 
CULAMA    EX  PRESS  A,  11.  Sp. 

(J9.  25-40  mm.  Head  grey,  freely  mottled  with  black  and 
fuscous,  in  9  a  tuft  on  crown  more  decided  reddish  colour.  Palpi 
short,  porrected,  fuscous,  lighter  on  terminal  joint.  Antennae  in 
(J  pectinate,  pectinations  narrow  towards  apex  and  base,  in  9 
simple.  Thorax  grey,  freely  mottled  with  black  and  fuscous,  in 
9  decidedly  reddish  tint.  Abdomen  very  hair}^,  grey,  in  g 
deeply  mottled  with  black  and  fuscous,  in  9  sparsely  with  black, 
but  freely  with  fuscous  and  reddish.  Forewings  costa  slightl}' 
wavy,  hindmargin  rounded,  grey,  diffused  with  darker  gre}^  fus- 
cous, and  slaty  scales;  and  marked  with  lines  and  short  bars  of 
velvety  black.  Forewings  with  costal  edge  whitish,  crossed  with 
bars  of  black,  about  twelve  in  number,  extending  across  the  wing 
as  wavy  denticulate  lines,  irregular  in  contour,  and  at  places 
faint,  at  others  deep  black  and  diffused;  in  outer  third  of  wing 
there  are  short  black  cross  bars  which  join  the  transverse  lines 
and  form  an  irregular  network,  more  or  less  clearly  defined:  cilia 
grey,  with  a  dark  basal  line.  Hindwings  deep  grey  to  black;  cilia 
dark  grey,  with  lighter  basal  and  median  lines. 

Allied  to  C.  cnhginosa,  Walk.,  a  common  southern  species,  but 
rather  rare  in  Brisbane.  It  is  scarcely  half  the  size,  and  is  more 
strongly  barred  with  transverse  lines. 


BY    THOMAS    P.    LUCAS.  "  247 

Brisbane;  found  in  larval  state  by  Mr.  Illidge,  our  most  care- 
ful observer. 

Family  PYSALIDIDiE, 

Pyralidinse. 

E  U  D  A  I  M  O  N  I  S  M  A,    n.g. 

Head  with  adpressed  hairs,  a  conspicuous  tuft  of  hairs  from 
between  the  eye  and  palpus,  on  either  side,  radiating  outwards. 
Tongue  well  developed.  Antennae  finely  serrate.  Maxillary 
palpi  short,  truncate.  Labial  palpi  adpressed  to  tongue,  straight, 
drooping,  first  and  second  joints  hairy,  terminal  joint  short, 
acuminate.  Forewings  with  veins  7  and  8  from  a  point,  8  and  9 
stalked,  10  and  11  out  of  7.  Hindwings  with  vein  6  free, 
approaching  7  before  middle,  7  and  8  stalked. 

Allied  to  ^Enogenes. 

EuDAiMONiSMA  Batchelorella,  nsp. 

(^9-  1^-15  nim.  Head  woolly  white,  with  two  short  frontal 
tufts  tipped  with  fuscous-black.  Palpi  white,  edged  with  dark 
fuscous,  terminal  joint  short,  deep  fuscous.  Antennae  simple, 
ochreous-white.  Thorax  white,  with  a  narrow  line  anteriorly 
and  a  small  patch  on  dorsum  light  fuscous.  Abdomen  white  with 
a  few  light  fuscous  scales  on  free  edge  of  segments.  Legs  white; 
first  pair  dark  fuscous  at  joints.  Forewings  woolly  white  sparsely 
suffused  with  fuscous  scales,  and  lined  and  bordered  with  rich 
chestnut-fuscous;  costa  straight,  apex  bowed,  hindmargin  straight, 
inner  half  obliquely  bowed.  Forewings  with  chestnut  band  along 
costa.  finely  edged  and  irregularly  interrupted  with  white ;  a 
circular  line  of  chestnut  from  i  costa  to  J  inner  margin,  twice 
angled  posteriorly  and  deepl}'-  in  middle  anteriorl}';  a  discal  spot, 
not  touching  costa  beyond  |,  prolonged  and  narrowed  obliquely 
to  median  vein;  a  fine  wavy  dentate  circular  line  from  |  costa, 
becoming  parallel  with  hindmargin  through  middle  third  of  wing, 
thence  bent  at  right  angles  anteriorly,  again  bent  sharply  on 
itself,  and  again  angled  to  |  inner  margin;  a  blotch  from  anal 


248  NEW    SPECIES    OF    QUEENSLAND    LEPIDOPTERA, 

angle  is  suffused  across  this  last  line  to  discal  spot;    a  subhind- 

marginal  line,  and  a  hindmarginal  line  deep  chestnut  brown,  form 

a  band  with  a  fine  white  line  between  :  cilia  fuscous,  darker  at 

base.     Hindwings  as  forewings,  a  lunular  discal  spot   opposite 

and  near  to  J  costa;  a  circular  and  w^avy  line  from  |   costa   to 

opposite  J  inner  margin;  hindmarginal  line  thinning  out  at  anal 

angle,  not  so  dark  as  the  same  line  in  forewings  ;  cilia  whitish, 

with  basal  row  of  chestnut  dots. 

Mr.  Batchelor,  a  collector  by  occupation,  has  bred  this  insect 

freely,  feeding  on  a  wild  vine,  Lyonsia  reticulata.     It  spins  a  web 

in  a  leaf,  or   webs  the  leaf   to  the  tree   against  which  the  vine 

grows.       It  is  a  green  caterpillar,  with   two  subdorsal  rows  of 

j^ellow  dots. 

Brisbane. 

Family  XYLORYCTID^. 

Cryptophaga  confundens,  n.sp. 

(J 9.   28-45  mm.     Head  grey,  mottled  with  slaty-grey  and  light 
fuscous.     Palpi  slaty -gre}^,  first  and  second  joints  lighter  on  upper 
side.     Antennae  dark  grey  or  black,  with  fine  grey  annulations. 
Thorax  grey,  diffused  with  slaty-grey  and  darker  slate-bluish  dots. 
Abdomen    fuscous-grey,    freely   splashed   with   whitish-grey   and 
black,  in  a  band  along  base  of  each  segment,  but  metallic-coppery 
dusted  with  silver  hairs  on  free  half  of  segments  ;  in  some  speci- 
mens the  basal  band  is  crossed  by  rich  black  lines  edged  with 
grey   or    white.       Legs   woolly-grey,    with   rich   blackish-fuscous 
bands.       Forewings    oblong,    costa   gently   arched,    hindmargin 
slightly  concave  in  costal  half,  rounded  on  inner  half;  vein  2  from 
before  h',  gi'ey,  more  or  less  diffused  with  slaty-grey  (and  in  some 
specimens  sparingly  with  red  coppery  tint),  with  white  scales,  and 
richly  and  variously  marked  with  rich  black  bars  and  lines;  a  rich 
black  band  at  base,  more  or  less   constant  and  becoming  more 
definite  and  constant  along  inner  margin  to  \,  a  rich  band  from 
middle  of  base  obliquely  toward  costa,  thinning  out  or  suffusing 
with  another  from  J  to  before  ^;  a  band  transversely  from  ^i^  costa 
to  within  \  inner  margin  opposite  |;  half  way  between  this  line 


BY    THOMAS  P.     LUCAS.  249 

and  base  of  wing  is  an  irregular  more  or  less  parallel  line,  in  some 
specimens  stopping  short,  in  others  running  irregularly  to  inner 
margin,  and  in  others  one  or  two  other  irregular  lines  still  nearer 
the  base  of  wing;  a  conspicuous  wavy  zigzag  band  from  costa  to 
^  inner  margin  prominently  arngled  outwards  before  and  over 
middle,  and  thence  sending  a  line  at  an  angle  towards  base  of 
wing;  a  rich  wide  band  of  black  in  some  specimens  along  median 
fold,  in  others  this  is  absent;  a  suffusion  of  white  beyond  outer 
transverse  line,  shaded  more  or  less  towards  costa  with  slaty- 
grey,  in  some  specimens  absent,  in  others  only  as  a  lunar  mark 
over  inner  half  of  wing ;  a  white  zigzag  subhindmarginal  line 
becoming  lost  before  inner  margin,  and  in  many  specimens  alto- 
gether absent :  cilia  grey,  lined  with  black  and  fuscous.  Hind- 
wings  fuscous,  lighter  towards  the  base;  cilia  light  buff,  fuscous 
at  base. 

Brisbane;  in  stumps  and  second  growth  of  felled  Eucah'pts. 

A  most  variable  species.     Allied  to  C.  epigramma,  Meyr. 

Catoryctis  truncata,  n.sp. 

^.  20  mm.  Head  ochreous-white;  palpi  ochreous-white,  ter- 
minal joint  diffused  and  sparingly  dotted  with  fuscous;  antennae 
dark  fuscous;  thorax  ochreous;  abdomen  ochreous,  with  a  band 
of  blackish-fuscous  at  base  of  segments;  legs  fuscous.  Forewings 
elongate,  costa  arched,  hindmargin  obliquely  rounded,  ochreous, 
with  white  bands  and  blackish  fuscous  lines  and  bands.  Fore- 
wings  with  four  longitudinal  bands  of  ground  colour,  the  first 
along  costa,  thins  out  towards  apex  and  base,  and  is  bounded  by 
a  subcostal  silver-white  line  from  base  to  beyond  ^  costa;  the 
second  line  is  median  and  is  suffused  with  silvery- white;  the  third 
line  is  immediately  before  the  inner  margin  and  the  fourth  is 
inner  marginal,  both  are  from  near  base  of  inner  margin  and  thin 
out  towards  apex;  the  space  between  first  and  second  lines  is  deep 
blackish-fuscous,  and  is  divided  by  eight  lines  of  ground  colour, 
running  obliquely  outwards  from  median  to  costa,  the  apical  ones 
gradually  become  shorter;  the  second  space  between  the  second 
and  third  bands  is  blackish-fuscous  and  is  divided  in  hind  marginal 


250  NEW    SPECIES    OF    QUEENSLAND    LEPIDOPTERA. 

third  by  four  ground  colour  longitudinal  parallel  lines  :  cilia  dark 
fuscous.  Hindwings  light  grey  shaded  with  darker  fuscous ; 
veins  darker;  cilia  gre3ash-fuscous,  with  a  broad  darker  band 
near  base. 

May  Orchard,  Brisbane;  one  specimen  at  light. 

Allied  to  C.  subparallela,  Meyr.  (?),  and  C.  nono-lhiea,  Lucas. 

T  H  E  I  0  s  I  A,   n.g. 

Head  with  adpressed  hairs.  Ocelli  absent.  Tongue  well 
developed.  Antennae  moderate,  simple,  finely  ciliated,  basal  joint 
elongate,  with  moderate  pecten.  Maxillary  palpi  ver}^  short, 
adpressed  to  tongue.  Labial  palpi  very  long,  recurved,  first  joint 
short,  second  very  long,  with  adpressed  hairs,  third  joint  linear, 
smooth.  Thorax  hairy,  epaulettes  and  collar  with  very  long  hairs. 
Abdomen  moderate.  Tibiae  hairy,  spurs  long.  Forewings  with 
veins  3  and  4  from  a  point,  6  free,  7  and  8  stalked,  9  from  7. 
Hindwings  with  vein  2  from  J  cell,  3  and  4  from  angle,  5  free,  (> 
and  7  short  stalked,  8  concurrent  with  7  for  ^  from  base. 

Comes  near  to  Ptilostibes,  Meyr. 

Theiosia  .etheria,  n.sp. 

9-  28  mm.  Head  and  palpi  reddish-ochreous;  antennae  fuscous, 
basal  joint  ochreous;  thorax  ochreous,  wdth  a  rich  chestnut  band 
behind  collar;  abdomen  whitish-ochreous;  legs  ochreous,  tarsal 
joints  of  hind  pair  fuscous.  Forewings  with  costa  gently  rounded, 
hindmargin  rounded,  reddish-ochreous,  sparsely  speckled  with 
black  dots,  and  marked  with  white,  grey- black,  and  chestnut. 
Forewings  with  a  kidney-shaped  rich  chesenut-coloured  discal 
blotch,  extending  over  middle  third  of  wing,  beyond  |  costa;  this 
is  bordered  on  costal  edge  by  a  fine  white  line;  and  this- is  again 
enclosed  by  a  broad  band  of  smoky  black,  which  extends  from 
base  of  wing  to  opposite  ^  costa,  and  is  again  enfolded  by  a  broad 
white  suffusion,  reaching  to  costa  at  base,  and  gradually  receding 
from  costa  at  J,  leaving  a  band  of  ground  colour  for  costal  border 
to  apex;  the  white  is  freely  speckled  with  black  dots;  a  subhind- 


BY    THOMAS    P.    LUCAS.  251 

marginal  line  of  black  dots,  and  a  hindmarginal  reddish-brown 
line  :  cilia  reddish-ochreous,  diffusing  to  reddish-fuscous  at  apex 
and  apical  hind  margin.  Hind  wings  light  ochreous;  cilia  light 
ochreous. 

May  Orchard,  Brisbane;  bred. 

Xylohycta  heliojMacula,  Lower. 

I  have  bred  this  species  from  larvae  feeding  on  a  species  of 
mistletoe  (Loranthus)  in  Brisbane.  The  three  or  four  previously 
known  species  were  captured  at  light. 


252 


A  REVISION  OF  THE  GENUS  JOTOXOMUS  (FAMILY 
CARABIDjE:  subfamily  FER02^INI). 

By  Thomas  G.  Sloane. 

JVoto7iomus  was  founded  b}'-  Baron  de  Chaudoir  (Bull.  Soc.  Imp. 
Nat.  Mosc.  1865,  ii.  p.  83)  as  a  subgenus  of  Feronia;  but  in  the 
present  paper  it  is  assumed  to  have  full  generic  value.  Strictly 
speaking  it  seems  to  be  a  subgenus  of  Pterostichus,  as  are  also 
C eratofevonia*  Sarticus  and  Rhahdotus;  if  we  regard  all  these  as 
entitled  to  full  rank  as  genera,  and  it  is  certainly  convenient  to 
do  so,  they  may  be  tabulated  as  follows  : — 

Mentum  with  sinus  parallel  on  sides Cera tofornia 

Mentum  with  sinus  obliquely  divergent. 

Prothorax  with  border  narrow,  not  forming  an  explanate 
border  to   the  lateral  basal  impressions,  these  never 

spreading  to  lateral  border    Xotononius. 

Prothorax  with  border  explanate  posteriorly. 

Lateral  basal   impressions   of    prothorax   broad,  deep, 

foveiform Sarticiis. 

Posterior  part  of  prothorax  transversely  depressed  (this 

depressed  part  reaching  the  border  laterally) PiJiahdotHS. 

JVotono7nus  may  be  briefly  diagnosed  as  follows  : — 
Apterous.  Mentum  with  sides  of  sinus  oblique.  Prothorax 
with  border  narrow  and  not  forming  an  explanate  margin  to  the 
lateral  basal  impressions.  Elytra  bordered  on  base,  striate  ; 
interstices  not  costate  or  carinate,  first  with  an  abbreviated  stria 
at  base,  third  punctiferous.  Posterior  femora  not  dilatate  towards 
apex,  or  channelled  on  lower  side. 

*  M.  Tschitseherine  has  recently  formed  this  new  subgenus  to  include 
Feronia  (PterosticJius J  rerjalu,  Casteln.,  and  P.  pJnjhircJius  SI. 


BY    THOMAS    G.    SLOANE.  253 

Attention  ma}^  be  drawn  to  the  following  features  which  vary, 
and  which  are  useful  in  aiding  the  determination  of  species  : — 
Head :  size,  convexity,  length  of  mandibles,  &c. 
Prothorax  :   shape,    convexity,   comparative   width   of  base   and 
apex;  disc,  if  opaque  or  nitid;  sides  sinuate  or  not  before 
base,  form  of  basal  angles,  nearness  of  anterior  angles  to 
head;  position  of  posterior  marginal  puncture  (an  important 
classificatory  character)  ;  width  of   border,  especially  near 
base;  etc. 
Elytra:  shape,  convexity,  sinuosities  of  apex,  striation;  interstices 
(opaque  or  nitid),  convexity,  punctures  of  third,  form  of  eighth 
(a  valuable  classificatory  character);  humeral  angles  dentate 
or  not;  reflexedness  of  lateral  border  near  base. 
Frosternum    and    mesosternum :    shape    of    intercoxal    declivity 
(this  seems  a  constant  and  useful  character  for  grouping  the 
species  in  Notonomus). 
Metastermim :  episterna  long,  or  short. 

The  habitats  of  the  species  of  Notonomus  are  among  the  heavily 
timbered  districts  of  South  Eastern  Australia.  The  range  of  the 
genus  extends  from  about  the  Grampian  Mountains  in  Western 
Victoria,  along  the  coastal  districts  of  Eastern  Australia  as  far 
north  as  the  Burnett  River.  Only  one  species  is  authentically 
known  to  me  from  north  of  the  Burnett  River,  A  single  species 
{N.  mediosulcatus,  Chaud.),  not  closely  allied  to  any  other,  is 
found  in  South  Western  Australia.  Probably  owing  to  their 
being  apterous,  and  frequenting  heavily  timbered  mountainous 
districts,  the  species  seem  usually  restricted  in  their  range.  No 
species  from  north  of  the  Clarence  River  is  known  to  me  as  being- 
found  as  far  south  as  Sydney,  nor,  as  far  as  I  know,  is  any  species 
from  Sydney  to  be  found  about  Melbourne. 

Dejean  first  described  species  belonging  to  the  genus  Notonomus 
(viz.,  three  species  in  the  year  1828).  Then  followed  Count  de 
Castelnau  with  two  species  {Trigonotoma  violacea  in  1834,*  and 

*  Chaudoir  reported  after  inspection  of  the  type  (Ann.  Soc.  Ent,  Belg.  xi., 
p.  162)  that  T.  violacea,  Casteln.,  was  a  species  of  Notonomus,  and  this  has 
recently  been  confirmed  by  M.  Tschitscherine  (Hor.  Soc.  Ent.  Koss.  1902, 
XXXV.,  p.  534). 


254  EEVISION  OF  THE  GENUS  yOTONOMUS, 

Poecilus  maryinatus  in  1838);  and  Xewman  with  one  species 
{Feroniaj)hilipjn  =  F.  chalyhea,  Dej.).  Nothing  further  was  done 
till  1865,  when  Baron  de  Chaudoir  and  M.  Motschulsky,  inde- 
pendently of  one  another,  and,  strange  to  say,  in  the  same  journal 
(Bull.  Mosc.  1865),  described  a  considerable  number  of  new 
species.  Chaudoir's  paper  contained  fourteen  new  species 
(of  these  one,  N.  discoderus,  proved  synonymous  with  H.  marginaius, 
Casteln. ).  Motschulsky  described  four  species  as  new,  but  three 
of  these  were  anticipated  by  Chaudoir.  Castelnau's  great 
paper  on  the  Australian  Carabidse  was  read  before  the  Royal 
Society  of  Victoria  on  May  13th,  1867,  and  was  published 
in  Vol.  viii.  of  that  Society's  Transactions  in  1868;  in  it  he  pro- 
posed three  hundred  and  thirt3^-seven  species  as  new,  including 
thirty-eight  referable  to  Notonoiiius.  Castelnau's  work  was  done 
without  a  knowledge  of  what  Chaudoir  and  Motschulsky  had 
done,  and,  mainly  owing  to  this,  the  names  of  twenty-one  of  his 
species  of  Notononius  have  now  become  synonyms.  In  1871  Sir  W. 
Macleay  published  descriptions  of  five  species;  of  these  it  seems 
that  only  one  will  stand.  Chaudoir  (who  evidently  liecame 
possessed  of  Dejean's  types)  went  over  the  Castelnau  Collection, 
now  the  propert}'-  of  the  Museo  Civico  di  Genova,  and,  in  the 
Annals  of  that  Institution  for  1874  (Vol.  vi.),  published  what  he 
called  a  "Supplement"  to  his  essay  of  1865.  Knowing  at  that 
time  all  his  own,  Dejean's,  and  Castelnau's  species  (excepting 
Feronia  wilcoxi,  Casteln.,  F.  striaticollis,  Casteln.,  and  F.  rufi- 
palpis,  Casteln.,  the  types  of  which  he  could  not  find),  he  was 
able  to  settle  the  synonymy  of  Notonomus  authoritatively.*  This 
was  fortunate  for  subsequent  workers,  for  it  would  have  been 
practically  a  hopeless  task  for  anyone  without  the  data  which 
Chaudoir  possessed,  because  both  Chaudoir's  and  Castelnau's 
descriptions  of  the  species  of  Notonomus  are  generally  very  faulty, 
being  too  brief,  and  often  consisting  of  merely  a  few  lines  of  a 


*  I  do  not  feel  sure  that  Chaudoir  knew  all  Motschulsky's  species  in  nature, 
though  he  treats  of  them  as  confidently  as  if  he  did,  but  without,  as  far  I 
know,  saying  that  he  knew  them. 


BY    THOMAS    G.    SLOANE.  255 

comparison  with  some  other  poody  described  species  of  the  author. 
Besides  arranging  the  synonymy,  Chaudoir,  in  his  "Supplement," 
redescribed,  or  made  notes  on  many  of  Castelnau's  species,  and 
described  four  new  species.  In  1878  Chaudoir  made  his  last 
contribution  to  the  genus  by  the  description  of  N.  joar  allelomorph  a, 
which  I  believe  to  be  only  a  form  of  N.  opulentus^  Casteln.;  and 
in  the  same  year  H.  W.  Bates  also  described  a  single  species  from 
Tasmania.  Nothing  further  was  done  till  1889,  when  I  described 
two  new  species.  Since  that  date  Mr.  Rainbow  (Rec.  Aust. 
Mus.  1899)  described  N.  montanus,  but,  as  this  name  had  already 
been  used  in  the  genus  by  Castelnau,  albeit  Castelnau's  name  has 
become  a  synonym,  it  cannot  stand.  This  makes  forty-five  valid 
species  described  up  to  the  present  time.  In  the  present  paper  I 
add  twenty-seven,  making  a  total  of  seventj^-two;  no  doubt  there 
are  many  more  species  yet  to  be  discovered. 

In  undertaking  the  revision  of  the  genus,  I  have  had  to  make 
out  the  species  of  Dejean,  Chaudoir  and  Motschulsky  from  the 
descriptions  (except  a  few  that  are  well  known  in  Australian 
collections),  as  well  as  most  of  Castelnau's.  I  have  not  had 
Dejean  and  Motschulsky's  descriptions  before  me,*  but  have  relied 
on  Chaudoir's  notes  to  identify  their  species.  In  all  cases  where 
I  have  made  out  species  from  descriptions,  I  have  indicated  that 
such  was  the  case  in  my  notes.  The  identification  of  many  species 
has  proved  diflicult,  but  it  has  been  the  work  of  a  number  of 
years,  and  has  been  gone  over  many  times  in  the  endeavour  to 
arrive  at  a  correct  determination  of  each  species. 

I  offer  a  tabulation  of  the  species  known  to  me  in  which  an 
attempt  is  made  to  arrange  the  species  in  a  fairly  natural 
order,  or  at  least  what  seems  to  me  a  natural  order.  But, 
seeing  that  the  species  of  a  large  genus  in  their  relationships 
towards  one  another  resemble  the  branches  of  a  tree  springing 
from  one  trunk,  rather  than  a  continuous  chain,  it  is  impossible 
to  place  them  in  a  linear  series  by  the  aid  of  a  dichotomous  table 
without  species   being   separated   sometimes  from   nearly  allied 


*  Excepting  that  of  Xeuropates  pristonycJioides,  Motsch. 


256  EEVISION  OF  THE  GENUS  XOTO^OMCS, 

congeners.  In  lettering  the  subdivisions  of  the  table  I  have 
used  italics  for  general  characters,  Roman  capitals  for  divisions 
that  may  be  regarded  as  groups  of  species,  and  Greek  letters  for 
minor  subdivisions  ;  by  bearing  this  in  mind  an  idea  may  be 
formed  of  the  gronjjs  into  which  I  consider  the  species  might  be 
lotted. 

The  following  species  are  unknown  to  me  in  nature,  or  at  least 
I  have  been  unable  to  identify  any  of  them  among  the  species  I 
have  had  before  me  : — N.  violaceus,  Casteln.  (of  which  I  have  not 
seen  the  description,  so  that  it  is  quite  unknown  to  me),  iV. 
phillipsi,  Casteln.  (see  under  i\".  strzleckianus,  and  postea,  p.  278), 
JV.  depressipennis,  Chaud.,  and  iV.  striaticollis,  Casteln.  (see  under 
i\^.  rugosicoUis,  SI.,  and  J\^.  discorimosus^  SI.,  postea,  pp.  305,  306), 
N.  ingratus,  Chaud.,  N.  subii'idescens,  Chaud.,  N.  darlingi, 
Casteln.,  N.  tvilcoxi,  Casteln.  N.  simidans,  Chaud., i^'^.  resplendens, 
Casteln.,  are  species  to  which  I  have  allotted  separate  headings 
under  which  my  ideas  as  to  their  resemblances  and  affinities  may 
be  found;  I  cannot  think  that  any  of  these  species  can  be  con- 
specitic  with  any  species  for  which  I  propose  a  new  name  in  the 
present  paper. 

Table  of  Species  known  to  me. 

a.  Metasternum  with  episterna  short. 
h.  Elytra  fully  and  strongly  striate. 

c.  Intercoxal  declivity  of  prosternum  rounded,  usually  narrow  in  middle. 
cl.  Elytra  with  third  interstice  normally  bearing  more  than  two  punc- 
tures, the  posterior  puncture  on  apical  declivity. 
e.  Elytra  with  basal  border  joining  lateral  border  at  humeral  angle 
without  any  interruption. 
f.  Elytra  with   eighth   interstice  not   (or   very  lightly)   convex, 
decidedly  wider  than  ninth  on  basal  half. 
(1.  Pro  thorax  with  sides  sinuate  before  the  base. 
A.  Prothorax  with  lateral  basal  spaces  convex. 

(B.  Prothorax  with  posterior  marginal  puncture  (  N.frenchi. 

in  lateral  channel  before  basal  angles I  N.croesus. 

/3/3.  Prothorax  with  posterior  marginal  puncture  at  basal 
angle  {^  with  joints  of  anterior  tarsi  not  dilatate  or 

squamulose  beneath) N.  lyeroni. 

AX.  Prothorax    with    lateral    basal    spaces    depressed    (not 
convex) iV.  accedens. 


BY    THOMAS    G.    SLOANE. 


257 


17 


gg.  Prothorax  with  sides  meeting  base  without  any  sinuosity. 
B.  Prothorax  but  little  narrowed  to  base,  much  wider  across 
base  than  apex, 
y.  Form    depressed,    prothorax    suh- j  N.  poUtnlus. 

obliquely  narrowed  to  base    \N.  tuhericauda. 

77.  Form  rather  narrow,  convex;  prothorax  with  sides 

evenly  rounded  ^.  kosciuskianus. 

BB.  Prothorax  decidedly  narrowed  posteriorly,  base  not,  or 
very  little  wider  than  apex, 
c.  Prothorax  with  basal  angles  widely  obtuse. 
D.  Prothorax  with  posterior  marginal  puncture  at  basal 
angle  on  border,  or  the  border  interrupting  mar- 
ginal channel  to  receive  it, 
S.  Prothorax  with  sides  narrowed   to  base    in  an 
even   curve,   basal  angles    quite    rounded   off 

A^.  rainhoioi. 

8d.  Prothorax  with  sides  a  little  obliquely  narrowed 
to    base,   basal  angles    obtuse    but    marked 

A^.  gippsiensis. 

DD.  Prothorax  with  posterior  marginal  puncture  before 
the  basal  angle. 
E.  Head  large;    eyes   deeply   inclosed   at   base   by 
greatly  developed  orbits,  not  prominent;  orbits 

swollen  below  eyes a^.  satrapa. 

EE.  Head  moderate;  eyes  not  deeply  enclosed  at  base, 
rather    prominent;     orbits     not    unusually 

swollen  below  eyes a^.  arthuH. 

cc.  Prothorax  with  basal  angles  marked  (posterior  mar- 
ginal puncture  at  basal  angle), 
r.  Head  large ;  border  of  prothorax  ending  at  basal 
angles,  posterior  marginal  puncture  at  basal  angle 

but  not  on  border ^V.  vmelUri. 

FF.  Head  not  large,  prothorax  with  border  extending 
on  to  base  on  each  side  and  dilatate  at  basal 
angles  to  receive  posterior  marginal  puncture. 
G.  Prosternum  not  bordered  along  anterior  margin. 

e.  Colour  black N.  macoyi. 

ee.  Colour  cupreous A\  besii. 

GG.  Prosternum  with  narrow  continuous  border 
along  anterior  margin  (colour  deep  black) 
N.  hoicitti. 


258  EEVISION  OF  THE  GENUS  NOTONOMUS, 

ff.  Elytra  with  eighth  interstice  narrow  and  very  convex 
(similar  in   form   to   ninth).      Form  elongate,  head  very 

large N.  hakeioelli. 

ee.  Elytra  with  point  of  junction  of  basal  and  lateral  borders  at 

humeral  angle   interrupted   (the   basal  border  more  or  less 

raised  and  prominent.) 

H.  Prothorax    with    lateral    border   wide,   increasing    in   width 

posteriorly. 

C'  Prothorax  cordate,  narrower  at  base  than  at  apex;  posterior 

marginal  puncture  at  basal  angle  , .  .X  Jietcheri. 

iC.  Prothorax  subquadrate,  wider  at  base  than  at  apex;  posterior 
marginal  puncture  on  inner  side  of  lateral  channel  distant 
from  basal  angle. 
rj.  Head  rather  large,    metallic ;    posterior   femora   lightly 

swollen  in  middle N.  strzelecldanus. 

r]T].  Head  small,  black;  posterior  femora  strongly  dilatate  in 

middle  on  lower  side  {^) N.  froggatti. 

HH.  Prothorax  with  lateral  border  narrow,  not  wider  towards  base. 
6.  Elytra  with  eighth  interstice  net  convex  near  base. 

I.  Prothorax   of   a   gilt   brassy  colour,   elytra   purple    (liery 

towards  sides) ,  N.  hassi, 

a.  Colour  of  upper  surface  green X.  gippslandicns. 

66.  Elytra  with  eighth  interstice  roundly  convex. .  .A^.  variicolUs. 
dd.  Elytra  with  third  interstice   bipunctate,  the   posterior   puncture 
before  apical  declivity. 

I.  Prothorax  longer  than  broad,  greatly  narrowed  and  sinuate  on 

sides  posteriorly xV.  a nf/ustihasis. 

II.  Prothorax  broader  than  long,  sides  not  sinuate  before  base. 

J.  Prothorax  with  lateral  border  narrow,  elytra  with  lateral  border 
narrow  at  base. 
K.  Elytra  with  interstices  very  convex,  eighth  and  ninth  very 
narrow;  prothorax  metallic  (  $  with  apical  sinuosities  of 

elytra  very  strongly  developed) N.  eacisijjennis. 

KK.  Elytra  with  interstices  lightly  convex,  eighth  wider  than 

ninth;  colour  black X  Jdngi. 

jj.  Prothorax  with  lateral  border  widely  reflexed,  especially 
towards  base;  elytra  with  lateral  border  widely  carinate  at 
base. 

X.  Elytra  with  a  cupreous  margin JV.  marginatm. 

XA.  Elytra  without  a  metallic  margin   X.  ligdneyenHs. 

cc.  Intercoxal  declivity  of  prosternum  flat — often  wide  in  middle  and 
bordered  laterallv. 


BY    THOMAS    G.    SLOANE.  259 

h.  Elytra  with   third    interstice    normally  bipunctate,  the   posterior 
puncture  before  apical  declivity. 
i.  Prothorax  with  posterior  marginal  puncture  on  border  at  basal 
angle. 
K.  Metasternum  with  median  longitudinal  channel  not  extending 
to  apex, 
/x.  Elytra  with   tenth    interstice   short   and   feebly  developed, 
(lateral  border  widely  reflexed  at  base,  eighth  interstice 

much  wider  than  ninth  on  basal  half) N.  australasia. 

fxfi.  Elytra  with  tenth  interstice  convex,  long  (extending  on  to 
anterior  half  of  elytra). 
p.  Basal  border  of  elytra  strongly  dentate  at  humeral  angles, 
(eighth    and     ninth    interstices    narrow  and    convex) 

iV".  incrassatus. 

vv.  Basal  border  of  elytra  a  little  raised  but  not  dentiform  at 
humeral  angles. 
I-  Elytra  with  third  interstice  bipunctate;  prothorax  short, 

basal  impressions  oval  N.  leai. 

^^.  Elytra  with   third   interstice  tripunctate  ;    prothorax 
rather  long,  basal    impressions    long  and   narrow 

N.  lateralis. 

KK.  Metasternum  deeply  longitudinally  channelled  from  base  to 
apex. 
L.  Prothorax  short,    cordate;    elytra  with 
eighth    and    ninth    interstices  wide ; 

colour  metallic 

LL.  Prothorax  long;  elytra  depressed,  interstices  sharply  convex, 

eighth  and  ninth  narrow;  colour  black N,  liiagerus. 

ii.  Prothorax  with  posterior  marginal  puncture  not  placed  on  lateral 
border. 

M.  Pronotum  nitid  (though  usually  finely  striolate). 
j.  Prothorax  with  sides  sinuate  before  base. 

77.  Head  and  prothorax  black;  elytra  obscurely  coloured,  with 

fiery  cupreous  margin N.  queenslandicus. 

7777.  Head  and  prothorax  metallic;  elytra  with  green  margin, 
p.  Prothorax    with     base    hardly    wider   than    apex  ( $ ) 

N.  n  itidicoUis. 

pp-  Prothorax  with  base  evidently  much  wider  than  apex 

($) N.  latibasis. 

jj.  Prothorax  with  sides  not  sinuate  posteriorly. 

I.  Form  narrow;  prothorax  long,  not  wider  at  base  than  at 
apex ;  elytra  with  eighth  interstice  much  wider  than 
ninth  anteriorly,  very  narrow  posteriorly — opposite  the 
elongate  convex  tenth  interstice N.  suhopacus. 


N.  triplogenioides. 
N.  jervisensis. 


260  EE VISION  OF  THE  GENUS  X0T0X03[rS, 

II.  Form  wide;  prothorax  transverse,  wider  across  base  than 
apex;  elytra  with  eighth  interstice  a  little  wider  than 
ninth,  not  unusually  narrowed  posteriorly,  tenth  inter- 
stice short,  feebly  developed N.  violaceomarginatus. 

MM.  Pronotum  opaque,  closely  transversely  striolate. 
m.  Elytra  with  all  discoidal  interstices  equally  smooth. 
cr.  Prothorax  with  sides  sinuate  before  the  base. 

r.  Elytra  with  eighth  interstice  wide N.  riigosicollis. 

TT.  Elytra  with  eighth  interstice  narrow...  A'',  depressipennis. 
era:   Prothorax  with  sides  not  sinuate  posteriorly  (elytra  with 

eighth  interstice  narrow,  convex) N.  discoriinosus. 

vim.  Elytra    with     alternate    interstices     of     disc     foveolate 

N.  opacicollis. 

hli.  Elytra  with  third   interstice  normally  bearing  more  than  two 

punctures,  the  posterior  puncture  normally  on  apical  declivity. 

n.  Prothorax  with  posterior  marginal  puncture  at  basal  angle  on 

lateral  border  (border  usually  dilated  to  receive  puncture). 

N.  Elytra  with  basal  border  joining  lateral  border  at  humeral 

angle  without   any  interruption.      Elytral   striae   crenulate 

N.  amahilis, 

NN.  Elytra  with  point  of  junction  between  basal  and  lateral  borders 
at  humeral  angle  interrupted  (the  basal  border  more  or  less 
raised), 
o.  Size  rather  small ;   prothorax  with  lateral  border  narrow ; 
elytra  with  border  narrowed  to  base. 
V.  Head  and  prothorax  black. 

0.  Form  not  elongate,  third  interstice  of  elytra  normally 
with  more  than  two  punctures  and  not  swollen 
posteriorly. 

X-  Elytra  viridescent,  lightly  convex N.  sphodroides. 

XX'  Elytra  purple,  depressed  N.  dyscoloides. 

^<^.  Form  elongate;  third  interstice  of  elytra  bipunctate 
and  swollen  at  posterior  puncture... A",  victoriensis. 

vv.  Head    and    prothorax    seneous.      (Form    elongate) 

iV.  opnlentus. 

00.  Size  large ;  prothorax  with  lateral  border  wide  and  erect 
posteriorly;  elytra  with  lateral  border  widely  carinate  at 
base, 
•v//--  Prothorax  with  sides  sinuate  before  base,  basal  angles 

sharply  marked N.  (Oteomicans. 

yj^^'  Prothorax  with  sides  not  sinuate  posteriorly,  basal 
angles  obtuse N.  colossus. 


BY    THOMAS    G.    SLOANE.  261 

nil.  Prothorax   with    posterior  marginal    puncture    not   on   basal 

border. 

p.  Prothorax  cordate;  head  large;  elytra  with  eighth  interstice 

narrower  than  ninth  (size  small,  colour  black )...xV.  rufipalpis. 

pp.  Prothorax  hardly  narrowed  to  base;  head  small;  eighth  elytral 

interstice  wider  than  ninth  (size  large). 

0).  Form  depressed;  elytra  with  humeral  angles  strongly  dentate 

(colour  Eeneous) K.  cques. 

(0(o.  Form  convex;  elytra  with  basal  border  only  a  little  raised 
at  humeral  angle.     (Elytra  dark,  with  green  margin) 

N.  spenceri. 

bb.  Elytra  with  striae  1-4  deeply  impressed,  5-7  obsolete... iV.  mediosnlaitm. 
bbb.  Elytra  with  striae  obliterated,  or  faint. 
p.  Elytra  with  humeral  angles  dentate. 

q.   Size  large;  prothorax  transverse,  lightly  and  obliquely  rounded  to 

base;  black N.  moUi^tus. 

qq.  Size  moderate;  prothorax  strongly  angustate  to  base,  with  sides 

lightly  sinuate;  elytra  virescent iV.  Ixershawi. 

pp.  Elytra  with  humeral  angles  not  dentate. 

r.  Prothorax  widest  before  middle,  a  little  narrowed  to  base;  elytra 
with  basal  border  a  little  raised  at  point  of  junction  with  lateral 

border;  elytra  chalybeous  X.  chalijbeus. 

rr.  Form  convex;  prothorax  widest  about  middle,  sides  strongly  and 
evenly  rounded ;   elytra  with   basal   border  joining  lateral  at 

humeral  angles  without  interruption ;  colour  black N.  gravis. 

aa.  Metasternum  with  episterna  elongate. 

Q.  Elytra  with  humeral  angles  dentate iV".  miles. 

QQ.  Elytra  with  basal  border  joining  lateral  border   at  humeral  angles 
without  interruption  .,   N.  lemeuri. 

iSToTONOMUS    FRENCHI,    ll.sp. 

9.  Depressed,  oval.  Head  very  large;  mandibles  long,  decus- 
sating; labrum  emarginate:  prothorax  subcordate;  sides  strongly 
sinuate  and  straight  before  base;  basal  angles  obtuse,  widely 
bordered  at  summit  ;  posterior  marginal  puncture  in  lateral 
channel  distant  from  base :  elytra  lightly  striate,  sixth  and  seventh 
stride  fainter ;  interstices  depressed,  third  4-punctate,  tenth 
moderately  developed;  humeral  angles  rounded.  Head  and  pro- 
thorax of  a  gilt  brassy  colour,  prothorax  with  cupreous  reflections 


262  EEVISION  OF  THE  GENUS  NOTOXOM  US, 

in  middle  of  disc;  el3^tra  purple  with  coppery  reflections  (the  disc 
darker  and  less  metallic  than  the  sides);  under  surface  black,  legs 
reddish-piceous. 

Prothorax  broader  than  long  (4-9  x  5-8  mm.),  widest  before  the 
middle,  strongly  narrowed  to  base;  sides  rounded  on  anterior 
two-thirds,  strongly  sinuate  about  posterior  fifth  and  meeting 
base  at  right  angles;  apex  truncate,  angles  wide,  obtuse;  base 
truncate,  very  lightly  and  widely  emarginate  in  middle,  narrower 
(4-3  mm.)  than  apex  (4-5  mm.);  border  wide;  lateral  basal  im- 
pressions elongate,  wide;  lateral  basal  spaces  depressed  behind 
posterior  marginal  puncture.  Elytra  truncate-oval  (12x7*5  mm. ), 
widest  behind  middle,  depressed  on  disc;  sides  lightly  rounded; 
lateral  apical  sinuosities  wide,  shallow;  basal  border  raised  at 
humeral  angles,  but  joining  lateral  border  without  interruption; 
lateral  border  widely  reflexed  and  carinate  near  base.  Intercoxal 
declivity  of  prosternum  round  and  narrow  in  middle;  of  meso- 
sternum  deeply  concave.     Length  21,  breadth  7"5  mm. 

Hab. — Vic.  :  Southern  Gippsland  (French). 

Allied  to  iV.  crcesus,  Casteln.,  but  differing  by  its  broader  and 
more  depressed  form;  head  larger;  prothorax  more  transverse, 
more  ampliate  at  widest  part,  the  spaces  between  the  basal  im- 
pressions and  the  sides  flatter;  elytra  of  a  much  more  coppery- 
purple  on  the  sides.  The  type  specimen  has  the  fifth  interstice 
of  the  elytra  unipunctate.  The  male  will  probably  prove  to  have 
the  interstices  of  the  elytra  more  convex  and  the  basal  angles  of 
the  prothorax  more  marked  as  in  N.  crif^suif. 

N'ote.  —  A  specimen  received  from  Mr.  French,  as  from  Victoria, 
is  of  narrower  and  more  elongate  form  (18-5  x  6 -5  mm);  prothorax 
much  more  strongly  narrowed  to  base,  with  sides  more  strongly 
sinuate;  elytra  of  a  steel  blue  colour  with  faint  purple  reflections, 
head  and  prothorax  with  coppery  reflections.  It  probably  repre- 
sents a  closely  allied,  but  distinct,  species,  but  more  specimens  of 
both  forms  than  are  before  me  would  be  required  to  pronounce  a 
confident  opinion  on  this. 


BY    THOMAS    G.    SLOANE.  263 

NoTONOMUS  CRCESUS,  Castelnau. 

Feronia  (^Pterostichus)  crresus,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  viii.  p.  209  ;  F.  {Pterostichus)  plutus,  Casteln.,  I.e.  :  JV. 
peroni,  Casteln.;  Chaud.,  Ann.  Miis.  Civ.  Genova,  1874,  vi.  p.  588. 

Prothorax  and  head  of  a  gilt  coppery  colour;  elytra  purplish- 
black  with  cupreous  tinge  on  base  and  lateral  declivities;  legs 
dark  piceous.  (^)  Head  large  (3-7  mm.  across  eyes);  mandibles 
long,  decussating;  labrum  lightly  and  widely  emarginate;  eyes 
prominent.  Prothorax  broader  than  long  (4'1  x  5-1  mm.), 
narrowed  to  base;  sides  lightly  rounded  on  anterior  two-thirds, 
strongly  sinuate  about  posterior  fourth  and  meeting  base  at  right 
angles;  base  truncate,  narrower  (3*5  mm.)  than  apex  (3-7  mm.); 
basal  angles  rectangular,  obtuse  at  summit;  lateral  border  wide,  re- 
flexed;  posterior  marginal  puncture  in  lateral  channel  distant 
from  basal  angle;  median  line  lightly  impressed;  lateral  basal 
impressions  wide.  Elytra  oval  (10*5  x  63  mm.),  lightly  convex; 
lateral  apical  sinuosities  well  marked;  shoulders  rounded;  basal 
border  joining  lateral  border  without  interruption;  striae  strongly 
impressed;  interstices  lightly  convex,  third  5-punctate;  ninth 
stria  and  tenth  interstice  well  developed.  Intercoxal  declivity 
of  prosternum  rounded;  of  mesosternum  concave.  Anterior  tarsi 
with  three  basal  joints  widely  and  obliquely  dilatate  and  squamu- 
lose  beneath. 

(9)  Prothorax  with  basal  angles  more  obtuse ;  elytra  more 
depressed;  sixth  and  seventh  striae  much  weaker  than  others; 
interstices  depressed,  fifth  with  a  single  puncture  behind  middle. 
Length  18*5,  breadth  6*3-6*5  mm. 

Hah. — Victoria. 

The  specimens  on  which  the  above  description  is  founded  were 
given  to  me  by  the  late  Mr.  W.  Kershaw,  of  Melbourne.  They  seem 
to  represent  the  form  on  which  Castelnau  bestowed  the  name 
Feronia  plutus.  Two  specimens  {$)  sent  to  me  by  Mr.  C.  French, 
as  from  Mount  Baw  Baw,  Gippsland,  seem  to  represent  Feronia 


264  REVISION  OF  THE  GENUS  NOTOXOMUS, 

croesics,  Casteln.;  these  agree  with  the  description  given  above, 
except  that  the  prothorax  is  a  little  less  strongly  sinuate  on  the 
posterior  part  of  the  sides,  the  posterior  angles  more  obtuse,  the 
elytra  of  a  more  subviridescent  bronzed  colour. 

NoTONOMUS  PERONi,  Castelnau. 

Feronia  {Pterostichus)  peronii,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  viii.  p.  209;  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874,  vi. 
p.  588. 

Elliptical-oval,  depressed.  Upper  surface  of  a  metallic-bluish 
or  greenish  tinge,  elytra  often  flushed  with  purple;  legs  piceous- 
red,  tibiae  and  tarsi  darker  than  femora;  antennae  dark  piceous. 
Head  large.  Prothorax  quadrate-cordate  (3-9  x  4'8  mm.),  about 
as  wide  at  base  as  at  apex  (3-6  mm.);  sides  rounded  on  anterior 
two-thirds,  straight  posteriorly;  basal  angles  rectangular;  posterior 
marginal  puncture  placed  at  l^asal  angles  in  lateral  channel. 
Elytra  oval  (10-5  x  63  mm.),  a  little  ampliate  (especially  in  ^) 
behind  middle,  strongly  striate;  lateral  apical  sinuosities  well 
developed,  stronger  in  9  than  in  $\  basal  border  arcuate  on 
posterior  margin,  meeting  lateral  l^order  at  humeral  angles  with- 
out interruption;  interstices  lightly  convex  in  ^,  flat  in  9,  third 
3-  or  4-punctate.  ^  with  anterior  tarsi  not  dilatate  or  squamulose 
beneath.  $  with  one,  9  with  two  setigerous  punctures  on  each 
side  of  apical  ventral  segment.     Length  15-20,  breadth  5-7 '7  mm. 

Hah. — Victorian  Mountains,  Ferntree  Gully  (Sloane). 

I  apply  the  name  N.  pei^onl  to  this  species  because  in  shape  of 
prothorax  and  colour  it  agrees  with  Castelnau's  description  of 
that  species.  It  seems  to  be  the  most  widely  spread  species  in 
the  mountainous  districts  of  Eastern  Victoria,  and  it  is  hardly 
possible  that  it  could  not  have  been  known  to  Castelnau.  It  is 
remarkable  to  find  in  this  species  alone  of  the  genus  Notonomus 
the  anterior  tarsi  in  the  $  neither  dilatate  nor  squamulose 
beneath. 


BY    THOMAS    G.    SLOANE.  26') 

NoTONOMUS  ACCEDENS,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  92;  Ann.  Mus.  Civ.  Geneva,  1874, 
p.  581;  Iieit7'02mtes  2^ristonyc/ioides,  Motsch.  (1),  Bull.  Mosc.  1865, 
ii.  p.  264. 

Elliptical,  depressed.  Black.  Head  smooth,  convex  (3-1  mm. 
across  eyes);  frontal  impressions  well  marked,  diverging 
backwards  ;  eyes  hemispherical,  very  prominent.  Prothorax 
broader  than  long  (3'6  x  4*3  mm.),  subcordate,  widest  before 
middle,  hardly  narrower  at  base  (3  mm.)  than  at  apex  (3-1  mm.); 
disc  depressed;  sides  rounded  on  anterior  two- thirds,  subsinuate 
before  base;  apex  emarginate;  anterior  angles  obtuse,  slightly 
advanced;  base  truncate;  basal  angles  obtuse  but  rather  sharply 
marked,  lateral  border  narrow,  ending  at  basal  angles;  posterior 
marginal  puncture  a  little  before  the  basal  angles,  not  on  border; 
median  line  strongly  impressed,  reaching  almost  to  apex;  lateral 
basal  impressions  wide,  lateral  basal  spaces  flat.  Elytra  rather 
obovate,  widest  behind  middle  (9  x  5-8  mm.),  sides  lightly  rounded, 
rather  sharply  inturned  towards  humeral  angles;  lateral  apical 
sinuosities  very  wide  but  decidedly  marked  ;  humeral  angles 
rounded;  basal  border  meeting  lateral  border  without  any  inter- 
ruption; disc  strongly  striate;  seventh  stria  slightly  impressed; 
interstices  depressed,  third  2-punctate  (often  3-punctate);  ninth 
stria  and  tenth  interstice  feebly  developed.  Intercoxal  declivity 
of  presternum  rounded,  narrow  ;  of  mesosternum  concave. 
Anterior  tarsi  in  ^  lightly  dilatate,  not  squamulose  on  sides  of 
joints  beneath.     Length  14*5-17,  breadth  5-6"3  mm. 

Ilab. — Vic.  :  Upper  Yarra  (French),  Yarragon  and  Otway 
Ranges  (Sloane). 

Allied  to  N.  politulns,  Chaud.,  but  differing  by  the  prothorax 
strongly  angustate  to  the  base,  which  is  not  wider  than  the  aj^ex. 
It  is  a  common  species  in  Southern  Victoria;  the  form  from  the 
Otway  Ranges  has  a  viridescent  tinge  on  the  elytra,  and  might 
be  regarded  as  a  variety.  I  consider  JVeuropates  pristonychoides, 
Motsch.jtobe  synonymous  with  /\^.  accedens,heca,u.se  Motschulsky's 


266  EEVISION  OF  THE  GENUS  NOTONOMUS, 

description  is  applicable  to  it,  while  it  does  not  at  all  suit  the 
species  {N.  angustihasis,  SI.)  to  which  Chaudoir  has  applied  it. 
The  habitat  given  by  Motschulsky  lends  support  to  this  view, 
viz.,  neighbourhood  of  Port  Phillip,  which  is  the  habitat  of  N. 
accedens,  but  not  of  N.  anyicstibasis. 

^OTONOMUS  POLiTULUS,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  86;  Ann.  Mus.  Civ.  Genova,  1874, 
p.  581;  Feronia  (Pterostichus)  tasmanica,  Casteln.,  Trans.  Roy. 
Soc.  Vict.  1868,  p.  210. 

9.  Oval,  depressed.  Black,  shining.  Head  moderate  (3-1  mm. 
across  eyes),  smooth,  convex;  eyes  prominent,  almost  hemispheri- 
cal. Prothorax  broader  than  long  (3-8  x  4*7  mm.),  wider  across 
base  (3-7  mm.)  than  apex  (3-3  mm.),  widest  about  middle,  sub- 
depressed,  lightly  declivous  to  sides,  depressed  towards  middle  of 
base;  sides  lightly  rounded;  apex  lightly  emarginate;  anterior 
angles  obtuse,  not  close  to  head  ;  base  widely  emarginate  in 
middle;  basal  angles  obtuse;  posterior  marginal  puncture  a  little 
before  basal  angles  on  the  explanate  margin,  not  on  border; 
median  line  lightly  impressed;  lateral  basal  impressions  rather 
wide,  long,  shallow,  lateral  basal  spaces  slightly  convex.  Elytra 
ovate  (10-3  X  6  3  mm.),  subdepressed  ;  lateral  apical  sinuosities 
almost  obsolete;  humeral  angles  rounded;  basal  border  joining- 
lateral  border  without  interruption  ;  striae  strongly  impressed, 
seventh  light,  ninth  distinct;  interstices  depressed,  not  convex  on 
apical  declivity,  third  2-punctate  (sometimes  3-punctate),  tenth 
moderately  developed  towards  apex.  Intercoxal  declivity  of 
prosternum  rounded;  of  mesosternum  concave.  Length  17, 
breadth  6*3  mm. 

Hah. — Tasmania. 

Though  N.  politulus  seems  to  have  the  third  interstice  of  the 
elytra  more  often  with  two  than  with  three  punctures,  I  have 
thought  it  more  allied  to  y.  kosciusklanus,  SI.,  and  kindred 
species  than  to  N".  king  I,  Chaud.,  and  other  typical  species  of  the 
group  characterised  by  their  bipunctate  third  elytral  interstice. 


BY    THOMAS    G.    SLOANE.  267 

Another  reason  for  so  placing  it  is  that  the  position  of  the 
posterior  puncture  near  the  extremity  of  the  third  elytral  inter- 
stice seems  to  me  of  more  classificatory  value  in  Notononius  than 
the  mere  number  of  punctures  on  the  interstice  which  is  a 
variable  character  in  some  species.  N.  accedens,  Chaud.,  a  closely 
allied  species,  has  the  punctures  of  the  third  interstice  varying  in 
number  and  placed  as  in  N.  politidus.  It  may  be  noted  that  in 
all  these  species  the  presence  of  the  posterior  puncture  and  its 
position  on  the  interstice  are  constant. 

ISToTONOMUS  TUBERicAUDA,  Bates. 

Cist.  Ent.  1878,  ii.  p.  323. 

From  Bates's  description,  the  place  of  this  species  is  evidently 
near  /V.  politidus.  I  append  Bates's  Latin  diagnosis,  and  would 
quote  from  the  note  he  has  attached  to  it  the  following  : — "  The 
third  [elytral]  interstice  has  two  large  punctures,  the  posterior  of 
which  (near  the  apex),  is  the  centre  of  a  dilatation,  slightly 
elevated  in  the  (J,  but  raised  into  a  prominent  tubercle  in  the 

9-" 

"  ^y.  polltulo  (Chaud.),  affinis,  elongatus,  niger  politus;  thorace 
fere  quadrato,  angulis  posticis  subrectis;  elytris  apice  distincte 
sinuatis,  humeris  baud  dentatis,  supra  fortiter,  simpliciter  striatis, 
interstitiis  prope  apicem  angustioribus,  tertio  excepto  dilatato  et 
in  ^  valde  tuberoso.  Long.  16  mm.  South  or  Central  Tasmania 
(Simson)," 

NOTONOMQS    KOSCIUSKIANUS,   n.Sp. 

Oval.  Head  small,  convex;  prothorax  rounded  on  sides,  wider 
at  base  (3  5  mm.)  than  apex  (3  mm.);  basal  angles  rounded; 
posterior  marginal  puncture  distant  from  base  and  a  little  distant 
from  lateral  border :  elytra  oval,  strongly  striate ;  interstices 
convex,  third  3-  or  4-punctate.     Black. 

Head  3  mm.  across  eyes;  frontal  impressions  hardly  marked; 
eyes  prominent,  hemispherical.  Prothorax  broader  than  long 
(4  X  4*7  mm.),  lightly  convex,  not  depressed  along  middle  of  base; 
sides  strongly  and  evenly  rounded,  a  little  more  gently  narrowed 


2 68  REVISION  OF  THE  GENUS  NOTOyOMUS, 

anteriorly  than  posteriorly ;  apex  lightly  emarginate ;  anterior 
angles  obtuse,  close  to  head;  base  truncate;  basal  angles  not 
marked;  lateral  margins  a  little  explanate  at  basal  angles;  border 
narrow;  median  line  lightly  impressed;  lateral  basal  impressions 
long,  rather  wide;  lateral  basal  spaces  rather  wide  and  slightly 
convex.  Elytra  oval  (10  x  5*7  mm.),  lightly  convex;  sides  rounded; 
lateral  apical  sinuosities  wide,  almost  obsolete;  basal  border  not 
arcuate  on  posterior  margin,  joining  lateral  border  without  the 
slightest  interruption;  lateral  border  narrow;  ninth  stria  well 
marked  on  apical  third ;  tenth  interstice  feebly  de\eloped. 
Intercoxal  declivity  of  prosternum  rounded  ;  of  mesosternum 
lightly  concave.     Length  15-17,  breadth  o"2-5'7  mm. 

^^6.— N.S.W.  :  Mount  Kosciusko  (Fletcher). 

The  affinity  of  this  species  is  to  JV.  politulus,  Chaud.,  from  which 
it  is  readily  distinguished  by  being  narrower,  much  more  convex, 
the  prothorax  more  rounded  on  sides  and  at  basal  angles,  the 
interstices  of  the  elytra  more  convex,  Szc.  In  facies  it  resembles 
JV.  rainbotvi,  SL,  but  is  smaller;  prothorax  wider  at  base,  lateral 
basal  impressions  more  elongate  and  not  approaching  nearly  to 
the  lateral  border;  posterior  marginal  punctures  placed  a  con- 
siderable distance  in  front  of  the  basal  angles  (not  on  the  border, 
but  a  little  away  from  it,  on  the  flattened  marginal  part).  The 
elytra  are  very  similar  to  those  of  iV.  rainbovn.  The  position  and 
number  of  the  punctures  on  the  third  interstice  vary. 

NOTONOMUS    RAINBOWI,  n.sp. 

N.  montanus,  Rainbow,  Rec.  Aust.  Mus.  1899,  iii.  p.  149. 

Elliptical-oval,  robust,  convex.  Head  convex,  eyes  very  promi- 
nent; prothorax  convex,  rounded  on  sides  and  at  basal  angles: 
elytra  convex,  strongly  striate;  interstices  convex,  third  3-  or 
4-punctate;  humeral  angles  not  marked.  Intercoxal  declivity  of 
prosternum  rounded  ;  of  mesosternum  concave.  Black  ;  legs 
piceous. 

Head  smooth,  convex  between  eyes  (4-2  mm.  across  eyes); 
frontal  impressions  light;  eyes  almost  hemispherical,  much  more 
prominent  than  posterior  part  of  orbits — these  protuberant,  sloping 


BY    THOMAS    G.    SLOANE.  269 

sharply  to  head.  Prothorax  broader  than  long  (5-3  x  6  mm.), 
convex,  not  depressed  along  middle  of  base,  a  little  narrower 
(3-9  mm.)  at  base  than  at  apex  (4*2  ram.);  sides  strongly  rounded, 
hardly  more  gently  posteriori}"  than  anteriorly;  apex  truncate: 
anterior  angles  not  marked,  close  to  head;  base  roundly  truncate; 
basal  angles  not  marked;  border  narrow,  thick,  a  little  wider 
towards  base;  posterior  marginal  puncture  on  margin  at  basal 
angle;  median  line  light;  lateral  basal  impressions  wide,  short. 
Elytra  oval  (1-i  x  7-8  mm.),  convex;  sides  rounded;  lateral  apical 
sinuosities  light,  wide;  basal  border  joining  lateral  border  at 
shoulders  without  the  slightest  interruption ;  lateral  border 
narrow;  lateral  channel  giving  off  a  ninth  stria,  and  so  forming 
a  well  developed  tenth  interstice  on  posterior  half  of  elytra. 
Length  23,  breadth  7-8  mm. 

^a6.— N.S.W.  :  Mount  Kosciusko  (Fletcher). 

Closely  allied  to  and  resembling  N".  gippsiensis,  Casteln.,  but 
more  convex  and  differing  by  having  the  prothorax  more  strongly 
and  evenly  rounded  on  the  sides,  the  basal  angles  quite  rounded 
off,  the  posterior  marginal  puncture  placed  on  the  border,  &c. 
This  is  the  species  which  Mr.  Rainbow  described  as  iV.  montamcs, 
but  as  that  name  was  used  in  the  genus  Notonomus  by  Castelnau 
it  is  necessary  to  change  it,  even  although  Castelnau's  name  has 
sunk  to  the  rank  of  a  synonym. 

Notonomus  gippsiensis,  Castelnau. 

Feronia  (Pterostichus)  gippsiensis,  Casteln.,  Trans.  Roy.  Soc. 
Vict.  1868,  p.  208 ;  N.  gippsiensis,  Chaud.,  Ann.  Mus.  Civ. 
Genova,  1874,  p.  577. 

9.  Robust.  Black,  shining ;  legs  piceous-red.  Head  large 
(4-5  mm.  across  eyes);  eyes  (with  orbits)  strongly  protuberant. 
Prothorax  subdepressed,  subcordate  (5-8  x  6*8  mm.) ;  base  and 
apex  of  equal  width  (4*8  mm.);  sides  rounded,  lightly  narrowed 
to  base;  basal  angles  obtuse;  posterior  marginal  puncture  placed 
at  basal  angle  but  not  on  border.  Elytra  oval  (14  x  8-5  mm.), 
subdepressed ;    humeral    angles    rounded ;    basal   border  joining 


270  EEVISION  OF  THE  GENUS  XOTOXO.VUS, 

lateral  border  without  interruption;  strite  strongly  impressed; 
interstices  depressed,  third  4-punctate.  Intercoxal  declivity  of 
prosternum  narrowed  and  rounded  in  middle;  of  mesosternum 
•concave.     Length  25,  breadth  8-5  mm. 

Hab. — Victorian  Mountains. 

The  description  given  is  founded  on  a  specimen  given  to  me  by 
Mr.  C.  French  which  I  compared  with  a  specimen  in  the  Howitt 
Collection  ticketed  Feronia  gippsiensis,  Casteln.,  and  found  iden- 
tical. Specimens  (9)  taken  by  me  at  Marysville,  Victoria,  have 
a  very  faint  metallic  tinge  on  the  prothorax,  and  seem  conspecific 
with  A^.  s.atrapa  of  Chaudoir's  Supplement,  but  are  evidentl}^  not 
the  iV.  satrapa  of  Castelnau. 

NoTONOMUS  SATRAPA,  Castcln.  (var.  *?). 

Feronia  {Pterostichus)  satrapa,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  p.  208;  N.  satrapa,  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874, 
vi.  p.  578. 

Head  large;  prothorax  cordate,  posterior  angles  not  marked: 
elytra  ovate,  widest  about  posterior  third,  a  little  narrowed  to 
base;  third,  fifth  and  seventh  interstices  punctate.  Legs  piceous- 
red;  ^  with  upper  surface  of  a  bronzy  tint;  9  black  with  a  faint 
greenish  tinge  on  upper  surface. 

5.  Head  large  (4-8  mm.  across  eyes);  vertex  convex;  front 
depressed,  declivous  on  each  side  to  eyes;  frontal  impressions 
distinct,  wide;  eyes  (with  orbits)  convex,  protuberant.  Prothorax 
cordate  (5-2  x  6'3  mm.);  sides  rounded,  strongly  narrowed  to  base, 
subsinuate  posteriorly  (at  posterior  marginal  puncture) ;  base 
much  narrower  (3-6  mm.)  than  apex  (4*8  mm.);  border  narrow, 
narrower  at  basal  angles;  median  line  well  marked;  lateral  basal 
impressions  wide,  not  long,  reaching  base ;  posterior  marginal 
puncture  in  lateral  channel,  distant  from  base.  Elytra  lightly 
convex,  oboval  (14x8-5  mm.),  widest  about  posterior  third,  a 
little  narrowed  to  base;  humeral  angles  not  marked;  basal  border 
joining  lateral  border  without  any  interruption;  striae  strongly 
impressed;  interstices  depressed,  third  with  about  five  punctures, 


BY    THOMAS    G.    SLOANE.  271 

fifth  with  two  punctures  on  apical  third  (sometimes  a  third  one 
further  forward),  seventh  with  three  or  four  punctures.  Inter- 
coxal  declivity  of  prosternum  narrowed  and  rounded  in  middle; 
of  mesosternum  concave.  (J  with  prothorax  less  narrowed  to 
base  (apex  4*5,  base  3*8  mm.);  sides  less  ampliate  at  widest  part, 
not  subsinuate  posteriorly  (5-2  x  6  mm.);  elytra  not  ampliate  at 
posterior  third,  not  narrowed  to  base  (13  x  7*2  mm.);  interstices 
convex.     Length  21-5-23,  breadth  7"2-8-5  mm. 

Hab. — Vic.  :  Eastern  Gippsland  (French). 

This  was  one  of  the  species  taken  by  Mr.  C.  French  during  the 
expedition  of  the  Victorian  Field  Naturalists'  Club  to  Mount 
Goonmurk,  in  November,  1889.  It  seems  to  me  to  be  N'.  satrapa, 
Casteln.,  (perhaps  a  variety  differing  by  its  submetallic  lustre  and 
the  presence  of  punctures  on  the  fifth  interstice  of  the  elytra).  I 
cannot  accept  Chaudoir's  treatment  of  ISf.  satrapa  in  his  'Supple- 
ment '  as  satisfactory;  he  evidently  had  not  the  true  Feronia 
satrapa  of  Castelnau  before  him. 

NoTONOMus  ARTHURi,  Sloane. 

Proc.  Linn.  Soc.  N.S.W.  1889  (2),  iv.  p.  1294. 

Elliptical-oval.  Head,  prothorax  and  under  surface  black ; 
elytra  violaceous;  legs  piceous-red.  Head  not  large;  front  strongly 
biimpressed;  eyes  convex,  prominent.  Prothorax  subcordate,  a 
little  narrower  at  base  than  apex ;  posterior  angles  rounded ; 
posterior  marginal  puncture  before  basal  angles  on  inner  side  of 
marginal  channel ;  border  narrow,  hardly  widened  near  base. 
Elytra  oval,  strongly  striate  ;  interstices  convex,  third  3-  or 
4-punctate,  eighth  wider  than  ninth  on  basal  half,  tenth  well 
developed  on  posterior  third  ;  humeral  angles  rounded ;  basal 
border  meeting  lateral  border  without  interruption.  Intercoxal 
declivity  of  prosternum  rounded  and  narrowed  in  middle;  of 
mesosternum  concave.     Length  17-19,  breadth  6'2-7*2  mm. 

Z^a6.— N.S.W.  :  Mount  Wilson  (Olliff),  Wentwqrth  Falls  and 
Mount  Tomah  (Fletcher). 

A  distinct  species,  the  true  affinity  of  which  seems  to  be  towards 
N.  kosciuskianus,  SI.,  and  allies.     In  facies  and  appearance  it  is 


272  EEVISION  OF  THE  GENUS  ^^OTOXOML^S, 

like  iV.  variicollis,  Chaud.,  and  N.  excisipennis,  SI.  ;  from  iV. 
variicollis  it  is  at  once  distinguished  by  the  basal  angles  of  the 
prothorax  not  marked,  and  the  posterior  marginal  puncture  not 
placed  at  the  basal  angle.  N.  excisipennis  is  thoroughly  differen- 
tiated from  F.  arthuri  by  its  unusually  narrow  ninth  elytral 
interstice. 

NOTONOMUS    MUELLERI,  n.sp. 

Oval,  robust.  Head  large ;  prothorax  subquadrate,  lightly 
narrowed  to  base;  basal  angles  marked;  posterior  marginal  punc- 
ture at  extremity  of  lateral  channel  just  before  basal  angle; 
elytra  striate;  interstices  depressed,  third  3-punctate  on  apical 
two-thirds;  humeral  angles  rounded.  Black,  nitid;  legs  piceous- 
red. 

Head  large  [4  mm.  {^)  to  4*3  mm.  (9)  across  eyes],  convex; 
frontal  impressions  weak;  eyes  with  orbits  reniform,  convex, 
much  more  prominent  in  $  than  in  9.  Prothorax  broader  than 
long  (4'5  X  5-6  mm.),  depressed  towards  base,  very  little  narrower 
at  base  (4  mm.)  than  at  apex  (4-2  mm.);  sides  lightly  rounded, 
obliquel}'  (not  roundly)  narrowed  to  base  without  sinuosity;  base 
widely  truncate;  basal  angles  well  marked,  obtuse;  bordernarrow, 
terminating  at  basal  angles ;  median  line  light ;  lateral  basal 
impressions  wide,  shallow,  straight  and  parallel  at  bottom. 
Elytra  truncate-oval  (11-5  x  6*7  mm.), depressed;  base  wide;  lateral 
apical  sinuosities  well  developed,  wide ;  basal  border  joining 
lateral  border  without  any  interruption;  tenth  interstice  feebly 
developed.  Intercoxal  declivity  of  prosternum  rounded;  of  meso- 
sternum  lightly  concave;  metasternum  not  channelled  to  apex. 
Length  17-21,  breadth  6-7-8 mm. 

Hah. — N.S. W.  :  Mount  Kosciusko  (specimens  from  Mr.  W.  W. 
Froggatt). 

A  distinct  species;  in  facies  it  resembles  N.  i^eroni,  Casteln., 
but  differs  by  colour,  prothorax  not  sinuate  on  sides  near  base 
and  with  basal  angles  obtuse;  ^  with  three  basal  joints  of  anterior 
tarsi  broadly  dilatate  and  squamulose  beneath,  tfec.  Its  affinity 
seems  towards  N.  gippsiensis,  Casteln. 


BY    THOMAS    G.    SLOANE.  273 

NOTONOMUS    MACOYI,  n.sp. 

(J.  Elongate-oval,  conv^ex.  Head  moderate;  prothorax  broader 
than  long,  narrowed  to  base;  posterior  angles  marked,  but  wide  : 
elytra  oval,  strongly  striate;  humeral  angles  rounded;  interstices 
convex,  third  4-punctate.  Black;  under  surface  piceous-black, 
legs  and  antennse  piceous-red. 

Head  rather  large  (4-1  mm.  across  eyes);  frontal  impressions 
distinct,  short,  divergent  backwards;  e3^es  protuberant,  postocular 
part  of  orbits  large.  Prothorax  broader  than  long  (4  x  4*7  mm.), 
widest  a  little  before  middle,  roundly  narrowed  to  base;  apex 
(3-5  mm.)  wider  than  base  (3-2  mm.);  sides  lightly  rounded,  falling 
on  base  in  a  gentle  oblique  curve;  base  narrow,  emarginate  in 
middle;  basal  angles  lightly  marked,  obtuse;  posterior  marginal 
puncture  placed  on  border  at  basal  angles;  lateral  border  narrow, 
wider  towards  base,  passing  round  basal  angle  on  each  side  of 
base:  median  line  strongly  impressed;  lateral  basal  impressions 
wide,  deep,  parallel,  linear  at  bottom.  Elytra  oval  (9x5-6  mm.), 
convex;  lateral  apical  sinuosities  wide,  but  decided;  basal  border 
joining  lateral  border  without  interruption;  ninth  stria  and  tenth 
interstice  well  developed  towards  apex;  interstices  convex,  not 
carinate  on  apical  declivity,  eighth  wider  than  ninth  on  basal 
half,  anterior  puncture  of  third  placed  about  basal  fifth.  Inter- 
coxal  declivity  of  prosternum  narrow  and  rounded  in  middle;  of 
mesosternum  very  lightly  and  widely  concave.  Length  16-7, 
breadth  5 -4-5 -6  mm. 

Hah. — Vic.  :  Marys  villa  (Sloane). 

In  facies  resembling  N.  kiyigi,  Chaud.,  but  differing  by  its  more 
transverse  prothorax,  third  interstice  of  elytra  4-punctate,  &c. 
Two  specimens  {$)  from  the  same  locality  are  in  my  collection;  one 
has  the  prothorax  more  evenly  and  more  lightly  rounded  on  the 
sides,  and  therefore  less  ampliate  (3-9  x  4-5  mm.)  than  the  type. 

NOTONOMUS    BESTI,  n.Sp. 

Elliptical-oval.     Head  rather  small,  eyes  very  prominent;  pro- 
thorax subquadrate,  lightly  narrowed  to  base;  basal  angles  obtuse; 
18 


tl  BR  A  R 


274  REVISION  OF  THE  GENUS  NOTONOMITS, 

posterior  marginal  puncture  on  border  at  basal  angle  :  elytra  oval; 
humeral  angles  rounded;  striae  deep;  interstices  convex,  third  5- 
or  6-punctate.  Dark  metallic  copper  colour  with  a  rather  purple 
tinge,  head  more  brassy,  margin  of  elytra  dark;  under  surface 
black;  legs  piceous,  excepting  anterior  pair  piceous-red. 

Head  small  (3-5  mm.  across  eyes);  frontal  impressions  distinct, 
narrow,  divergent  posteriori}^;  eyes  almost  hemispherical,  much 
more  prominent  than  orbits;  posterior  part  of  orbits  protuberant, 
sloping  sharply  to  head.  Prothorax  broader  than  long  (44  x 
4-8  mm.),  widest  before  middle,  a  little  wider  across  base  (3 -6 mm.) 
than  apex  (3-4  mm.)  ;  sides  lightly  rounded,  gently  narrowed 
without  sinuosity  to  base;  basal  angles  obtuse  but  marked;  base 
lightly  emarginate;  lateral  border  narrow  anteriorly,  much  wider 
and  reflexed  near  base,  thickened  to  receive  posterior  marginal 
puncture  at  basal  angles;  median  line  well  marked;  lateral  basal 
impressions  wide,  deep,  linear  at  bottom.  El3'tra  oval  (9-5  x 
5*8  mm.),  lightly  convex;  lateral  sinuosities  of  apex  well  marked; 
basal  border  hardly  raised  at  humeral  angles,  joining  lateral  border 
with  just  the  slightest  interruption;  ninth  stria  and  tenth  inter- 
stice well  developed.  Intercoxal  declivity  of  prosternum  narrow 
and  rounded  in  middle;  of  mesosternum  lightly  concave.  Length 
18-5,  breadth  5*8  mm. 

Hab. — Victorian  Mountains,  Track  from  "Wood's  Point  to  the 
Yarra  (Best;  Coll.  Sloane). 

Allied  to  JV.  macoyi,  SL,  but  differing  by  colour;  head  smaller, 
eyes  more  prominent.  Compared  with  JV.  gipjMandicus,  Casteln., 
(as  identified  by  me)  the  colour  is  different,  the  eyes  more 
prominent;  the  prothorax  more  transverse,  with  posterior  angles 
less  marked,  &c. 

Note  (1). — A  specimen  (9)  given  to  me  by  Mr.  C.  French, 
ticketed  "Victoria,"  is  larger  (19-5x7  mm.);  head  larger;  pro- 
thorax more  transverse  (4-5  x  5-4  mm.),  more  roundly  ampliate  at 
widest  part;  elytra  more  convex,  interstices  very  lightly  convex 
except  at  apex.  Elytra  blackish-purple  (not  metallic);  prothorax 
and  head  gilded  copper  colour,  otherwise  agreeing  in  colour  with 


BY    THOMAS    G.    SLOANE.  275 

the  typical  form.     It  seems  too  near  iV.  besti  to  be  considered  a 
distinct  species. 

J^ote  (2). — A  specimen  {^),  from  the  northern  slopes  of  the 
Buflfalo  Mountain,  near  Porpunkah,  is  in  my  collection,  which, 
though  closely  resembling  JV.  besti,  is  perhaps  a  distinct  species. 
The  differences  I  note  are: — Colour  brassy;  prothorax  more 
rounded  on  sides,  more  strongly  narrowed  to  base,  basal  angles  a 
little  more  marked,  sides  hardly  subsinuate  before  base;  basal 
border  of  elytra  joining  lateral  border  without  the  slightest 
interruption;  mesosternum  not  concave  between  coxae. 

NOTONOMUS    HOWITTI,   n.Sp. 

9.  Oval,  robust.  Head  not  large ;  prothorax  broader  than 
long,  a  little  narrowed  without  sinuosity  to  base ;  posterior 
marginal  puncture  at  basal  angle  outside  lateral  channel :  elytra 
strongly  striate;  humeral  angles  rounded;  interstices  depressed, 
third  3-punctate,  eighth  wider  than  ninth.     Black. 

Head  convex  (3  mm.  across  eyes);  frontal  impressions  short, 
distinct,  divergent  posteriorly;  eyes  convex,  prominent,  strongly 
inclosed  at  base.  Prothorax  broader  than  long  (3-5  x  4*2  mm.); 
base  and  apex  of  equal  width  (3  mm.);  sides  lightly  rounded, 
meeting  base  in  a  gentle  curve;  base  very  lightly  emarginate  in 
middle,  almost  truncate  on  each  side  (curving  forward  very 
slightly  to  basal  angles);  basal  angles  well  marked,  obtuse;  lateral 
border  narrow,  thick,  a  little  wider  and  more  reflexed  towards 
base,  passing  round  basal  angles,  thickened  at  basal  angle  to 
receive  posterior  marginal  puncture;  median  line  lightly  impressed; 
lateral  basal  impressions  deep,  not  elongate,  space  between  them 
rather  depressed.  Elytra  oval  (8*5  x  5  mm.),  convex ;  sides 
rounded  ;  lateral  apical  sinuosities  lightly  marked  ;  basal  and 
lateral  borders  meeting  at  humeral  angle  without  the  slightest 
interruption ;  ninth  stria  and  tenth  interstice  well  developed. 
Prosternum  bordered  along  anterior  margin;  intercoxal  declivity 
rounded  and  narrowed  in  middle;  mesosternum  lightly  concave 
between  coxae.     Length  14,  breadth  5  mm. 


276  REVISION  OF  THE  GENUS  NOTOXOMUS, 

Hab.—Yic.  :  Wood's  Point  to  Yarra  Track  (D.  Best:  ColL 
Sloane;  unique). 

Kesembling  N.  rufipaljns,  Casteln.,  but  very  distinct  :  by 
head  smaller ;  prothorax  with  border  wider  and  not  the  least 
sinuate  posteriorly,  the  posterior  marginal  puncture  not  in  lateral 
channel  but  on  the  widened  border  at  each  basal  angle;  elytra 
with  interstices  depressed,  eighth  wider  than  ninth;  mesosternum 
concave  between  coxae,  &c. 

NOTONOMUS   BAKEWELLI,  n.sp. 

Elongate-oval,  robust.  Head  large ;  prothorax  subcordate; 
basal  angles  subrectangular :  elytra  oval,  strongly  striate;  inter- 
stices convex,  third  4-punctate,  eighth  narrow  and  convex 
throughout  its  whole  length.  Black,  a  faint  bronzy  tinge  on 
prothorax  near  basal  impressions,  a  ver}^  faint  blackish-green 
tint  on  elytra. 

Head  large  (3*8  mm.  across  eyes),  wide  across  occiput,  convex; 
frontal  striae  strongly  impressed,  wide,  extending  to  cl3'peus;  eyes 
convex,  not  prominent;  posterior  part  of  orbits  as  long  as  eyes, 
sloping  gently  to  head.  Prothorax  broader  than  long(4-l  x  5  mm.), 
wider  at  apex  (3-7  mm.)  than  at  base  (3'3  mm.);  sides  lightly 
rounded  on  anterior  three-fourths,  lightly  sinuate  before  base; 
apex  emarginate  in  middle;  base  truncate;  basal  angles  strongly 
marked,  obtuse  at  summit;  posterior  marginal  puncture  placed  at 
basal  angle  outside  lateral  channel;  border  narrow,  thick,  reflexed, 
thickened  at  basal  angle  to  receive  marginal  puncture.  Elytra  oval 
(10  X  6  mm.),  rather  depressed  on  disc;  sides  lightly  rounded,  lateral 
sinuosities  of  apex  almost  obsolete;  basal  border  joining  lateral 
border  at  humeral  angles  without  interruption;  interstices  convex, 
tliird,  fifth  and  seventh  wider  than  others,  eighth  narrow,  about 
same  width  as  ninth  on  basal  half,  tenth  interstice  obsolete 
except  at  apical  extremity.  Prosternum  longitudinally  channelled 
towards  base,  intercoxal  declivity  rounded  and  rather  wide  in 
middle.     Length  18,  breadth  6  mm. 

Hah  — Victoria  (Coll.  Sloane;  unique). 


BY    THOMAS    G.    SLOANE.  277 

I  am  indebted  to  Mr.  C.  French  for  this  fine  species,  which  in 
general  appearance  resembles  i\^.  variicollis,  Chaud.,  but  differs 
by  head  larger  and  wider  across  occiput,  ej^es  less  prominent, 
postocular  part  of  orbits  longer  and  rising  more  gently  from 
head,  eyes  more  distant  from  buccal  fissure,  the  inter\ening 
space  more  bulged;  prothorax  shorter,  more  strongly  narrowed  to 
base  ;  border  thicker  and  widened  at  basal  angles  to  receive 
posterior  marginal  puncture;  elytra  less  convex,  basal  border  not 
raised  above  the  lateral  at  point  of  junction,  lateral  apical  sinuosi- 
ties much  weaker,  ninth  interstice  narrower;  intercoxal  part  of 
prosternum  strongly  channelled,  &lq. 

NOTONOMUS    PLETCHERI,    n.Sp. 

Robust,  convex.  Head  large ;  prothorax  subcordate,  with 
posterior  marginal  puncture  at  basal  angle:  elytra  oval,  strongly 
striate;  interstices  convex,  third  3-  or  4-punctate,  tenth  moderately 
developed;  humeral  angles  dentate.  Upper  surface  rather  metallic; 
prothorax  nitid,  greenish,  or  of  a  beautiful  coppery  colour;  head 
of  same  colour  as  prothorax  (but  fainter);  elytra  greenish,  often 
flushed  with  a  purple  or  cupreous  tinge — ninth  interstice  and 
margin  usually  a  dark  copper  colour;  under  surface  and  legs 
piceous-black. 

Head  large  (4-4-5  mm.  across  eyes);  frontal  impressions  distinct; 
eyes  convex,  rather  prominent;  postocular  part  of  orbits  large. 
Prothorax  broader  than  long  (5  x  5*9  mm.),  a  little  narrower  at 
base  (3-9  mm.)  than  at  apex  (4  mm.)  ;  sides  rounded,  curving 
gently  to  base  without  sinuosity;  basal  angles  decidedly  marked, 
obtuse  at  summit;  border  wide,  reflexed,  thickened  at  basal  angles 
to  receive  posterior  marginal  puncture;  median  line  deep;  lateral 
basal  impressions  deep,  reaching  base  in  full  depth.  Elytra 
truncate-oval  (11x6-8  mm.),  convex,  strongly  rounded;  lateral 
apical  sinuosities  deep,  wide;  eighth  interstice  wider  than  ninth 
on  basal  half;  basal  border  strongly  raised  and  obtusely  dentate 
at  humeral  angles.  Intercoxal  declivity  of  prosternum  narrow 
and  rounded  in  middle;  of  mesosternum  lightly  concave.      9  with 


278  KEVISION  OF  THE  GENUS  NOTOXOMUS, 

head  larger  than  ^,  and  with  lateral  curve  of  prothorax  usually 
more  oblique  posterior^.     Length  17-22,  breadth  5-7-7'5  mm. 

/{ab. — N.S.W.  :  Springwood  (Sloane ;  common);  Sydney 
(Froggatt,  Lea,  Sloane;  rare);  Thornleigh  (Froggatt). 

iV.  Jietcheri  has  the  facies  of  N.  ceneomicans,  Chaud.,  and 
resembles  that  species  in  the  wide  borders  of  prothorax  and  elytra, 
and  the  form  of  the  lateral  basal  impressions  of  the  prothorax,  but 
may  be  distinguished  at  once  by  having  the  intercoxal  declivit}'-  of 
the  prosternum  narrow  and  rounded — not  flat  and  margined 
laterally.  It  is  more  nearly  allied  to  N.  variicollis,  Chaud.,  from 
which  it  differs  by  its  less  elongate  form;  head  larger;  prothorax 
much  wider,  more  widely  margined;  elytra  wider,  more  rounded  on 
sides,  humeral  angles  strongly  dentate,  lateral  border  wider  and 
more  reflexed. 

NOTONOMUS   STRZELECKIANUS,  n.Sp. 

Var.  (?)  JV.  (Plerostichus)  j^^'^Uipsii,  Casteln.,  Trans.  Roy.  Soc. 
Vict.  1868,  p.  212. 

Elongate-oval,  robust.  Head  large;  prothorax  subquadrate,  a 
little  narrowed  to  base;  basal  angles  obtuse,  but  marked  (some- 
times almost  rectangular);  posterior  marginal  puncture  distant 
from  base,  in  lateral  channel :  elytra  oval,  convex,  strongly  striate; 
interstices  convex,  third  5-punctate.  9  usually  with  six  setigerous 
punctures  near  apical  margin  of  last  ventral  segment.  Head 
black,  with  vertex  bronzy;  prothorax  of  a  dark  coppery-bronze, 
usually  lighter  and  more  brassy  laterally  ;  elytra  dark,  of  a 
bronzy  or  purplish  tint;  legs  piceous. 

Head  large  (3-3  mm.  across  eyes);  frontal  impressions  strongly 
marked,  wide,  parallel;  eyes  convex,  moderately  prominent,  post- 
ocular  part  of  orbits  nearly  the  length  of  eyes,  rising  gently  from 
head  ;  prothorax  broader  than  long  (4  1  x  4*7  mm.)  ;  lightly 
narrowed  to  base,  wider  across  base  (3-6  mm.)  than  apex  (3*3  mm.); 
sides  lightly  rounded,  gently  and  obliquely  narrowed  to  base 
without  sinuosity ;  border  wide  and  strongly  reflexed  behind 
anterior  marginal  puncture;  median  line  well  marked;  lateral 
basal  impressions  deep.      Elytra  truncate-oval   (9-5  x  54  mm.), 


BY    THOMAS    G.    SLOANE.  279 

lightly  rounded  on  sides;  lateral  apical  sinuosities  well  developed; 
basal  border  strongly  raised  and  dentate  at  humeral  angles;  tenth 
interstice  feebly  developed  near  apex.  Intercoxal  declivity  of 
prosternum  narrow,,  and  rounded  in  middle  ;  of  mesosternum 
concave.     Length  15-17'5;  breadth  5-5-8  mm. 

Hah. — Vic.  :  Baw  Baw  Mountain  (sent  to  me  by  Mr.  C. 
French). 

The  following  features  in  combination  distinguish  this  species: 
— {a)  The  posterior  marginal  puncture  of  prothorax  distant  from 
basal  angles;  (6)  the  elytra  with  strongly  dentate  humeral  angles, 
and  pluripunctate  third  interstice.  These  characters  associate  it 
with  N.  eques,  Casteln.,  from  which  it  differs  by  its  smaller  size, 
more  convex  elytra,  prothorax  more  strongly  narrowed  to  base, 
prosternum  with  intercoxal  declivity  rounded,  mesosternum 
decidedly  concave  between  coxae.  The  third  elytral  interstice  is 
wider  than  the  others,  and  has  the  posterior  puncture  placed  on 
its  outer  side,  which  is  swollen  just  at  this  puncture, 

N.  (Pterostichus)  phillipsi,  Casteln.  This  name  is  too  near  iV. 
philipjn,  Newm.,  a  previously  described  species,  though  now  a 
synonym.  The  treatment  of  iV^.  phillipsi  by  both  Castelnau  and 
Chaudoir  is  unsatisfactory.  Chaudoir's  note  suggests  to  my  mind 
that  the  specimen  ((J)  which  he  found  in  the  Castelnau  collection 
was  not  that  which  had  served  Castelnau  for  his  description, 
where  no  mention  is  made  of  the  green  border  of  the  elytra  noted 
by  Chaudoir,  and  the  measurement  given  is  too  large  for  the 
specimen  Chaudoir  examined.  A  specimen  ((J)  of  the  size  and 
colour  of  that  described  by  Chaudoir  as  J^,  phillipsi,  Casteln., 
has  been  given  to  me  by  Mr.  C.  French,  who  took  it  during  the 
trip  of  the  Victorian  Field  Naturalists'  Club  to  Eastern  Gippsland 
in  the  year  1889;  this  form  differs  from  typical  specimens  of  J\^. 
strzeleckianus  by  its  smaller  head;  elytra  with  a  green  margin, 
interstices  a  little  more  convex,  third  not  swollen  where  the 
posterior  puncture  is  placed — this  punture,  too,  is  not  on  the  side 
of  the  interstice  but  in  the  middle.  I  hardly  think  this  form 
can  be  considered  as  specifically  distinct  from  iV.  strzeleckianus, 
so  have  placed  it  under  that  species,  at  least  provisionally. 


280  REVISION  OF  THE  GENUS  NOTOXOMUS, 

NOTONOMUS    FROGGATTI,    11. sp. 

(J.  Oval,  depressed.  Head  small;  prothorax  subquadrate,  much 
wider  at  base  (3'7  mm.)  than  at  apex  (3-1  mm.);  posterior  angles 
obtuse;  posterior  marginal  puncture  distant  from  basal  angle  : 
elytra  rather  depressed,  strongly  striate;  interstices  convex,  third 
5-punctate,  tenth  feebly  developed  near  apex.  Black,  with  faint 
greenish  reflections  near  lateral  basal  impressions  of  prothorax; 
lateral  channel  of  elytra  greenish;  legs  piceous-red. 

Head  small  (3  mm.  across  eyes),  transversely  impressed  on  each 
side  above  base  of  eyes;  front  depressed,  distinctly  biimpressed. 
Prothorax  broader  than  long  (4*2  x  4*9  mm.);  sides  lightly 
rounded,  greatly  narrowed  in  a  gentle  curve  to  apex,  lightly  and 
obliquely  narrowed  to  base;  basal  angles  wide,  obtuse,  a  little 
marked  ;  border  wide  and  strongly  reflexed  behind  anterior 
marginal  puncture;  median  line  lightly  impressed;  lateral  basal 
impressions  narrow,  short,  shallow,  not  reaching  base  in  full 
depth.  Elytra  truncate-oval  (9-3  x  5 -6 mm.), lightly  convex,  rather 
depressed  on  disc;  sides  lightly  rounded,  hardly  narrowed  to  base; 
lateral  sinuosities  of  apex  well  developed,  wide ;  basal  border 
strongly  raised  and  dentate  at  humeral  angles.  Intercoxal 
declivity  of  prosternum  narrow,  rounded  in  middle;  of  mesoster- 
num  deepl}''  concave.  Posterior  femora  strongly  dilatate  in  middle 
(at  apex  of  trochanters).     Length  16,  breadth  5*6  mm. 

Hab. — N.S.W.  :  Mount  Kosciusko  (given  to  me  by  Mr.  W. 
W.  Froggatt). 

Allied  to  N.  strzeleckianus,  SI.,  but  differing  by  colour,  smaller 
head;  prothorax  more  transverse,  narrower  at  apex,  much  wider 
at  base;  elytra  more  depressed,  lateral  border  more  widely  reflexed 
at  base;  posterior  femora  more  dilatate,  with  lower  side  swollen 
in  middle,  &c. 

NoTONOMUs  BASSi,  n.sp. 

(J.  Oval,  rather  elongate,  subdepressed.  Head  dark  sub- 
metallic-green;  prothorax  of  a  gilt  cupreous  colour;  elytra  purple- 
black  on  disc  and  border,  metallic-purple  towards  sides;  under 
.surface  black;  legs  piceous. 


BY    THOMAS    G.    SLOAN E.  281 

Head  small  (2-6  mm.  across  eyes),  convex  ;  eyes  prominent. 
Prothorax  hardly  broader  than  long  (3-6  x  3*7  mm.),  widest  before 
middle;  base  and  apex  narrow,  of  equal  width  (2-6  mm.);  sides 
lightly  rounded  anteriorly,  decidedly  and  obliquely  narrowed  to 
base ;  anterior  angles  close  to  head  ;  base  lightl}'  and  widely 
emarginate  in  middle,  truncate  on  each  side;  basal  angles  sharply 
marked,  almost  rectangular;  posterior  marginal  puncture  at  basal 
angles  in  lateral  channel;  lateral  basal  impressions  elongate;  lateral 
border  fine,  narrow,  obsoletely  subsinuate  just  before  basal  angles. 
Elytra  truncate-oval  (8-2x5  mm.),  widest  behind  middle,  a  little 
narrowed  to  base;  lateral  apical  sinuosities  well  developed,  wide; 
basal  border  with  posterior  margin  evenly  and  lightly  arcuate; 
humeral  angles  lightly  raised  (not  dentiform)  at  point  of  junction 
of  basal  and  lateral  borders;  strise  deep;  interstices  convex, 
third  4-punctate,  eighth  wide  and  depressed  on  basal  half,  ninth 
with  umbilicate  punctures  wide  apart  on  sides,  tenth  very  short, 
only  developed  near  extremity.  Intercoxal  declivity  of  proster- 
num  rounded;  of  mesosternum  concave.  Length  14,  breadth 
5  mm. 

Hah. — Yic.  :  Yarragon  (Sloanej. 

N.  bassi  differs  from  N.  gippslanclicus  (as  identified  by  me)  by 
colour;  facies  less  elongate,  less  parallel,  less  convex;  prothorax 
more  ampliate  at  widest  part,  with  basal  angles  more  sharply 
marked;  elytra  with  basal  border  more  raised  above  the  lateral 
border  at  point  of  junction,  and  with  the  inner  humeral  angle 
acute  and  far  more  sharply  marked. 

One  of  my  Yarragon  specimens  {$)  has  the  prothorax  evidently 
wider  (3*7  x  4  mm.),  with  base  wider  (2-9 mm.)  than  apex  (2  8 mm.), 
but  presents  no  other  differences  from  the  description  given 
above. 

NoTONOMUS  GiPPSLANDicus,  Castelnau  (?). 

Feronia  (Pterostichus)  gippslandica,  Casteln.,  Trans.  Roy.  Soc. 
Yict.  1868,  viii.  p.  211;  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874, 
vi.  p.  581. 


282  EEVISION  OF  THE  GENUS  XOTONOMUS, 

^.  Elongate-oval.  Green,  nitid ;  border  of  prothorax  and 
elytra  black,  suture  and  base  of  elytra  blackish;  under  surface 
and  legs  black,  tibiae  piceous. 

Head  small  (3  mm.  across  eyes),  vertex  transversely  impressed; 
eyes  prominent.  Prothorax  about  as  long  as  wide  (4-2  x  4*2  mm.), 
lightly  narrowed  to  base;  apex  truncate,  hardly  as  wide  (2'7  mm.) 
as  base  (2-8  mm.) ;  sides  lightly  rounded,  obliquely  narrowed 
without  sinuosity  to  base;  basal  angles  well  marked,  obtuse  at 
summit;  posterior  marginal  puncture  in  lateral  channel  at  basal 
angle ;  border  narrow;  median  line  lightly  impressed ;  lateral 
basal  impressions  elongate,  deep.  Elytra  oval  (9-2  x  5-5  mm.), 
lightly  convex;  sides  lightly  rounded;  lateral  sinuosities  of  apex 
strongly  developed ;  shoulders  rounded  ;  basal  border  a  little 
raised  above  lateral  border  at  humeral  angles  (these  not  the  least 
dentiform) ;  striae  deep ;  ninth  distinct  near  apex ;  interstices 
lightly  convex,  third  3-  or  4-punctate,  tenth  interstice  only 
shortly  developed  near  apex.  Intercoxal  declivity  of  prosternum 
rounded;  of  mesosternum  concave.     Length  16,  breadth  5-5  mm» 

Hab. — Vic.  :  Fernshaw  (Sloane),  Lillydale  (W.  Kershaw). 

I  ascribe  the  name  N.  gippslaiidicus,  Casteln.,  to  this  species 
from  the  applicability  of  the  original  description  to  it.  Chaudoir 
thought  it  only  a  local  variety  of  N.  opulentus,  Casteln.,  but  if  I 
am  right  in  my  identification  of  N.  opulentus  the  species  are 
quite  distinct. 

]}^ote. — A  specimen  (9)  from  the  mountains  at  the  source  of 
the  Ovens  River,  Victoria,  has  been  sent  to  me  by  the  Rev.  Thos. 
Blackburn.  Compared  with  the  species  I  take  to  be  A^.  gipps- 
landicus,  Casteln.,  it  is  duller  coloured;  prothorax  smaller  and 
narrower  (3-6  x  3*6  mm,);  elytra  more  oval  (8*5  x  5-3  mm.),  and, 
proportionately  to  the  prothorax,  wider ;  humeral  angles  still 
more  obsolete.  Length  15,  breadth  5*3  mm.  An  examination  of 
a  number  of  specimens  of  these  two  forms  would  be  required  to 
determine  if  each  is  entitled  to  rank  as  a  species. 


BY    THOMAS    G.    SLOANE.  28^ 

NoTONOMUS  VARiicoLLis,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  86;  Aim.  Mus.  Civ.  Geneva,  1874,  vi. 
p.  577  :  Feronia  (Pterostichus)  comes,  Casteln.,  Trans.  Roy.  Soc. 
Vict.  1868,  viii.  p.  207. 

Ellongate-oval,  robust,  convex.  Head,  prothorax,  under  surface 
and  legs  black,  shining;  elytra  purple  or  bronzy-green,  suture  and 
margin  black. 

Head  moderate  (3-6  mm.  across  eyes);  frontal  impressions  well 
marked.  Prothorax  a  little  broader  than  long  (4'7  x  5  mm.), 
very  lightly  narrowed  to  base ;  sides  lightly  rounded,  usually 
hardly  subsinuate  before  base;  base  usually  a  little  wider  (4  mm.) 
than  apex  (3-6  mm.);  basal  angles  well  marked,  subrectangular 
(obtuse  at  summit);  posterior  marginal  puncture  just  free  from 
border  at  basal  angle  ;  border  fine,  narrow.  Elytra  oval 
(10*5  X  6  mm.);  strongly  striate;  interstices  convex,  third  4-  or 
5-punctate.     Length  16-21,  breadth  5 -5-7 '5  mm. 

Hah. — N.S.W.  :  Bulli,  Wollongong,  Kiama,  Burrawang 
(Sloane). ' 

I  place  the  following  forms  as  varieties  : — 

Var.  A.     Wholly  black,  basal  angles  of  prothorax  obtuse. 

Hab. — Sassafras  (given  to  me  by  the  late  A.  S.  Olliff). 

Var.  B.     Wholly  black,  head  a  little  larger. 

Hab. — Jervis  Bay  (Fletcher). 

NoTONOMus  ANGUSTiBASis,  Sloane. 

Notonomus  pristonychoides,  Chaud.,  {nee  Motschulsky),  Ann. 
Mus.  Civ.  Genova,  1874,  vi.  p.  588. 

Elongate,  robust.  Black,  elytra  dark  purple.  Head  long, 
narrow  (3*3  mm.  across  eyes),  cylindrical  ;  antennae  long, 
slender.  Prothorax  longer  than  broad  (5  x  4*6  mm.),  of  equal 
width  at  base  and  apex  (3*3  mm.),  widest  before  middle, 
strongly  narrowed  and  sinuate  on  sides  posteriorly;  sides  lightly 
rounded  on  anterior  two- thirds,  lightly  narrowed  to  apex,  meeting- 
base  at   right    angles ;    anterior   angles    close    to   head,   obtuse 


'284  REVISION  OF  THE  GENUS  yOTOXOMUS, 

bordered ;  base  truncate,  sloping  forward  a  little  on  each  side 
from  basal  impressions  to  angles;  basal  angles  rectangular,  obtuse 
at  summit ;  border  narrow,  thick,  reflexed,  not  passing  basal 
angles;  posterior  marginal  puncture  distant  from  base  in  lateral 
channel;  lateral  basal  impressions  long,  narrow,  turning  lightly 
outward  anteriorly.  Elytra  narrowly  oval  (11  x  6*5  mm.);  base 
narrow;  sides  rounded;  lateral  apical  sinuosities  strong;  basal 
border  strongly  raised  and  obtusely  prominent  at  humeral  angles, 
posterior  margin  forming  an  even  curve  between  humeral  angles; 
lateral  border  narrow,  reflexed,  very  narrow  near  base  ;  striae 
deep;  interstices  convex,  third  2-punctate,  ninth  convex  towards 
base,  tenth  not  developed.  Intercoxal  declivity  of  prosternum 
narrow  and  a  little  rounded  in  middle;  of  mesosternum  very 
lightly  and  widely  concave.  Legs  long;  posterior  femora  long, 
narrow.     Length  17 "5-2 1-5,  breadth  5 -5-7 "3  mm. 

Hah. — N.S.W.  :  Dunoon,  Richmond  River  (Helms),  Ourimbah 
and  Narrara  (Sloane,  in  Juty). 

Yar.  ?  Black  ;  wider ;  prothorax  not  longer  thp-n  broad 
{4-7  X  4-8  mm.);  elytra  wider  and  more  oval  (10*7  x  6-7  mm.). 
Length  19  mm.     This  is  perhaps  a  distinct  species. 

Hab. — Q.  :  Wallangarra  (W.  Kershaw). 

The  species  to  which  I  have  given  the  name  iV.  angustibasis 
is  an  isolated  and  very  distinct  species.  It  seems,  from 
Chaudoir's  description,  to  be  that  which  he  regarded  as 
Neuropates  pi^istofiychoides,  Motsch.,  and  I  have  seen  it  in  the 
Macleay  Collection  under  the  name  iV.  tvilcoxi,  Casteln.,  l)ut  it 
certainly  does  not  agree  with  Motschulsky's  description  of  Neuro- 
pates pristonyclwides,  nor  with  Castelnau's  description  of  N. 
wUcoxi.  To  my  mind  Motschulsky's  description  of  JV.  pristony- 
choides  is  not  applicable  to  any  form  of  N.  angustibasis  which  I 
have  seen,  nor  have  I  seen  any  specimen  so  small  as  7  x  2J  lines, 
nor  with  three  punctures  on  the  third  interstice  of  the  elytra; 
but  I  identify  without  misgivings  as  JV.  pristonychoides,  Motsch., 
a  Victorian  species  which  I  unhesitatingly  refer  to  N.  accedens, 
Chaud.,  {vide  supra,  p.  265). 


BY    THOMAS    G.    SLOANE.  285 

The  description  given  above  is  founded  on  a  specimen  ((J)from 
Dunoon.  I  took  it  plentifully  at  Ourimbah  and  Narrara;  the 
form  from  these  localities  is  more  elongate  and  more  convex, 
with  prothorax  and  elytra  proportionately  more  elongate  and  less 
ampliate  at  widest  part  than  specimens  from  Dunoon,  but  I  have 
not  thought  it  sufficient!}'-  distinct  to  receive  a  separate  name. 

NOTONOMUS    EXCISIPENNIS,  n.Sp. 

Elliptical-oval.  Head  moderate  ;  prothorax  subcordate  ; 
posterior  angles  rounded;  posterior  marginal  puncture  a  little 
before  base  in  lateral  channel  :  elytra  truncate-oval,  strongly 
striate;  interstices  strongly  convex,  third  2-punctate,  eighth  ver}^ 
narrow.  Under  surface  and  legs  piceous-black ;  head  black, 
sometimes  with  a  bronzy  reflection;  prothorax  metallic-purple, 
brighter  towards  sides,  nearly  black  on  disc;  elytra  of  a  dark 
purple  tint,  often  viridescent  towards  suture;  purple  on  lateral 
declivities,  more  obscure  near  lateral  margins. 

^.  Head  rather  large  (3-3  mm.  across  eyes);  front  a  little 
depressed,  lightly  biimpressed ;  eyes  convex,  prominent.  Pro- 
thorax broader  than  long  (3-8  x  4-7  mm.),  a  little  narrowed  to 
base,  hardl}'  narrower  at  base  (3-1  mm.)  than  apex  (3.2  mm.); 
sides  lightly  rounded,  not  sinuate  before  base ;  apex  truncate, 
angles  widely  obtuse;  base  truncate,  rounded  on  each  side  near 
angles,  these  not  marked;  lateral  border  line,  narrow,  continued 
evenly  round  basal  angles  to  middle  of  lateral  basal  impressions; 
median  line  strongly  impressed;  lateral  basal  impressions  strongly 
impressed,  parallel,  not  long.  Elytra  truncate-oval  (9*5  x  5 -5 mm.), 
lightly  rounded  on  sides;  humeral  angles  rounded;  basal  border 
meeting  lateral  border  without  any  interruption  at  humeral  angles; 
lateral  sinuosities  of  apex  strongly  developed.  Intercoxal  declivity 
of  prosternum  rounded  ;  of  mesosternum  concave.  Anterior 
femora  thick  and  greatly  swollen  in  middle. 

9-  Head  a  little  larger  than  in  (J;  space  between  ninth  inter- 
stice and  border  of  elytra  very  wide  and  obliquely  truncate  pos- 
teriorly, the  lateral  sinuosities  of  apex  being  exceptionally 
strongly  developed.     Length  15-5-18,  breadth  5-6*2  mm. 


"286  EEVISION  OF  THE  GENUS  NOTONOMUS, 

Hah. — N.S.W.  :  Ourimbah  (Sloane;  in  scrub,  3rd  July). 

A  distinct  species,  resembling  in  facies  and  colour  N.  arthuri^ 
SI.,  but  easily  to  be  distinguished  by  the  el3^tra  having  the  third 
interstice  bipunctate  (sometimes  a  third  puncture  is  present), 
•eighth  interstice  ver}^  narrow;  lateral  apical  sinuosities  strongly 
developed,  especially  in  the  ^. 

NoTONOMus  KiNGi,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  86. 

Elliptical-oval,  convex.  Black.  Head  moderate  (3-2  mm.  across 
eyes),  convex;  frontal  impressions  light,  narrow,  divergent  back- 
wards; eyes,  with  orbits,  reniform.  Prothorax  as  long  as  broad 
(4- 5  X  4-5  mm.),  widest  before  middle;  apex  (3-4  mm.)  wider  than 
base  (3'1  mm.);  sides  lightly  rounded,  not  sinuate  before  base; 
basal  angles  rounded,  hardly  marked;  posterior  marginal  puncture 
just  before  basal  angles,  not  on  border;  median  line  well  marked; 
lateral  basal  impressions  narrow,  rather  long,  reaching  base  in 
full  depth.  Elytra  oval  (10  x  6  mm.);  humeral  angles  not  marked; 
basal  border  joining  lateral  border  Avithout  any  interruption  at 
humeral  angles;  lateral  sinuosities  of  apex  well  developed;  striae 
deep;  interstices  convex,  third  2-punctate  (sometimes  3-punctate); 
ninth  stria  and  tenth  interstice  well  developed.  Intercoxal 
declivity  of  presternum  narrowed  and  a  little  rounded  in  middle; 
of  mesosternum  vertical  and  very  lightly  concave.  Length  15-18, 
breadth  5-6  mm. 

//a6.— N.S.W.  :  Maitland  (Froggatt),  Armidale  (Lea),  Glen 
Innes  (Sloane). 

Chaudoir's  note  on  ^V.  kingi  is  so  brief  as  to  be  unworthy  of 
the  name  of  a  description,  being  merely  five  lines  of  comparison 
with  N.  variicollis,  Chaud.,  in  which  the  colour  is  not  stated; 
nothing  is  added  under  N.  kingi  in  his  'Supplement,'  except 
that  it  was  unknown  to  Castelnau ;  but  in  his  note  in  the 
'  Supplement '  on  N.  satanas,  Casteln.,  he  compares  that  species 
with  iV.  kingi  (which  suggests  the  inference  that  N.  kingi  is  a 
black  species),  and,  amongst  other  characters,  notes  that  N. 
satanas  has  the  third  intei'stice  of  the  elytra  punctate,  the  same 


BY    THOMAS    G.    SLOAXE.  287 

as  iV.  kingi;  Castelnau  described  the  elytra  of  N.  satanas  as 
having  the  third  interstice  bipunctate.  By  the  aid  of  the  data 
that  N.  kifigi  is  a  black  species  with  the  third  elytral  interstice 
bipunctate,  assisted  by  Chaudoir's  comparative  remarks  with  JV. 
variicoUis  and  ^Y.  satanas,  I  arrive  at  the  conclusion  that  the 
name  is  applicable  to  the  species  for  which  I  have  used  it  above. 
It  is  a  distinct  species;  compared  with  N.  jyolitulus,  Chaud.,  it  is 
more  elongate,  more  convex,  with  the  posterior  puncture  of  the 
third  elytral  interstice  not  nearly  so  near  to  the  apex. 

Note. — Specimens  from  New  England  have  the  prothorax  a 
little  shorter;  sides  subsinuate  before  base;  basal  angles  more 
marked;  the  elytra  with  basal  border  raised  above  lateral  border 
at  humeral  angles.  It  may  be  a  distinct  species,  but  I  hesitate 
to  consider  it  so  from  the  specimens  available  to  me. 

NoTONOMUS  INGRATUS,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  87;  Ann.  Mus.  Civ.  Genova,  1874,  vi. 
p.  582. 

I  venture  to  place  N.  ingratus,  Chaud.,  between  iV.  kingi, 
Chaud.,  and  N.  marginatus,  Casteln.  It  is  unknown  to  me  in 
nature.  In  the  original  diagnosis  it  is  described  as  having  the 
prothorax  a  little  longer  and  less  rounded  on  the  sides,  the  elytra 
longer  and  less  convex  than  N.  marginata,  Casteln.;  the  inter- 
stices slightly  and  equally  convex;  the  shoulders  very  slightly 
dentate ;  from  the  '  Supplement '  we  learn  further  that  it  is 
rather  near  N.  politulus,  Chaud.,  coloured  the  same,  with  a  more 
marked  iridescent  reflection  on  the  elytra;  prothorax  narrower, 
almost  as  long  as  broad.     Length  18  mm. 

Hab. — Southern  Australia. 

NoTONOMUS  MARGINATUS,  Castelnau. 

Poecilus  marginatus,  Casteln.,  Hist.  Nat.  Ins.  1838,  i.  p.  105; 
Notonomus  discoderus,  Chaud.,  Bull.  Mosc.  1865,  ii.  p.  87;  Ann. 
Mus.  Civ.  Genova,  1874,  vi.  p.  584 ;  Feronia  (Pterostichus) 
hunteriensis,  Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  213. 

Resembling  N.  australasice,  Dej.,  but  differing  by  the  prothorax 
more  brightly  cupreous  towards  the  sides,  elytra  with  a  bright 


288  REVISION  OF  THE  GENUS  NOTOXOMdS, 

metallic-cupreous  margin,  basal  angles  of  prothorax  a  little  more 
obtuse;  basal  border  of  elytra  not  raised  above  lateral  border  at 
humeral  angles  (in  N.  ausiralasice  it  is  slightly  raised  above  the 
lateral  border  at  point  of  junction);  prosternum  with  intercoxal 
declivity  narrowed  and  rounded  in  middle;  mesosternum  decidedly 
concave  between  cox£e.     Length  15-5-16,  breadth  5-8x6  mm. 

Hab.—N.S.W.  :  Hunter  River  (Froggatt). 

Two  specimens  (9)  in  my  collection  without  exact  locality 
have  the  elytra  as  convex  as  in  iV.  austral asice,  Dej.,  and  these  I 
look  upon  as  typical  of  the  species. 

Var.  N.  luculentus. — I  suggest  this  varietal  name  for  a 
form  from  the  Blue  Mountains  which  has  the  9  with  the 
prothorax  longer  and  more  narrowed  to  the  base ;  elytra 
more  depressed,  with  lightly  convex  interstices;  (J  with  elytra 
depressed,  interstices  very  convex.  Length  13-5-16  mm. 
Excepting  for  the  bright  cupreous  margin  of  the  elytra,  N. 
luculentus  resembles  the  species  I  have  named  N.  sydneyensis  so 
closely  that  I  cannot  indicate  other  differences,  though  I  belie^■e 
them  to  be  two  closely  allied  but  distinct  species.  Length 
13-5-16  mm. 

Hah. — Springwood  (Sloane). 

NOTONOMUS    SYDNEYENSIS,  n.Sp. 

Oval,  depressed.  Head  not  large  (2-8  mm.  across  eyes);  pro- 
thorax broader  than  long,  not  sinuate  on  sides  before  base;  basal 
angles  obtuse  :  elytra  truncate-oval,  stronglj'^  striate;  interstices 
convex,  strongly  convex  in  ^,  lightly  so  in  9,  third  2-punctate, 
eighth  and  ninth  very  convex  :  intercoxal  declivity  of  prosternum 
rounded;  of  mesosternum  concave.  Head  black  (sometimes  with 
faint  seneous  tinge),  prothorax  and  elytra  dark  purple  (sometimes 
greenish);  prothorax  nitid;  elytra  rather  nitid  in  $,  subopaque 
in  9;  under  surface  black;  tarsi  and  palpi  piceous. 

Head  convex;  frontal  impressions  well  marked.  Prothorax 
transverse  (3-4  x  4*2  mm.);  sides  lightly  rounded,  a  little  more 
obliquely  narrowed  to  base  than  to  apex;  base  (3  mm.)  hardly 
wider  than   apex   (2-9  mm.),  emarginate  in  middle,  arcuate   on 


BY    THOMAS    G.    SLOANE.  289 

sides  ;  apex  emarginate;  basal  angles  obtuse,  hardly  marked; 
border  wide,  especially  towards  base,  passing  round  basal  angles 
on  to  sides  of  base;  posterior  marginal  puncture  on  the  border  at 
basal  angle ;  median  line  strongly  impressed  ;  lateral  basal 
impressions  rather  long,  deep,  parallel.  Elytra  ovate  (8x5  mm,), 
depressed  on  disc,  declivous  in  a  gentle  slope  to  apex,  strongly 
sinuate  on  each  side  of  apex;  basal  border  with  posterior  margin 
arcuate  and  raised  above  plane  of  elytra,  joining  lateral  border 
without  interruption  at  humeral  angles ;  lateral  border  widely 
reflexed,  carinate  near  base;  eighth  interstice  convex,  narrow  (but 
wider  than  ninth  on  basal  half),  ninth  very  narrow  in  ^,  and 
interrupted  on  inner  side  by  the  umbilicate  punctures,  tenth 
feebly  developed  just  in  front  of  the  apical  sinuosities.  Length 
12-5-16,  breadth  4-3-5-8  mm. 

Hab. — N.S.W.  :  Manly  (Fletcher),  Narrara  and  Ourimbah 
(Sloane;  plentiful  in  heavily  timbered  brush  land). 

Allied  to  iV.  marginatus,  Casteln.,  of  which  it  may  be  a  variety 
without  the  bright  border  to  the  elytra.  Compared  with  what  I 
consider  the  typical  form  of  A",  marginatus.  the  9  has  the  pro- 
thorax  proportionately  longer  and  narrower  at  the  base ;  the 
elytra  more  depressed  with  more  convex  interstices,  the  eighth 
especially  being  narrower  and  more  convex.  I  cannot  differentiate 
it  from  the  species  from  Spring  wood  which  I  have  placed  under 
N.  7yiarginatus  as  var.  luculentus,  except  by  the  absence  of  the 
bright  cupreous  lateral  channel  of  the  elytra,  but  I  believe  it  to 
be  a  distinct  species. 

NoTONOMus  AUSTRALASIA,  Dejeau. 

Feronia  australasice,  Dej.,  Spec.  iii.  p.  277;  Chaud.,  Bull.  Mosc. 
1865,  ii.  p.  87;  Ann.  Mus.  Civ.  Genova,  1874,  vi.  p.  583:  Feronia 
(Oniaseus)  mitchelli,  Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii. 
p.  216. 

Oval,  robust.    Upper  surface  of  a  bronzy,  or  bronzy-black  colour, 

sometimes  viridescent;  under  surface  and  legs  black,  or  piceous- 

black.     Head  large  (3-3  mm.  across  eyes).     Prothorax  transverse 

(4x5  mm.),  lightly  rounded  on  sides,  wider  at  base  (3*7  mm.) 

19 


'290  EEVISION  OF  THE  GENUS  NOTOXOMUS, 

than  apex  (3'5  mm.);  posterior  angles  obtuse,  a  little  marked; 
posterior  marginal  puncture  on  border  at  basal  angle ;  lateral 
channel  and  border  wide.  Elytra  truncate-oval  (9  x  55  mm.); 
humeral  angles  hardly  marked;  striae  deeply  impressed,  minutely 
crenulate  (more  perceptibly  so  in  9);  interstices  convex  (strongly 
so  in  (J,  lightly  so  and  rather  opaque  in  9),  third  2-punctate, 
eighth  wider  than  ninth  on  basal  half  (hardly  convex  on  basal 
half  in  9),  ninth  lightly  convex,  wider  posteriorly,  tenth  distinct 
before  lateral  basal  sinuosity.  Intercoxal  declivity  of  prosternum 
flat;  of  mesosternum  flat.     Length  15-17,  breadth  5-2-5-8  mm. 

Hub. — I*^.S  W.  :  Sydney  District  [Ermington,  Penshurst,  Ryde, 
Carlton,  Auburn  (Sloane)],  Grose  Valley  (Fletcher). 

Castelnau  gives  the  range  of  the  species  as  "  Queensland, 
N.S.  Wales  and  Victoria,"  but  I  have  seen  it  only  from  the 
Sydney  District.     The  synonymy  is  on  the  authorit}^  of  Chaudoir. 

NoTONOMUS  INCRASSATUS,  Chaudoir. 
Ann.  Mus.  Civ.  Genova,  1874,  p.  583. 

Robust,  oval.  Black,  elytra  often  of  a  bronzy  or  purplish 
tint.  Head  rather  large  (3-2  mm.  across  eyes).  Prothorax 
lightly  transverse  (3-8  x  4*3  mm.)  widest  before  middle,  lightly 
narrowed  to  base,  lightly  convex;  apex  and  base  of  equal  width 
(3-4  mm.);  basal  angles  marked,  but  obtuse  at  summit;  posterior 
marginal  puncture  placed  on  inner  side  of  border  at  basal  angles; 
border  narrow,  often  subsinuate  just  beside  basal  angles;  lateral 
basal  impressions  oval  (rather  long).  Elytra  truncate-oval 
(8x5  mm.),  deeply  striate;  interstices  convex,  third  2-punctate, 
eighth  narrow,  convex,  ninth  narrower  than  eighth,  interrupted 
on  inner  side  by  umbilicate  punctures,  tenth  well  developed, 
narrow,  convex,  extending  forward  to  about  anterior  third.  Inter- 
coxal declivity  of  prosternum  flat;  of  mesosternum  flat.  Length 
13-16,  breadth  4'5-5-6mm. 

Hab. — N.S.W.  :  Sydney,  Carlton,  Auburn,  Appin,  Springwood, 
BuUi  and  Wollongong  (Sloane). 


BY    THOMAS    G.    SLOANE.  291 

iV.  incrassatus  is  easily  identified  by  prothorax  with  basal  angle 
marked;  elytra  with  humeral  angles  dentate,  eighth  and  ninth 
interstices  narrow,  tenth  interstice  elongate.  It  differs  from  N. 
australasice,  Dej.,  by  the  features  mentioned  above,  and  also  by 
—  ia)  the  border  of  the  prothorax  much  narrower,  less  reflexed 
and  hardly  developed  on  sides  of  base,  (6)  border  of  elytra  much 
narrower  near  the  base.  It  is  a  species  of  variable  width;  the 
form  which  I  look  upon  as  the  typical  one  is  that  from  the 
vicinity  of  Sydney,  which  is  wholly  black,  and  has  the  border  of 
the  prothorax  subsinuate  at  the  basal  angles.  Specimens  from 
Springwood  and  Appin  have  the  border  meeting  the  base  without 
any  sinuosity,  which  causes  the  basal  angles  to  be  obtuse,  though 
marked.  Specimens  from  Bulli  (where  it  is  common  on  the 
heavily  wooded  mountain)  have  the  elytra  purple,  and  are  of 
narrower  form  than  those  found  about  Sydney.  A  specimen 
from  Wollongong  is  of  still  more  elongate  form  (13-5  x  4-5  mm.), 
and  has  the  prothorax  and  head  more  elongate,  eyes  less  pro- 
tuberant and  more  strongly  inclosed  behind  (the  convexity  and 
prominence  of  the  eyes  differ  in  other  species  besides  N.  incras- 
satus). 

NOTONOMUS    LEAI,   n.sp. 

Robust,  convex,  subparallel.  Head  thick  (2*8  mm.  across  eyes), 
convex;  prothorax  subquadrate,  rounded  on  sides;  basal  angles 
obtuse;  posterior  marginal  puncture  on  border  at  basal  angle; 
lateral  basal  impressions  wide,  deep,  foveiform;  elytra  truncate- 
oval,  lightly  striate;  interstices  hardly  convex,  third  2-punctate, 
eighth  narrow  and  lightly  convex,  ninth  very  narrow  and  convex. 
Black;  elytra  subnitid,  often  with  a  purple  or  greenish  tinge, 
hardly  more  opaque  in  9  than  in  (J. 

Head  wide  and  convex  between  eyes;  front  lightly  biimpressed 
between  antennae;  eyes  convex,  not  prominent.  Prothorax  convex, 
broader  than  long  (3*7  x  4*5  mm.),  widest  about  middle;  sides 
lightly  and  evenly  rounded,  decidedly  narrowed  to  apex,  meeting 
base  without  any  sinuosity;  apex  hardly  emarginate;  anterior 
-angles  obtuse,  not  prominent;   base  (3*5  mm.)  wider  than  apex 


292  REVISION  OF  THE  GENUS  NOTOXOMUS, 

(3-2  mm.),  truncate  in  middle,  curving  decided!)'- forward  on  each 
side;  basal  angles  very  obtuse,  hardly  marked;  border  thick,  very 
narrow  anteriorly,  wider  towards  base;  median  line  lightly  im- 
pressed ;  lateral  basal  impressions  wide,  short,  deep.  Elytra 
narrow  ((S-5  x  4*8  mm.),  convex;  sides  very  lightly  rounded;  lateral 
sinuosities  of  apex  light;  scutellar  striole  ver}'-  short;  basal  border 
truncate  on  posterior  margin,  obtuse  and  hardly  the  least  raised 
above  the  narrow  lateral  border  at  their  point  of  junction;  ninth 
interstice  very  narrow,  interrupted  on  inner  margin  by  umbilicate 
punctures;  tenth  interstice  long,  convex,  extending  forward  to 
basal  third.  Tntercoxal  declivity  of  prosternum  flat;  of  meso- 
sternum  widely  and  lightly  concave.  Length  13-17,  breadth 
4-5-5-5  mm. 

Ilab. — N.S.W.  :  Blue  Mountains  [Katoomba  and  Mount  Vic- 
toria (Sloane),  Wentworth  Falls  and  Grose  Valley  (Fletcher)]. 

It  probably  comes  nearest  to  iV.  incrassatus,  Chaud.,  amongst 
described  species,  but  differs  by  prothorax  with  basal  angles 
widely  obtuse;  lateral  basal  impressions  deeper,  wider,  more  oval; 
elytra  with  interstices  hardly  convex,  humeral  angles  not  marked, 
&c.  From  N.  austi^alasice,  Dej.,  it  differs  by  its  form  narrower 
and  more  convex;  head  smaller;  prothorax  shorter,  less  narrowed 
to  base,  sides  more  rounded  to  apex  and  base,  posterior  angles 
less  marked,  lateral  basal  impressions  shorter  and  wider;  elytra 
less  deeply  striate,  interstices  flatter,  eighth  and  ninth  narrower, 
humeral  angles  not  marked,  tkc.  It  may  be  distinguished  from 
N.  lateralis,  SI.,  by  prothorax  more  transverse,  with  lateral  channel 
wider,lateral  basal  impressions  deeper  and  wider,  basal  angles 
more  widely  rounded;  elytra  with  third  interstice  bipunctate  (the 
posterior  puncture  further  from  apex),  eighth  interstice  narrower, 
&c. 

NOTONOMDS    LATERALIS,   SloanC. 

Proc.  Linn.  Soc.  KS.W,  1889  (2),  iv.  p.  1295. 

One  of  my  original  type  specimens  (9)  is  before  me;  it  has  the 
intercoxal  declivity  of  the  prosternum  narrowed  and  a  little 
rounded  in  middle,  of  the  mesosternum  decidedl}^  concave;  elytra 


BY    THOMAS    G.    SLOANE.  293 

flat  on  disc  and  abruptly  declivous  on  base  and  apex,  tenth  inter- 
stice convex  and  very  long — extending  forward  to  the  basal  sixth 
of  the  elytra — the  margin  wide  and  flat.  A  second  specimen 
(without  anterior  legs,  but  which  seems  from  the  quadripunctate 
apical  ventral  segment  to  be  the  9)  has  been  given  to  me  by  Mr. 
J.  J.  Fletcher  as  from  Mount  Tomah ;  its  dimensions  are : — 
16x5  mm.,  head  3*2  mm.  across  eyes,  prothorax  4  x  4*65  mm., 
elytra  9x5  mm.;  it  has  the  elytra  lightly  convex,  with  eighth 
interstice  much  wider  than  ninth  on  basal  half,  ninth  very  narrow 
and  convex,  tenth  strongly  developed  and  extending  forward  to 
basal  sixth;  lateral  channel  wide;  border  narrow,  hardly  reflexed, 
lightly  sinuate  on  each  side  of  apex;  intercoxal  declivity  of  pro- 
sternum  flat,  of  mesosternum  wide  and  not  concave;  ventral  seg- 
ments not  transversely  impressed  laterally. 

After  seeing  this  specimen  I  conclude  that  the  flattened  elytra, 
with  sides  abrupt,  margin  wide  and  non-sinuate  laterally  at  apex, 
as  shown  by  the  type  specimen  (9),  may  be  merely  characteristic 
of  an  aberrant  or  deformed  specimen. 

ffab.—^.S.W.:  Blue  Mountains  [Mount  Wilson  (Olliff), 
Mount  Tomah  (Fletcher)]. 

« 

NoTONOMUS  TRiPLOGENioiDEs,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  85;  Feronia  {Pte.rostichiis)  duca/is, 
Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  207. 

^.  Large,  robust,  convex.  Upper  surface  cupreous,  prothorax 
and  lateral  margin  of  elytra  usually  of  a  brighter  and  more  fiery 
copper  colour;  under  surface  and  legs  piceous-black.  Head  large 
(4-7mm.  across  eyes).  Prothorax  transverse-cordate (5 -4  x  7'4mm.); 
base  and  apex  of  equal  width;  sides  lightly  sinuate  before  base; 
posterior  angles  obtuse  but  marked;  posterior  marginal  puncture 
at  basal  angle  on  inner  side  of  border  and  outside  lateral  channel; 
lateral  basal  impressions  short,  deep.  Elytra  truncate-oval 
(13-5  X  8  mm.),  convex;  lateral  apical  sinuosities  light;  basal 
border  raised  and  dentate  at  humeral  angles;  strise  deep,  minutely 
crenulate  at  bottom,  ninth  well  marked  towards  apex;  interstices 


294  EEVISION  OF  THE  GENUS  NOTO^'OMUS, 

convex,  third  2-punctate,  tenth  well  developed  towards  apex. 
Intercoxal  declivity  of  prosternum  wide,  fiat  (or  rather  concave); 
of  mesosternum  strongly  concave;  metasternum  deeply  longitu- 
dinally channelled  in  middle  (this  median  channel  extending  to 
apex).  Posterior  femora  with  lower  side  strongl}''  dilatate  above 
apex  of  trochanters.     Length  24,  breadth  7 '3-8  mm. 

Hah. — N.S.W.  :  Bulli,  Wollongong  and  Kiama  (Sloane). 

NOTONOMUS    JERVISENSIS,  n.sp 

2-  Only  differs  from  N.  triplogenioides,  Chaud.,  by — {a)  colour 
(prothorax  black  on  disc,  fier}'-  copper  near  margins;  elj^tra  almost 
black,  with  obsolete  cupreous  tinge,  lateral  margins  of  a  fiery 
copper  colour);  {h)  prothorax  with  posterior  angles  much  more 
widely  obtuse,  the  antebasal  sinuosity  of  the  sides  obsolete. 
Length  24,  breadth  8  mm.  (head  4'75  mm.;  prothorax  5-5  x  7mm.). 

/^ft6._N.S.W.  :  Jervis  Bay  (Fletcher;  Coll.  Sloane). 

It  is  possibly  a  well  marked  variety  of  iV.  triplogenioides,  but, 
even  if  this  be  the  case,  seems  entitled  to  rank  as  a  named  variety. 

NOTONOMUS    LIRAGERUS,   n.sp. 

Elongate,  depressed.  Prothorax  hardly  broader  than  long, 
lightly  narrowed  to  base;  basal  angles  rectangular;  posterior 
marginal  puncture  at  basal  angle  on  border  :  elytra  depressed, 
deeply  striate;  interstices  very  convex,  third  2-punctate.  Black, 
nitid. 

Head  small,  oval  (3*2  mm.  across  ej^es),  convex,  faintly  biim- 
pressed  between  antennae — the  impressions  extending  on  to 
clypeus;  eyes  convex,  rather  prominent.  Prothorax  almost  as 
long  as  broad  (4'6  x  4*7  mm.),  not  ampliate  at  widest  part,  a  little 
wider  across  base  (3*3  mm.)  than  across  apex  (3-2  mm.);  sides 
lightly  rounded,  shortl}^  and  lightly  sinuate  before  base;  apex 
truncate;  anterior  angles  not  marked,  close  to  head;  base  emar- 
ginate  in  middle,  sloping  lightly  forward  on  each  side;  lateral 
border  narrow  on  anterior  third,  wide  and  reflex ed  on  posterior 
two-thirds,  extending  round  basal  angles  to  lateral  impressions; 
median  line  rather  strongly  impressed,  reaching  base;  lateral  basa 


BY    THOMAS    G.    SLOANE.  295 

impressions  deep,  narrow,  parallel.  Elytra  narrow,  truncate-oval 
(10  X  6  mm.),  depressed;  sides  lightly  and  evenly  rounded,  lateral 
apical  sinuosities  well  developed;  basal  border  raised  and  forming 
a  strong  obtuse  projection  at  humeral  angles;  lateral  border  wide, 
reflexed,  narrow  just  behind  humeral  angles;  interstices  sharply 
raised,  seventh  wider,  more  convex  than  others  and  forming 
lateral  margin  of  disc,  eighth  and  ninth  narrow,  convex,  tenth 
well  developed  in  front  of  lateral  sinuosities.  Intercoxal  declivity 
of  prosternum  flat;  of  mesosternum  hardly  at  all  concave;  meta- 
sternum  longitudinally  channelled  in  middle  from  base  to  apex. 
Lateral  parts  of  abdomen  and  episterna  of  metasternum  rugulose- 
punctulate.     Length  18,  breadth  6  mm. 

Hah. — N.S.W.  :  Port  Macquarie  (Sloane). 

A  distinct  species  ;  it  is  perhaps  nearest  to  N'.  angustihasis, 
Sloane,  but  dijffers  decidedly  by  pro  thorax  less  narrowed  and 
more  shortly  sinuate  posteriorly,  posterior  marginal  puncture  at 
the  basal  angle;  elytra  flatter,  interstices  more  strongly  convex, 
ninth  narrower;  posterior  femora  wider  and  more  dilate  in  middle. 

NoTONOMUS  suBiRiDESCENS,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  85;  Ann.  Mus.  Civ.  Genova,  1874,  vi. 
p.  576. 

Unknown  to  me.  If  it  has  the  intercoxal  declivity  of  the  pro- 
sternum  flat  its  position  would  seem  to  be  near  N.  liragp.rus,  SI.; 
if  rounded,  it  would  come  near  .V.  kingi,  Chaud.,  but  this  does 
not  appear  to  me  likely  to  be  its  place.  From  Chaudoir's  original 
description  and  the  note  in  his  'Supplement '  I  select  the  following 
characters  as  diagnostic  of  iV.  sahiridescens  : — Compared  with  N. 
mneomicam — prothorax  scarcely  shorter  than  the  breadth;  upper 
surface  more  convex;  lateral  margin  evidently  finer  and  narrower 
anteriorly,  and  forming  on  the  contrary  a  wider  border  as  it 
approaches  the  posterior  angles  :  elytral  interstices  more  convex 
— particularly  towards  the  apex — eighth  and  ninth  decidedly 
narrower  throughout  their  whole  length;  third  bipunctate. 
Colour  black.     Length  23  mm. 

Hah. — Southern  Australia  (Melbourne;  Coll.  Laferte). 


296  REVISION  OF  THE  GENUS  NOTOyOMUS, 

]^.  suhiridescens  may  resemble  JV.  liragerus,  SI.,  but  I  do  not 
think  it  can  be  identical  on  account  of  its  larger  size  and 
apparently,  from  Chaudoir's  description,  it  should  have  the  pro- 
thorax  more  convex,  and  the  elytra  more  rounded  on  sides. 

NOTONOMUS    QUEENSLANDICUS,   n.sp. 

Elongate-oval,  robust.  Head  small ;  prothorax  subcordate ; 
sides  sinuate  before  base;  basal  angles  subrectangular;  posterior 
marginal  puncture  placed  a  little  before  base  on  inner  side  of 
lateral  channel;  elytra  deeply  striate;  interstices  in  ^  convex,  in 
9  depressed;  humeral  angles  strongly  dentate.  Black;  elytra  of 
a  bronzy  subviolaceous  tint,  with  ninth  interstice  and  lateral 
channel  of  a  bright  brassy  green  or  copper  colour;  under  surface 
and  legs  black;  antennae  piceous-black. 

Head  oval  (2-3  mm.  across  eyes);  vertex  lightly  transversely 
impressed;  frontal  impressions  deep,  divergent  posteriorly.  Pro- 
thorax  rather  small,  broader  than  long  (3  x  3-5  mm.),  depressed 
towards  base,  faintly  transversely  striolate;  sides  lightly  arcuate, 
more  strongly  narrowed  to  apex  than  base,  strongly  sinuate 
before  base;  apex  (2-5  mm.)  narrower  than  base  (2-7  mm.),  emar- 
ginate;  anterior  angles  distant  from  head,  obtuse;  base  emarginate 
in  middle,  lightly  arcuate  on  each  side;  basal  angles  well  marked, 
obtuse;  median  line  reaching  from  base  to  apex;  lateral  basal 
impressions  deep,  narrow,  elongate;  lateral  basal  spaces  lightly 
convex,  depressed  near  basal  margin.  Elytra  oval  (7"5  x  4 -3 mm.), 
truncate  at  base,  widely  and  lightly  sinuate  on  each  side  of  apex, 
lightly  convex,  strongly  declivous  to  apex,  abruptly  declivous  on 
sides ;  basal  border  with  posterior  margin  curving  forward 
externally,  sharply  raised  and  forming  a  strong  obtuse  prominence 
at  humeral  angles;  third  interstice  wider  than  two  inner  ones, 
.sixth  narrower  than  fifth  and  seventh,  narrowed  to  base,  seventh 
very  wide  on  basal  third,  eighth  and  ninth  narrow,  convex,  equal 
in  width  on  basal  third,  inner  margin  of  ninth  interrupted  by 
the  umbilicate  punctures,  tenth  obsolete.  Intercoxal  declivity  of 
prosternum  flat  and  bordered  laterally;  of  mesosternum  wide, 
hardly  concave.     Length  13,  breadth  4*3  mm. 


BY    THOMAS    G.    SLOANE.  297 

Hah. — Southern  Queensland  (Law;  Coll.  Sloane). 

Closely  allied  to  N.  nitidicoUls,  Chaud.,  with  which  it  agrees 
in  all  general  features — as  antennae,  labrum,  borders  of  prothorax 
and  elytra,  &c.;  but  differing  by  colour;  9  with  eyes  more  promi- 
nent; elytra  more  convex,  with  flatter  interstices,  scutellar  striole 
shorter,  lateral  sinuosities  of  apex  weaker. 

NoTONOMUS  NiTiDicoLLis,  Chaud. 

Bull.  Mosc.  1865.  ii.  p.  8S ;  Feronia  (Pterostichus)  master^si, 
Casteln.,  Trans.  Roy.  Soc.  Vict.  viii.  p.  211;  F.  {Ft.)  impressi- 
collis,  Casteln.,  I.e.,  p.  212;  IV.  viridicinctiis,  Macl.,  Trans.  Ent 
Soc.  N.S.W.  1871,  ii.  p.  108;  K  angustipennis,  Macl.,  I.e.,  109. 

Elongate-oval,  robust.  Head  small  :  prothorax  subcordate, 
sinuate  on  sides  before  base;  basal  angles  subrectangular;  posterior 
marginal  puncture  a  little  before  the  base  on  inner  side  of  lateral 
channel :  elytra  oval,  deeply  striate;  third  interstice  2-punctate, 
eighth  narrow,  convex;  humeral  angles  strongly  dentate.  Black; 
head  and  prothorax  nitid  with  metallic  reflections  of  a  brassy  or 
purplish  tinge;  elytra  subopaque,  bronzy  purple,  eighth  and  ninth 
interstices  more  metallic,  the  wide  lateral  channel  sometimes 
brassy  green. 

Head  small  (2*2  mm.  across  eyes);  vertex  lightly  transversely 
impressed;  front  biimpressed,  the  impressions  rather  deep,  sharply 
defined,  extending  backwards  from  clypeal  punctures  and  diver- 
gent posteriorly;  eyes  convex,  subprominent.  Antennae  not  long, 
joints  short  towards  apex.  Labrum  lightly  longitudinally  im- 
pressed in  middle,  rounded  on  anterior  margin.  Prothorax  rather 
small,  broader  than  long  (3  x  3*5  mm.),  depressed  towards  base, 
faintly  transversely  striolate  (but  nitid);  sides  lightly  and  almost 
equally  rounded,  shortly  but  decidedly  sinuate  before  base;  apex 
(2-4  mm.)  hardly  as  wide  as  base  (2*5  mm.),  emarginate;  anterior 
angles  marked,  obtuse;  base  emarginate  in  middle,  truncate  on 
each  side;  angles  rectangular,  obtuse  at  summit;  lateral  border 
narrow,  a  little  wider  and  more  decidedly  reflexed  on  basal  half; 
median  line  deep,  extending  from  base  almost  to  apex;  lateral 


298  REVISION  OF  THE  GENUS  XOTOXOMCS, 

basal  impressions  deep,  narrow,  elongate,  extending  almost  half 
the  length  of  prothorax,  gradually  shallower  forward ;  lateral 
basal  spaces  depressed  (hardl}'-  the  least  convex).  Elytra  oval 
(7*4  X  4-3  mm.),  strongly  sinuate  on  each  side  of  apex,  subdepressed 
on  disc,  abruptly  declivous  on  sides,  strongly  declivous  to  apex; 
basal  border  with  posterior  margin  curving  strongly  to  the  dentate 
humeral  angles  ;  lateral  border  reflexed,  narrowed  near  base ; 
three  inner  interstices  hardly  convex  on  disc,  third  wider 
(especially  at  posterior  marginal  puncture)  than  first  and  second, 
fifth,  sixth  and  seventh  convex,  sixth  narrower — especially  near 
base — seventh  very  wide  on  basal  third,  eighth  and  ninth  convex, 
very  narrow  (of  about  equal  width  on  basal  half),  inner  margin 
of  ninth  decidedly  interrupted  by  the  umbilicate  punctures,,  tenth 
slightly  developed  near  apical  sinuosities.  Intercoxal  declivity 
of  presternum  flat  and  bordered  on  each  side;  of  mesosternum 
wide  and  hardly  concave.     Length  12-5,  breadth  4-3  mm. 

Hab. — Q.:  Burnett  RiverDistrict(Spencer),  Gayndah (Masters), 
Brisbane  (Macleay  Coll.). 

I  have  seen  a  specimen  in  the  Macleay  Museum  ticketed 
"Brisbane"  which  I  identify  as  S.  nitidicoUis,  Chaud.,  and  a 
comparison  of  this  Brisbane  specimen  with  specimens  of  JV. 
vlridicinctus,  MacL,  and  N.  angustipemiis,  Macl.,  convinces  me 
that  all  three  names  belong  to  one  species.  The  types  of  both 
of  these  species  of  Macleay  have  been  carefully  examined  by  me, 
and  a  specimen  (9)  in  my  collection  (absolutely  identical  with  IV. 
angustipennis  from  comparison  with  t3^pe)  from  the  Burnett 
River  has  been  used  for  my  description  of  this  species.  The 
specimen  in  the  Maclea}'  Collection  which  I  consider  typical  of 
N.  nitidicoUis  has  the  margin  of  the  elytra  of  a  dull  coppery 
bronze;  in  all  other  respects  it  agrees  with  the  specimen  I  have 
used  for  my  description. 

NOTONOMUS    LATIBASIS,   n.sp. 

9.  Oval,  depressed:  prothorax  wide,  depressed;  base  (3  mm.) 
evidently  wider   than   apex  (2*7  mm.);   sides  shortly  subsinuate 


BY    THOMAS    G.    SLOANE.  299' 

before  base ;  bcasal  angles  obtusely  subrectangular ;  posterior 
marginal  puncture  placed  a  little  before  the  base  on  inner  side  of 
lateral  channel:  elytra  truncate-ov-al,  deeply  striate;  third  inter- 
stice 2-punctate,  eighth  narrow,  convex;  humeral  angles  feebly 
dentate.  Head  black  with  a  purple  tinge ;  prothorax  nitid, 
purple-black,  violaceous  towards  sides;  elytra  dark  purple,  some- 
what opaque,  lateral  channel  bright  brassy  green;  under  surface 
and  legs  piceous. 

Head  oval  (2-5  mm.  across  eyes);  vertex  lightly  transversely 
impressed ;  front  strongly  biimpressed,  impressions  divergent 
posteriorly;  eyes  convex,  subprominent.  Prothorax  broader  than 
long  (3-4  X  4  mm.),  widest  just  before  the  middle,  depressed,  very 
faintly  transversely  striolate;  sides  lightly  arcuate,  more  strongly 
and  roundly  narrowed  anteriorly  than  posteriorly;  apex  emai'- 
ginate ;  anterior  angles  rather  prominent,  obtuse;  base  wide, 
emarginate  in  middle,  sloping  lightly  forward  to  angles  on  each 
side;  lateral  border  widely  reflexed,  narrower  in  front  of  anterior 
marginal  puncture;  median  line  deep,  reaching  from  base  almost 
to  apex;  lateral  basal  impressions  long,  parallel,  narrow  at  bottom 
(extending  almost  half  the  length  of  prothorax,  becoming  faint 
anteriorly);  lateral  basal  impressions  a  little  raised  (hardly  convex). 
Elytra  truncate-oval  (7*6  x  4-8  mm.),  wide  and  truncate  at  base, 
strongly  sinuate  on  each  side  of  apex,  depressed,  lightly  declivous 
to  apex,  abruptly  declivous  on  sides;  basal  border  curving  very 
little  forward  to  humeral  angles  on  posterior  margin,  raised  into 
an  obtuse  subprominent  projection  at  point  of  junction  with  lateral 
border;  this  widely  reflexed,  narrow,  close  to  base;  three  inner 
interstices  very  lightly  convex  on  disc,  third  much  wider  than 
first  and  second  at  posterior  puncture,  fifth,  sixth  and  seventh 
convex,  sixth  narrower  (especially  near  base),  seventh  very  wide 
on  basal  third,  eighth  narrow,  a  little  wider  than  ninth  on  basal 
half,  tenth  narrow  and  feebly  developed  near  apical  sinuosity. 
Intercoxal  declivity  of  prosternum  flat,  bordered  laterally' ;  of 
mesosternum  wide,  hardly  concave.     Length  13,  breadth  48  mm. 

Hah. — Southern  Queensland  (two  specimens  [9]  brought  from 
Burnett  River  District  by  Professor  W.  B.  Spencer;  Coll.  Sloane). 


30O  REVISION  OF  THE  GENUS  ^'OTONOMUS, 

I  have  compared  my  specimens  with  the  type  specimens  of  /Y. 
cyaneocinctus,  Macl.,  and  have  considered  it  a  distinct  but  closely 
allied  species  on  account  of  the  juxtabasal  sinuosity  of  the  sides 
of  the  prothorax;  this,  however,  is  a  feature  that  requires  more 
study  to  determine  its  true  value  in  the  genus  Notonomus.  It  is 
nearly  allied  to  N.  nitidicollis,  Chaud.,  with  which  it  agrees 
generally  in  shape  of  head,  antennae,  labrum,  interstices  of  elytra, 
ct:c.,  but  differs  by  the  shape  of  the  prothorax  (which  is  of  wider 
form  with  the  sides  less  rounded,  less  narrowed  to  both  apex  and 
base,  much  wider  across  base);  the  elytra  more  depressed,  more 
lightly  declivous  to  apex,  more  lightly  rounded  on  sides,  less 
deeply  striate,  interstices  less  convex. 

NoTONOiMUS  viRiDiLiMBATUS,  Castelnau. 

Feronia  i^Pterostichus)  viridilimbata,  Casteln. ,  Trans.  Roy.  Soc. 
Vict.  1868,  viii.  p.  215;  F.  (Pt.)  viridimaryinata,  Casteln.,  l.c.^ 
p.  216;  N.  viridilimbatus,  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874, 
vi.  p.  584 ;  N.  cyaneocincfus,  Macl.,  Trans.  Ent.  Soc.  N.S.W., 
1871,  ii.,  p.  108. 

I  offer  the  following  translation  of  Chaudoir's  note  on  N.  viri- 
dilimbatus, Casteln.,  =  iV.  viridimarginatus,  Casteln.  :  Length 
12*5-15  mm.  Resembling  most  closely  N".  nitidicollis,  from  which 
it  differs  by  the  posterior  angles  of  the  prothorax  which  are  sub- 
obtuse,  not  preceded  by  a  sinuosity,  and  not  forming  any  lateral 
prominence.  Its  habitat  is  the  same,  and  perhaps  it  will  be  found 
to  be  only  a  variety. 

I  have  seen  the  types  of  J^.  cyaneocinctus,  Macl.,  in  the  Aus- 
tralian Museum,  Sydney,  and  have  the  following  note,  made  with 
these  type  specimens  before  me: — Differs  from  S.  viridicinctusi 
Macl.,  by  the  sides  of  the  prothorax  not  the  least  sinuate  before 
the  base;  the  elytra  have  the  shoulders  dentate;  the  form  of  the 
prothorax  is  intermediate  between  A\  violaceomarginatus,  Macl., 
and  iV.  viridicinctus,  Macl.,  (=i\r.  nitidicollis,  Chaud.),  being 
more  narrowed  behind  than  in  N.  violaceomarginatus,  and  less  so 
than  in  N.  viridicinctus. 


BY    THOMAS    G.    SLOANE.  301 

Comparing  Chaudoir's  note  on  ]\\  inridilimhatus  with  my  note 
on  X.  cyaneoclnclus,  leaves  but  little  doubt  in  m}'  mind  that  these 
two  must  be  conspecific. 

NoTOxoMUS  DARLiNGi,  Casteln. 

Feronia  (Pterostichus)  darlingi,  Casteln.,  Trans.  Roy.  Soc.  Yict. 
1868,  viii.  p.  216  ;  N.  darlings  Chaud.,  Ann.  Mus.  Civ.  Genova, 
1874,  vi.  p.  591. 

Unknown  to  me  in  nature,  but  the  following  features  from 
Chaudoir's  note  on  it  in  his  '  Supplement '  seem  to  thoroughly 
differentiate  it,  and  should  render  it  easy  of  recognition.  Much 
resembles  iV^.  nitidicoUis,  but  the  elytral  striae  are  densely  and 
distinctly  crenulate;  head  larger;  prothorax  a  little  less  elongate, 
wider  anteriorly,  more  rounded  on  sides,  which  are  similarly 
sinuate  before  the  base,  basal  angles  equally  sharp;  elytra  wider, 
interstices  more  depressed,  third  similarly  punctate,  eighth  not 
so  narrow.  Head  of  an  obscure  copper  colour,  prothorax  metallic- 
violet,  elytra  obscure  violet,  with  the  two  lateral  interstices  and 
the  marginal  channel  of  a  shining  coppery-purple-violet.  Length 
15  J  lines. 

Hab. — Q.  :  Pine  Mountains  (Masters  ^o?e  Castelnau). 

NoTONOMUS  SUBOPACUS,  Chaudoir. 

Ann.  Mus.  Civ.  Genova,  1874,  vi.  p.  586. 

Elongate.  Upper  surface  of  a  dark  bronze  colour;  head  black 
with  slight  bronzy  reflections;  prothorax  metallic;  elytra  rather 
nitid  in  ^,  opaque  and  darker  in  9. 

Head  rather  large,  wide  (2 -7  mm.  across  eyes),  a  little  con- 
stricted on  each  side  behind  eyes ;  front  depressed,  faintly 
biimpressed.  Prothorax  a  little  broader  than  long(315  x  3'6mm.), 
widest  before  middle,  not  ampliate  at  widest  part,  of  equal  width 
at  base  and  apex  (2-6  mm.);  disc  lightly  transversely  striolate; 
sides  lightly  arcuate,  lightly  narrowed  to  apex,  lightly  narrowed 
to  base  in  an  even  curve  without  any  sinuosity;  anterior  angles 
obtuse,  not  prominent;  base  emarginate  in  middle;  basal  angles 


302  KEVISION  OF  THE  GENUS  A'OTOyOMUS. 

marked,  but  obtuse;  lateral  border  and  lateral  channel  narrow; 
posterior  marginal  puncture  placed  a  little  before  base  on  inner 
side  of  lateral  channel;  median  line  deep,  with  a  subfoveiform 
impression  in  its  course  a  little  before  base;  lateral  basal  impres- 
sions rather  short,  wide,  deep.  Elj^tra  truncate-oval  (Tw  x 
4*3  mm.),  lightly  declivous  to  apex,  sides  lightly  rounded;  apex 
sinuate  on  each  side  (strongly  so  in  9);  basal  border  a  little  raised 
and  obtusely  subdentate  at  humeral  angles;  lateral  border  narrow; 
stride  deep;  interstices  strongly  convex  in  g,  lightly  so  in  9,  third 
2-punctatej  hardly  wider  than  second,  sixth  narrowed  at  base, 
eighth  much  wider  than  ninth  on  basal  half,  ninth  narrow,  convex, 
interrupted  on  inner  side  by  the  umbilicate  punctures,  tenth  well 
developed,  elongate  (extending  forward  to  about  middle  of  elytra), 
narrow,  convex.  Intercoxal  declivity  of  prosternum  flat ;  of 
mesosternum  hardly  concave.     Length  13-5,  breadth  4*3  mm. 

Hab. — Clarence  River  (Lea). 

Allied  to  iV.  nitidicollis,  Chaud.,  but  differing,  intej^  alia,  by 
its  facies  narrower,  more  elongate,  more  convex  ;  head  more 
-depressed .  between  eyes;  prothorax  less  ampliate  at  widest  part, 
less  emarginate  at  apex,  sides  without  juxtabasal  sinuosities;  elytra 
less  strongly  declivous  to  apex,  less  strongly  dentate  at  humeral 
angles,  third  interstice  narrower  at  beginning  of  apical  declivity, 
eighth  much  wider  on  basal  half,  tenth  well  developed,  lateral 
border  narrower.  It  differs  from  iV^.  queenslandicus,  SI.,  and  iV^. 
latibasis,  SI.,  by  the  same  characters. 

j}^ote. — N.  tvilcoxi,  Casteln.  (Trans.  Roy.  Soc.  Vict.  1868,  viii. 
p.  211).  Although  Chaudoir  has  considered  that  JV.  subopacus 
could  not  be  J\^.  loUcoxi  because  Castelnau's  description  of  the 
prothorax  of  N.  wilcoxi  did  not,  in  his  opinion,  at  all  agree  with 
the  prothorax  of  N.  subopacKs,  yet  I  should  say  that  the  only 
discrepancy  is  the  "  rather  acute  "  posterior  angles  of  i\^.  wilcoxi^ 
which  may  mean  that  the  prothorax  should  have  the  sides  lightly 
sinuate  near  the  base,  and  this  feature  may  not  be  of  sufficient 
importance  to  justify  keeping  the  species  distinct.  I  should 
-expect  that  xY.  subopacus  might  pro^•e  to  be  only  a  variety  of  J\^. 


BY    THOMAS    G.    SLOAN E.  303 

wilcood.     This  is  a  point  which  collecting  on  the  Clarence  River 
should  eventually  settle. 

Hab. — Clarence  River  (Jide  Castelnau). 

NOTONOMUS    VIOLACEOMARGINATUS,   Maclcay. 

Trans.  Ent.  Soc.  N.S.W.  1871,  ii.  p.  108;  Notonomus  jiuiyii^rei- 
pe7inis,  Macl.,  Lc,  p.  107. 

9.  Oval.  Prothorax  subquadrate ;  sides  not  sinuate  before 
base;  basal  angles  obtuse;  posterior  marginal  puncture  placed  a 
little  before  base  on  inner  side  of  marginal  channel :  elytra 
truncate-oval,  deeply  striate;  interstices  depressed,  third  2-punc- 
tate;  humeral  angles  lightly  dentate.  Head  and  prothorax  of  a 
greenish-black ;  el3^tra  opaque,  purple-black,  lateral  channel 
cyaneous;  under  surface  and  legs  black. 

Head  moderate  (3*1  mm.  across  eyes),  wide  between  eyes;  front 
shortly  and  distinctly  biimpressed.  Prothorax  subquadrate 
(4-2x5  mm.),  widest  about  middle,  depressed ;  sides  lightly 
rounded,  gently  narrowed  to  base  in  an  even  curve;  apex  lightly 
emarginate,  narrower  (3-4  mm.)  than  base  (3-8  mm.);  anterior 
angles  widely  obtuse,  distant  from  head;  base  emarginate  in 
middle,  sloping  forward  on  each  side  in  a  gentle  curve;  median 
line  reaching  from  base  to  near  apex;  lateral  basal  impressions 
elongate,  narrow  at  bottom.  Elytra  truncate-oval  (10x6  mm.), 
strongly  sinuate  on  each  side  of  apex,  rather  depressed  on  disc, 
lightly  declivous  to  apex,  abrupt  on  sides;  basal  border  very 
lightly  curved  on  posterior  margin,  a  little  raised  and  slightly 
prominent  at  humeral  angles  ;  lateral  channel  wide ;  border 
narrow,  reflexed ;  sixth  interstice  hardly  narrower  than  fifth 
except  near  base,  seventh  wide,  eighth  a  little  wider  than  ninth 
on  basal  half,  tenth  well  developed,  narrow,  elongate  before  apical 
sinuosity.  Intercoxal  declivity  of  prosternum  flat ;  of  meso- 
sternum  flat.     Length  16*5,  breadth  6  mm. 

Hab. — Q.  :  Gayndah  (Masters  and  Spencer). 

I  have  compared  the  specimen  (9),  on  which  the  above  descrip- 
tion is  founded,  with  Macleay's  types  of  N.  piLVpureipennis  and 


304  EEVISION  OF  THE  GENUS  XOTONOMUS. 

iV.  violaceomarginatus,  and  found  it  identical  with  the  latter.     iV. 
purpureipe7i7iis  is  founded  on  the  (J. 

NOTONOMUS    RUGOSICOLLIS,  n.sp. 

(J,  Elliptical.  Protliorax  opaque,  densely  shagreened  and 
transversely  striolate;  sides  lightly  sinuate  before  basal  angles; 
posterior  marginal  puncture  in  lateral  channel  near  basal  angle : 
elytra  deeply  striate;  interstices  strongly  convex,  third  narrower 
than  second  on  basal  third,  2-punctate;  basal  border  prominent 
at  humeral  angles.  Head  and  prothorax  black  (a  faint  purple 
tinge  near  sides  of  prothorax);  elytra  bronzy-black,  submetallic 
towards  sides,  metallic  behind  basal  border,  ninth  interstice  and 
lateral  channel  of  a  bright  brassy  colour;  under  surface  black; 
legs  piceous-black. 

Head  not  large  (2-5  mm.  across  eyes),  subdepressed  between 
eyes,  these  prominent.  Prothorax  broader  than  long  (3  x  3 -6 mm), 
depressed  on  disc;  sides  lightly  rounded,  gently  narrowed  to  apex, 
very  lightly  narrowed  to  base  ;  anterior  angles  obtuse  ;  base 
(2-7  mm.)  wider  than  apex  (2-5  mm.),  emarginate  in  middle; 
basal  angles  rectangular;  lateral  channel  wide;  median  line  deep, 
reaching  from  base  almost  to  apex;  lateral  basal  impressions  long, 
narrow,  deep;  posterior  part  of  space  between  basal  impressions 
longitudinally  rugulose  but  not  transversely  striolate.  Elytra 
truncate-oval  (7*7  x  4*35  mm.),  lightly  and  widely  convex,  lightly 
but  decidedly  convex  on  each  side  of  apex;  sides  lightly  rounded; 
basal  border  arcuate  on  posterior  margin,  strongly  raised  and  sub- 
dentate  at  humeral  angles;  lateral  channel  wide;  lateral  border 
narrow,  reflexed,  not  narrower  near  base;  sixth  interstice  narrowed 
at  base,  eighth  much  wider  than  ninth  on  basal  half,  about  as  wide 
as,  but  much  more  depressed  than,  discoidal  interstices  at  middle 
of  length,  ninth  narrow,  convex,  interrupted  on  inner  margin  by 
umbilicate  punctures,  tenth  shortly  but  decidedly  developed 
before  lateral  apical  sinuosity.  Intercoxal  declivity  of  prosternum 
flat;  of  mesosternum  wide,  flat.     Length  12-5,  breadth  4*35  mm. 

Hah. — N.S.W.  :  Clarence  River  (Lea;  Coll.  Sloane;  unique). 


BY    THOMAS    G.    SLOANE.  305 

This  species  has  a  superficial  resemblance  to  iV.  subopacus, 
Chaud.,  but  differs  conspicuously,  inter  alia,  hy  the  prothorax 
rugulose-striolate  and  sinuate  on  the  sides  near  the  base.  Its 
position  is  intermediate  between  N.  opacicoUis,  Chaud.,  and  iV^. 
disco riniosus,  SI. ;  from  JV.  opacicoUis  it  differs  greatly  by  its 
smaller  size,  narrower  form,  less  transverse  prothorax,  non-foveolate 
elytral  interstices,  &c.;  from  iV^.  discorimosus  it  differs  by  its 
prothorax  more  elongate,  with  sides  sinuate  before  the  base, 
posterior  angles  strongly  marked;  elytra  with  the  eighth  interstice 
wide  as  in  N.  opacicoUis.  From  N.  striaticoUis,  Casteln.,  a  species 
not  known  to  me  in  nature,  I  think  it  distinct,  because  the  pro- 
thorax has  the  sides  sinuate  before  the  base,  and  the  basal  angles 
rectangular.  It  evidently  cannot  be  iT.  depressij^^mis,  Chaud., 
from  the  description  of  which  it  offers  conspicuous  differences  as 
under : — (a)  lateral  basal  impressions  of  prothorax  longer  and 
quite  as  deep  as  in  iV.  opacicoUis;  (b)  elytra  not  flatter;  (c)  six 
inner  interstices  not  the  least  depressed  (in  (J);  (d)  eighth  inter- 
stice wide  on  basal  half  (not  very  narrow  and  convex). 

NOTONOMUS    DISCORIMOSUS,    n.sp. 

(J.  Oval.  Prothorax  transverse,  with  sides  strongly  and  evenly 
rounded :  elytra  truncate-oval,  deeply  striate;  interstices  convex, 
third  2-punctate,  eighth  narrow;  humeral  angles  dentate.  Head 
black;  prothorax  opaque,  black  with  a  faint  viridescent  tinge  on 
disc,  becoming  metallic  near  sides;  elytra  subnitid,  dark  purple, 
becoming  aeneous  near  base  and  on  lateral  declivities,  ninth  inter- 
stice and  lateral  margin  of  a  bright  brassy  colour.  (9  with  elytra 
not  opaque,  of  a  uniform  bronzed  purple  with  lateral  channel 
green. )  # 

Head  not  large;  eyes  convex,  prominent.  Prothorax  depressed, 
transverse  (3-3  x  4  mm.);  wider  across  base  (2*7  mm.)  than  apex 
(2-5  mm.);  upper  surface  transversely  striolate  and  longitudinally 
shagreened ;  sides  evenly  rounded,  narrowed  posteriorly  in  an 
even  curve  without  any  antebasal  sinuosity;  apex  emarginate; 
base  lightly  and  widely  emarginate  in  middle;  basal  angles  obtuse; 
20 


306  REVISION  OF  THE  GENUS  NOTOyOMUS, 

lateral  border  and  channel  wide;  posterior  marginal  puncture 
foveiform,  placed  a  little  distance  from  basal  angle  on  inner  side 
of  marginal  channel;  median  line  strongly  impressed,  reaching 
from  base  almost  to  apex;  lateral  basal  impressions  short,  narrow, 
deep,  uniting  with  lateral  channel  at  base.  Elytra  lightly  convex, 
oval  (7 '5  X  4  mm.),  evenly  and  decidedly  rounded  on  sides,  widely 
sinuate  on  each  side  of  apex;  lateral  channel  and  border  wide. 
Intercoxal  declivity  of  prosternum  flat;  of  mesosternum  vertical 
and  lightly  concave.  Femora  short,  stout;  posterior  femora  very 
thick  in  middle.     Length  13,  breadth  4-6  mm. 

Hab. — N.S.W.  :  Richmond  River  (Froggatt  and  Lea). 

Var.  A.  Smaller:  prothorax  less  strongly  rounded  on  sides 
and  more  narrowed  to  base;  basal  angles  more  marked:  elytra 
more  depressed  on  disc;  interstices  opaque  in  both  sexes:  colour 
duller;  elytra  purple-black,  margined  with  green. 

Hab. — Tweed  River  (Lea). 

Differs  evidently  from  N.  depressijyennis,  Chaud.,  by  prothorax 
shorter  and  without  the  juxtabasal  sinuosities  of  sides.  I  have 
thought  that  it  could  not  be  N.  striaticoUis,  Casteln.,  because  of 
its  short,  broad  thorax,  not  "  considerabl}'  longer  than  broad." 

jSTotgngmus  opacicollis,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  88;  Orbitus  purpuripennis,  Motsch., 
Ball.  Mosc.  1865,  ii.  p.  248;  Feronia  ( Pterostichus)  purpureo- 
limbata,  Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  214  ; 
Var.  F.  (Pi.)  impressipennis,  Casteln.,  I.e.,  p.  214. 

Black,  elytra  with  metallic  border.  Prothorax  roughly 
shagreened  and  transversely  striolate,  transverse  (4-3  x  5-4  mm.), 
much  wider  at  base  (4-2  mm.)  than  apex  (3  5  mm.);  posterior 
angles  rectangular ;  posterior  marginal  puncture  in  marginal 
channel  at  basal  angle.  Elytra  deeply  striolate;  interstices  nitid 
and  convex  in  ^  (sometimes  rather  metallic),  depressed  and  opaque 
in  9,  third  2-punctate,  second,  fourth  and  sixth  wide  and  foveolate 
on  apical  third,  eighth  wider  than  seventh  at  basal  third,  ninth 
narrow,  convex,  with  inner  side  interrupted  by  large  umbilicate 


BY    THOMAS    G.    SLOANE.  307 

punctures ;  humeral  angles  dentate.  Intercoxal  declivity  of 
prosternum  flat;  of  mesosternum  flat.  Length  15-17,  breadth 
6-6  8  mm. 

Hah. — "Clarence  River"  {fide  Castelnau),  Richmond  River 
(Helms). 

The  only  species  known  with  the  interstices  of  the  elytra  foveo- 
late.  The  synonymy  given  is  on  the  authority  of  Chaudoir  (Ann. 
Mus.  Civ.  Genova,  1874,  vi.,  p.  585). 

NoTONOMUS  AMABiLis,  Casteluau. 

Ferotiia  {Pterostichus)  amahilis,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1863,  viii.  p.  215;  N.  amahilis,  Chaud.,  Ann.  Mus.  Civ.  Genova, 
1874,  vi.  p.  590. 

Elliptical-oval.  Head  small  (2*8  mm.  across  eyes);  pro  thorax 
subcordate;  sides  shortly  subsinuate  before  basal  angles :  elytra 
oval,  deeply  crenulate-striate;  third  interstice  3-  or  4-punctate, 
ninth  convex;  humeral  angles  not  marked.  Head  black,  some- 
times with  greenish  metallic  tinge;  prothorax  with  wide  metallic 
margins  of  brassy  green  becoming  cupreous-purple  internally, 
disc  dark  with  purple  reflections;  elytra  dark  purple,  with  lateral 
channel  bright  metallic  brassy-green;  under  surface  black;  pro- 
sternal  episterna  iridescent;  legs  piceous-black,  tarsi  and  palpi 
piceous-red;  antennae  piceous-brown. 

Ej'es  convex,  rather  protuberant;  postocular  part  of  orbits  not 
large,  rising  from  head  in  a  carve  almost  continuous  with  the 
eyes.  Labrum  lightly  emarginate.  Apical  joints  of  antennae 
short.  Prothorax  broader  than  long  (3-5  x  4  mm.),  widest  a  little 
before  the  middle;  apex  emarginate,  a  little  wider  (2-8  mm.)  than 
base  (2-7  mm.);  sides  lightly  rounded,  lightly  narrowed  to  apex, 
decidedly  narrowed  in  a  gentle  curve  posteriorly,  shortly  sinuate 
before  basal  angles  (in  9  the  sinuosity  often  obsolete);  anterior 
angles  obtuse,  widely  bordered;  basal  angles  strongly  marked  but 
obtuse;  base  emarginate  in  middle,  curving  decidedly  forward  on 
each  side;  lateral  border  rather  widely  reflexed,  hardly  narrower 
towards  apex  than  towards  base;  median  line  strongly  impressed. 


308  EE VISION  OF  THE  GENUS  NOTONOMUS, 

reaching  base;  lateral  basal  impressions  deep,  narrow,  elongate. 
Elytra  oval  (8x5  mm.)  ;  lateral  apical  sinuosities  strongly 
developed ;  base  narrow ;  basal  border  joining  lateral  border 
without  any  prominence  at  humeral  angles;  interstices  strongly 
and  roundly  convex,  eighth  about  same  width  as  ninth  on  basal 
half;  umbilicate  punctures  of  ninth  interstice  large,  interrupting 
inner  margin;  lateral  channel  finely  crenulate  near  inner  side 
along  course  of  ninth  stria ;  lateral  border  widely  reflexed. 
Intercoxal  declivity  of  prosternum  rather  narrow  in  middle,  flat, 
bordered  on  each  side  near  peduncle;  of  mesosternum  widely  and 
lightly  concave.     Length  14-15,  breadth  4 '5-6  mm. 

Hab, — N.S.W.  :  Springwood  and  Ourimbah  (Sloane),  Merim- 
bula  (fide  Castelnau). 

NoTONOMUS  SPHODROIDES,  Dejean. 

Feronia  sphodroides,  Dej.,  Spec.  iii.  p.  236  :  i\".  sjyhodroideSy 
Chaud.,  Bull.  Mosc.  1865,  ii.  p.  91;  Ann.  Mus.  Civ.  Genova,  1874, 
vi.  p.  587. 

Elliptical-oval,  depressed.  Prothorax  truncate-cordate;  basal 
angles  rectangular;  posterior  marginal  puncture  at  basal  angle : 
elytra  decidedly  sinuate  on  each  side  of  apex,  striate;  interstices 
lightly  convex  in  ^,  hardly  convex  in  9?  third  3-punctate  ; 
humeral  angles  a  little  raised.  Black,  nitid;  elytra  steel  blue; 
legs  piceous-black. 

Head  convex,  not  large  (2*6  mm.  across  eyes);  front  lightly 
biimpressed ;  eyes  convex,  prominent,  lightly  inclosed  at  base; 
postocular  part  of  orbits  meeting  head  in  a  curve  continuous  with 
that  of  the  eyes.  Prothorax  broader  than  long  (3-3  x  3*8  mm.), 
depressed  (especially  towards  base),  a  little  wider  at  base 
(2*8  mm.)  than  apex  (2*6  mm.);  sides  lightly  narrowed  to  apex, 
gently  narrowed  to  base,  lightly  sinuate  before  basal  angles;  base 
lightly  emarginate  in  middle ;  basal  angles  strongly  marked, 
rectangular  with  summit  obtuse;  lateral  border  narrow  anteriorly, 
wider  posteriorly,  dilated  at  basal  angles  to  receive  posterior 
marginal  puncture;  median  line  finely  but  rather  deeply  impressed, 


BY    THOMAS    G.    SLOANE.  309 

reaching  nearly  to  base;  lateral  basal  impressions  shallow,  sulci- 
form  at  bottom,  parallel;  lateral  basal  spaces  flat  near  basal 
margin.  Elytra  truncate-oval  (7*2  x  4*7  mm.);  basal  border  a 
little  raised  above  lateral  border  at  point  of  junction;  ninth  stria 
well  marked;  tenth  interstice  well  developed  posteriorly.  Inter- 
coxal  declivity  of  prosternum  flat;  of  mesosternum  wide,  very 
lightly  concave.     Length  13,  breadth  4*7  mm. 

Hab. — Vic.  :  Dandenong  Ranges  (French),  Ferntree  Gully 
(Sloane). 

I  place  this  species  under  N.  sphodroides,  Dej.,  because,  except 
for  the  absence  of  any  purple  tinge  on  the  prothorax,  a  specimen 
(9)  from  the  Dandenong  Ranges  given  to  me  by  Mr.  C.  French, 
differs  from  the  species  which  I  regard  as  A^.  dyscoloides,  Motsch., 
by  similar  characters  to  those  which  are  said  by  Chaudoir  to 
differentiate  N.  sphodroides  from  N.  dyscoloides.  The  following  is 
the  substance  of  Chaudoir's  note  on  these  differences  in  his 
*  Supplement,'  p.  587  : — Prothorax  more  rounded  on  the  anterior 
part  of  the  sides,  these  hardly  sinuate  behind,  and  falling  a  little 
obliquely  on  the  base,  forming  with  it  an  obtuse  angle,  the  summit 
of  which  is  lightly  rounded.  The  elytra  are  more  oval,  with  the 
shoulders  rounded  as  well  as  the  sides,  the  upper  surface  percep- 
tibly more  convex,  not  flattened  on  the  disc;  the  intermediate 
strise  still  more  lightly  marked;  the  elytra  are  not  violet,  but  of 
a  greenish  blue.  I  do  not  place  much  reliance  on  the  slightly 
more  protuberant  eyes,  nor  the  form  of  the  lateral  basal  spaces 
of  the  prothorax  mentioned  by  Chaudoir;  his  remark,  that  he 
had  confused  this  species  with  N.  dyscoloides,  indicates  how  closely 
these  species  are  allied,  but  I  concur  in  thinking  them  distinct^ 
Only  the  9  was  known  to  Chaudoir;  I  have  both  ^  and  9;  the 
description  I  have  given  is  founded  on  the  (J. 

XOTONOMUS    DYSCOLOIDES,  Motschulsky. 

Neuropates  dyscoloides,  Motsch.,  Bull.  Mosc.  1865,  ii.  p.  263; 
Feronia  (Pterostichus)  victorice,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  viii.  p.  210;  F.  (Pi.)  semiviolacea,  Casteln.,  I.e.,  p.  212;  F. 
(Pt.)  subvilis, Casteln., I.e.,  p.  21 3;  Feronia  (A^otonomus) dyscoloides, 
Chaud.,  Ann.  Mus.  Civ.  Genova,  1874,  vi.  p.  587. 


310  EEVISION  OF  THE  GENUS  NOTOyOMUS, 

Elliptical,  upper  surface  depressed.  Head  and  prothorax  black, 
nitid;  elytra  violaceous,  with  sutural  interstices  dark.  Head  not 
large  (2*8  mm.  across  ej^es);  eyes  convex,  prominent.  Prothorax 
broader  than  long  (3-4x4  mm.),  truncate-cordate,  subdepressed; 
sides  sinuate  posteriorly;  basal  angles  rectangular;  base  (3'1  mm.) 
hardly  wider  than  apex  (3  mm.);  border  narrow  anteriorly,  widely 
reflexed  towards  base,  extending  on  sides  of  base  to  lateral 
impressions;  posterior  marginal  puncture  at  basal  angle  on  an 
internal  dilatation  of  border;  lateral  basal  impressions  long, 
parallel;  lateral  basal  spaces  almost  flat.  Elytra  truncate-oval 
(8 "2  X  5  mm.),  depressed  on  disc,  strongly  declivous  to  apex,  deeply 
striate;  interstices  lightly  convex,  third  3-punctate,  eighth  and 
ninth  convex,  eighth  wider  than  ninth  on  basal  half;  basal  border 
hardly  raised  above  lateral  border  at  point  of  junction.  Inter- 
coxal  declivity  of  prosternum  wide,  flat;  of  mesosternum  wide, 
vertical,  hardly  at  all  concave.     Length  14*5,  breadth  5  mm. 

Hah. — Victoria  (W.  Kershaw). 

Judging  by  the  synonymy  given  above  (on  the  authority  of 
Chaudoir)  N.  dyscoloides  is  a  very  variable  species.  I  take  the 
form  on  which  my  description  is  founded  to  be  Feronia  semivio- 
lacea,  Casteln.,  and  I  should  have  thought  F.  victories  from  des- 
cription to  have  been  synonymous  Avith  iV^.  sphodroides,  Dej.  I 
have  not  seen  Motschulsky's  description,  but  have  identified  the 
species  from  Castelnau's  diagnoses  and  Chaudoir's  note. 

NoTONOMUS  siMULANS,  Chaudoir. 

Ann.  Mus.  Civ.  Genova,  1874,  vi.  p.  586. 

I  have  not  identified  this  species.  The  following  is  a  translation 
of  the  too  brief  description  : — Length  12-5  mm.  It  much 
resembles  A^.  dyscoloides  in  facies,  but  is  differently  coloured.  Of 
a  very  brilliant  black  as  if  polished;  elytra  coppery,  with  the 
margin  black;  e3''es  more  prominent;  elytra  a  little  less  oval,  more 
declivous  to  tlie  apex,  more  strongly  striate;  interstices  more 
convex,  with  thrr.e  ov  four  large  punctures  on  the  third. 

Hah. — Southern  Australia. 


BY    THOMAS    G.    SLOANE.  311 

NOTONOMUS  VICTORIENSIS,  n.sp. 

Elon^ate-oval,  subdepressed.  Head  not  large :  prothorax  a 
little  broader  than  long;  sides  sinuate  posteriorly;  basal  angles 
rectangular;  posterior  marginal  puncture  at  basal  angle:  elytra 
long,  strongly  striate;  interstices  convex  in  (J,  depressed  in  9, 
third  2-punctate;  humeral  angles  shortly  dentate.  Black,  nitid; 
elytra  with  a  greenish  tinge;  sides  of  abdomen  piceous;  epimera 
of  metasternum  reddish;  legs  piceous-red. 

(J.  Head  oval  (3  mm.  across  eyes);  front  feebly  biimpressed. 
Prothorax  broader  than  long  (3-7  x  4-1  mm.),  depressed;  apex  and 
base  of  equal  width  (3  mm.);  sides  lightly  rounded  on  anterior 
four-fifths,  strongly  and  shortly  sinuate  posteriorly,  meeting  base 
at  right  angles ;  border  narrow,  thickened  at  basal  angles  to 
receive  posterior  marginal  puncture;  median  line  distinct;  lateral 
basal  impressions  elongate,  rather  shallow.  Elytra  truncate-oval 
(8-7  X  4-8  mm.),  lightly  convex;  sides  lightly  rounded  (subparallel) 
lateral  apical  sinuosities  very  feebly  developed;  basal  border  very 
little  raised,  obtusely  but  decidedly  dentate  at  humeral  angles; 
third  interstice  swollen  at  beginning  of  apical  declivity  (position 
of  posterior  puncture),  eighth  wider  than  ninth  on  basal  half, 
extending  almost  to  base,  tenth  well  developed  posterior^. 
Intercoxal  declivity  of  prosternum  wide  in  middle,  hardly 
rounded  ;  of  mesosternum  lightly  concave.  Posterior  tibiae  a 
little  curved. 

9.  Elytra  more  rounded  on  sides,  less  deeply  striate;  sixth  and 
seventh  striae  very  lightly  impressed  on  basal  half;  interstices 
depressed.     Length  13-15,  breadth  4 -7-5 -2  mm. 

Hah. — Vic.  :  Eastern  Gippsland  (taken  by  Mr.  C.  French 
during  the  visit  of  the  Victorian  Field  Naturalists'  Club  to  Mount 
Goonmurk,  November,  1889). 

Allied  to  N.  dyscoloides,  Motsch.,  but  of  narrower  form;  pro- 
thorax narrower,  more  elongate;  third  interstice  of  elytra  swollen 
at  posterior  puncture;  intercoxal  declivity  of  prosternum  not  flat 
and  margined;  posterior  tibiae  slightly  curved;  legs  and  under 
surface  more  reddish-piceous. 


312  REVISION  OF  THE  GENUS  yOTOSOMUS, 

Note. — One  specimen  ((J)  is  less  elongate;  prothorax  a  little 
shorter;  elytra  more  metallic  in  colour  (bronzy  or  brassy  purple), 
with  more  convex  interstices,  puncture  of  third  interstice  near 
apex,  eighth  not  reaching  the  base.  Length  13,  breadth  4-7  mm. 
I  should  expect  N.  simulans,  Chaud.,  to  resemble  this  form  in 
colour,  but  to  be  a  different  species  intermediate  between  N. 
victoriensis  and  N.  dyscoloides. 

NoTONOMUS  OPULENTUS,  Castehi.;  Chaudoir. 

Feronia  (Fterostichus)  opulenta,  Casteln.,  Trans.  Roy.  Soc. 
Vict.  1868,  viii.  p.  210;  N.  opulentus,  Chaud.,  Ann.  Mus.  Genova, 
1874,  vi.  p.  580;  Feronia  (Pterostichus)  auricollis,  Casteln.,  /.c, 
p.  211;  C?)  N.  parallelomorpha,  Chaud.,  Bull.  Mosc.  1878,  liii. 
p.  73. 

Elongate.  Prothorax  and  posterior  part  of  head  aeneous  (pro- 
thorax  sometimes  of  a  purplish  brassy  colour);  elytra  dark  violet; 
under  surface  black,  nitid;  episterna  of  prosternum  iridescent; 
legs  black.  Head  small  (2-6  mm.  across  eyes),  oval,  convex  ; 
frontal  impressions  distinct,  sharpl}^  defined,  short,  diverging 
backwards;  eyes  convex,  not  protuberant.  Prothorax  of  equal 
length  and  breadth  (3-8  x  3*8  mm.),  widest  before  middle,  not  the 
least  ampliate  at  widest  part,  wider  across  base  (3  mm.)  than 
apex  (2*7  mm.),  depressed  across  basal  third;  sides  lightly  and 
obliquely  narrowed  to  base,  shortly  subsinuate  before  base  (the 
posterior  sinuosity  sometimes  obsolete);  anterior  angles  obtuse, 
not  marked;  base  emarginate  in  middle,  truncate  on  each  side; 
basal  angles  rectangular  (sharply  marked);  border  narrow,  a  little 
more  widely  reflexed  posteriorly  than  anteriorly,  dilated  at  basal 
angles  to  receive  posterior  marginal  puncture,  extending  on  each 
side  of  base  to  lateral  basal  impressions — these  deep,  elongate, 
narrow,  sulciform  at  bottom  ;  lateral  basal  spaces  depressed. 
Elytra  long  (8x4*8  mm.),  subdepressed,  strongly  declivous  to 
apex,  truncate  at  base;  sides  lightly  rounded,  gently  narrowed  to 
base;  apical  sinuosities  well  developed;  basal  border  raised  into 
a  short,  strong  subdentiform  prominence  at  each  humeral  angle; 


BY    THOMAS    G.    SLOANB.  313 

lateral  border  reflexed,  rather  wide  on  middle  of  sides,  very 
narrow  near  base;  striae  deep;  interstices  convex,  very  strongly  so 
on  apical  declivity,  third  wide,  usually  3-punctate  (rarely  a  fourth 
puncture  present  about  basal  fourth,  rarely  only  2-punctate), 
eighth  a  little  wider  than  ninth  on  basal  half  and  reaching  to 
base,  ninth  narrow,  interrupted  on  inner  margin  by  the  umbilicate 
punctures,  tenth  well  developed  towards  apex,  short,  narrow 
(subcarinate).  Intercoxal  declivity  of  prosternum  wide,  flat;  of 
mesosternum  flat,  hardly  concave.  Fourth  and  fifth  ventral 
segments  very  lightly  and  widely  transversely  impressed;  apical 
segment  in  ^  6-punctate.  9  with  first  joint  of  anterior  tarsi 
wide  (not  dilatate),  clothed  beneath  with  a  thick  brush  of  whitish 
squamulse.     Length  11-7-15-5,  breadth  4-1-5  mm. 

Hah.  — Yic.  :  Upper  Yarra  (Best),  South  Gippsland  (French). 
•  A  distinct  species  varying  greatly  in  size  and  appearance.  Mr. 
Masters  has  informed  me  that  it  is  to  his  knowledge  Fernnia 
opuleiita,  Casteln.,  and  I  had  arrived  at  the  same  conclusion  from 
Castelnau's  too  brief  description.  I  also  think  that  Chaudoir's 
description  of  JV.  parallelomorpha  is  founded  on  a  form  of  iV". 
opulentus;  one  of  the  specimens  in  my  collection  agrees  closely  with 
his  description,  even  having  three  punctures  on  the  third  interstice 
of  one  elytron  and  two  on  the  other.  If  I  am  right  in  my  identi- 
fication of  N.  parallelomor'pha,  Queensland  is  not  its  habitat.  I 
place  Feronia  auricollis,  Casteln.,  as  a  synonym  of  N.  opulentus 
on  Chaudoir's  authority. 

NoTONOMus  .ENEOMiCANs,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  84;  Feronia  (^Pterosticlius)  lapeyrousei, 
Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  213. 

9.  Robust,  convex.  Prothorax  cupreous  (very  dark  on  disc, 
bright  tov/ards  sides);  head  black,  with  faint  cupreous  tinge; 
elytra  black,  with  faint  metallic  tinge,  ninth  interstice  and  margin 
cupreous,  inflexed  margins  with  an  aeneous  tinge.  Head  large 
(4-5 mm. across  eyes).  Prothorax  truncate-cordate (5 -15  x  6-5 mm.), 
a  little  wider  across  base  (4*7  mm.)  than  apex  (4-5  mm.);  sides 


314  EEVISION  OF  THE  GENUS  ^'OTONOMUS, 

lightly  sinuate  just  before  base ;  basal  angles  subrectangular; 
posterior  marginal  puncture  on  border  at  basal  angle;  lateral 
basal  impressions  strongly  impressed,  uniting  with  lateral  channel 
at  posterior  extremity.  Elytra  truncate-oval  (13x8-3  mm.), 
lightly  convex;  basal  border  strongly  raised  and  obtusely  dentate 
at  humeral  angles;  lateral  border  widely  reflexed  from  humeral 
angle  to  apical  sinuosity;  strise  deep,  minutely  crenulate  at  bottom; 
interstices  convex,  third  4-  or  5-punctate,  tenth  short,  well 
developed  towards  apex.  Tntercoxal  declivity  of  prosternum  flat, 
bordered  on  sides;  of  mesosternum  lightly  concave.  Length  23, 
breadth  8*3  mm. 

Hab. — N.S.W.  :  Ash  Island,  Hunter  River  (Jide  Castelnau). 

My  specimen  was  given  to  me  by  Mr.  A.  M.  Lea  as  from  the 
Manning  River. 

Var.  A.  Form  more  elongate ;  prothorax  more  decidedly 
narrowed  to  base,  and  with  posterior  lateral  sinuosities  longer 
and  more  marked  ;  elytra  less  convex.  Head  blackish-green; 
prothorax  blackish-green  on  disc,  bright  green  towards  sides; 
elytra  black,  with  ninth  interstice  and  lateral  margin  bright 
green;  inflexed  margins  blackish-green.  Length  21-25,  breadth 
7-2-8-5  mm. 

Hab. — Narrara  and  Ourimbah  (Sloane  ;  plentiful  in  damp 
heavily  timbered  places  in  July,  1896). 

NoTONOMUS  COLOSSUS,  n.sp. 

(J.  Large,  robust,  convex.  Head  large  (4-7  mm.  across  eyes); 
prothorax  truncate-cordate,  not  sinuate  on  sides  before  base; 
basal  angles  rounded;  posterior  marginal  puncture  on  border  at 
basal  angle :  elytra  strongly  striate ;  interstices  convex,  third 
5-punctate;  basal  border  not  dentate  at  humeral  angles.  Head 
black,  with  a  very  faint  bronzy  tinge;  prothorax  black,  cupreous 
towards  sides,  with  a  greenish  tinge  near  lateral  channel  (disc 
with  slight  cupreous  reflections);  elytra  bronzy  black,  ninth  inter- 
stice and  margin  of  a  dark  copper  colour. 

Front  lightly  biimpressed;  eyes  large,  convex;  postocular  part 
of  orbits  small,  but  rising  sharply  from  head.     Prothorax  broader 


BY    THOMAS    G.    SLOANE.  315 

than  long  (5-5  x  7  mm.),  about  same  width  (4-7  mm.)  at  base  and 
apex;  sides  strongly  rounded,  meeting  base  in  an  even  curve; 
border  wide,  reilexed;  lateral  basal  impressions  short,  deep,  uniting 
with  lateral  channel  at  posterior  extremity.  Elytra  truncate- 
oval  (13x8  mm.);  lateral  apical  sinuosities  wide;  lateral  border 
wide  and  sharply  reflexed  from  humeral  angle  to  apical  sinuosity; 
eighth  interstice  wider  than  ninth  on  basal  half,  tenth  short,  well 
developed  towards  apex.  Intercoxal  declivity  of  prosternum  flat, 
but  not  bordered  on  sides;  of  mesosternum  concave.  Length  24, 
breadth  8  mm. 

A  second  specimen  (9)  with  prothorax  narrow^er  (5-5  x  6*5 mm.), 
more  narrowed  to  base  (4-3  mm.). 

iTa^.— N.S.W.  :  Tamworth  (Musson). 

Closely  resembling  N.  (eneomicans,  Chaud.,  but  I  think  distinct. 
The  following  differences  may  be  noted  : — Prothorax  with  sides 
more  evenly  rounded  and  meeting  the  base  in  a  curve  without 
any  sinuosity,  basal  angles  obtuse;  basal  border  of  elytra  with  its 
posterior  margin  much  less  arcuate  (sloping  far  less  decidedly 
forward  to  the  humeral  angles),  and  only  a  little  raised  above  the 
lateral  border  (not  dentate)  at  point  of  junction. 

NoTONOMUS  RUFiPALPis,  Castelnau. 

Feronia  (Omaseus)  riifipaljns,  Casteln,,  Trans.  Roy.  Soc.  Yict. 
1868,  viii.  p.  216. 

Elliptical-oval,  robust,  black.  Head  large  (3*4  mm.  across  eyes), 
convex;  frontal  impressions  strongly  impressed,  divergent  poste- 
riorly; eyes  convex,  not  prominent  (eyes  with  orbits  reniform); 
postocular  part  of  orbits  greatly  developed,  protuberant  but  not 
so  prominent  as  eyes.  Prothorax  smooth,  broader  than  long 
(3 '9  X  4  5  mm.),  widest  before  middle,  lightly  narrowed  to  base; 
apex  (3-4  mm.)  broader  than  base  (3-2mm.);  sides  lightly  rounded; 
base  lightly  emarginate;  basal  angles  a  little  marked,  obtuse; 
lateral  border  thick,  narrow,  wider  posteriorly,  lightly  subsinuate 
just  before  basal  angles,  extending  round  basal  angles;  posterior 
marginal  puncture  in  the  wide  lateral  channel  at  basal  angle; 


316  EE VISION  OF  THE  GENUS  ^^OTONOMUS, 

median  line  strongly  impressed;  lateral  basal  impressions  narrow, 
rather  long;  lateral  basal  spaces  convex.  Elytra  truncate-oval 
(8-5  X  5-2  mm.),  rather  depressed  on  disc,  abruptly  declivous  on 
lateral  declivities;  deeply  striate;  sides  lightly  rounded:  lateral 
sinuosities  of  apex  feeble;  shoulders  rounded;  basal  border  slightly 
raised  above  lateral  border  at  humeral  angles  (hardly  subdentate); 
interstices  convex,  third,  fifth  and  seventh  a  little  wider  than 
others,  third  4-punctate,  eighth  narrow,  not  as  wide  as  ninth; 
tenth  interstice  and  ninth  stria  well  developed.  Prosternum  not 
bordered  on  anterior  margin;  basal  declivity  narrow  in  middle, 
bordered  on  sides:  mesosternum  wide,  not  concave  between  coxaa. 
Posterior  tibiae  lightly  incurved  near  apex.  Length  15,  breadth 
5  2  mm. 

Hab. — Victoria. 

A  distinct  species.  I  believe  it  to  be  Omaseus  o'l/Jipalpis , 
Casteln.,  the  only  apparent  differences  from  Castelnau's  descrip- 
tion being  that  it  is  a  little  larger,  and  the  third  interstice  of  the 
elytra  has  four — not  five — punctures  in  both  my  specimens  ; 
neither  difference  appears  to  me  of  specific  value. 

NoTONOMUS  EQUEs,  Castelnau. 

Feronia  (Pterostichus)  eques,  Casteln.,  Trans.  Roy.  See.  Vict. 
1868,  viii.  p.  208;  aY.  eques,  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874, 
vi.  p.  578. 

Elliptical,  parallel,  depressed.  Upper  surface  cupreous;  pro- 
thorax  and  lateral  channel  of  elytra  brighter  than  disc  of  elytra; 
under  surface,  inflexed  margins  of  elytra  and  legs  piceous-black. 
Head  not  large  (4  mm.  across  eyes);  front  strongly  biimpressed 
behind  clypeus;  eyes  convex,  but  not  prominent;  postocular  parts 
of  orbits  rising  gently  from  head,  not  protuberant.  Prothorax 
quadrate  (5-6  x  5-8  mm.),  widest  before  middle,  wider  across  base 
(4*7  mm.)  than  apex  (4*4  mm.)  ;  sides  very  lightly  rounded 
anteriorly,  gently  narrowed  posteriorly  and  meeting  base  obliquely 
without  perceptible  sinuosity;  basal  angles  obtuse,  but  marked; 
base  emarginate  in  rbiddle;  posterior  marginal  puncture  placed 


BY    THOMAS    G.    SLOANE.  317 

considerably  before  the  base  on  inner  side  of  lateral  channel — 
this  wide  and  flat,  especially  towards  base.  Elytra  truncate-oval 
(12-5  X  6-3  mm.),  depressed  on  disc,  deeply  striate;  interstices 
convex,  finely  shagreened,  third  5-punctate,  eighth  wider  than 
ninth  and  extending  almost  to  base,  tenth  feebly  developed; 
lateral  apical  sinuosities  strongly  developed;  basal  border  raised 
and  strongly  dentate  at  humeral  angles;  lateral  border  widely 
reflexed.  Intercoxal  declivity  of  prosternum  flat;  of  mesosternum 
widely  and  lightly  concave.  Posterior  femora  lightly  dilatate  in 
middle.  9  with  three  setigerous  punctures  on  each  side  of  apical 
ventral  segment.     Length  19-22,  breadth  6*3-7 '3  mm. 

Hah. — N.S.W.  :  lUawarra  ;  Burrawang  (Sloane;  common  in 
iSTovember). 

N.  eques  is  easily  identified  by  its  large  size,  flattened  dorsal 
surface  and  brassy  colour. 

NOTONOMUS    SPENCERI,  n.sp. 

9.  Oval,  robust,  convex.  Head  small :  prothorax  subquadrate, 
with  the  angles  rounded;  base  wide;  posterior  marginal  puncture 
distant  from  basal  angle :  elytra  oval,  strongly  striate;  interstices 
convex,  third  5-punctate.  Black;  head  with  a  slight  greenish 
tinge;  prothorax  bright  green  towards  sides;  elytra  with  green 
margin  (ninth  interstice  and  lateral  channel  green);  under  surface 
black;  legs  piceous. 

Head  small  (3*2  mm.  across  eyes);  frontal  impressions  distinct, 
wide,  parallel;  eyes  prominent.  Prothorax  broader  than  long 
(4  X  4*7  mm.),  convex;  sides  lightly  rounded,  narrowed  to  apex 
(3 -2  mm.)  in  an  even  curve,  very  lightly  narrowed  to  base 
(3-6  mm.)  in  a  gentle  curve;  basal  angles  widely  rounded;  border 
wide  and  reflexed  behind  anterior  marginal  puncture;  median 
line  lightly  impressed;  lateral  basal  imjDressions  deep,  narrow. 
Elytra  oval  (9*8  X  5-4  mm.),  convex;  sides  evenly  rounded;  lateral 
apical  sinuosities  well  developed;  humeral  angles  rounded;  basal 
border  a  little  raised,  subdentate  at  point  of  junction  with  lateral 
border;  tenth  interstice  feebly  developed  towards  apex.     Inter- 


318  REVISION  OF  THE  GENUS  XOTONOML^S, 

coxal  declivity  flat  and  margined  on  sides;  of  mesosternum  lightly 
concave.     Length  16,  breadth  5-4  mm. 

ffab. — Vic.  :  Eastern  Gippsland  (C.  French). 

This  species  must  be  closely  allied  to  -^^.  resplendevs,  Casteln., 
but  is  not  large  enough  nor  sufficiently  brightly  coloured  to  be 
that  species  ;  it  also  seems  to  have  the  prothorax  too  short.  It 
differs  from  JV.  strzeleckianus,  SI.,  by  head  smaller;  prothorax  less 
narrowed  to  base,  with  basal  angles  far  more  rounded;  elytra 
more  convex,  with  shoulders  much  less  marked,  and  with  the 
external  angle  of  the  basal  border  hardly  subdentate;  basal 
declivity  of  prosternum  wide,  flat  and  bordered  laterally.  From 
iV^.  froggatti,  SI.,  it  differs  by  its  more  convex  form;  prothorax 
more  evenly  rounded  on  sides,  with  basal  angles  more  rounded 
and  less  marked;  elytra  not  so  wide  at  base,  with  shoulders 
rounded  and  not  strongly  dentate;  form  of  basal  declivity  of 
prosternum;  posterior  femora  mueh  less  dilatatc,  tfec. 

NoTONOMUS  RESPLENDENS,  Castelnau. 

Feronia  (Pterostichus)  resplendens,  Casteln.,  Trans.  Roy.  Soc. 
Yict.  1868,  viii.  p.  213;  Chaud.,  Ann.  Mus.  Civ.  Genova,  1874,  vi. 
p.  579. 

I  do  not  know  this  species.  It  evidently  should  be  placed  near 
N.  eques^  Casteln.,  from  which  it  differs,  according  to  Chaudoir, 
inter  alia,  by  prothorax  a  little  more  elongate,  as  long  as  broad; 
posterior  marginal  puncture  placed  further  forward  near  the 
lateral  border;  elytra  with  shoulders  more  rounded,  and  with  the 
tooth  at  the  extremity  of  the  basal  border  obsolete,  interstices  a 
little  more  convex.     Length  19-20  mm. 

Hah. — N.S.W.  :  Merimbula  {fide  Castelnau). 

When  in  Sydney,  after  having  finished  my  notes  on  Noionomus, 
I  found  in  the  Macleay  Museum  the  following  undescribed  species, 
which  is  thoroughly  differentiated  from  all  others  by  the  rib-like 
interstices  of  the  elytra  separated  by  wide  opaque  sulci. 


BY    THOMAS    G.    SLOANE.  319 

NOTONOMUS    OPACISTRIATIS,   11. sp. 

Oval,  rather  depressed.  Head  elongate,  with  a  decided  trans- 
verse impression  across  vertex;  men  turn  with  sinus  oblique:  pro- 
thorax  subcordate ;  sides  sinuate  before  base ;  basal  angles 
rectangular  ;  posterior  marginal  puncture  on  border  at  basal 
angle ;  lateral  basal  impressions  short,  deep,  foveiform  :  elytra 
with  wide  opaque  striae;  interstices  strongly  convex,  nitid,  eighth 
and  ninth  narrow  and  close  together,  third  2-punctate.  Inter- 
coxal  declivity  of  prosternum  wide,  flat;  of  mesosternum  wide, 
not  concave  ;  metasternal  episterna  short.  Black;  antennae  and 
tarsi  piceous-red. 

Head  smooth;  frontal  impressions  linear,  deep,  oblique;  eyes 
convex,  subprominent;  posterior  part  of  orbits  swollen,  nearly  as 
long  as  and  a  little  less  prominent  than  eyes.  Prothorax  smooth, 
nitid,  broader  than  long  (4  x  4*5  mm,),  widest  a  little  before 
middle;  apex  and  base  of  equal  width  (3  mm.);  sides  lightly 
rounded,  shortly  and  rather  sharply  sinuate  just  before  base; 
anterior  margin  truncate;  anterior  angles  close  to  head;  base 
truncate;  border  narrow,  widest  towards  base;  lateral  channel 
narrow,  uniting  with  lateral  basal  impressions  posteriorly.  Elytra 
truncate-oval  (8-5  x  5-5  mm.),  lightly  and  widely  sinuate  on  each 
side  of  apex;  interstices  1,  3,  5  wider  than  others;  lateral  channel 
opaque;  striole  at  base  of  first  interstice  short,  deep;  lateral 
border  narrow  near  base;  basal  border  decidedly  raised  at  humeral 
angles  (subdentate).  Ventral  segments  punctate— first  strongly 
so,  three  apical  ones  with  a  finely  punctulate  wide  depressed  space 
on  each  side.     Length  16,  breadth  5*5  mm. 

Hah. — Q.  :  Cairns  (Froggatt;  two  specimens,  (J9). 

NoTONOMUS  MEDiosuLCATUS,  Chaudoir. 

Chaud.,  Bull.  Mosc.  1865,  ii.  p.  86  ;  Ann.  Mus.  Civ.  Genova, 
1874,  vi.  p,  590;  Adetiini  punctata,  Casteln.,  Trans.  Roy.  Soc. 
Vict.  1868,  viii.  p.  157;  Feronia  {Omaseus)  occideiitalis,  Casteln. ^ 
I.e.,  p.  220  ;  Fer.  (^Omaseus)  satanas,  Casteln.,  (*?)  I.e.,  p.  221 ;  N. 
mediosiUcaius,  Sloane,  Proc.  Linn.  Soc.  N.S.W.  1898,  xxiii.  p.  478. 


320  REVISION  OF  THE  GENUS  XOTOSOMUS, 

Oval,  convex.  Black;  elytra  often  of  a  greenish  or  purple 
colour.  Head  oval,  convex.  Pro  thorax  broader  than  long 
(4  X  4-5  mm.),  rounded  on  sides,  a  little  more  strongl}'-  so  poste- 
riorly than  anteriorly;  basal  angles  widel}^  rounded,  not  the  least 
marked;  posterior  marginal  puncture  on  edge  of  border;  lateral 
basal  impressions  deep,  foveiform.  Elytra  oval  (9  x  5'3  mm.); 
four  inner  striae  strongly  impressed,  5-7  obsolete,  third  2-punctate. 
Prosternum  with  anterior  margin  bordered.  Intercoxal  declivity 
rounded.     Length  14-16,  breadth  4*7-5 -3  mm. 

Hab. — S.W.  Australia  :  Swan  River,  Pinjarrah,  Donnybrook 
and  Bridgetown  (Lea),  Albany  (Masters). 

iV^.  mediosidcatus  has  no  affinity  to  any  other  species.  It  is  an 
isolated  species,  and  varies  greatly. 

NoTONOMUS  MOLESTUS,  Chaudoir. 

Feronia  [Notonomus)  molesta,  Chaud.,  Bull.  Mosc.  1865,  ii. 
p.  89  ;  Ternox  ohsoletus,  Motsch.,  I.e.,  p.  268 ;  Per.  (Percus) 
lacustris,  Casteln.,  Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  222. 

Robust,  convex.  Black,  shining.  Elytra  obsoletely  striate; 
third  interstice  depressed,  2-punctate  on  posterior  half;  humeral 
angles  strongly  dentate.     Length  23,  breadth  $  7*5,  ^  8*2  mm. 

Hab. — Vic.  :  Grampian  Mountains  (Best). 

Differs  from  N.  chalybeus,  Dej.,  by  its  larger  size;  more  robust 
form;  prothorax  more  strong^  narrowed  to  base,  more  rounded 
on  sides,  with  lateral  basal  impressions  broader  and  less  elongate; 
elytra  with  humeral  angles  strongly  dentate.  From  N.  gravis, 
Chaud.,  It  differs  by  its  larger  size;  prothorax  with  sides  less  evenly 
rounded  (oblique  towards  base),  basal  angles  less  rounded;  elytra 
strongly  dentate  at  humeral  angles,  more  strongly  sinuate  on 
each  side  of  apex. 

NOTONOMUS    KERSHAWI,  n.Sp. 

Elliptical,  subdepressed.  Prothorax  truncate-cordate,  sub- 
sinuate  on  sides  posteriorly;  basal  angles  subrectangular:  elytra 
obsoletely  striate  on  disc;  interstices  not  raised;  humeral  angles 


IJY    THOMAS    G.    SLOANK.  321 

strongly  toothed.  Nitid;  head,  prothorax  and  under  parts  black; 
elytra  virescent,  metallic. 

Head  hirge,  convex  (3-2  mm.  across  eyes).  Prothorax  subcordate 
(3*5  X  -l:*l  mm.),  narrower  at  base  (3-2  mm.)  than  apex  (3-5  mm.); 
sides  lightly  rounded  anteriorly,  obliquely  narrowed  to  base,  sub- 
sinuate  just  before  base ;  anterior  margin  lightl}^  emarginate  ; 
anterior  angles  obtuse,  bordered;  base  truncate;  basal  angles  sub- 
rectangular,  obtuse  at  summit;  border  narrow;  posterior  marginal 
puncture  a  little  within  basal  angle  ;  lateral  basal  impressions 
narrow  at  bottom,  elongate.  Elytra  oval  (9  2  x  5  5  mm.),  widest 
behind  middle,  a  little  narrowed  to  base;  basal  border  lightly  arcuate 
on  posterior  margin,  obtusely  but  decidedly  raised  above  lateral 
border  at  humeral  angles;  strise  on  disc  very  finely  impressed,  on 
lateral  declivit}^  obsolete,  eighth  strongly  impressed;  third  inter- 
stice 2-punctate  on  apical  half.     Length  15,  breadth  5*5  mm. 

Hab. — Vic.  :  Near  the  mouth  of  the  Gellibrand  River  (Sloane). 

Allied  to  K.  molefhts,  Chaud.,  but  differing  by  its  smaller  size, 
less  robust  and  convex  form;  prothorax  proportion atel}^  longer, 
less  rounded  on  the  sides,  with  basal  angles  more  rectangular,  etc. 

NoTONOMUS  CHALYBEUS,  Dejean. 

Feronia  chalyhea,  Dej.,  Spec.  iii.  p.  234;  F.  j^hilippi,  Newm., 
EntomoL,  1842,  p.  401;  Fer,  (JVoionojuus)  chal7/bea.  Chaud.,  Bull. 
Mosc.  1865,  ii.  p.  90  ;  Fe7\  [Percus)  bipuiicfafa,  Casteln.,  Trans. 
Roy.  Soc.  Yict.  1868,  p.  221. 

Elongate-oval,  robust.  Black,  elytra  chalybeous.  Head  rather 
large  (3*8  mm.  across  eyes) ;  frontal  impressions  well  marked, 
divergent  posteriorly;  eyes  prominent;  postocular  part  of  orbits 
two-thirds  the  size  of  eyes,  rising  sharply  from  head.  Prothorax 
broader  than  long  (4-5  x  5-5  mm.),  widest  before  middle,  lightly 
narrowed  to  base;  sides  lightly  rounded,  narrowed  anteriorly  in 
a  curve  and  posteriorly  subobliquely;  base  wider  (4-1  mm.)  than 
apex  (3-8  mm.);  truncate,  curving  a  little  forward  on  each  side  near 
angles;  basal  angles  obtuse,  lightly  marked ,  posterior  marginal 
puncture  interrupting  border  at  basal  angles;  lateral  basal  im- 
21 


322  EEVISION  OF  THE  GENUS  yOTOXOMUS, 

pressions  wide,  parallel ;  a  marginal  channel  extending  from 
posterior  extremity  of  basal  impressions  and  curving  round  the 
basal  angles  at  a  little  distance  from  the  border.  Elytra  truncate- 
oval  (lO'O  X  6'6  mm.),  widest  behind  middle,  a  little  narrowed  to 
base,  feebly  striate;  striae  obsolete  towards  sides  (excepting  eighth 
and  ninth;;  third  interstice  2-punctate  on  posterior  half,  ninth 
seriate-punctate;  basal  border  lightly  arcuate  on  posterior  margin, 
not  dentate  at  humeral  angles;  lateral  border  reflexed,  lightly 
carinate  near  base.  Intercoxal  declivity  of  presternum  rounded; 
of  mesosternum  concave.     Length  17-21,  breadth  5-7-7 "5  mm. 

Hab. — Yic.  :  Dandenong  (French),  Ferntree  Gull3^and  Lilydale 
(  Sloan  e). 

Chaudoir  is  the  authorit}^  for  placing  Feronia  hipunctata^ 
Casteln.,  under  N.  ehalybeus,  Dej.,  as  a  synonym,  but  I  have  never 
seen  any  specimen  so  small  as  13  mm.,  which  Chaudoir  gives  as 
the  size  of  Dejean's  type  specimen.  Judging  from  Newman's 
description,  I  consider  Feronia  biptmctata,  Casteln.,  to  be  con- 
specific  with  Feronia  philipjn,  Newm. 

Var.  (?)  otwaijensis.  Specimens  from  the  Otway  Forest  have 
the  elytra  a  beautiful  golden  green,  and  the  stride  quite  obsolete 
on  the  disc.     It  seems  a  variety  that  requires  a  name. 

Hab. — Depths  of  the  Otway  Forest,  between  Cape  Otway  and 
the  mouth  of  the  Gellibrand  River  (Sloane). 

NoTONOMUs  GRAVIS,  Chaudoir. 

Bull.  Mosc.  1865,  ii.  p.  90;  Feronia  (^Percus)  montana,  Casteln., 
Trans.  Roy.  Soc.  Vict.  1868,  viii.  p.  221. 

Elongate-oval,  robust,  ver}^  convex.  Black,  nitid.  Head 
moderate;  frontal  impressions  well  marked,  divergent  posteriorly; 
eyes  prominent.  Prothorax  short,  transverse  (-4  x  4'6  mm.), 
widest  about  middle;  sides  evenly  rounded;  base  and  apex  of 
equal  width  (3*2  mm.):  basal  angles  roundly  obtuse;  lateral  border 
thick,  lateral  channel  wide  and  explanate  at  basal  angles;  lateral 
basal  impression  wide.  Elytra  oval  (9*5  x  5-5  mm.),  very  faintly 
striate;  the  stride  usually  obsolete  towards  the  sides  (excepting 


BY    THOMAS    G.    SLOAN E.  323 

eighth);  third  interstice  2-piinctate  on  apical  half,  ninth  seriate- 
punctate;  basal  border  truncate  on  posterior  margin,  and  joining 
lateral  border  at  humeral  angles  without  the  slightest  interrup- 
tion. Intercoxal  declivity  of  prosternum  rounded  ;  of  meso- 
sternum  concave.     Length  16-19,  breadth  5 -5-6  6  mm. 

Hab.—Yic.  :  Melbourne  (Sloane),  Geelong  (J.  F.  Mulder), 
Ballarat  (Froggatt). 

Differs  from  N.  chalyhaus^  Dej.,  by  head  smaller;  prothorax 
much  more  convex  and  differently  shaped  — the  widest  part  at  the 
middle,  and  the  sides  equally  rounded  to  apex  and  base —  the 
apex  narrow,  with  the  angles  not  so  wide  and  nearer  the  head, 
the  basal  angles  far  less  marked,  lateral  basal  impressions  shorter 
and  deeper;  elytra  more  convex,  with  the  border  of  both  base 
and  sides  less  raised,  the  lateral  border  being  much  narrower  and 
not  carinate  near  the  base,  etc.  All  my  specimens  are  black, 
without  any  metallic  tinge. 

NoTONOMUS  aiiLES,  Castehi. 

Trans.  Roy.  Soc.  Viot.  1868,  viii.  p.  208;  Chaud.,  Ann.  Mus. 
Civ.  Geneva,  1874,  vi.  p.  582. 

Elongate,  rather  convex.  Head  oval:  prothorax  hardly  broader 
than  long;  basal  angles  subrectangular;  posterior  marginal  punc- 
ture at  basal  angle  on  inner  side  of  border  :  elytra  oval,  deeply 
striate;  interstices  strongly  convex,  eighth  and  ninth  narrow; 
humeral  angles  dentate;  lateral  border  very  narrow,  but  carinate 
near  base:  prosternum  with  basal  declivity  flat  and  bordered; 
metasternal  episterna  elongate.  Black ;  elytra  (excepting  first 
interstice  and  margin)  dark  purple. 

Head  large  (3  mm.  across  eyes),  elongate,  convex;  eyes  small, 
depressed,  deeply  imbedded  in  orbits;  genjis  tumid  behind  eyes 
and  buccal  fissure;  each  lateral  setigerous  puncture  of  clypeus 
connected  by  a  feeble  sulcus  ;  with  a  fine  puncture  at  each 
lateral  angle  of  clypeal  suture.  Prothorax  narrow,  very  little 
onger  than  broad  (3-7  x  3*9  mm.),  widest  (not  ampliate)  about 


324  EEVISION  OF  THE  GENUS  SOTONOMCS, 

middle,  very  lightly  narrowed  anteriorly,  lightly  and  obliquely 
(with  short  light  prebasal  sinuosity)  narrowed  posteriorly;  apex 
emarginate,  anterior  angles  rather  prominent,  very  close  to  head; 
base  (2-9  mm.)  narrower  than  apex  (3-1  mm.),  hardly  emarginate 
in  middle,  curving  forward  on  each  side;  border  narrow,  thick, 
almost  obsolete  on  sides  of  base;  median  line  not  deep,  a  subfovei- 
form  depression  on  its  course  between  lateral  basal  impressions — 
these  oval,  deep,  wide  at  bottom;  lateral  basal  spaces  convex. 
Elytra  oval  (8-5  x  4*8  mm.),  widest  about  middle,  narrowed  to  base, 
lightly  convex;  lateral  declivities  not  abrupt;  lateral  sinuosities 
of  apex  strongly  developed;  basal  border  projecting  strongly  at 
humeral  angles,posterior  margin  raised  and  truncate;  lateral  border 
narrow,  thick,  not  reflexed  on  posterior  part  of  sides;  eighth  stria 
wide  and  deep;  three  inner  interstices  continuing  to  apex  in  full 
width,  third  with  two  or  three  punctures  on  apical  half,  tenth 
feebly  and  shortly  developed  near  apex.  Posterior  femora  com- 
pressed, dilatate  (slightly  so  in  9,  strongl}'  so  in  (J)  on  lower  side 
above  apex  of  trochanters ;  posterior  tibiae  slightl}^  arcuate  ; 
posterior  tarsi  thick,  not  long;  joints  much  shorter  than  usual  in 
Notonomus.  Length  13-16,  breadth  4-3-5-2  mm. 
^«5._N.S.W. :  Kiama  (Sloane). 

An  isolated  species  distinguished  at  once  from  all  others  of  tlie 
genus,  except  N.  lesueuri,  Casteln,,  by  its  elongate  metasternal 
episterna.  It  is  allied  to  N.  lesueuri  by  all  characters  of  a 
generic  nature,  but  is  readily  separated  by  its  less  cylindrical 
shape,  elytra  strongly  dentate  at  humeral  angles,  ttc.  Six  speci- 
mens are  before  me,  two  of  which  have  the  third  interstice  of 
each  elytron  3-punctate ;  this  proves  the  presence  of  a  third 
puncture  to  be  more  than  a  mere  accidental  occurrence.  The 
eyes  are  more  prominent  in  some  specimens  than  in  others.  I 
report  this  species  as  N.  miles,  Castelnau,  on  the  authority  of  Mr. 
Masters,  who  has  informed  me  that  his  specimens  so  named  were 
received  by  him  from  Count  de  Castelnau.  Chaiidoir  evidently 
overlooked  the  elongate  mestasternal  episterna  when  examining 
N.  miles. 

[Printed  off  October  14th,  1902.1 


f.L.S.N.S.W.,    1902. 


P.L.S.N.S.W.,    1902. 


P.L.S.N.S.W..    1902. 


Fig.  1. 

EUCALYPTUS    WOOLLSIAN/! 


bUCALiPTUS    CONICA. 


P.LSN.SW.    1902 


PI.  xi 


EUCALYPTUS      MELANOPHLOIA,  F.,.Af. 


P.L.S.N.S.W..    1902. 


BACTERIUM    EUCALYPTI,    )l.  Sp. 


BY   THOMAS    G.    SLOANE.  325 

NoTONOMUS  LESUEURi,  Casteliiau. 

Feronia  {Pterostichus)  lesueurii,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  viii.  p.  210;  Fer.  lesueuri,  Chaud.,  Ann.  Mus.  Civ.  Genova, 
1874,  vi.  p.  596. 

Elongate,  narrow,  subcylindrical.  Black,  elytra  with  a  bronzy 
or  purplish  tinge.  Head  oval,  convex  (3-5  across  eyes) ;  eyes 
with  orbits  reniform,  depressed.  Prothorax  as  long  as  broad 
(4-5  X  4-55  mm.),  convex,  widest  (not  ampliate)  at  middle,  wider 
across  base  (3-5  mm.)  than  apex  (3-3  mm.);  sides  lightly  arcuate, 
very  lightly  narrowed  anteriorly,  a  little  more  strongly  so  in  an 
even  curve  posteriorly ;  apex  truncate ;  basal  angles  widely 
rounded;  border  narrow,  not  extending  on  to  the  sides  of  base, 
posterior  marginal  puncture  on  border  at  basal  angle;  median 
line  fine,  lateral  basal  impressions  oval.  Elytra  oval  (10  x  5*3 mm.); 
very  lightly  rounded  on  sides,  convex,  deeply  striate;  interstices 
lightly  convex,  more  strongly  so  towards  apex,  three  inner  ones 
reaching  apex  in  full  width,  third  2-punctate,  eighth  narrow, 
convex,  narrower  than  ninth,  tenth  well  developed  before  apical 
sinuosities,  narrow,  not  long;  humeral  angles  not  marked;  basal 
border  not  raised  at  humeral  angles,  joining  lateral  border  with- 
out any  interruption;  lateral  border  very  narrow,  thickened  but 
not  rellexed  posteriorly.  Intercoxal  declivity  of  prosternum  flat, 
bordered;  of  mesosternum  flat:  metasternal  episterna  eloyigate. 
Length  16-5-19,  breadth  4-75-5-8  mm. 

Hah. — N.S.W.  :   Burrawang  (Sloane;  common  in  November). 

A  distinct  species,  isolated  by  its  elongate  metasternal  episterna, 
obtuse  basal  angles  of  prothorax,  humeral  angles  of  elytra  not 
dentate,  &c. 

■■'•  •^■ 


22 


326 


CONTRIBUTIONS  TO  A  KNOWLEDGE  OF  AUS- 
TRALIAN ENTOZOA. 

No.  ii. — On  a  new  species  of  Distomum  from  the  Sawfish 
Shark,  Pristiojyhorus  cirratus,  Lath. 

By  S.  J.  Johnston,  B.A.,  B.Sc,  Economic  Zoologist, 
Technological  Museum,  Sydney. 

(Plate  xiii.) 

I  am  indebted  to  Professor  Haswell  for  three  specimens  of 
this  Trematode,  which  were  found  in  the  body  cavity  of  the 
Sawfish  Shark,  PristiojyJiorus  cirratus,  Lath.,  at  Sydney.  I  have 
examined  several  specimens  of  the  same  host  since  receiving  those 
from  Professor  Haswell,  but  have  found  no  Distomes,  though  in 
the  body  cavity  of  one  specimen  a  number  of  eggs  were  found 
which  very  probably  belong  to  the  same  species,  for  the  eggs  with 
their  very  thick  shells  are  characteristic  of  the  species :  the 
worms  may  have  escaped  through  the  abdominal  pores  when  the 
fish  was  caught.  This  Avorm  has  a  very  extensible  neck;  fasten- 
ing itself  by  the  ventral  sucker  it  stretches  its  neck  out  for  more 
than  an  inch,  longer  than  the  body  itself. 

DiSTOMUM    PRISTIOPHORI,    sp.n. 

Body  elongated,  pointed  at  the  anterior  end,  and  rounded  at 
the  posterior,  flattened  dorsoventrally,  with  rounded  sides;  head- 
lobe  triangular.     Average  length  25  mm.,  breadth  6  mm. 

Oral  sucker  deep,  cup-shaped  and  small ;  subterminal,  the 
opening  being  on  the  ventral  aspect;  diameter  1*5  mm.  Ventral 
sucker    orbicular,    sessile,    with    deep    cavity,    larger    than    oral 


BY    S.    J.    JOHNSTON.  327 

sucker;  diameter  2*5  mm.  Both  suckers  very  muscular,  devoid 
of  hooks  or  lobes  of  any  kind. 

Common  genital  aperture  situated  near  the  middle  line  about 
half-way  between  the  oral  and  ventral  suckers.  Excretory  aper- 
ture situated  at  the  extreme  posterior  end.  Laurer's  canal 
opening  on  the  dorsal  surface  about  the  middle  of  the  body. 

Cuticle  with  a  transversely  striated  appearance  owing  to  fine 
closely  arranged  grooves  running  round  the  worm. 

Alimentary  canal  simple;  mouth  situated  at  the  base  of  the 
oral  sucker,  leading  into  a  well  developed,  muscular  pharynx; 
behind  the  pharynx  the  canal  immediately  divides  into  the  two 
limbs  of  the  intestine,  the  oesophagus  being  practically  non- 
existent. Limbs  of  the  intestine  unbranched,  but  not  quite 
simple,  being  thrown  into  bays  and  folds  throughout  their  length; 
they  terminate  in  blind  sac-like  ends  at  the  posterior  end  of  the 
body.  Walls  of  the  intestine  thick,  lined  by  long  columnar  cells, 
nucleated  at  the  base,  of  a  glandular  character. 

Excretory  system  very  well  developed,  consisting  of  two  main 
canals  which  run  forwards  and  unite  in  front  of  the  oral  sucker. 
In  the  anterior  half  of  the  body  these  excretory  vessels  run 
laterally;  but  some  distance  behind  the  testes  they  bend  inwards 
and  meet  about  the  middle  line  to  form  a  single  median  vessel, 
which  opens  into  a  large  excretor}^  vesicle  in  the  posterior  end  of 
the  body.  A  number  of  constrictions  occur  along  the  excretory 
canals.  The  excretory  vesicle  opens  on  the  exterior  through  a 
very  short  canal,  ending  in  the  excretory  pore.  The  excretory 
canals  were  partly  filled  up  with  crystals  belonging  to  the  cubical 
system,  and  which  had  become  stained  by  eosin;  some  envelope 
crystals  of  calcium  oxalate  were  present. 

Testes  two,  situated  just  posterior  to  the  middle  of  the  bod}-, 
ovoid  in  shape;  lying  almost  on  the  same  level.  The  undeveloped 
sperms  give  their  interior  that  follicular  appearance  usual  in 
Trematodes,  but  there  is  also  a  large  number  of  fully  developed 
sperms  present.  Each  vas  deferens,  passing  dorsal  to  the  uterus 
and  ventral  sucker,  joins  its  fellow  at  the  origin  of  the  vesicula 
seminalis,  which  is  large  and  pear-shaped,  and  filled  with  fully 


328  AUSTRALIAN    ENTOZOA,  NO.   II., 

developed  spermatozoa  with  small  round  heads  and  long  tails. 
The  vesicula  seminalis  is  continued  as  the  ejaculatoryduct  which 
traverses  the  penis  to  open  on  the  exterior.  The  vesicula  semi- 
nalis and  the  proximal  part  of  the  ejaculatoryduct  are  surrounded 
by  a  strong  muscular  sac  ;  the  somewhat  considerable  space 
between  the  walls  of  the  vesicula  seminalis  and  this  sac  are 
traversed  by  muscular  and  connective  tissue  fibres  which  join  the 
walls  of  each.  Between  the  fibres  lie  a  number  of  glandular 
cells  representing  a  prostate  gland. 

The  male  and  female  ducts  open  close  together  into  the  common 
genital  chamber,  which  is  of  some  size,  and  which  opens  on  the 
exterior  on  the  ventral  surface  on  the  middle  line  and  half-way 
between  the  two  suckers. 

The  ovary  is  an  ovoid  body  of  solid  appearance  lying  just 
anterior  to  the  testes.  In  the  walls  of  the  distal  part  of  the 
oviduct  the  shell  glands  are  situated,  and  this  part  of  the  oviduct 
constitutes  the  ootype.  Into  the  end  nearest  the  ovary  Laurer's 
canal  opens;  about  its  middle  the  main  duct  of  the-  vitelline 
glands. 

The  uterus  is  a  very  long  and  narrow  tube  which  is  thrown 
into  a  considerable  number  of  folds  between  the  ovary  and 
ventral  sucker;  skirting  the  side  of  the  ventral  sucker  it  runs  for- 
wards to  open  alongside  the  male  opening  into  the  common  genital 
chamber.  There  is  no  vagina.  The  uterus  is  full  of  eggs,  which 
are  characterised  by  a  very  thick  chitinous  shell.  The  average 
longitudinal  diameter  of  the  eggs  is  0'077  mm.,  transverse 
0*062  mm.;  the  thickness  of  the  shell  is  0*008  mm.  The  eggs 
consist  of  the  ovum  and  three  or  four  large  vitelline  cells. 

The  vitelline  glands  are  a  number  of  small,  rounded,  grape-like 
follicles  which  communicate  with  one  another  by  small  ducts.  A 
main  lateral  duct  on  each  side,  about  the  level  of  the  ovary,  opens 
into  a  median  duct;  these  median  ducts  meet  together  and  open 
into  the  ootype  by  a  single  opening.  The  follicles  are  situated 
laterally,  in  the  middle  third  of  the  body,  and  are  grouped  round 
the  intestine  on  each  side  of  the  body.  The  vitelline  mass 
consists  of  large  nucleated  cells  of  an  albuminous  character. 


BY    S.    J.    JOHNSTON.  329 

Laurer's  canal,  a  canal  with  muscular  walls,  opens  on  the 
exterior  about  the  middle  of  the  dorsal  surface,  opposite  the 
middle  of  the  ovary.  It  skirts  round  the  right  side  of  the  ovary 
and  opens  into  the  proximal  end  of  the  ootype.  In  its  interior 
a  number  of  sperms  can  be  seen.  The  canal  is  quite  large  enough 
to  be  used  in  copulation. 

The  most  characteristic  features  of  the  worm  are  its  size,  the 
character  and  position  of  the  suckers,  the  folded  but  unbranched 
intestine,  the  ovoid  shape  of  the  ovary  and  testes  and  their 
situation,  the  great  length  of  the  uterus,  the  grape-like  vitelline 
glands,  and  the  well  developed  excretory  system. 

The  simple  nature  of  the  intestine,  the  absence  of  hooks  or 
lobes  from  the  suckers,  the  almost  total  obliteration  of  the 
oesophagus,  and  the  absence  of  a  retractile  telescopic  tail  part 
indicate  the  position  of  this  species  to  be  in  Dujardin's  subgenus 
Brachylaimus. 

In  external  characters,  as  general  shape,  size,  character  and 
position  of  the  suckers  and  transversely  striated  appearance,  it 
closely  resembles  D.  veliporlum,  Creplin,  found  in  the  American 
barn-door  skate,  Eaja  kevis,  in  Prionodon  milberti,  and  in  Hexar- 
chus  g7'lseus.  In  its  internal  anatomy  it  shows  a  fairly  close 
resemblance  to  D.  tereticolle,  Rud.  The  excretory  system  is  very 
marked  and  very  similar  in  each.  The  limbs  of  the  intestine  in 
D.  tereticolle  are  without  the  folds  shown  in  D.  irristio'pliori. 
They  both  have  the  uterus  long,  slender  and  much  folded,  and  a 
similar  structure,  arrangement  and  amount  of  development  in  the 
vitelline  glands;  in  this  new  species  the  ovary  and  testes  are 
situated  much  nearer  together. 

The  figures  for  the  plate  were  drawn  by  my  wife. 


EXPLANATION  OF  PLATE  XIIL 

Distomum  pristiophori. 

Fig.  1. — View  of  the  whole  animal  (  x  6). 

Fig.  2.  —Transverse  section  through  the  middle  of  the  ventral  sucker  (  x  15) 
Fig.  .3. — Transverse  section  through  posterior  end  showing  the  large  excretory 
vesicle  (  x  20). 


330  AUSTRALIAN    ENTOZOA,     NO.  II. 

Fig.  4.— Transverse  section  through  the  vesicula  seminalis  and  its  surround- 
ing sac  ( X  50). 

Fig.  5. — Transverse   section   through   the   anterior   end   showing  common 
genital  opening  and  chamber  (  x  15). 

Fig.  6. — Portion  of  the  worm  showing  termination  of  genital  ducts  (  x  20). 

Fig.  7.— Transverse   section    through    the   ovary   showing   Laurer's  canal, 
ootype  and  vitelline  duct  (  x  20). 
All,  except  fig.  1,  drawn  with  the  camera  lucida. 

Reference  letters. 

c.g.c,  common  genital  chamber. — c,  cirrus. — ej.d.,  ejaculatory  duct. — 
ex.c,  excretory  canal. — ex. v.,  excretory  vesicle. — g.p.,  genital  aperture. — 
int.,  intestine. — L.C,  Laurer's  Canal. — m.s.,  muscular  sac. — oo.,  ootype. — 
0.$.,  oral  sucker. — ov.,  ovary. — ovid.,  oviduct. — 2)Ji.,  pharnyx. — pr.g.,  pro- 
state gland. — s.g.,  shell  glands. — ^i  cO  t,^  testes. — ut.,  uterus. — v.d.,  vas 
deferens. — v.g.,  vitelline  glands. — vit.d.,  vitelline  duct. — r.s.,  ventral  sucker. 
v.i^em.,  vesicula  seminalis. 


I 


331 


NOTES  ON  AUSTRALIAN  RHOPALOCERA.LYC.ENIDjE, 

Part  i. 
By  G.  a.  Waterhouse,  B.Sc,  B.E.,  F.E.S. 

(Plate  xiv.) 

It  is  my  intention  in  this  series  of  papers  to  publish  from  time 
to  time  notes  on  the  generic  position  of  Australian  species 
referred  to  the  Lyccenid(e,  so  that  our  knowledge  of  this  family 
may  be  brought  more  into  line  with  that  of  other  parts  of  the 
world,  particularly  of  the  Indo-Malayan  region.  At  the  present 
time  our  genera  are  large  and  unwieldy,  containing  many  very 
dissimilar  forms,  for  which  there  are  suitable  genera  at  hand  in 
which  to  place  them.  The  system  of  classification  used  will  be 
that  given  by  the  late  Mr.  L.  de  Niceville  in  his  '  Butterflies  of 
India,  Burmah  and  Ceylon.  Part  iii.  Lyccenidce,''  to  whom  I  am 
also  indebted  for  many  valuable  generic  notes  on  Australian 
specimens  which  have  been  submitted  to  him.  In  addition  to 
my  own  collection  of  Australian  and  foreign  Lyccenidce  I  have 
had  access  to  the  very  fine  collection  in  the  Macleay  Museum. 
In  addition  I  intend  giving  notes  on  life-histories,  distribution 
and  synonyms,  as  well  as  descriptions  of  any  new  species  it  may 
be  my  good  fortune  to  discover. 

Megisba  nigra,  Miskin. 

Lycctiiia  nigra,  (9)  Misk.,  Proc.  Linn.  Soc.  N.S.  Wales,  1890, 
V.  (2),  36. 

Miskin  described  the  female;  I  now  give  a  description  of  the 
male. 


332    •  AUSTRALIAN  RUOPALOCfJRA  :  LYC.ENID.E, 

^.  Upper  side  black,  with  white  central  areas.  Foreiving 
with  costa  almost  straight,  apex  acute  (much  more  so  than  in  9), 
outer  margin  nearly  straight  (convex  in  ^) ;  three  subcostal 
nervules,  first  subcostal  entirely  free  from  costal  nervure,  a  black 
costal  margin,  apex  and  outer  margin,  the  medial  conical  white 
patch  extending  obliquely  from  near  middle  to  inner  margin; 
base  brown.  Hindwing  coloured  as  in  fore  wing,  except  that  the 
central  white  patch  extends  from  near  middle  to  costa,  forming  a 
continuation  of  the  white  patch  of  the  fore  wing. 

Underside  silvery-white,  with  light  black  lunular  external 
margins.  Forewing,  pale  black  costa  and  outer  margin  a  pale 
brown  streak  at  end  of  cell,  and  four  black  costal  spots.  Hind- 
ioing  with  black  outer  margin,  three  subbasal  spots,  an  apical  and 
an  anal  black  spot. 

The  distinction  between  the  sexes  is  that  the  apex  is  more 
acute,  the  outer  margin  of  the  forewing  and  the  anal  angle  more 
pronounced  in  the  male  than  in  the  female;  otherwise  the  sexes 
are  similar.  This  species  should  correctly  be  referred  to  the  genus 
Megisba,  Moore,  of  which  the  type  is  J/,  malaya,  Horsf.  The 
insects  of  this  genus,  which  probably  contains  a  single  very 
variable  species,  may  be  anything  from  wholly  black  to  white 
with  black  margins  on  the  upperside;  while  on  the  underside  the 
markings  are  much  more  constant,  but  are  usually  darker  and 
larger  according  as  the  white  of  the  upperside  decreases  in  size. 
The  species  is  represented  by  individuals  with  or  without  a  tail 
to  the  hindwing.  The  expanse  in  both  sexes  is  about  20  mm. 
Though  convinced  that  the  Australian  species  is  only  one  of  the 
many  forms  of  the  type  of  the  genus,  I  prefer,  however,  for  the 
present  not  to  sink  Miskin's  name ;  of  all  the  forms  which 
de  Niceville  sinks  under  M.  malaya  ours  is  closest  to  M.  albidisca, 
Moore.* 

Though  the  species  is  one  of  our  blackest  Lycaenids,  yet  it  is  the 
tvhitest  form  of  the  genus,  and  the  name  nigra  is  rather  out  of 
place.     J/,   malaya  is   recorded   all   through   the   Indo-Malayan 

*  Journ.  Asiat.  Soc.  Bengal,  Vol.  liii.  Pt.  ii,,  p.  21,  1884. 


BY    G.    A.    WATERHOUSE.  333 

region,  and   the  Australian   form   has   been   taken   at  Card  well 
(Miskin)  and  Cairns  {J'^.  E.  Turner). 

HOLOCHILA    CYPROTUS,  OUiff. 

Chrysophanus  cyprotus^  OIL,  Proc.  Linn.  8oc.  N.8.W.  x.  716, 
1885;  Holochila  cyprotus,  Misk.,  Syn.  Cat.  Rhop.  Aust.  p.  64, 
1891;  H.  purpurea^  Grose  Smith  k  Kirby,  Rhop.  Exot.  pi.  x., 
figs.  11  ik  12  {$),  Pt.  39,  p.  7,  1896. 

A  pair  of  this  species  determined  by  the  late  Mr.  Oil  iff  is  in 
ni}^  collection;  and  on  looking  through  the  '  Rhopalocera  Exotica ' 
I  found  an  excellent  figure  of  the  male  under  the  name  of  H. 
purpurea  from  Sydney  and  Moreton  Bay.  The  localities  agree, 
and  also  the  description  and  figure;  and  there  is  no  doubt  in  my 
mind  that  H.  purpurea  is  the  same  insect.  The  confusion  was 
caused  no  doubt  by  H.  cyprotus  being  originall}'  placed  in  the 
genus  Chrysophanus,  to  which  it  certainly  does  not  belong.  This 
species  is  very  local;  and  I  have  found  only  one  brood,  and  that 
appeared  in  Sydney  very  early  in  the  season,  and  was  on  the  wing- 
only  a  short  time  The  females  are  much  rarer  and  of  slower 
flight  than  the  males.  In  Brisbane  Mr.  R.  Illidge  has  taken 
larvae  feeeding  on  Jacksonia  sp.,  but  as  I  know  of  no  Jacksonia 
nearer  to  Sydney  than  Blacktown,  there  must  be  some  other 
food-plant. 

Loc. — Sydney,  Como,  and  Katoomba,  N".S.W.;  Brisbane  and 
Rockhampton,  Q. 

Holochila  helenita,  Semper. 

Journ.  Mus.  Godeff.  xiv.,  p.  162,  1879;  Holochila  androdus, 
Misk.,  Proc.  Linn.  Soc.  N.S.W.  1890,  v.  (2),  p.  41;  If.  suhargentea, 
Grose  Smith  &l  Kirby,  Rhop.  Exot.  Hoi  i.,  Pt.  38,  fios.  9  c<c  10 
{S\  1896. 

This  species  for  many  years  has  passed  under  Miskin's  name  of 
androdus.  By  all  the  authors  mentioned  above,  this  species  is 
compared  with  the  allied  form,  H.  absimilis,  Felder.  Semper  says 
the  male  is  greenish  and'  shining,  w^ith  a  dark  arrows-shaped 
shadow  standing  out  along  the  veins  in  the  centre  of  the  fore  wings: 


334  AUSTRALIAN  RHOPALOCERA  :  LYC.fJNID.E, 

in  the  female  the  white  areas  are  larger  than  in  absimilis,  and  in  the 
hindwing  the  white  area  extends  to  the  costa.  Thus  it  will  be 
seen  that  when  describing  II.  androdus,  Miskin  was  only  renaming 
an  old  species.  Miskin  does  not  mention  the  arrow-shaped 
shadow  on  the  forewing,  but  one  of  his  localities,  Cape  York,  is 
the  same  as  Semper's.  Regarding  //.  suhargentea,  Mr.  R.  E. 
Turner,  who  has  seen  both  the  type  specimen  of  androdus  and 
the  figure  of  snhargentea,  says  that  they  are  identical.  The  figure 
is  a  very  good  one,  and  shows  the  arrow-shaped  shadow  on  the 
forewing,  a  character,  however,  not  mentioned  in  the  description. 
The  female  is  not  figured,  and  the  locality  given  for  the  male  is 
Cape  York. 

Mr.  Turner,  who  has  caught  many  specimens  of  this  species 
both  at  Cairns  and  Cape  York,  informs  me  that  the  markings  on 
the  underside  vary,  but  that  those  of  the  upperside  are  constant. 
A  careful  examination  of  the  descriptions  convinces  one  that  both 
Miskin,  and  Grose  Smith  and  Kirby  described  a  much  older  form. 
It  should  also  be  noted  that  the  locality,  Cape  York,  is  given  by 
all  three  writers. 

Loc. — Cairns,  Cooktown,  and  Cape  York,  Q. 

Ogyris,  West  wood. 

Gen.  Diurn.  Lepid.  ii.,  472  (1850-1852);  Miskin,  Proc.  Linn. 
8oc.  X.S.W.  1890,  V.  (2),  23. 

The  three  following  species  of  this  beautiful  Australian  genus 
have,  b}^  all  writers,  been  greatly  confused.  The  insects  are  all 
of  a  brilliant  silvery-blue  on  the  upperside;  and  on  account  of 
their  at  one  time  great  rarity  they  are  usually  poorly  represented 
in  collections.  Also  authors  knowing  only  one  species,  have  tried 
to  make  it  fit  described  species  with  little  success.  The  difficulty 
is  cleared  up,  however,  when  we  recognise  that  there  are  not  one 
or  two  species,  as  authors  have  variousl}''  supposed,  but  three;  and 
that  the  commonest  and  best  known,  is  the  one  that  as  yet  has 
not  been  named.     1  give  the  three  descriptions  below. 


BY    G.    A.    WATERHOUSE,  335 

Ogyris  orcetes,  Hewitson. 
Cat.  Lye.  Brit.  Mus.  p.  3,  t.  1,  figs.  12-13  (9),  1862. 
(Plate  xiv.,  figs.  1-2.) 

J".  Upperside  silvery-blue  with  a  faint  black  edging;  the 
blue  has  a  slight  purplish  tint  which  distinguishes  it  from  the 
males  of  the  two  following  species.  The  apex  also  is  more  acute 
and  the  outer  margin  straighter. 

Underside  grey.  Forewing  with  cell  crossed  by  five  white 
transverse  bars;  the  first  small  basal,  between  the  second  and 
third  darker  grey,  between  the  fourth  and  fifth  almost  black  and 
a  dark  bar  external  to  cell.  The  dark  discal  band  straight,  from 
costa  to  first  median  nervule;  a  dark  submarginal  band.  Hind- 
wing  with  several  irregular  brown  spots  surrounded  with  darker 
brown  and  dark  brown  submarginal  sufiusions. 

The  male  may  be  distinguished  from  the  males  of  the  two 
following  species  by  the  totally  different  shade  of  blue,  the  very 
small  brown  margins,  the  more  acute  apex  to  the  forewing;  and 
on  the  underside  the  discal  band  is  straight,  nor  can  I  find  any 
trace  of  blue  in  the  cell  as  is  found  in  the  males  of  both  the  other 
species.  The  specimen  referred  to  this  species  by  Miskin"^  is  in 
reality  the  male  of  0.  heivitsoni. 

2'  Upperside.  Foreiving  silvery-blue,  costal  margin  brown, 
widening  beyond  the  cell,  apex  and  outer  margin  broadly  brown, 
a  dark  bar  at  end  of  cell  and  costa  near  apex  marked  with  two 
white  splashes;  the  apex  is  acute  and  outer  margin  nearly  straight 
as  in  male.  Hindiving  silvery-blue,  with  brown  outer  margin, 
narrowest  at  the  terminations  of  the  veins,  fringes  white. 

U  n  d  e  r  s  i  d  e  as  in  male,  but  the  white  bars  of  the  cell  are 
often  bordered  with  silvery-blue.  This  may  be  distinguished  from 
the  females  of  the  two  following  species  by  the  absence  of  scarlet 
in  the  cell  and  the  straight  discal  band  of  the  underside.  The 
upperside  is  almost    identical   with    that  of   the   female  of    0. 

''  Proc.  Linn.  Soc.  N.S.W.  1890,  v.  (2),  26. 


336  AUSTRALIAN  RHOPALOCERA  :  LYC.EyiD.E, 

hewitsoiii,  but  has  a  more  acute  apex  and  a  straighter  outer 
margin. 

Hewitson  gives  an  excellent  figure  of  this  sex  only.  Though 
Miskin  correctly  determined  a  female  specimen  from  Mackay  for 
Mr.  R.  E.  Turner,  yet  he  says  that  the  Australian  Museum  con- 
tained a  female.  But  the  specimens  of  0.  orrjutes  have  only  been 
there  for  about  two  years;  and  I  have  no  doubt  he  mistook  a 
male  of  0.  amaryllis  for  the  female  of  0.  oroetes,  as  he  was  doubt- 
ful of  his  determination.  Mr.  F.  D.  Dodd,  of  Townsville,  has 
bred  some  fine  examples  of  this  species,  from  which  m}^  descriptions 
are  mainly  taken;  they,  however,  do  not  show  the  blue  in  the 
cell  on  the  underside  as  shown  in  Hewitson's  figure,  but  this  is 
the  only  variation.  The  recognition  marks  of  this  species  are  the 
acute  apex  and  straight  outer  margin  to  the  forewing ;  and 
the  straight  discal  dark  band  on  the  underside  of  the  forewing. 
A  further  recognition  mark  of  the  male  is  the  purplish-blue  tint 
on  the  upperside;  and  of  the  female  the  absence  of  scarlet  in  the 
cell  of  the  underside  of  the  forewing. 

Loc. — Moreton  Bay  (Hewitson),  Mackay  (li.  E.  Turner), 
Townsville  (Dodd). 

This  species  appears  to  be  confined  to  Queensland. 

Ogyris  AMARYLLIS,  Hewitson. 
Cat.  Lye.  Brit.  Mus.  p.  3,  t.  1,  figs.  5-6  (9),  1862. 
(Plate  xiv.,  figs.  3-4.) 

(J.  Upperside  dark  silvery  morpho-blue.  Forewing  with 
a  linear  black  costal  margin  and  a  broad  black  outer  margin,  and 
two  white  costal  splashes  near  apex.  Hindwing  with  a  black 
outer  margin,  narrowest  at  the  terminations  of  the  veins. 

U  n  d  e  r  s  i  d  e.  Forewing  with  outer  marginal  \  gre}^  rest  of 
the  wing  almost  black;  cell  crossed  by  five  white  transverse  bars 
sprinkled  with  blue,  two  basal,  very  small,  two  central  and  one 
near  end  of  cell,  beyond  end  of  cell  a  short  white  transverse  bar; 
the  discal  band  is  black  and  only  well  defined  towards  the  costa, 
but  towards  the  middle  merges  into  the  inner  marginal  half  of 


.    BY    G.    A.    WATEHHOUSE.  337 

the  wing.  Hindwing  brown,  with  the  pattern  of  the  markings  as 
in  the  previous  species,  but  much  darker,  with  a  ver}^  dark  central 
blotch  and  a  dark  blotch  towards  outer  margin. 

This  no  doubt  is  what  Miskin  determined  to  be  0.  orcetes  (9),  as 
it  was  the  only  specimen  at  all  like  that  sex  that  was  in  the 
Australian  Museum  when  he  saw  the  specimens.  He  says  it 
agrees  "pretty  well"  with  Hewitson's  figure  of  0.  orcetes  (9).  It 
is  a  much  darker  blue  than  any  except  its  female,  and  the  black 
outer  margin  is  much  broader  than  in  either  of  the  males  of  the 
other  species,  being  as  broad  as  the  outer  margins  of  the  females 
of  0.  onetes  and  0.  heivitsoni. 

9.  Upperside.  Foretoing  dark  silvery-blue,  costal  margin 
black,  narrow  until  the  end  of  cell,  where  it  broadens  out  to  the 
width  of  the  cell ;  apex  broadly  black,  and  outer  margin  very 
broadly  black;  two  white  splashes  on  costa  near  apex,  the  blue  and 
black  occupying  about  equal  areas.  Hindwing  dark  blue,  with  a 
broad  black  outer  margin  and  a  black  line  at  end  of  cell. 

Underside,  Foreioing  darker  than  in  male,  but  otherwise 
as  in  male,  except  that  the  areas  of  the  cell  between  the  first  and 
second,  and  third  and  fourth  white  bars  are  brilliant  scarlet  and 
the  white  bars  are  more  sprinkled  with  blue  scales.  Hindiving 
as  in  male,  but  much  darker. 

Hewitson's  figure  of  the  female  of  this  species  is  excellent,  and 
there  can  be  no  mistaking  it.  Miskin  is  also  correct  in  his  iden- 
tification. In  the  possession  of  scarlet  in  the  cell  of  the  underside 
it  resembles  the  female  of  0.  hewitsoni;  but  then  the  uppersides 
of  the  two  species  are  totally  different.  The  recognition  marks 
of  this  species  are  the  broad  black  outer  margins  on  the  upperside; 
and  on  the  underside  the  black  discal  band  starts  from  near  the 
costa,  proceeds  a  short  distance,  and  then  is  lost  in  the  black 
lower  half  of  the  forewing. 

This  appears  to  be  a  rare  species;  Hewitson  has  recorded  it 
from  Moreton  Bay,  and  I  have  taken  it  on  the  Richmond  River. 


338  AUSTEALIAN  RHOPALOCERA  :  LYC.EXID.E, 

Ogyris  heavitsoni,  n.sp. 
(Plate  xiv.,  figs.  5-8.) 

(J.  Upper  side  pale  silvery-blue,  Avith  very  narrow  black 
margins,  widest  at  apex  of  forewing,  and  with  two  white  splashes 
on  eosta  near  apex.     Hindwing  with  a  linear  black  outer  margin. 

Underside.  Foreicing  grey,  much  darker  at  the  end  of 
cell,  which  is  crossed  by  five  white  bars;  in  southern  specimens 
these  bars  are  freely  sprinkled  with  blue  scales,  which  are 
generally  absent  in  northern  specimens;  a  white  patch  beyond 
the  cell  and  an  irregular  dark  brown  discal  band  beginning  near 
costa  and  extending  to  the  first  submedian  nervule,  the  edges  of 
the  band  very  irregular.  Hindwing  after  the  general  pattern  of 
0.  oroetes  {$),  but  the  markings  are  more  distinct. 

This  is  a  very  variable  species,  but  I  do  not  find  any  variation 
in  the  direction  of  either  of  the  other  species.  The  male  is  nearest 
to  that  of  0.  orcetes,  but  the  tint  of  blue  is  lighter,  the  apex  not 
so  acute,  and  the  outer  margin  of  forewing  is  more  convex.  It 
is  the  species  that  Miskin*  put  as  the  male  of  0.  orcHes.  Speci- 
mens bred  by  Mr.  Dodd  at  Townsville  do  not  show  blue  scales  on 
the  underside,  and  are  lighter  than  specimens  from  Victoria  or 
South  Australia. 

9-  Upper  side.  Foreioing  pale  silvery-blue,  with  brown 
costal  margin;  apex  and  outer  margin  much  wider  than  in  the 
male;  two  white  costal  splashes  near  apex,  a  dark  bar  marking 
the  end  of  cell.  Hindicing  silvery-blue,  with  dark  brown  outer 
margin;  costa  near  apex  broadly  dark  brown. 

Underside.  Forewing  as  in  male,  but  with  the  areas  of 
cell  between  first  and  second,  and  third  and  fourth  white  bars  a 
brilliant  scarlet.     Hindwing  as  in  male. 

This  was  known  to,  and  described  by  Miskinf  as  the  male  of 
0.  aynaryllis.     It  has  the  same  scarlet  underside  in  the  cell  as  0. 

*  Proc.  Linn.  Soc.  N.S.W.  1890,  v.  (2),  p.  2.5. 
t  Proc.  Linn.  Soc.  N.S.W.  1890,  v.  (2),  p.  26. 


BY    G.    A.    AVATERHOUSE.  339 

amaryllis,  but  the  uppersides  are  different.  It  is  also  a  variable 
species,  northern  forms  being  lighter  and  larger. 

In  .Spry  and  Anderson's  '  Victorian  Butterflies '  this  species  is 
figured  under  the  name  of  0.  amaryllis. 

It  is  by  far  the  commonest  of  the  three  species,  and  I  have 
specimens  from  all  the  Australian  States  but  Tasmania.  It  is  in 
both  sexes  somewhat  like  0.  ormtes  on  the  upperside,  and  like  0. 
amaryllis  on  the  underside.  The  recognition  marks  may  l^e 
taken  as  the  irregular  discal  band  on  the  underside  of  the  fore- 
wing;  and  the  pale  silvery-blue  colour. 

Great  confusion  has  been  caused  regarding  the  three  silvery- 
blue  species  mainly  through  want  of  sufficient  material.  All 
three  species  are  from  Queensland.  0.  orcetes  appears  to  be 
confined  to  that  State.  0.  amaryllis  ranges  into  the  Xorthern 
Rivers  of  New  South  Wales;  while  0.  hewitsoni  is  found  all 
through  Australia.  Miskin's  great  mistake  was  in  failing  to 
recognise  that  it  was  only  the  females  that  had  scarlet  in  the  cell 
on  the  underside  of  the  fore  wing,  and  thus  making  his  0. 
amaryllis  consist  of  the  female  of  0.  hewitsoni  as  the  male,  and 
the  true  0.  amaryllis  as  the  female.  Likewise  he  made  0.  orcetes 
comprise  two  different  males.  It  is  curious  to  note  that  the 
underside  of  the  hindwing  is  almost  the  same  in  the  three  species 
except  in  the  intensity  of  the  brown  colouration. 

Ogyris  .enone,  n.sp. 

(Plate  xiv.,  fig.  9.) 

/J.  Upperside.  Forewing  with  costa  convex,  apex  acute; 
outer  margin  straight,  slightly  concave  in  the  middle ;  inner 
margin  straight :  between  costa  and  subcostal  nervure  a  dark 
margin,  grey  at  base,  and  then  black  to  the  apex,  where  it  is 
widest ;  outer  margin  black,  about  the  same  width  as  costal 
margin;  a  prolongation  of  the  black  costal  margin  downwards 
occurs  at  the  end  of  cell;  a  few  pale  blue  scales  are  sprinkled  on 
the  black  costal  margin  near  the  apex;  the  rest  of  the  wing  is 
pale  silvery-blue;  where   the  nervules  approach  the  black   outer 


340  AUSTEALIAN  RHOPALOCERA  :  LYC.KNID.E, 

margin  they  are  defined  with  black.  Hindwing  pale  silvery-blue, 
with  a  broad  black  costal  margin;  outer  margin  black  and  linear, 
with  well  defined  grey  cilia;  where  the  nervules  meet  the  linear 
margin  they  are  defined  with  black;  anal  angle  of  wing  produced 
into  a  blunt  tail.  Abdominal  fold  well  developed.  The  wing  is 
longer  than  broad,  being  produced  towards  anal  angle,  and  the 
extremities  of  the  median  nervules  are  marked  by  prolongations 
of  the  outer  margin. 

Underside.  Forewing  grey,  lighter  on  the  lower  half  of 
the  wing;  cell  with  a  brown  spot  towards  the  base  small  and 
surrounded  by  white;  a  central  black  spot  surrounded  with  white, 
which  is  plentifully  covered  with  blue  scales,  giving  a  perfectly 
blue  border  to  the  spot  when  viewed  directly;  end  of  cell  marked 
with  a  black  bar,  bordered  internally  with  white,  covered  with 
blue  scales,  and  externally  principally  with  white;  from  this  bar 
in  an  oblique  direction  basally  to  the  inner  margin  descends  a 
black  bar  wholly  external  to  the  cell,  and  marked  off  into  spots 
by  the  median  nervure  and  nervules.  The  discal  band  consists 
first  of  a  series  of  three  small  brown  spots  surrounded  by  white, 
divided  by  the  nervules:  next  two  larger  darker  spots  bordered 
with  white,  covered  with  a  few  blue  scales  internally;  these  two 
spots  lie  between  the  discoidals,  and  the  lower  discoidal  and  first 
median  nervules;  between  the  first  and  second  median  nervules 
and  nearer  the  outer  margins  than  the  former  spots  lies  an  oval 
dark  spot,  and  obliquely  below  this  a  paler  spot.  The  general 
colour  of  the  wing  is  almost  white.  There  is  a  submarginal 
brown  band.  Hindiving  grey,  with  indistinct  markings  somewhat 
after  the  pattern  found  on  the  underside  of  the  hindwing  of  0. 
orwtes,  Hew.,  but  very  indistinct.  Along  the  disc  between  the 
nervules  is  a  series  of  five  inverted  "  fl3'ing  bird  "  marks,  the 
uppermost  being  furthest  from  the  outer  margin;  the  central 
portion  of  the  outer  margin  is  broadly  marked  with  rich  brown, 
as  is  also  the  anal  lobe. 

This  magnificent  species  is  described  from  a  single  specimen  ((J) 
caught  at  Cooktown  during  the  present  year  by  Mr.  R.  E.  Turner, 
who  has  kindly  allowed  me  to  describe  it.     In  shape  it  is  almost 


BY    G.    A.    WATERIIOUSE.  341 

identical  with  the  male  of  0.  genoveva,  Hew.,  while  the  silvery- 
blue  colour  reminds  one  of  the  three  preceding  species,  but  the 
colour  is  a  much  paler  silvery-blue,  and  also  the  anal  angle  is 
produced.  The  underside  is  much  paler  in  colour,  and  the  blue 
in  the  cell  is  more  pronounced  than  in  any  of  the  species  hitherto 
found.     The  female  of  this  species  is  at  present  unknown. 

Ogyris  ianthis,  Waterhouse. 

Proc.  Linn.  Soc.  N.S.W.  1900,  52-54,  PL  i.  figs.  1-4. 

My  description  of  this  species  was  taken  from  seven  specimens 
(<?  6,  9  1),  all  of  which  had  been  caught  on  the  wing.  Mr.  F. 
Brown  has  lately  discovered  larv?e  and  pupae  of  this  rare  species, 
so  that  I  am  now  enabled  to  give  a  fuller  description,  especially 
of  the  female. 

(J.  This  sex  shows  very  little  variation  from  the  original 
description.  The  margins  of  bred  specimens  are  on  the  upperside 
quite  black.  Among  themselves  all  the  bred  specimens  show  no 
variation  except  as  to  size,  the  average  size  being  about  34  mm., 
and  they  vary  from  30-36  mm. 

9.  As  mentioned  in  my  former  description  the  type  female  is 
slightly  smaller  than  the  average  size,  and  I  have  specimens 
varying  from  33  to  42  mm. 

Upperside.  ForpAving  :  The  central  area  in  bred  specimens 
is  a  rich  yellow  (much  deeper  than  the  type),  and  below  the  apex 
along  the  costa  is  often  a  variable  yellow  splash.  The  end  of  cell 
is  markedly  black.  Hindwlng  is  A^ery  variable,  hardly  two  speci- 
mens being  alike  in  regard  to  the  size  and  position  of  the  central 
orange  suffusion;  this  may  be  merely  absent  as  in  the  type,  or 
may  occupy  quite  half  of  the  wing;  sometimes  it  only  shows  in 
the  cell,  and  often  separate  from  the  central  spot  is  a  splash  of 
orange  below  the  subcostal  nervule. 

Underside.  Forewiyrg  as  in  type,  but  with  the  blue  margins 
to  the  spots  more  prominent.  Hindiving  with  a  white  line 
extending  from  the  middle  of  costa  to  basal  portion  of  abdominal 
23 


342  AUSTRALIAN  RHOPALOCERA  :  LYC.ENJD^E. 

margin ;  and   a  similar  semicircular  white    line    on   abdominal 
margin  towards  anal  angle. 

This  insect  is  the  connecting  link  between  0.  ahrota  and  the 
silvery-blue  species  of  the  genus.  The  types  of  this  species  have 
been  deposited  in  the  Australian  Museum. 


EXPLANATION  OF  PLATE  XIV. 

Fig.  1. — Ogyris  oroetes,  Hew.  {^),  Townsville,  Q. 

Fig.  2.-      ,,  „         ,,     (?) 

Fig.  3. — Ogyris  amarylUs,  Hew.  (<?),  Eichmoncl  Kiver,  N. S.W. 

Fig.  4.-      „  „  „     ($) 

Fig.  5. — Ogyris  hetvitsoni,  Waterh.  {^),  Townsville,  Q. 

Fig.  6.—     „  „  ,,         (?) 

Fig.  7.—     ,,  ,,  ,,         (<?),  Victoria, 

Fig.  8.—     „  „  ,,         (?) 

Fig.  9. — Ogyris  cenone,  Waterh.  [S],  Cooktown,  Q. 


343 


NOTES    AND    EXHIBITS. 


I 


Mr.  Froggatt  exhibited  samples  of  wheatshowing  the  different 
ways  in  which  the  grain  weevil  (^Calandria  oryza,  Linn.)  and  the 
grain-moth  larva  {Plodia  interpunctella,  Hubn.),  damage  stored 
wheat.  Also  three  samples  of  wheat  taken  from  the  same  bag  and 
kept  under  similar  conditions,  two  only  of  which  showed  damage 
by  weevil.  Likewise  cuttings  of  apple  wood  from  the  Ryde 
district  showing  curious  red  blotches  and  spots  caused  by  the 
presence  of  San  Jose  Scale  {Aspidiotus  perniciosus,  Comst.). 

Mr.  H.  S.  Mort  exhibited  a  specimen  of  Cerithium  dubium, 
Sowerb)^,  from  Port  Jackson,  a  new  record  for  New  South  Wales; 
and  one  of  Typhis  philippensis,  Watson,  also  from  Port  Jackson. 

Mr.  Waterhouse  exhibited  Megisba  nigra,  Misk.,  ((J 9)  from 
Cairns,  Q.;  Holochila  cyprotus,  Olliff,  ((J 9)  from  Sydney,  these 
specimens  having  been  determined  by  the  late  Mr.  Olliff;  H. 
helenita,  Semper,  ((J 9)  f  1*001  Cairns  ;  Ogyris  oroetes,  Hew., 
((J 9)  from  Townsville,  Q.;  0.  amaryllis,  Hew.,  ((J 9)  from  Rich- 
mond River;  0.  hewitsoni,  n.sp.,  ((J9)  fi'o^ii  Victoria  and  Towns- 
ville. 

Mr.  Johnston  showed  sections  and  a  mounted  specimen  of  the 
Distoynitm  described  in  his  paper. 

Mr.  Stephen  exhibited  a  very  interesting  collection  illustrating 
the  ethnology  of  Pleasant  Lsland  (0°  32'  S.  lat.,  165°  E.  long.). 

Mr.  North  exhibited  a  nest  and  two  eggs  of  the  Buff-sided 
Robin,  Paecilodryas  cerviniventris,  Gould,  together  with  a  skin  of 
the  bird,  procured  on  the  17th  January,  1902,  near  the  Daly 
River,  Northern  Territory,  S.A.  The  nest  is  a  thick-rimmed 
cup-shaped  structure,  externally  formed  of  thin  plant-stalks, 
scales  of  bark,  rootlets,  and  fine  spiral  vine  tendrils,  the  inside 
being  lined  with  a  few  dried  grass  stems,  and  a  quantity  of  black 


344  NOTES    AND    EXHIBITS. 

vegetable  fibre  resembling  horse-hair.  It  averages  externally  3 
inches  in  diameter  b}-  If  inches  in  depth,  the  inner  cup  measuring 
H  inclies  in  diameter  b}^  1:^  inches  in  depth.  The  eggs  are  oval 
in  form,  the  shell  being  close-grained,  smooth  and  lustreless. 
They  are  of  a  faint  \^ello\vi«h-green  ground  colour,  one  specimen 
having  a  band  of  confluent  blotches  around  the  thicker  end  of 
pale  ochraceous-brown  and  dull  chestnut-red,  intermingled  with  a 
few  underlying  spots  of  light  purplish-red;  the  other  has  the 
markings,  which  are  of  a  rich  purplish-red,  smaller  and  more 
evenly  distributed  over  the  surface  of  the  shell,  also  a  few  under- 
lying spots  of  much  paler  shades  of  the  same  colour.  Length 
(A)  0-75  X  0-58  inch;  (B)  0-72  x  0*6  inch.  The  specimens  were 
kindly  lent  for  description  by  Mr.  Charles  French,  Junr.,  Assistant 
Government  Entomologist,  Melbourne. 

Mr.  Maiden  exhibited,  on  behalf  of  Mr.  W.  V.  Fitzgerald, 
specimens  of  the  plants  described  as  new  species  in  the  latter's 
paper. 

Mr.  Maiden  exhibited  specimens  of  new  species  of  plants 
described  by  Prof.  K.  Schumann,  of  Berlin,  from  the  Bismarck 
Archipelago  (New  Britain).  Also  blanched  leafless  dead  stems 
of  Lepidium  riiderale,  Linn.,  "Wild  Mustard,"  from  Brewarrina. 
Cows  eat  the  plant  in  this  state  and  give  good  yields  of  milk, 
but  when  it  is  green  they  will  not  touch  it.  It  is  assumed  that 
in  the  state  as  exhibited  the  stems  had  lost  the  acridity  which 
the  fresh  plant  possesses.  Also  specimens  from  Dr.  Hugo  de 
Vries  of  Amsterdam,  of  his  Dipsacas  sylvestris  torsus,  being 
twisted  stems  showing  the  torsion  of  secondar}^  branches  to  left 
and  right. 

Mr.  Fletcher  showed  a  series  of  original  drawings  made  during 
the  memorable  journey  of  Charles  Darwin  in  H.M.S.  "Beagle," 
Captain  Fitzroy,  during  the  years  1831-36.  Also  the  MS.  journal 
of  the  artist,  the  late  Mr.  Syms  Covington,  which  with  the 
exception  of  one  break  was  kept  regularly  throughout  the  voyage; 
likewise  some  literary  relics  of  Darwin.     Mr.   Covington  accom- 


[ 


NOTES    AND    EXHIBITS.  "  345 

panied  Mr.  Darwin  as  his  personal  attendant,  and  helped  to 
collect  zoological  specimens,  particularly  birds.  Some  years  after 
the  return  of  the  "  Beagle,"  Mr.  Covington  emigrated  to  New 
South  Wales,  where  he  spent  the  rest  of  his  days.  The  exhil)ition 
of  this  very  interesting  series  of  relics  of  the  historic  \oyage 
became  possible  by  the  perspicacity  of  Mrs.  C.  W.  Morgan,  of 
Pambula,  and  the  kindness  of  Mr.  Syms  Covington,  Junr.  Mrs. 
Morgan  readily  appreciated  the  interest  which  these  mementos 
had  from  a  naturalist's  standpoint;  whereupon  Mr.  Codington 
placed  them  at  her  disposal.  With  a  view  to  their  preservation, 
and  to  their  becoming  better  known  and  accessible  to  naturalists, 
Mrs.  Morgan  had  accordingly,  on  Mr.  Covington's  behalf,  pre- 
sented them  to  the  Society. 

Mr.  Covington  well  remembers,  when  a  boy,  assisting  his 
father  to  collect  barnacles  for  transmission  to  the  author  of  the 
"Monograph  on  the  Subclass  Cirripedia";  and  the  Australian 
species  recorded  from  Twofold  Bay  in  the  second  volume  of  this 
work  were  derived  from  this  source.  The  drawings  exhibited 
comprised  views  of  Porto  Praya,  Bahia,  the  Abrolhos  islets,  the 
entrance  to  Rio,  the  entrance  to  the  Santa  Cruz  River,  Woollya 
Cove  showing  the  Beagle's  boats,  the  three  wigwams  and  the 
embryonic  garden  prepared  for  the  Fuegians  whom  Capt.  Pitzroy 
here  restored  to  their  native  land  after  a  sojourn  in  England, 
Coquimbo,  Tahiti,  Bay  of  Islands,  N.Z.,  the  entrance  to  the 
River  Derwent,  Tasmania,  showing  the  lighthouse,  King  George's 
Sound,  W.A.,  Keeling  Islands,  St.  Helena,  and  Pernambuco; 
also  some  illustrations  of  the  national  costumes  then  in  fashion 
at  Lima. 

The  exhibit  aroused  much  interest;  and  a  hearty  vote  of  thanks 
was  accorded  to  Mrs.  Morgan  and  Mr.  Covington  for  their 
valuable  addition  to  the  Society's  collection  of  original  drawings 
of  historic  interest. 


346 


WEDNESDAY,  AUGUST  27th,   1902. 


The  Ordinaiy  Monthly  Meeting  of  the  Society  was  held  in 
the  Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening,  August  27th,  1902. 


Mr.  J.  H.  Maiden,  F  L.S.,  &c..  President,  in  the  Chair. 


Mr.  H.  W.  Potts,  F.C.S.,   F.L.S.,   Hawkesbury  Agricultura 
College,  Richmond,  N.S.W.,  was  elected  a  Member  of  the  Society 


The  Donations  and  Exchanges  for  the  month,  amounting  to  49 
Vols.,  51  Parts  or  Nos.,  1  Bulletin,  2  Reports,  3  Pamphlets,  and  3 
Miscellanea,  received  from  34  Societies,  etc.,  and  4  Individuals, 
were  laid  upon  the  table. 


34^ 


ON    A    NEW    CRYPTOCARYA    FROM     LORD    HOWE 

ISLAND,  TOGETHER  WITH  NOTES  ON  OTHER 

PLANTS  FROM  THAT  ISLAND. 

By  J.  H.  Maiden. 

(Plate  XV.) 

LAURINE.S:. 

Cryptocarya  Gregsoni,  n.sp. 

In  these  Proceedings  for  1898  (p.  138,  pi.  iii.)  I  described  and 
figured  the  "  Black  Plum  "  of  Lord  Howe  Island  so  far  as  I  was 
able  to  do  so  from  leaves  and  fruits  only.  Since  then  I  have 
obtained  flowers,  and  am  able  to  supplement  the  description  in 
the  following  words  : — 

Flowers  in  a  very  short  apparently  terminal  bracteate  panicle, 
not  exceeding  the  generally  crowded  leaves.  Pedicels  very  short 
and  thick,  angular  or  flattened,  as  is  also  the  whole  inflorescence 
and  young  branches,  covered  with  minute  rust-coloured  hairs. 
Calyx-segments  scarcely  1  line  long,  rust-coloured,  hairy  inside 
and  outside.     Glands  large,  free  from  the  stamens,  sessile. 

The  plant  is  hence  a  Cryptocarya,  and  I  have  pleasure  in 
naming  it  after  my  friend  Jesse  Gregson,  Esq.,  of  Newcastle, 
who  has  actively  assisted  my  botanical  work  for  many  years. 

TILIACEJE. 

No  species  of  Elaeocarpus  has  hitherto  been  recorded  from 
Lord  Howe  Island,  but  I  have  obtained  specimens  of  a  species 
from  a  plant  stated  to  be  a  shrub  growing  on  the  top  of  Mount 
Gower.     A  leaf  and  fruit  are  figured  herewith  (PL  xv.),  but  no 


348  A    NEW    CRYPTOCARYA    FROM    LORD    HOWE    ISLAND, 

flowers  are  available.     I  give  a   description  of  the  leaves  and 
fruits  for  general  information. 

Leaves  narrow-oval,  2-2^  inches  long,  remotely  denticulate  in 
the  upper  half,  narrowed  into  a  petiole  about  ^  inch  long, 
coriaceous,  with  prominent  principal  veins,  but  not  very  promi- 
nently reticulate,  glabrous  when  full  grown.  Inflorescence  in 
short  cymes  crowded  on  the  base  of  the  last  year's  shoots.  Drupe 
ovate,  rather  above  j  inch  long,  bluish  (*?),  the  putamen  only 
slightly  wrinkled  but  distinctly  ribbed  in  the  one  fruit  examined. 

SYMPLOCACEiE. 
Symplocos  CANDELABRUM,  Brand. 

The  following  is  taken  from  Brand's  Revision  of  the  Symplo- 
cacese  (p.  39),  in  Engler's  '  Pflanzenreich  ' : — ■ 

"  49.  S.  candelabrum,  Brand,  n.sjD.  [S.  Staiuelii,  Hemsl.  in  Ann- 
of  Bot.  X.  (1896)  242,  non  F.  Muell.).— Arbor.  Folia  coriacea, 
8-10  cm.  longa,  2|-3  cm.  lata,  oblongo-elliptica,  subintegra,  in 
petiolum  3  cm.  longum  attenuata.  Racemi  glabri,  pedicellis 
longitudine  calycis;  bracteae  inaequales,  lanceolato-acutae,  cili- 
atae;  cal3^cis  lobi  tubo  subbreviores;  stamina  c.  25  (ex  1  flore) 
longitudine  corollae,  ovarium  glabrum  ;  caetera  spec,  praec* 
Fructus  ignotus. 

"Australien:  Lord  Howe  Island  (Maiden;  Herb.  Berlin). — 
Bliiht  im  April." 

Hemslej^'s  reference  is  "  *?.  Staioelli,  F.  Muell.  Fragm.  v.  p.  60, 
ix.  p.  77;  Benth.  Fl.  Austral,  iv.  p.  292  (sub  siyicata)  "  (Ann.  Bot. 
X.  242). 

MYRSINACE^. 

In  the  recent  monograph  of  the  Myrsinacea}  by  Prof.  Carl 
Mez  in  Engler's  '  Pflanzenreich,'  I  observe  that  it  is  stated  that 
there  is  no  true  3Iyrsine  in  Australia,  the  species  hitherto  referred 
to  that  genus  belonging  to  the  genus  Rapanea,  Aubl.  I  will  deal 
with  this  matter  on  a  subsequent  occasion,  only  treating  it  now 


S.  gracilis,  Brongn.  &  Gris,  etc. 


BY    J.    H.    MAIDEN.  349 

as  far  as  the  Lord  Howe  Island  flora  is  concerned.     The  genera 
Myrdne  and  Rapanea  are  chiefly  distinguished  by  the  following 
characters  : — 
Et  lilamenta  et  stylus  manifesta;  albumen  ruminatum.     Stigma 

magnum,  florum  ?  labosum Myrsme. 

Antherae  sessiles  et  stylus  nullus  albumen  saepius  laeve Ecqyanca. 

Following  are  the  previous  references  to  Lord  Howe  Myrsin- 
acese  : — 

Myrsine  crassijolia,  R.Br.,  M.  2)laiystigma,  F.v.M.,  both  quoted 
in  Muell.  Fragm.  (ix.  77);  and  other  references  given  byHemsley 
(Ann.  Bot.  x.  242). 

Mez  does  not  admit  Rapanea  crassifoUa^  Mez  {^Myrsine  crassi- 
folia,  R.Br.),  to  the  list  of  Lord  Howe  plants,  although  he  states 
(op.  cit.  365)  that  it  occurs  in  Norfolk  Island.  He,  however, 
recognises  two  species  of  Myrsine  from  Lord  Howe  Island,  viz., 
Rapanea  myrtilliyia,  Mez,  n.sp.,  and  R.  ijiloiystigma^  Mez  {Myrsine 
platystigraay  F.v.M.).  He  states  that  in  herbaria  the  two  species 
are  mixed.  The  matter  is  of  such  importance  that  1  transcril)e 
Mez's  descriptions  of  his  new  species  here  : — 

"  56.  R.  myrtillina,  Mez,  n.sp.  Frutex  humilis  ramulis  gla- 
berrimis,  crassiusculis,  valde  verrucosis.  Folia  petiolis  1-2  mm. 
longis  stipitata,  glaberrima,  integerrima,  oblongo-lanceolata, 
±  4  mm.  lata,  basin  versus  persensim  acuta  apice  rotundata, 
rigida,  margine  zona  maxime  incrassata  cincta,  laevia,  subtus 
punctis  lineisque  brevibus  creberrimus  permanifestis  insignia. 
Inflorescentiae  e  ramulis  brevissime  verruciformibus,  valde  dissitis 
formatae  perpauci  (1-2-)  florae,  non  nisi  fructiferae  cognitae. 
Bacca  globosa  ±  4  mm.  diam.  metiens,  punctulis  atris  marmorata, 
pedicello  brevi  (1-1,  5  mm.)  crassoque  stipitata,  sepalorum  lobis 
margine  glabri  triangularibus,  acutis,  atropunctatis  insidens. 

"  Australien  :  Lord  Howe's  Island  (Fullagar). 

''Nota.      In  herbariis  cum  sequenti  commixta." 

"57.  R.  j^lntystigma  (F.  Muell.),  Mez  [Myrsine  platystiyma,  F. 
Muell.  Fragm.  viii.  1873,  48). — Ramuli  crassiusculi,  tortuosi, 
glaberrimi.  Folia  glaberrima,  petiolis  usque  ad  7  mm.  longis 
stipitata,  elliptica,  basi  breviter  acuta  apice  rotundata,  usque  ad 


350  A    NEW    CRYPTOCARYA    FROM    LORD    HOWE    ISLAND. 

22  mm.  lata,  rigidiuscule  coriacea,  margine  ipso  satis  incrassata, 
costis  suberectis  subtus  tenuissime  filiformi-prominulis,  punctis 
intrafoliaceis  magnis  raris,  praesertim  prope  basin  dissitis,  in 
foliis  junioribus  subtus  prosilientibus  praedita,  supra  densissime 
punctulis  minutissimis  atris  obsita.  Inflorescentiae  e  ramulis 
verruciformibus  formatae,  1-4  florae,  ad  3  mm.  longae,  pedicellis 
brevissimis  (vix  1  mm.  longis),  crassis,  glabris;  flores  2-2,  5  mm. 
longi,  glabri;  sepala  subpatenti-pateriformia,  ad  J  connata,  lobis 
squamiformi-ovatis,  apice  paullo  fimbriatis,  punctis  perpaucis 
(1-2)  nigris  praeditis;  petala  permanifeste  ad  J  longit.  connata, 
late  elliptica,  apice  rotundata,  atro-punctata;  antherae  (in  flore 
(J)  quam  petala  paullo  tantum  breviores,  latissime  ovatae,  apice 
rotundatae  et  hie  puncto  magno  atro  subdorsali  praeditae;  ovarium 
in  flore  (J  ovoideum;  stigmate  magno  loboso. 

"  Australien  :  Lord  Howe's  Island  (Fullagar),  an  der  Kuste  des 
trop.  Australiens  (F.  Bauer). — Kult.  im  Bot.  Gart.  Sidney." 

Mez  states  the  difference  between  the  two  species  briefly  as 
follows  : — 

Folia  +  13  mm.  longa,  oblongo-lanceolata,  lineolata myrtillina. 

Folia  +  50  mm.  longa,  elliptica,  baud  lineolata jdaty.^tigvia. 

ORCHIDE^. 

Dendrobium  gracilicaule,  F.V.M.,  var.  Howeanum,  Maiden, 
these  Proceedings,  1899,  382. 

D.  gracilicaule,  F.v.M.,  (Fragm.  ix.  78,  as  a  Lord  Howe  Island 
plant). 

Dr.  A.  B.  Rendle  {Journ.  Bot.  xxxix.  197,  June,  1901)  describes 
Dendrohium  (Stachyobium)  Jonesii  from  Gerald  town,  Johnstone 
River,  Queensland,  and  surmises  that  it  may  be  identical  with  my 
Lord  Howe  Island  plant.  The  latter  is  very  closely  allied  to  D. 
Jonesii  and  seems  to  differ  from  it  merely  in  the  colour  of  the 
flowers,  which  is  more  greenish-yellow  in  the  Lord  Howe  plant, 
and  the  transverse  purple  stripes  of  the  labellum  of  D.  Jonesii  are 
onlj^  faintly  marked  in  the  other  plant. 

I  am  inclined  to  think  that  the  proper  view  is  to  look  upon  the 
orchid  in  question  as  a  form  of  D.  gracilicaule,  but  the  matter  is 


i 


BY   J.    H.    MAIDEN.  351 

not  one  of  much  consequence,  and  Dr.  Rendle's  view  may  be  the 
better  one. 

LYCOPODIACEiE. 

Lycopodium  varium,  R.Br. 

With  reference  to  the  query  in  regard  to  this  plant  formerly 
expressed  by  me  (these  Proceedings,  1899,  p.  383),  the  following- 
letter  from  Mr.  Hemsley  settles  the  point  as  far  as  I  am  con- 
cerned : — 

"  With  regard  to  LycopoJiunn  varimn  being  a  native  in  Lord 
Howe  Island,  I  can  only  say  that  the  specimen  is  correctly  named, 
and  it  is  labelled  'Lord  Howe's  Island,  Fullagar,  Herb.  F. 
Mueller,  1877,'  in  the  late  Mr.  Bentham's  handwriting.  In  Fl. 
Austral,  (vii.  674),  C.  Moore  stands  as  the  collector,  but  there  is 
no  specimen  at  Kew  except  the  one  labelled  as  above  explained. 
This  was  taken  from  Mueller's  herbarium  by  Bentham,  when  he 
had  it  on  loan.  Perhaps  Mueller's  herbarium  contains  a  specimen 
from  Moore  as  well."  I  cannot  trace  a  specimen  collected  by 
Moore  so  far,  and  that  point  is  unimportant. 


EXPLANATION  OF  PLATE  XV. 

Ela^ocarpus  sj). 
Fig.  L — Leaf  and  petiole. 
Fig.  2.— Drupe. 

Fig.  3. — Longitudinal  view  of  putamen. 

Fig.  4. — View  of   putamen  from   above,  shewing  its  distinctly  ribbed 
character. 


352 


THE  ULCER  DISEASE  (BLACK  OPHTHALMIA?)  OF 
RAINBOW  TROUT. 

{Micrococcus  jjyoyenes.) 

By  R.  Grbig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

Society. 

During  the  months  of  February  and  March  of  this  year  a 
disease  appeared  among  the  rainbow  trout,  Sahyio  irideus,  in  the 
fish  hatcheiy  of  the  New  South  Wales  Government  at  Prospect. 
The  hatchery  adjoins  the  reservoir  of  the  Sydney  water  supply, 
and  under  normal  conditions  is  furnished  with  a  continuous  supply 
of  pure  water.  Owing,  however,  to  the  drought  the  water  in  the 
reservoir  fell  very  low,  and  the  tanks  in  the  hatchery  did  not 
receive  an  adequate  supply.  As  a  result  the  water  in  the  tanks, 
which  were  carrying  a  maximum  number  of  one,  two  and  three- 
year  old  trout,  became  foul,  and  there  can  be  little  doubt  that 
this  was  indirectly  responsible  for  the  outbreak  of  the  disease. 
When  the  supply  again  became  regular  the  disease  almost 
disappeared. 

The  disease  is  characterised  by  the  fishes  having  a  sluggish, 
aimless  movement  in  the  water,  in  which  they  generally,  keep 
near  the  surface.  The  snout  and  tail  become  whitish  from  their 
rubbing  the  snout  against  the  sides  of  the  cement  tank,  and  at 
other  times  resting  vertically  in  the  water  and  moving  the  tail 
to  and  fro  against  the  cement  tank  bottom.  When  the  eye 
becomes  affected  the  lens  whitens  and  the  eye  decays,  leaving  an 
empty  capsule.  The  epidermis  generally  becomes  dark  in  colour, 
and  this,  in  conjunction  with  the  atrophy  of  the  eye,  has  given 
rise  to  the  name  of  the  disease,  black  ophthalmia. 


IJY    R.    GREIG    SMITH.  353 

Other  fishes  develop  small  white  ulcers  on  various  parts  of  the 
epidermis.  Beneath  the  skin  and  under  the  sores  the  muscle  is 
hsemorrhagic.  The  scales  fall  off  from  these  places,  the  epidermis 
disappears,  and  the  muscle  becomes  exposed.  This  has  a  reddish 
colour,  while  the  epidermis  at  the  margin  for  about  one-sixteenth 
of  an  inch  is  white,  so  that  the  combination  of  colours  gives  the 
sore  a  striking  appearance.  Although  these  are  of  the  nature  of 
ulcers  there  is  no  appearance  of  pus.  It  is  possible  that  the 
black  ophthalmia  and  the  production  of  these  ulcers  are  due  to  two 
different  causes,  but  at  the  beginning  of  the  investigation  it  was 
believed  that  only  one  disease  was  affecting  the  trout. 

The  post-mortem  examination  of  many  carcases  that  I  received 
showed  that  the  body  organs  were  apparently  healthy,  and  that 
they  were  practically  sterile.  Neither  moulds  nor  algse  could  be 
detected  on  the  ulcers,  in  the  orbital  capsule  or  on  the  partially 
affected  eye.  The  muscles  and  organs  when  examined  in  the 
fresh  condition  were  free  from  parasites.  But  under  the  small 
unbroken  ulcers  coccoid  forms,  occurring  singly  and  in  groups, 
were  detected,  and  from  these  a  pure  culture  of  a  white  micro- 
coccus which  had  all  the  cultural  and  other  characters  of  Micro- 
coccus pyogenes,  (Rosenbach)  Migula,  was  obtained. 

In  mammals  this  bacterium  is  responsible  for  the  formation  of 
ulcers,  boils,  etc.,  containing  pus,  and  the  presence  of  pus  may  be 
said  to  be  the  only  difference  between  the  ulcers  of  the  trout  and 
those  of  the  higher  vertel^rates.  As  the  pyogenic  bacteria  are 
not  known  to  produce  pus  in  the  epidermal  lesions  of  fishes,  it 
seemed  probable  that  the  disease  would  be  reproduced  by  inocu- 
lating the  micrococcus  into  healthy  trout. 

To  test  this  point  Mr.  J.  A.  Brodie,  of  the  Department  of 
Fisheries,  enabled  me  to  infect  several  trout  at  the  Prospect 
hatcher}^  The  inoculations  took  place  some  time  after  the 
isolation  of  the  bacterium,  because  it  was  not  until  about  two 
months  had  elapsed  that  a  constant  supply  of  water  could  be 
obtained.  Six  healthy  trout  were  inoculated  with  a  suspension 
of  the  micrococci  in  normal  saline  under  the  epidermis  of  the 
posterior  region,  between  the  median  and  dorsal  lines.     After  the 


354  ULCER    DISEASE    OF    RAINBOW    TROUT, 

operation  the  fishes  were  put  into  metal  tubs,  fed  with  a  stream 
of  fresh  water  from  an  o^^erhead  tap,  and  the  tubs  were  protected 
by  nets  placed  over  them.  The  trout  were  left  in  charge  of  the 
caretaker,  who  had  instructions  to  forward  at  once  to  Sydney  the 
carcases  of  any  fish  that  might  die,  as  delay  in  doing  so  might 
seriously  influence  the  investigation. 

On  the  sixth  day  after  inoculation  one  of  the  trout  died,  and 
the  carcase  was  received  at  the  laboratory.  The  caretaker  had 
noted  that  it  had  been  sickly  on  the  previous  two  or  three  days. 
The  carcase  had  no  epidermal  lesions,  and  on  dissection  the 
organs  were  found  to  be  healthy.  The  blood  vessels  of  the 
stomach,  intestine  and  milt  were,  however,  much  congested,  and 
the  muscle  at  the  point  of  inoculation  was  soft  and  haemorrhagic. 
In  the  bacteriological  examination,  Micrococcus  pyogenes  was 
obtained  from  the  muscle  at  the  site  of  inoculation,  from  the 
spleen  and  from  the  kidne}^  The  muscle  on  the  reverse  side  and 
the  heart  blood  were  sterile.  Two  fishes  died  on  the  ninth  day, 
a  Saturday,  but  owing  to  a  misunderstanding  regarding  the 
necessity  of  a  speedy  despatch  and  the  arrival  at  the  laboratory 
on  Sunday,  the  carcases  were  not  sent.  A  fourth  trout,  which 
had  developed  a  w^hite  mark  on  the  head,  somehow  eluded  the 
protecting  net  and  got  out  of  the  vessel.  The  caretaker  noted 
that  after  death  the  mark  had  almost  disappeared.  It  was 
unfortunate  that  this  carcase  was  not  despatched  to  me.  A 
fifth  fish  died  eight  weeks  after  the  inoculation  with  the  external 
lesions  of  ophthalmia — viz.,  one  of  the  eyes  had  decayed — only 
the  capsule  remained.  The  epidermis  of  the  head  and  tail  had 
whitened  in  places,  but  no  ulcers  had  formed  on  the  body.  Upon 
dissection  the  organs  were  seen  to  be  healthy,  but  there  was  a 
considerable  congestion  in  the  blood  vessels  of  the  intestine  and 
body  cavity.  The  organs,  and  especially  the  milt,  were  stained  a 
deep  orange-yellow  by  the  bile.  Like  the  fishes  that  died  after 
natural  infection,  the  body  lesions  of  this  experimental  fish  were 
not  pronounced.  The  juices  of  the  various  organs,  the  blood  and 
portions  of  the  brain  were  examined  bacteriologically,  and  while 
putrefactive    bacteria   were    found,    Micrococcus  pyogenes   could 


BY    R.    GREIU    SMITH.  355 

neither  be  detected  nor  isolated.  This  is  precisely  what  occurred 
with  the  fishes  that  succumbed  naturally  to  the  disease.  It  was 
only  from  those  trout  which  had  body  sores  that  the  micrococcus 
could  be  obtained.  The  sixth  experimental  trout  was  alive  and 
apparently  healthy  at  the  time  of  writing. 

In  reviewing  the  infection  experiment,  it  is  seen  that  one  was 
lost,  thus  reducing  the  number  to  five.  Three  of  the  five  succumbed 
about  the  same  time,  and  since  the  inoculated  bacterium  was 
recovered  from  one,  it  is  probable  that  had  the  others  been  sent 
to  the  laboratory  the  organism  would  also  have  been  recovered 
from  them.  The  fourth  developed  ophthalmia  and  died,  while 
the  fifth  was  apparently  unaffected  by  the  inoculation. 

Since  the  infected  micrococcus  could  not  be  separated  from  the 
trout  that  succumbed  with  the  lesion  of  ophthalmia,  it  is  possible 
that  the  fish  might  have  died  quite  independently  of  the  inocula- 
tion. On  enquiry,  I  was  informed  by  Mr.  Brodie  that  about 
three  trout  had  died  weekly  from  ophthalmia  since  the  time  of 
the  inoculation,  and  that  epidermal  sores  had  never  been  observed 
after  the  height  of  the  disease.  As  the  tanks  were  carrying  about 
400  fishes,  this  means  a  death  rate  of  1  in  20  during  the  period 
that  elapsed  between  the  time  of  inoculation  and  the  death  of  the 
fourth  experimental  fish. 

So  far  as  the  infection  experiments  go,  the  micrococcus  was 
pathogenic,  although  it  did  not  produce  lesions  similar  to  those 
from  which  it  was  isolated.  This  can  be  explained  by  the 
experimental  fishes  being  in  more  healthy  surroundings.  Had 
the  water  been  as  unhealthy  as  that  in  the  tanks  during  the 
height  of  the  disease,  it  is  probable  that  the  ulcers  would  have 
developed.  Among  mammals  the  same  micrococcus  produces 
sores  when  the  vitality  of  the  animal  is  lowered  by  some  cause, 
and  since  it  obtains  among  the  higher  animals,  there  is  no  reason 
wh}^  it  should  not  also  hold  among  fishes.  Another  point  to  be 
remembered  is  that  by  the  time  a  constant  flow  of  water  could 
be  depended  upon  at  the  hatchery  about  two  months  had  passed, 
and  during  this  time  the  bacterium  may  have  lost  in  part  its 


356  ULCER    DISEASE    OF    RAINBOW    TROUT, 

ulcer-producing  function  while  under  artificial  conditions  of 
culture.  There  is,  therefore,  reason  to  believe  that  Micrococcus 
pyogenes  is  the  cause  of  the  disease  in  salmon  trout,  characterised 
by  the  production  of  ulcers,  and  •  that  its  specific  pathogenic 
action  is  influenced  by  an  unhealthy  or  low  condition  of  the  fish 
such  as  might  be  caused  by  an  insufficient  water  suppl}'-  and  the 
overcrowding  of  the  tanks. 

There  is  the  doubt  that  the  production  of  sores  and  ophthalmia 
are  induced  by  two  different  agents.  The  reasons  for  believing 
this  are  —(1)  One  carcase  seldom  shows  both  types  of  lesion;  (2) 
fishes  die  of  ophthalmia  at  times  when  no  ulcers  are  found  upon 
other  trout;  (3)  Micrococcus  pyogenes  could  not  be  isolated  from 
those  cases  of  ophthalmia  which  did  not  show  the  ulcers;  and  (4) 
the  ulcer  disease  is  known  in  other  localities,  and  in  describing 
the  disease  writers  make  no  mention  of  ophthalmia. 

In  a  report  upon  an  epidemic  among  brook  trout  {Salvelinus 
fontinalis)  on  Long  Island,  Calkins"^  describes  a  disease  which 
appears  to  be  identical  witli  the  ulcer  disease  of  rainbow  trout. 
His  photograph  of  the  diseased  brook  trout  shows  an  appearance 
precisely  similar  to  that  which  I  saw  on  the  rainbow  trout.  He 
did  not  examine  the  carcases  bacteriological^,  but  as  the  result 
of  many  observations  upon  sections  of  the  organs  of  diseased 
fishes  made  after  imbedding  in  paraffin,  Calkins  concluded  that 
the  disease  was  caused  by  a  new  sporozoon  which  he  named 
Lymphosporidium  truttce.  The  plates  which  are  reproduced  in 
his  paper  show  spores  2  ^  in  diameter,  and  he  mentions  sporo- 
zoites  0-5  /x  in  diameter.  The  micrococci  which  I  found  might 
simulate  these  sporozoites.  In  the  milt  of  diseased  rainbow 
trout  forms  were  met  which  at  first  sight  appeared  to  be 
Calkins'  spores,  but  these  proved  to  be  the  bodies  of  the  sperma- 
tozoa. The  similarity  between  Calkins'  spores  and  the  bodies  of 
the  spermatozoa  is   remarkable.     They  both   stain   deeply  with 


*  Fourth  Annual  Report  of  the  Commissioners  of  Fisheries,  Game  and 
Forests  of  the  State  of  New  York,  1898. 


BY    R.    GREIG    SMITH.  357 

nuclear  stains:  they  are  the  same  shape  and  the  same  size;  and, 
lastly,  they  both  have  the  cleft  at  the  broad  end,  which  Calkins 
says  is  the  first  stage  in  the  formation  of  the  sporozoites. 

Marsh*  also  writes  about  the  same  disease  among  brook  trout, 
and  mentions  that  Loch  Leven  trout  are  susceptible,  but  that 
rainbow  trout,  kept  in  ponds  by  the  side  of  the  Loch  Leven  trout, 
have  not  contracted  the  disease,  and  appear  to  be  immune.  The 
experience  at  Prospect  shows  that  the  immunity  must  have  been 
accidental. 


*  The  Fishing  Gazette,  xix  401,  417. 

24: 


358 


NOTES  ON  AUSTRALIAN  NEUROPTERA  AND  THEIR 
LIFE-HISTORIES. 

By  Walter  W.  Froggatt,  F.L.S. 

The  Neuroptera  treated  of  in  this  paper,  with  one  exception 
[Bittacus  australis),  all  belong  to  the  Family  Hemerohiidcp.  This 
has  been  subdivided  into  seven  Subfamilies  containing  a  number 
of  very  handsome  lace-wings,  of  which  the  ant-lions  are  probably 
the  best  known.  Although  nearly  fifty  years  have  elapsed  since 
most  of  the  Australian  species  were  described,  little  or  nothing 
has  been  recorded  about  their  habits  or  transformations;  yet 
several  species  are  common  in  the  neighbourhood  of  Sydney,  and 
the  funnel-shaped  pits  of  the  larval  ant-lions  are  common  in  any 
dry  sheltered  sandy  place.  I  am  not  aware  that  anj^one  has  kept 
the  larvae  and  bred  the  perfect  insects  so  that  the  species  could  be 
determined. 

All  our  known  species  have  been  described  from  dead,  shrivelled, 
and  sometimes  very  old  and  mutilated  specimens,  obtained  from 
collectors  without  any  definite  habitats;  and,  as  in  the  case  of 
many  of  Walker's  descriptions,  in  which  colour  and  size  are  the 
chief  characters  given,  when  the  living  or  freshly-captured  insects 
are  examined  their  characters  do  not  agree  always  with  those 
mentioned  in  the  original  descriptions. 

Through  the  kindness  of  Dr.  Sharp,  of  the  Cambridge  Uni- 
versity Museum,  to  whom  I  submitted  a  number  of  t3^pical 
Neuroptera  for  determination,  and  after  a  careful  examination  of 
the  specimens  in  the  Macleay  Museum,  in  conjunction  with  the 
descriptions  of  Leach  and  Walker,  I  have  been  able  to  get  my 
specimens  identified  and  named. 

During  the  last  few  years,  while  visiting  the  country  districts 
on  departmental  work,  I  have  had  opportunities  of  collecting  the 
larvce  of  some  of  these  insects  and  breeding  out  the  imagines. 


BY    WALTER    W.    FROGGATT.  359 

Accordingly,  in  addition  to  notes  on  life-histories,  I  liave  given 
brief  descriptions  of  some  of  the  species,  with  remarks  on  habitats 
and  distribution. 

BiTTACUS    AUSTRALIS,  Klug. 

Klug,  Abh.  Akad.  Wiss.  Berlin,  100,  11,  1836;  B.  corethrarncs, 
Ramb.,  Hist.  Nat.  Ins.  Neurop.  327,  1842. 

This  representative  of  a  world-wide  genus  has  an  extensive 
range  over  Australia  and  Tasmania.  It  is  very  common  in  the 
neighbourhood  of  S3'^dney  in  early  summer,  frequenting  Lepto- 
spermum  and  Melaleuca  bushes  when  in  bloom,  to  the  foliage  of 
which  the  insects  cling  by  the  two  fore  pairs  of  legs,  letting  the 
long  hind  ones  hang  loosely  down  behind  ready  to  strike  at  any 
incautious  insect  that  may  come  within  kicking  distance.  In 
general  appearance  they  closely  resemble  thickset  crane  flies  or 
*' daddy  longiegs  ";  and  apparently,  by  mimicking  these  harmless 
Diptera,  they  are  enabled  not  only  to  catch  them,  but  to  beguile 
other  helpless  creatures  within  reach  of  their  deadly  hind  legs. 

General  colour  reddish-brown  marked  with  black;  wings  narrow, 
fuscous,  thickly  covered  with  black  nervures;  stigma  of  the  same 
colour  near  the  tips.  Head  long,  slender,  and  turned  downwards 
in  front,  forming  a  regular  beak  furnished  with  two  sharp  jaws. 
Legs  long,  cylindrical,  lightly  clothed  with  hairs,  and  of  a 
curious  annular  structure  extending  into  the  tarsi,  the  most 
remarkable  point,  however,  being  the  form  of  the  hind  legs. 
Femora  thickened  like  the  hind  leg  of  a  grasshopper;  the  tibiae 
long  and  slender,  with  two  very  long  slender  spines  standing  out 
at  the  extremity;  tarsi  composed  of  five  large  joints  fringed  on 
the  undersurface  with  fine  spines,  the  last  joint  curving  round  to 
a  sharp  point  (taking  the  place  of  the  tarsal  claw);  in  the  living 
insect  the  whole  of  the  undersurface  of  the  tarsi  is  covered  with  a 
sticky  sponge-like  process.  When  the  tarsi  touch  an  insect  they 
curl  round,  the  terminal  claw  closing  over  between  the  apical 
spines  of  the  tibiae. 

I    have    frequently  taken    home    specimens    of    Bittacus    and 
enclosed  them  in  a  glass  jar  into  which  house  flies  were  introduced. 


360  AUSTRALIAN  NECROPTERA, 

These  were  very  soon  captured  by  the  neuropters  striking  out  with 
their  hind  legs;  a  fly  enfolded  on  the  stick}^  spiny  tarsi  was  held 
in  a  vice-like  grip,  while  the  leg  curled  round  under  the  head; 
and  in  a  ver}^  short  time  it  was  sucked  dry  and  dropped  on  the 
bottom  of  the  jar. 

F.  Brauer  has  given  an  account  of  the  insect-catching  habit  of 
the  common  European  species  {Bittaciis  tipvlariufi,  Fabr.).^ 

Glenurus  pulchellus,  Rambur. 

Myrmehon  pulchellus,  Ramb.,  Hist.  Nat.  Ins.  Neurop.  408, 
1842. 

This  is  the  commonest  Australian  species  of  ant-lion,  and  has 
a  'wide  range  over  New  South  Wales  and  Queensland.  The 
perfect  insects  are  generally  found  resting  on  grass  stalks  or  twigs 
in  open  forest  country,  with  their  wings  folded  down  over  the 
body.  When  disturbed  they  flit  along  with  a  very  awkward 
motion  as  if  the  wings  were  too  big  for  the  body,  so  that  they 
are  very  easily  run  down  and  captured. 

Length  of  bod}'-  1  inch;  expanse  of  wings  2^  inches. 

General  colour  chocolate-brown,  mottled  and  marbled  with 
light  brown;  face  below  the  eyes,  a  line  between  them,  under- 
surface  of  head,  thorax,  and  base  of  abdomen  3'ellow;  antenme 
dark  reddish-brown,  at  the  base  and  near  the  tip  marked  with 
dull  orange.  Legs  light  brown,  mottled  with  black.  Forewings 
semitransparent,  thickly  covered  with  a  fine  network  of  nervures, 
mottled  and  shaded  with  black,  forming  a  distinct  spot  at  the 
stigma;  all  the  nervures  clothed  with  little  hairs,  finest  along  the 
front  margin;  hind  wings  similar  in  form,  with  the  apical  portion 
deeply  blotched  with  chestnut-brown,  enclosing  a  double  or  con- 
fluent white  spot,  with  another  small  spot  at  the  tip. 

The  larva  of  this  species  is,  I  believe,  the  common  ant-lion 
found  forming  funnel-shaped  pits  in  sand  under  the  shelter  of 


*  Bauer,  "  Ueber  die  Lebensweise  cles  liittanis  f//)»Zrt>•//^^•,"  Fabi.,  Verb, 
z.-b.  Ver.  in  Wien,  iii.,  1853,  p.  151. 


BY    AVALTER    W.    FROGGATT.  361 

logs,  caves,  etc.,  at  the  bottom  of  which  it  hides,  and  preys  upon 
the  ants  that  ma}'^  fall  into  the  pit-fall. 

Glenurus  falsus,  Walker. 

Myrmeleon  falsus,  Walk.,  Brit.  Mus.  Cat.  Neurop.  393,  1853. 

This  species  was  described  from  a  specimen  collected  at  Port 
Stephens,  N.S.W.  It  is  not  so  common  as  the  former  species, 
but  specimens  of  it  are  to  be  found  about  Sydney  in  midsummer. 
It  differs  from  G.  jndchellus  in  having  the  parallel  markings  on 
the  forewings  much  more  defined  and  dai^ker,  and  only  one  black 
irregularly  rounded  blotch  near  the  tip  of  the  hind  wings.  The 
general  colour  is  also  much  darker  without  any  yellow  markings. 
In  structure,  size  and  form  of  wings  there  is  very  little  difference, 
except  that  the  wings  are  a  little  more  slender  and  pointed  at  the 
tips. 

Glenurus  erythrocephalus,  Leach. 

Myrmeleon  erytlirocephahini,  Leach,  Zool.  Misc.  i.  70,  pi.  xxx., 
1814. 

This  distinctive  lace-wing  was  also  figured  and  briefly  described 
in  Griffith's  '  Animal  Kingdom  '  (Insects,  ii.  1832,  pi.  105,  p.  327). 
I  have  a  specimen  collected  at  Grafton,  Richmond  River,  that 
agrees  with  the  wing-markings  of  this  last  figure;  but  Dr.  David 
Sharp,  Curator  of  the  Cambridge  Museum,  to  whom  a  duplicate 
specimen  was  forwarded,  thinks  it  is  a  distinct  species.  I  have 
examined  several  in  the  Macleay  Museum,  and  find  that  the 
black  spots  and  markings  are  very  variable;  in  the  hindwings  of 
one  specimen  the  wing  on  one  side  bears  five  spots  and  its  fellow 
only  three. 

Length  of  body  1|;  expanse  of  wings  2^  inches. 

Head,  prothorax,  centre  of  meso-  and  metathorax  and  thighs 
rich  reddish-orange;  antennte,  tibiee  and  tarsi  black;  eyes  large, 
hemispherical,  projecting  on  the  sides,  slate-grey;  remainder  of 
thorax  and  abdominal  segments  greyish-brown.  Wings  large, 
hyaline;  forewings  ornamented  with  three  parallel  rows  of  black 
spots,  the  first  and  second  consisting  of  9-10  and  the  third  of  two 


362  AUSTRALIAN  ^EUROPTERA, 

blotches;  hindwings  with  usiiall}'-  three  spots  towards  the  tip,  but 
the  markings  are  variable. 

I  have  never  taken  this  species  alive,  but  I  have  specimens 
from  Grafton,  Newcastle  and  Sj^dney.  It  will  probabl}-  require 
to  be  placed  in  another  genus  when  the  classification  is  worked  up, 
as  it  differs  from  the  typical  Glenurus.  Leach's  type  is  said  to 
be  in  the  Macleay  Museum  Collection. 

Glenurus  fundatus,  Walk. 

Myrmeleon  fiLiidatua,  Walk.,  Brit.  Mus.  Cat.  Neurop.  320,  1853. 

This  is  our  largest  species  of  ant-lion,  measuring  from  an  inch 
to  half  as  long  again  in  the  body;  and  with  an  expanse  of  over 
four  inches  across  the  wings. 

It  has  a  very  wide  range  over  Australia;  Walker  records  it 
from  Port  Curtis,  Q.,  Western  Australia,  and  Tasmania.  Mr. 
Masters  informs  me  that  he  has  taken  it  about  Sydney.  I  have 
specimens  from  Kalgoorlie,  W.A.,  and  a  fine  series  obtained  at 
Townsville,  N.Q.,  was  recentl}^  sent  to  me  by  Mr.  F.  A.  Dodd. 

General  colour  brown,  thickly  covered  w^ith  greyish  hairs, 
longest  and  densest  upon  thorax  and  legs;  face,  undersurface  of 
prothorax,  base  of  forelegs  and  wings  yellow  ;  antennae  grej^ 
Wings  hyaline;  hindwings  shortest  but  similar  to  forewings  in 
markings,  the  nervures  so  finely  marked  with  dark  brown  that 
they  have  a  greyish  tint. 

Glenurus  circuiter,  Walk. 

Myrmeleon  circuiter,  Walk.,  Brit.  Mus.  Cat.  Neurop.  400. 

This  is  one  of  the  most  distinctive  Australian  species  on  account 
of  the  rich  chestnut-coloured  blotches  and  spots  forming  a  varie- 
gated pattern  over  the  large  hyaline  wings,  both  pairs  of  which 
curve  round  at  the  apex  in  front,  and  are  distinctly  arcuate  on 
the  hind  margin  at  the  tips. 

The  species  has  a  wide  range.  There  are  specimens  in  the 
Macleay  Museum  labelled  Rockhampton,  Q.,  and  New  South 
Wales.     Mr.  Masters  informs  me  that  he  took  it  many  years 


BY    AVALTER    W.    FROGGATT.  363 

ago  ill  Sydney  gardens.  I  liave  captured  a  specimen  at  Bathurst, 
N.S.W. 

Length  of  body  IJ;  expanse  of  wings  3 J  inches. 

Head  and  prothorax  ochreous,  antennae  yellow,  legs  dull 
yellowish-brown;  meso-  and  metathorax black;  abdominal  segments 
brown  marked  with  yellow.  The  whole  insect  clothed  with  black 
hairs,  thickest  on  the  fangs  and  prothorax. 

SUPHALASCA    SABULOSA,  Walk. 

Ascalaphiis  sabulosits,  Walk.,  Brit.  Mus.  Cat.  Neurop.  427,  1853. 

This  insect  was  originally  described  from  Adelaide,  S.A.  My 
specimens  were  obtained  at  Condobolin,  N.S.W. 

Length  of  body  1 ;  expanse  of  wings  2  inches. 

General  colour  dark  brown,  with  face,  legs,  base  of  wings  and 
marks  on  abdominal  segments  yellow;  the  whole  insect  thickly 
clothed  w4th  fine  long  greyish  and  brown  hairs.  Eyes  large, 
rounded,  light  chestnut,  mottled  with  black.  Antennae  composed 
of  30  joints  of  equal  length  standing  straight  up  above  the  head; 
ver}'  slender,  with  a  hollowed  spoon-shaped  club  at  the  apex. 
Wings  of  uniform  length,  transparent,  with  a  clouded  parallel 
stripe  down  the  front  margin. 

Specimens  of  the  larvae  of  this  lace-wing  were  found  under  the 
bark  pulled  otf  the  trunk  of  a  dead  sheoak  (Casuarina)  at  Condo- 
bolin, N.S.W.,  on  the  9th  of  September.  They  were  resting 
against  the  trunk,  and  did  not  move  until  touched.  When  placed 
in  a  box  they  rested  against  the  side  in  a  similar  manner,  with 
the  head  pointing  upwards  and  the  ventral  surface  flattened 
against  the  wall,  apparently  never  moving  in  the  day  time;  nor 
did  they  eat  any  flies  or  other  live  insects  placed  in  the  box. 
They  remained  in  this  state  a  few  days  over  a  month,  when  one 
pupated,  forming  the  usual  round,  parchment-like  cocoon,  about 
I  inch  in  diameter,  from  which  the  insect  emei-ged  on  the  9th 
January. 

Larva. —  Length  of  body  J  inch,  with  the  projecting  jaws  ^  inch 
longer.     General  colour  dark   chocolate-brown,  slightly  mottled 


364  AUSTRALIAN  NEUROPTERA, 

with  lighter  brown,  the  whole  of  the  dorsal  surface  clothed  with 
fine,  flattened,  rosette-like  tufts,  with  finer  feather-like  ones  upon 
the  ventral  surface,  and  the  outer  margins  of  the  thoracic  and 
abdominal  segments  armed  with  spiny  processes.  Head  deeply 
arcuate  in  centre,  with  the  side  at  the  base  of  each  mandible 
swelling  out  and  rounded  to  the  back  of  the  head,  which  is 
flattened  on  the  summit  and  arcuate  behind.  ^Mandibles  very 
large,  curving  round  to  the  pointed  extremities,  with  the  outer 
edge  of  the  basal  portion  clothed  with  spiny  tubercles,  the  inner 
edges  furnished  with  three  stout  spines,  the  anterior  one  largest, 
the  spaces  between  them  covered  with  short  blunt  spines.  Eyes 
forming  a  rounded  space  containing  six  separate  lenses  covered 
with  spiny  tubercles.  Prothorax  forming  a  regular  neck  enclosed 
by  the  base  of  the  head;  dorsulum  and  mesonotum  forming  short 
narrow  folds  behind.  Abdomen  swelling  out  on  sides,  tapering 
at  apex  to  a  slender  pointed  tip.  Legs  long,  stout,  covered  with 
fine  spines;  tarsi  long,  furnished  with  stout  claws. 

8uPHALASCA  FLAViPES,  Leach. 

Ascalaphus  Jlavipes,  Leach,  Zool.  Misc.  i.  48,  pi.  xx,:  Walk., 
Brit.  Mus.  Cat.  Neurop,  420,  1853:  Bubo  Jlavipes,  Ramb.,  Hist. 
Nat.  Ins.  Neurop.  357,  1842;  Suphalasca  Jlavipes,  Lefebvre,  Mag. 
Zool.  1842. 

This  insect  is  about  the  same  size  as  the  preceding  species,  but 
differs  in  having  the  head  and  thorax  yellow  marked  with  brown, 
and  the  thick  tuft  of  hair  clothing  the  front  of  the  head  black 
instead  of  grey,  though  below  and  above  it  is  grey.  The  legs  are 
bright  yellow  except  the  base  of  the  thighs  and  the  tarsi,  which 
are  black.  The  wings  are  marked  with  a  bright  yellow  blotch 
that  looks  like  a  true  stigma  at  the  extremity  of  each  wing, 
running  into  the  termination  of  the  parallel  nervures. 

Leach  and  Walker  give  the  locality  as  New  Holland.  My 
.specimens  were  taken  at  Bathurst,  N.S.W.,  clinging  to  a  slender 
branch  of  a  scrub  tree,  with  the  wings  folded  down  the  1)ack. 


BY    WALTER    AV.    FROGGATT.  365 

SUPHALASCA   SUBTRAHENS,   Walk. 

Ascalaphus  subtrahens,  Walk.,  Brit.  Mus.  Cat.  Neurop.  430, 
1853. 

This  species  is  unknown  to  me.  It  is  a  darker-coloured  species 
than  S.  flavipes.     The  type  was  described  from  Adelaide,  S.A. 

Nymphes  myrmelionides,  Leach. 

Zool.  Misc.  i.  p.  102,  pi.  xlv.  1814. 

This  handsome  insect  has  a  wide  range  over  New  South  Wales. 
I  have  specimens  found  in  the  neighbourhood  of  Sydney,  New- 
castle and  Armidale;  also  in  Southern  Queensland.  The  type 
was  in  Mr.  Alexander  Macleay's  Collection,  and  was  recorded 
from  Australia,  Mr.  McLachlan  says  there  is  a  fine  series  in  the 
British  Museum. 

Length  of  body  1 ;  expanse  of  wings  3-J  inches. 

General  colour  reddish-brown,  abdomen  darker;  eyes  grey; 
antennas  black,  reddish  at  apex,  composed  of  72  short  rounded 
joints  covered  with  fine  hairs.  Wings  large,  with  the  extremity 
of  each  blotched  with  an  elongate  white  mark  enclosed  with 
brown. 

Larvae  were  obtained  at  Armidale  about  the  end  of  November, 
hiding  among  rubbish  or  clinging  to  overturned  logs,  so  well 
€oated  with  bits  of  dirt  that  only  the  front  of  the  head  and 
mandibles  were  exposed;  until  disturbed  they  remained  jDerfectly 
motionless,  but  moved  quickly  when  touched.  In  captivity  they 
took  no  food,  and  after  remaining  for  three  weeks  in  a  jar  three 
of  them  pupated,  forming  typical,  rounded,  parchment-like,  pupal 
cases.  From  the  situation  in  which  they  were  found  they  would 
probably  feed  upon  wood  ants. 

Larva. — Length  8  lines;  general  colour  brown,  marked  with 
darker  parallel  lines  on  the  dorsal  surface  of  the  thorax.  Head 
a,rcuate  in  front,  rounded  on  the  sides,  with  the  hind  margin 
forming  two  rounded  lobes  attached  to  the  thorax  b}^  a  slender 
neck.  Jaws  longer  than  head,  shaped  like  a  pair  of  calipers, 
with  slender  points  and  a  sharp  pointed  thorn  in  the  centre  of 


366  AUSTRALIAN  SEUROPTERA, 

the  inner  edge,  clothed  with  long  fine  hairs,  shortest  at  extremi- 
ties. Eyes  small,  composed  of  six  rounded  facets,  projecting 
beside  the  base  of  the  jaws.  Antennae  standing  out  straight 
above  eyes,  consisting  of  a  short  cylindrical  basal  joint  and  a  long 
filiform  appendage.  Thoracic  neck  finely  granulated  with  little 
star-like  spots;  prothorax  in  front  produced  into  two  lobes  pro- 
jecting on  either  side  of  the  head,  together  with  the  rest  of 
the  thorax  forming  a  shield- shaped  plate.  Abdomen  narrow  at 
base,  elongate-oval,  with  a  short  pointed  tubercule  standing  out 
in  centre  of  each  segment.  The  whole  of  the  dorsal  surface 
clothed  with  fine  downy  hairs,  longest  on  the  sides,  with  a  fringe 
of  fine  linger-like  appendages,  longest  behind  the  head,  shortest 
at  the  tip  of  the  abdomen,  each  fringed  with  fine  hairs  so  that 
they  look  like  feathers.  Ventral  surface  pale  ochreous,  flattened, 
clothed  with  fine  hairs,  thickest  on  the  abdominal  segment. 
Legs  long,  slender,  hairy;  tarsi  stout,  claws  large. 

PoRiSMUS  STRiGATUs,  Burm. 

Osinylus  strigatus,  Burm.,  Handb.  Ent.  ii.  984;  Ramb.,  Hist. 
Nat.  Ins.  Neurop.  415  :  Chrysopa  macuUpennis,  White,  Eyre's 
Travels,  Vol.  i.  Appendix,  p.  432,  pi.  iv.,  fig.  2. 

This  insect  has  a  wide  range  over  New  South  Wales  and 
Queensland,  and  in  the  early  summer  is  not  uncommon  in  the 
neighbourhood  of  Sydney.  I  once  took  about  two  dozen  at  rest, 
like  a  number  of  moths,  upon  the  trunk  of  a  large-white  gum-tree. 

Length  of  body  h',  expanse  of  wings  1^  inches. 

Head  red;  palpi  and  antenna  black,  the  latter  composed  of 
about  75  joints,  the  first  broad  and  oblong,  the  others  small, 
round  and  clothed  with  short  spiny  bristles;  upper  surface  of 
thoracic  and  abdominal  segments  black;  undersurface  and  legs 
brown,  except  the  fore  pair,  which  are  yellow.  Wings  rich 
])lackish-brown  with  bright  metallic  reflections;  forewings  mottled 
and  barred  with  pale  yellow  forming  a  row  of  four  transverse 
bars  on  the  base,  with  smaller  parallel  lines,  and  several  spots  and 
two  curved  lines  at  the  apex  ;  hindwings  with  the  basal  half 
and  extreme  tip  yellow. 


I 


BY    WALTER    W.    FKOGGATT.  36T 

PsYCHOPSis  MiMiCA,  Newman. 

Newni,,  Entomologist,  p.  415,  1840-2;  Hemerohius  mimictis^ 
Walk.,  Brit.  Mus.  Cat.  Neurop.  p.  279. 

This  beautiful  insect  when  at  rest,  with  its  wings  drooped  down 
on  the  sides  of  the  short  cylindrical  body,  the  head  tucked  under 
and  almost  hidden  from  view  by  the  long  hairs  springing  from 
the  front  of  the  thorax,  is  wonderfully  like  some  of  the  homop- 
terous  insects  belonging  to  the  genus  FJata. 

Length  of  body  |;  expanse  of  wings  2^  inches. 

General  colour  dull  pale  yellow,  thickly  clothed  with  buff- 
coloured  hairs-  Head  short,  turned  down  in  front.  Eyes  large, 
dark  green,  ocelli  in  line  between  the  eyes.  Antennae  short,, 
slender,  composed  of  35  short  cylindrical  joints  clothed  with  fine 
hairs.  Jaws  large,  reddish-brown.  Wings  large ;  forewings 
swelling  out  in  front,  rounded  at  the  tips,  somewhat  arcuate 
behind;  hindwings  more  slender  at  the  base,  but  rounded  at  apex;, 
both  pairs  of  wings  thickly  covered  with  a  network  of  fine 
nervures  fringed  with  fine  buff-brown  or  Avhite  hairs,  so  that  the 
whole  of  the  wing  surface  is  covered  with  downy  hairs,  and  thickly 
fringed  on  the  outer  margins.  Forewings  blotched  with  irregular 
red  and  brown  spots,  and  delicate  brown  and  black  lines  and 
wavy  markings;  hindwings  with  a  single  brown  blotch  in  the 
centre. 

Newman  described  the  type,  which  is  now  in  the  British 
Museum,  in  three  lines,  but  he  gave  a  Avoodcut  of  the  insect  on 
the  title-page  of  the  volume.  His  specimen  was  received  from 
Adelaide,  S.A.  Though  not  a  common  insect,  it  has  a  wide 
range  over  the  eastern  and  southern  portions  of  Australia. 

In  the  middle  of  December  a  living  female  was  sent  to  me  b}^ 
a  correspondent  at  Muswellbrook,  which  while  in  transit  laid  three 
bright  green  oval  eggs.  These  were  placed  in  a  glass  jar,  and 
three  weeks  later  two  hatched  out  and  the  larvse  were  found 
crawling  about  trying  to  escape.  When  placed  in  a  watch  glass- 
with  some  larval  psyllids  and  aphides,  they  immediately  seized 
these  with  their  long  slender  mandibles  and  soon  sucked  them 


368  AUSTRALIAN  NEUROPTERA, 

dry,  the  fluid  running  up  the  hollow  jaws  being  easily  observable 
with  a  lens.  In  spite  of  every  care  both  larvae  died  within  a 
few  weeks. 

The  following  description  of  the  larva  is  based  upon  very 
immature  material: — Head  broad,  oval,  rounded  behind;  man- 
dibles long,  slender,  curved  inwards;  palpi  3-jointed,  long,  slender, 
reaching  to  the  middle  of  the  mandibles.  Antennae  springing 
out  from  centre  of  the  head,  composed  of  11  slender  irregular 
joints,  with  a  bristle  at  apex.  Eyes  small,  ocelli  not  apparent. 
Prothorax  narrow,  overlapping  head;  legs  long,  slender,  with  the 
tarsal  claw  long,  furnished  with  a  long  white  hair  at  base  of  tarsi. 
Dorsal  surface  covered  with  small  browm  tubercules  and  short 
white  hairs,  those  upon  the  abdomen  forming  two  transverse  rows 
on  each  segment. 

PSYCIIOPSIS   CCELIVAGUS,  Walk. 

Hemerohius  cailivagus,^dA\:.^  Brit.  Mus.  Cat.  Neurop.  279,  1853. 

This  is  a  lighter-coloured  species  than  the  preceding,  and 
smaller. 

Length  3  J;  expanse  of  wings  15  lines. 

A  number  of  specimens  of  a  dull-coloured  species  are  in  the 
Macleay  Museum,  and  these,  Mr.  Masters  informs  nje,  were  at 
one  time  taken  in  the  gardens  about  Sydne}^  They  may  be 
referable  to  this  species. 

PsYciioPSis  INSOLENS,  McLachl. 

McLachlan,  Journ.  Ent.  ii.  114,  pi.  vi.,  fig.  3,  1863. 

This  is  a  common  species  in  Queensland;  there  are  a  number  of 
specimens  of  it  in  the  British  Museum.  The  type  is  in  the 
British  Museum,  and  was  collected  by  the  late  Mr.  Diggles  at 
More  ton  Bay,  Q.  It  is  somewhat  smaller  than  P.  mimica,  and 
differs  from  it  in  being  much  lighter-coloured,  without  the  rich 
colouration  and  tints  on  the  wino:s. 


( 


BY    WALTER    W.    FROGGATT.  369 

Chrysopa  ramburii,  Schneider. 

This  is  the  typical  form  of  this  extensive  genus.  It  is  found 
in  tlie  neighbourhood  of  Sydney,  and,  from  an  economic  point  of 
view,  is  one  of  our  most  useful  insect  friends,  for  wherever  mealy 
bugs  and  other  soft-bodied  scale-insects  appear  in  any  quantity, 
the  active  little  larvaj  of  the  Chrysopa  follow  them  up,  covering 
themselves  with  bits  of  dirt  and  the  remains  of  their  food,  so  that 
they  might  easily  be  mistaken  for  mealy  bugs  were  it  not  for 
their  projecting  jaws,  long  legs,  and  active  habits.  When  full- 
grown  they  form  the  usual,  rounded,  parchment-like  pupa-cases, 
which  are  attached  to  branches  among  the  living  mealy  bugs. 

The  perfect  insects  are  of  a  delicate  green  tint,  with  bright 
golden-tinted  eyes,  and  delicate  gauze-like  wings.  The}'-  often  fly 
to  light  on  summer  nights,  and  if  they  alight  upon  the  table 
produce  a  most  disagreeable  smell. 


SOME  RECORDS  OF  NEW  SOUTH  WALES  MOSSES. 

By  W.  Forsyth. 

[Title]. 


CENSUS  MUSCORUM  AUSTRALIENSIUM. 

A  CLASSIFIED  Catalogue  of  the  Frondose  Mosses  of  Australia 
AND  Tasmania,  collated  from  available  Publications  and 
Herbaria  Records,  by  the  Rev.  W.  Walter  Watts  and 
Thomas  Whitelegge,  F.R.M.S. 

Part  I. 

Issued  separately  as  a  Supplement  to  this  Part  of  the  Proceedings. 


370 


NOTES    AND    EXHIBITS. 


Mr.  D.  G.  Stead  exhibited  life-sized  photographs  of  a  very  large 
Black  Bream  {Chrysophrys  anstralU)  showing  the  fish  in  profile 
and  in  face- view.  The  fish  weighed  4  lbs.  14  ozs.,  and  was  of  the 
following  dimensions  : — total  length  19|,  height  of  bod}'  7,  thick- 
ness 2f,  girth  15|  inches. 

Mr.  Waterhouse  exhibited  some  noteworthy  specimens  of  Ehopa- 
locera  as  follows  : — Abisara  segecia,  Hew.  ((J9),  from  Cape  York; 
an  Erycinid,  new  to  Australia,  but  a  well-known  New  Guinea 
form;  Holochila  marqarita,  Semper  ((j9),from  Cape  York,  which 
has  passed  unnoticed  since  its  description;  Arliopala  wildei,  Misk. 
((^),  from  Cairns;  and  Cyaniris  tenelia,  Misk.  (J 9),  from  Cairns, 
hitherto  placed  in  the  genus  Lyccena. 

Mr.  Froggatt  showed  a  fine  series  of  neuropterous  insects  and 
their  larvae  in  illustration  of  his  paper. 

Mr.  A.  J.  Walkom  brought  under  notice  a  specimen  of  an 
undetermined  species  of  Pentamerus  from  Molong,  N.S.W., 
apparently  different  from  the  Australian  species  at  present 
recorded.  Additional  specimens  are  desirable,  and  to  that  end 
attention  is  called  to  it. 

Mr.  R.  Greig  Smith  exhibited  microscopic  preparations  of 
diseased  rainbow-trout. 

Mr.  Maiden  exhibited  the  Lord  Howe  plants  refei-red  to  in  his 
paper.  Also  a  number  of  Antarctic  plants  collected  by  Hooker 
filius  between  the  years  1839-43  when  naturalist  in  those  regions 
in  the  '-'Erebus"  and  "Terror"  Expedition.  The  sjDecimens 
were  presented  to  the  National  Herbarium,  Sydney,  by  the  Pro- 
fessors of  Botany  at  Glasgow  and  Edinburgh. 

Mr.  Fletcher  exhibited  a  few  coins — the  remnant  of  a  once 
much  larger  collection,  formed  by  the  late  Mr.  Syms  Co^  ington 
■during  the  cruise  of  H.M.S.  "  Beagle  "  (1832-36).  These  were 
kindly  forwarded  by  Mr.  Syms  Covington  of  Pambula,  per  favour 


NOTES    AND    EXHIBITS.  371 

of  Mrs.  C.  W.  Morgan  also  of  Pambula,  to  be  added  to  the  relics 
of  the  Voyage  of  the  "  Beagle  "  exhibited  at  the  last  Meeting  of 
the  Society. 

Mr.  Cheel  showed  a  small  but  very  choice  collection  of  about 
twenty  species  of  foliaceous  lichens  from  the  North  Island  of  New 
Zealand,  brought  together  by  Mr.  W.  Gardner,  of  Penshurst, 
during  a  recent  visit,  as  follows  : — 

Order  1.  Collemaceae. 

Tribe  Ephebe.e. 
Ephehe  pubescens,  Fr. — Waiora  Valley,  Wairakei. 

Order  ii.  Discocarpese. 

Series  i.   Diploblaste^. 
Tribe  Cladonie^e. 

Cladina  pycnoclada,  Pers. — Waiora  Valley,  Wairakei.  Speci- 
mens are  in  the  National  Herbarium,  Sydney,  labelled  "  New 
Zealand  :  collected  by  A.  F.  Wilson,  May  1887." 

Clathrina  retipora  (Labill.),  Wain. — Waiora  Valley,  Wai- 
rakei. Specimens  are  in  the  National  Herbarium,  Sydney, 
labelled  "Lake  Manoporire  :  collected  June  1902,  and  presented 
by  the  Hon.  J.  Hawkins  Smith,  M.L.C." 

Series    ii.    THAMNOPHYLLOBLASTEiE. 

Tribe  Usnee^. 
Usnea  harhata  (L.),  var.  hirta  (L.),  Fr. — Okoroire,  Auckland. 
U.  harhata,  var.  dasypoga,  Fr. — Tikitere  Lake,  Rotorua;  and 
Rotoiti  Lake,  Rotorua. 

U.  harhata,  var.  ceratina,  Ach. — Tikitere  Lake,  Rotorua. 
U.  cormita,  Flot. — Tikitere  Lake,  Rotorua;  and  Okoroire. 
U.  trichodea,  Ach. — Okoroire,  Auckland. 
U.  longissima,  Ach. — Rotoiti  Lake,  Rotorua;  and  Okoroire. 

Tribe  Ramaline^. 
Ramalina  leiodea,  N}^. — Rotoiti  Lake,  Rotorua;  Tikitere  Lake, 
Rotorua;  Arateatea  Rapids,  Wairakei;  and  Okoroire,  Auckland. 


372  NOTES    AND    EXHIBITS. 


Tribe  Sticte^e. 


Sticta  Freycineiii,  Del. — Arateatea  Rapids,  Wairakei. 

S.  Billardierii,  Del. — Arateatea  Rapids,  Wairakei.  Specimens 
are  in  the  National  Herbarium,  Sydney,  labelled  "  New  Zealand  : 
collected  by  Messrs.  J.  Gillespie  and  R.  J.  Potter." 

,S'.  Richardii,  Mont. — Rotoiti  Lake,  Rotorua. 

S.  rubella,  Hook,  et  Tayl. — Arateatea  Rapids,  Wairakei. 

S.  orygiiuea.  Ach. — Arateatea  Rapids,  Wairakei.  Specimens 
are  in  the  National  Herbarium,  Sydney,  labelled  " South  Dunedin 
(Mr.  S.  Jamieson);  New  Zealand  (Mrs.  Martin;  August  1890)." 

Tribe  Parmelie^. 

Parmelia perlata,  Ach. — Rotoiti  Lake,  Rotorua;  Waiora  Valley, 
Wairakei;  Okoroire,  Auckland. 

P.  perlata,  var.  cetrarioide>i,  Del. — Okoroire,  Auckland. 

P.  perfo7'ata,  Wulf. — Waiora  Valley,  Wairakei;  Hamuraua 
River,  Rotorua. 

P.  phi/sodes  (L.),  Ach. — Rotoiti  Lake,  Rotorua. 

P.  physodes,  var.  mundata,  Nyl. — Waiora  Valley,  Wairakei. 

P.  perlusa,  Schrank. — Waiora  Valley,  Wairakei. 

Tribe  Physce^e. 

IVielochistes  chrysophthalma  (Ij.),  Th.  Fr.  (Syn.  Physciachrysoph- 
thcdmctj  DC. — Okoroire,  Auckland. 


WEDNESDAY,  SEPTEMBER  24th,   1902. 

The  Ordinary  Monthly  Meeting  of  the  Society  was  held  in  the 
Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening,  September  24th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S.,  etc.,  President,  in  the  Chair. 

The  Donations  and  Exchanges  for  the  month,  amounting  to  18 
Vols.,  35  Parts  or  Nos.,  1  Bulletin,  3  Annual  Reports  (Admini- 
strative), 7  Pamphlets,  and  5  Miscellanea,  received  from  41 
Societies,  ikc,  and  2  Individuals,  were  laid  upon  the  table. 


373 


AUSTRALIAN  FUNGI,  NEW  OR  UNRECORDED. 

DECADES  I.-II. 

By  D.  McAlpine,  Corresponding  Member. 

Although  a  considerable  amount  of  attention  has  been  given 
to  the  Fungus-flora  of  Australia,  a  large  number  of  species  still 
remain  to  be  described,  at  least  among  the  micro-fungi. 

These  are  either  new  to  science  or  have  not  hitherto  been 
recorded  for  Australia,  and  since  numbers  of  such  are  continually- 
coming  under  my  notice,  it  is  desirable  to  render  the  descriptions 
available  for  other  workers. 

The  introduction  and  cultivation  of  economic  plants  is  no 
doubt  responsible  for  many  of  these,  but  the  native  vegetation  is 
likewise  rich  in  new  forms,  and  it  will  be  my  aim  to  record  the 
indigenous  species  as  well  as  those  which  are  already  known.  In 
these  two  decades  eleven  new  species  and  twelve  different  genera 
are  represented. 

1.    AnTHOSTOMELLA    DANTHONIiE,  n.Sp. 

Minute,  black,  gregarious  pustules  on  leaves  and  leaf-sheaths. 
Perithecia  covered,  densely  crowded  and  running  in  lines,  opening 
at  surface  by  pore,  surrounded  by  pale  ruddy  tissue,  and  occupy- 
ing entire  depth  of  leaf,  always  much  compressed.  Asci  cylindrical 
or  bulging,  slightly  pointed  and  rounded  at  apex,  tapering  into 
short  stalk  at  base,  8-spored,  140-150  x  19-20  /u,  but  may  reach  a 
breadth  of  28  [x  when  sporidia  are  partially  two-rowed.  Sporidia 
monostichous,  occasionally  distichous,  dirty  brown,  very  thick- 
walled,  elliptical,  20-26  x  15-17  n;  paraphyses  hyaline  individually 
but  pink  in  the  mass,  longer  than  asci,  very  slender  (0-5-1  /u)  with 
highly  refractive  granular  contents. 
25 


374  AUSTRALIAN    FUNGI, 

Ardmona,  Vic.  (Robinson,  284);  very  common,  on  both  surfaces 
of  the  leaf-blades  but  mostly  on  outer  or  lower,  and  sheaths  of 
Danthonia  penicillata,  F.v.M.,  Sept.  1899,  and  May,  1902. 

This  is  a  true  and  destructive  parasite,  causing  the  leaves  to 
become  dried  up  and  brittle.  The  black  elongated  pustules 
arranged  in  lines  give  the  appearance  of  'black  rust'  and  the 
aeneral  resemblance  to  a  rust  is  heightened  by  the  occurrence  of 
ruddy  spots,  which  is  characteristic  at  the  commencement  of  the 
development  of  perithecia.  While  Z>.  penicillata  was  severely 
attacked,  other  species  adjacent  were  quite  free. 

2.  Camarosporium  dolichi,  n.sp. 

Perithecia  gregarious,  erumpent,  black,  dark  golden-brown  b}'- 
transmitted  light,  globose,  membranaceous,  with  rotund  papillate 
mouth.  Sporules  olivaceous,  oblong,  4-septate,  muriform,  with 
median  septum  very  distinct;   16  x  9  /x. 

Armadale,  near  Melbourne,  Vic;  on  leaflets  of  Dolichos  lablab, 
L.;  July,  1901. 

3.  Camarosporium  nigricans,  n.sp. 

Perithecia  very  minute,  globose,  black  and  golden-brown  by 
transmitted  light,  numerous,  membranaceous,  seated  superficially 
on  mycelial  threads,  no  apical  pore  observed,  50-57  /x  diam. 
Sporules  olivaceous  to  golden-brown,  elliptical,  generally  3-septate 
and  constricted  at  septa,  with  occasionally  1-2  longitudinal  septa, 
13-15  X  61 /x. 

Croydon,  Vic;  on  apple  twigs;  June,  1902. 

It  differs  from  C.  mali,  Ell.  &,  Ev.,  in  the  much  smaller 
perithecia,  and  in  the  sporules  not  being  5-8  septate;  the  specific 
name  is  indicative  of  the  blackening  of  the  twigs  by  the  fungus. 

4.  Capnodium  (?)  CASUARiN^,  n.sp. 

Forming  black,  soot-like  incrustations  all  over  branches  and 
branchlets,  not  readily  separating.  Hyphse  creeping,  septate, 
branching,  fuliginous,  firm,  thick-walled.  Cerato-pycnidia  yellow- 
ish-green  to   dirty   green,    very   elongated,    irregularly    shaped. 


BY   D.  McALPINE.  375 

variously  branched,  curved,  enveloped  more  or  less  by  brown 
h3"phae,  may  be  swollen  towards  base  and  taper  gradually  towards 
apex,  or  may  be  of  equal  diameter  throughout,  opening  by  minute 
circular  mouth.     Pycnospores  minute,  oval,  hyaline,  3  x  1^/*. 

Richmond  River,  N.S.W.  (Baker);  on  branches  and  branchlets 
of  Casuari7ia  torulosa,  Ait.;  August,  1897. 

5.  Leptosph.eria  cann.e,  n.sp. 

Perithecia  scattered,  minute,  black,  erumpent,  depressed- 
globose,  membranaceous  to  coriaceous,  with  distinct  apical  pore, 
100-120  n  diam.  Asci  elongated-clavate,  attenuated  at  base, 
8-spored,  90  x  20-2 1 J /z;  paraphyses  as  long  as  or  longer  than  asci, 
numerous,  filiform.  Sporidia  distichous,  olivaceous,  fusoid,  obtuse 
at  each  end,  with  granular  contents,  3-4  septate,  usually  slightly 
curved  and  constricted  at  septa,  18  x  6J-8  /a. 

Armadale,  near  Melbourne,  Vic. ;  on  upper  surface  of  pale 
brown  withered  patches  of  leaves  of  Carina  indica,  L.;  July,  1902. 

It  differs  from  L.  musarum,  Sacc.  &  Berl.,  in  the  perithecia 
being  scattered,  the  longer  and  broader  asci,  and  the  distinct  and 
numerous  paraphyses.  The  leaves  of  Canna  are  often  much  torn 
by  the  wind,  and  large  patches  become  brown  and  withered,  so 
that  the  diseased  appearance  may  not  be  caused  by  the  fungus 
which  is  comparatively  rare.  It  was  associated  with  Pleosjyora 
herharum,  Pers. 

6.  Macrosporium  internUxM,  n.sp. 

Forming  variously  coloured,  velvety  patches  covering  the  pulp 
inside,  while  the  skin  of  the  Orange  is  healthy-looking.  M3^celium 
consisting  of  dense  masses  of  hyphse,  dark  green  collectively,  but 
passing  through  colourless  and  yellowish-green  phases.  Hyphse 
generally  closely  septate,  copiously  branched,  and  branches  often 
at  right  angles,  i\-'6\  jx  broad.  Conidia  amber-coloured  at  first, 
then  olivaceous,  stalked,  clavate,  5-8  transverse  septa,  and  usually 
more  or  less  constricted  at  septa,  with  longitudinal  septa  either 
single  or  double  in  each  segment,  variable  in  size,  37-75  x  17-20 /x. 

Sydney,  N.S.W.  C.  French,  Junr.);  on  Mandarin  Oranges; 
Sept.,  1900. 


376  AUSTRALIAN    FUNGI, 

This  is  evident!}^  a  serious  disease  of  the  Orange,  rotting  the 
entire  contents,  while  the  skin  looks  quite  healthy  at  first,  although 
ultimately  becoming  discoloured.  Mr.  French  observed  that  75% 
of  the  Oranges  examined  were  black  inside  (Internal  Black  Rot), 
while  the  outside  appearance  of  the  fruit  was  perfect.  There 
was  a  considerable  amount  of  'Scale'  on  the  skin,  which  undoubt- 
edly rendered  the  fruit  an  easy  prey  to  the  fungus.  This  mould 
varies  considerably  in  its  colour.  It  is  snow-white  at  first,  then 
greenish  and  blackish,  and  the  conidia  form  a  yellowish-green 
layer  at  first,  then  dark  olivaceous. 

7.    METASPH.ERIA    LEPIDOSPERMiE,  U.Sp. 

Perithecia  in  lines  close  together  on  stem,  often  on  inky-black 
patches,  punctiform,  minute,  black,  membranaceous,  with  pore, 
erumpent  through  stomata,  72  ^  diam.  Asci  very  delicate  and 
rupturing  as  soon  as  ripe,  cylindric-clavate,  rounded  and  thickened 
at  apex,  narrowing  towards  base,  and  somewhat  pointed  at  the 
end,  86-90  x  13-1 5  ju;  paraphyses  slender,  as  long  as  asci.  Sporidia 
biseriate,  elongated-fusoid  to  elongated-clavate,  slightly  curved, 
typically  3-septate,  and  medium  septum  most  distinct,  not  con- 
stricted at  septa,  at  first  hyaline  then  honey -3^ellow,  25-26  J  x  6  J-7/x. 

Werribee,  Yic.  (C.  French,  Junr.);  on  Lepidosperma  sp.;  May, 
1902. 

The  stems  were  conspicuously  discoloured  with  inky-black 
patches,  which  were  isolated  or  run  together,  and  either  partial^ 
or  entirely  surrounding  the  stem.  The  minute  perithecia  running 
in  lines  are  just  visible  to  the  naked  eye,  and  while  they  frequentl}^ 
occur  on  the  discoloured  stem  they  are  also  found  on  the  black 
patches.  Hundreds  of  plants  are  attacked  and  dying;  even  the 
young  green  shoots  are  affected,  and  die  off  before  reaching 
maturity. 

8.  Phoma  berberina,  Sacc.  (fc  Roum. 

Sporules  fusoid,  hyaline,  6-7  x  2-2J  /x. 

Malvern,  near  Melbourne,  Yic;  on  withered  portions  of  leaf  of 
Berherls  vulgaris,  L.;  Jul}',  1902. 


BY   D.  McALPINE.  377 

This  species  has  only  hitherto  been  recorded  on  twigs  of  the 
Barberry  in  France. 

9.  Phoma  strobiligexa,  Desm.,  var.  microspora,  Sacc. 

Near  Melbourne,  Vic;  on  scales  of  cones  of  Pinus  insignisy 
DougL;  July,  1902. 

10.  Phoma  PASSiFLORiE,  n.sp. 

Perithecia  numerous,  gregarious,  erumpent,  dark  brown,  seated 
near  the  margin  of  tawny  sunken  spots,  150-200  ^  diam.  Sporules 
broadly  elliptical,  rounded  at  both  ends,  hyaline  to  subhyaline, 
with  finely  granular  contents,  12-16  x  6-7  fx. 

New  South  Wales;  on  fruit  of  Passiflora  edulis,  Sims. 

11.  Phyllosticta  asclepiadearum,  AVest, 

Malvern,  Vic;  on  upper  surface  of  leaves  of  Hoi/a  carnosa, 
KBr.;  March,  1900. 

It  presents  similar  milk-white  blotches  as  in  Pleospora  asclepi- 
adearum. 

12.  Phyllosticta  coPROSMiE,  n.sp. 

Spots  greyish,  with  distinct,  narrow  dark  red  raised  margin, 
surrounded  on  both  sides  of  leaf  by  indefinite  ruddy  zone,  gradually 
becoming  paler  towards  centre  as  well  as  outside.  Perithecia 
gregarious,  on  upper  surface,  erumpent.  globose,  ruddy  brown  by 
transmitted  light,  with  slightly  papillate  mouth,  250-330  /^  diam. 
Sporules  pale  pink  in  mass,  individually  hyaline,  elliptic,  rounded 
at  both  ends,  6  x  2-2J  /x. 

Sunbury,  Vic  (C.  French,  Junr.);  on  leaves  of  Coprosma  sp.; 
July,  1901. 

13.  Phyllosticta  ERioBOXRYiE,  Thuem. 

Armadale,  Vic;  on  green  leaves  of  Loquat;  April,  1900. 

The  dry  greyish  brown  spots  extend  from  the  midrib  towards 
the  margin  between  the  lateral  veins,  and  cause  considerable 
damage  to  the  otherwise  green  leaf. 


378  AUSTRALIAN    FUNGI, 

14.    PflYLLOSTICTA    GROSSULARIiE,  SaCC. 

Armadale,  Vic;  on  Gooseberry  leaves;  Jan.,  1899. 
Towards  the  margin  of  the  leaves  large  areas  become  blotched 
by  the  spots  running  together,  and  the  leaves  soon  fall. 

15.  Phyllosticta  myrticola,  Speg. 

Orbost,  Vic.  (Pescott);  on  living  leaves  of  Eugenia  smithii, 
Poir.;  July,  1900. 

This  species  has  only  hitherto  been  recorded  on  the  living  leaves 
of  a  species  of  Eugenia  in  Brazil,  and  is  new  to  Australia. 

16.  Pleospora  asclepiadearum,  n.sp. 

Hyphfe  adjoining  perithecia  sooty-brown,  septate,  branched, 
twisting  and  anastomosing,  up  to  7-9  fx  broad.  Perithecia  some- 
what gregarious,  at  first  covered  by  epidermis,  then  partially 
naked,  minute,  black,  punctiform,  globose,  ostiolum  minute, 
190-225  fi  diam.  Asci  cylindrical-oblong,  rounded  at  apex, 
tapering  into  a  slightly  dilated  knob  at  point  of  attachment,  about 
100  X  17-19 /x:  paraphyses  hyaline,  about  length  of  asci,  slightly 
swollen  at  apex  (H  /x  broad),  stained  pale  red  by  potassium 
iodide-iodine.  Sporidia  distichous  usually  at  base  and  monosti- 
chous  at  apex,  dark  olive-green,  ellipsoid,  5-septate,  constricted 
at  septa,  generally  with  four  longitudinal  septa,  and  occasionally 
slightly  muriform,  28-30  x  10-12  ^. 

South  Australia  (Molineux);  on  leaves  of  Hoya  carnosa,  R.Br.; 
July,  1898. 

This  disease  has  a  very  characteristic  appearance.  The  upper 
surface  of  the  leaves  has  a  large  milk-white  blotch,  sometimes 
occupying  fully  one-half  the  entire  surface. 

F.  diiylosjwra,  Ell.  &  Ev.,  found  on  dead  stems  of  Asclejnas 
incarnata,  L.,  in  America,  has  much  broader  asci,  and  the  straw- 
coloured  to  brown  sporidia  may  be  7-septate. 

The  Phyllosticta  asclepiadearum^  West.,  is  probably  an  early 
stage  of  this  species,  and,  therefore,  I  have  given  it  the  same 
specific  name. 


BY  D.  McALPINE.  379 

17.  Ramularia  hordei,  n.sp. 

Forming  whitish  patches  not  very  conspicuous  on  brownish 
lenticular  spots,  which  may  run  together  and  become  dark  brown. 
Hyphse  ramifying  in  leaf,  septate,  branched,  and  bearing  at  surface 
the  conidia.  Conidia  straight  or  fusoid,  hyaline,  acute  or  blunt 
at  ends,  1-septate  when  mature,  not  constricted  at  septum, 
15-17x3^. 

New  South  Wales  (Farrer);  on  leaves  of  Barley;  Oct.,  1900. 

There  is  only  one  species  of  Ramularia  recorded  on  Graminese, 
viz.,  R.  graminiGola^  Peck,  on  living  leaves  of  Poa  serotiita,  Ehrh., 
but  the  conidia  are  25-35  /x  long,  and  spuriously  1-septate,  whereas 
in  this  species  they  are  decidedly  1-septate. 

18.  Ramularia  PRiMULiE,  Thuem. 

Armadale,  Vic;  on  leaves  of  Primula  sinensis^  Sabine;  July, 
1902. 

It  forms  large  pallid  areas,  with  more  or  less  orbicular  centres. 
The  spots  on  the  Primrose  leaves  are  very  common  in  the  winter 
months,  but  the  fructification  of  the  fungus  is  rarely  obtainable 
except  from  perfectly  fresh  specimens. 

19.  Septoria  STBLLARiiE,  Rob.  &  Desm. 

Armadale,  near  Melbourne,  Vic.  ;  on  languishing  leaves  of 
Stellaria  mediay  Cyrill;  July,  1900. 

The  spots  were  distinct  at  first,  and  then  became  confluent. 

20.  Valsa  ambiens  (Pers.),  Fr. 

Perithecia  usually  5-6  imbedded  in  stroma.  Spermogonial 
stage  [Cytospora  carphosperma)  with  sporules  escaping  in  tendrils 
of  a  whitish  to  yellowish  colour. 

Mordialloc,  Vic;  on  dead  bark  of  Apple;  July,  1901. 


380 


ON  A  NEW  SPECIES  OF  ARDISIA  FROM  NEW  SOUTH 

WALES. 

By  R.  T.  Baker,  F.L.S.,  Curator,  Technological  Museum, 

Sydney. 

(Plate  xvi.) 

Ardisia  racemosa,  sp.nov. 

A  handsome,  glabrous  tree  about  20  to  30  feet  high,  and  6  to 
9  inches  in  diameter,  with  a  smooth  grey  bark.  It  is  found  in 
deep,  rich  alluvial  gullies,  generally  in  groves  or  clusters  similar  to 
Drimys  dipetala  and  D.  aromatica.  Branchlets  terete.  Leaves 
measuring  up  to  5  inches  long  and  1  inch  wide,  ovate-elliptical, 
acuminate,  thin,  almost  membranous,  pale-coloured  on  the 
underside,  often  becoming  channelled  near  the  midrib  when 
dried;  edges  very  slightly  crenate,  and  in  parts  are  sometimes 
undulate.  Lateral  veins  iine,  oblique.  Oil  glands  numerous, 
fairly  distinct.  Flowers  in  short,  axillary,  occasionally  lateral 
loose  racemes;  pedicels  filiform,  about  4  lines  long.  Sepals  5, 
valvate,  about  J  line  long,  acuminate,  marked  (except  the  edges), 
with  dark  purple  spots,  glabrous,  edges  scarious.  Petals  5, 
about  1^  lines  long,  imbricated  and  contorted  in  the  bud, 
acuminate,  glabrous,  spotted  similarly  to  the  sepals,  edges  light- 
coloured  or  not  marked,  the  lower  half  of  the  inner  surface 
covered  with  dense  ferruginous  hairs.  Filaments  exceedingly 
short.  Stamens  opposite  the  petals.  Anthers  glabrous,  sessile 
or  nearly  so,  cordate-sagittate,  with  subulate  recurved  points; 
valves  opening  by  longitudinal  slits  from  apex  to  base.  Style 
longer  than  the  stamens,  subulate.     Fruits  not  seen. 

Timber  light-coloured,  close-grained,  moderately  hard,  but  on 
account  of  its  size  could  probably  only  be  used  for  turning,  tool 
handles,  &c. 


BY    R.    T.    BAKER.  381 

Hah. — Tumbulgum,  on  the  Tweed  River  (W.  Bauerlen);  in  deep, 
rich  gullies. 

This  tree  was  discovered  by  Mr.  Bauerlen  in  October,  1897,  at 
the  above-mentioned  locality,  where  it  is  associated  with  other 
plants  only  recently  added  to  the  known  flora  of  New  South 
Wales,  such  as  Freycinetia  excelsa,  F.v.M.,  Petermannia  cirrosa, 
F.V.M.,  Tecoma  Baileyana,  Maiden  and  Baker,  and  other  species. 

It  differs  principally  in  the  mode  of  inflorescence  from  the  three 
species  of  Ardisla  described  from  Australia.  A.  pseudo-jambosa, 
F.v.M.,  has  comparatively  large  terminal  panicles,  often  over  six 
inches  long,  and  with  an  induplicate-valvate  corolla;  small,  shining 
sepals  and  petals,  and  well  exserted  anthers — characters  that  do 
not  apply  to  this  species. 

A.  brevipedata,  F.v.M.,  differs  from  it  in  having  flowers  arranged 
in  umbels  on  very  short  peduncles,  and  in  having  numerous 
flowers,  with  imbricate,  broad  and  short  sepals,  and  imbricate 
petals. 

A.  2>ci'ChyrrhachiSy  F.v.M.,  differs  from  it  in  its  thick  rhachis, 
more  numerous  flowers  (up  to  40)  in  a  fascicle,  larger  leaves  and 
flowers. 

Its  nearest  affinity  is  A.  brevipedata,  and  in  botanical  sequence 
it  might  be  placed  between  that  species  and  pseudo-jambosa, 
F.v.M. 

The  leaves  of  all  four  species  are  very  similar  in  texture  and 
shape,  and  have  minute,  crenulate  and  repandulate  edges,  and 
all  dry  with  a  wrinkled  surface. 

The  name  A.  repandula,  F.v.M.  (Frag.  iv.  82),  was  attached  to 
leaf  specimens  only,  and  was  thought  by  Bentham  (B.Fl.  iv.  p  276) 
to  be  identical  with  A.  brevipedata,  F.v.M.  No  specimen  of  this 
species  now  appears  to  be  available. 

Baron  von  Mueller,  in  his  'Papuan  Plants,'  describes  two 
Ardisias,  A.  solanacea  var.  haplosciadea,  from  a  plant  discovered 
by  W.  Bauerlen  on  the  Strickland  River  in  1885,  and  A.  poran- 
thera  from  a  New  Guinea  plant  cultivated  in  the  Sydney  Botanic 
Gardens. 


382  A  NEW  SPECIES  OF  ARDISIA. 

In  the  '  Flora  von  Kaiser  Wilhelms  Land,'  by  K.  Schuhmann, 
A.  imperialis,  collected  by  M.  Hollrimg,  n.  256,  is  recorded.  It 
is  described  (Eng.  Jahrb.  ix.  213)  as  a  tree  about  9  to  18  feet 
high,  and  with  deep  rose-coloured  flowers. 

These  species  are  all  distinct  from  the  one  now  described  from 
New  South  Wales. 

I  beg  to  thank  Mr.  J.  G.  Luehmann,  F.L.S.,  Curator,  National 
Herbarium,  Victoria,  for  the  use  of  his  herbarium,  and  also  Mr. 
Bauerlen  for  assistance  in  working  out  this  species. 


EXPLANATION  OF  PLATE  XVL 

Fig.  1. — Twig  with  leaves  and  inflorescence. 

Fig.  2.— Bud. 

Pig.  3. — Expanded  flower  showing  disposition  of  anthers. 

Fig.  4.— Expanded  flower  showing  calyx. 

Fig.  5. — Anther  showing  lateral  opening  of  cell. 

Fig.  6. — Anther,  front  view. 

(All  the  figures  except  No.  1  enlarged. ) 


383 


THE  BACTERIAL  ORIGIN  OF  THE  GUMS  OF  THE 
ARABIN  GROUP. 

By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the 

Society. 

I. — The  Soluble  (Arabin)  Wattle  Gums. 
{Bacterium  acacice,  n.sp.) 

While  working  upon  the  gums  and  slimes  produced  by  some 
bacteria,  it  seemed  to  be  exceedingly  probable  that  a  few  of  the 
gums  which  occur  naturally  or  are  supposed  to  be  formed  during 
a  pathological  condition  of  the  plant,  might  have  a  bacterial 
origin.  Like  the  mucilages,  gums  and  slimes  formed  in  or  on  the 
higher  plants,  the  bacterial  slimes  are  of  a  varied  nature.  For 
example,  dextran  may  be  considered  as  a  dextrose  anhydride, 
levan  as  a  levulose  anhj^dride,  the  gums  of  Schardinger  and 
Adametz  as  galactose  anhydrides ;  the  bacterium  of  Marshall 
Ward  and  Reynolds  Green  produces  a  hemi-cellulose,  and  Brown's 
Bad.  xylinuni  has  a  cellulose  en\  elope.  The  bacterium  which  I 
separated  from  the  sugar-cane  forms  a  pentosan  slime. 

Perhaps  the  most  valuable  of  the  vegetable  gums  are  those  of 
the  arabinan-galactan  class  such  as  gum  arabic  and  wattle  gum; 
and  when  one  studies  the  distribution  of  this  kind,  it  appears  to  be 
quite  within  the  bounds  of  possibility  that  bacteria  have  more  to 
do  with  its  formation  than  would  at  first  appear.  Indeed  it  is 
extremely  probable.  In  the  first  place,  the  gum  exudes  from 
cracks*  or   from  punctures   or   wounds   made   by   insects      The 

*  "  Wattle  gum  exudes  chiefly  during  the  summer  season  from  fissures 
and  accidental  injuries  to  the  bark.  .  After  careful  observation  I  have  formed 
the  opinion  that,  as  a  very  general  rule,  it  is  a  pathological  product.  I  came 
to  this  conclusion  long  before  I  was  aware  of  Trecul's  observations  that 
Acacias  and  the  Kosacese  yield  their  gums  most  abundantly  when  sickly  and 
in  an  abnormal  state  caused  by  a  fulness  of  sap  in  the  young  tissues." — 
Maiden,  Pharm.  Jour.  [3]  xx.  (1890),  869. 


384  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

infection  of  the  plant  from  without  is  thus  indicated.  In  the 
second  place,  every  tree  does  not  produce  gum.  This  is,  I  think, 
the  strongest  argument  in  favour  of  the  mycological  origin  of  the 
substance.  If  gum  acacia  were  a  natural  or  even  pathological 
product  of  the  plant  itself,  one  would  expect  to  find  it  more 
uniforml3^  distributed  than  it  is.  It  is  not  always  even  uniformly 
distributed  over  the  tree;  some  branches  may  be  exuding  gum 
while  others  are  not.  The  localised  positions  of  gum-bearing 
Acaciae  are  in  accord  with  the  theory  that  gum  results  from  the 
action  of  agents,  such  as  bacteria,  introduced  by  insects  into  the 
tissues  of  the  plant  or  by  wind-borne  dust,  laden  with  bacteria, 
lodging  in  a  crack  or  wound. 

A  recently  gathered  specimen  of  wattle  gum,  which  I  obtained 
from  Mr.  J.  H.  Maiden,  Government  Botanist,  showed,  when 
examined  microscopically,  a  number  of  granules  which  might 
have  been  the  plasmolysed  remains  of  bacteria.  These  were  not 
evident  in  older  gums. 

Of  interest  also  in  this  connection  is  the  circumstance  that 
wattle  gum  has  nutritive  properties,*  and  that  O'Sullivan  found 
a  proteid  in  samples  of  Gedda  gum.  It  is  not  at  all  improbable 
that  this  proteid  substance  was  the  remains  of  bacterial  cells. 

It  would  be  useless  to  attempt  the  isolation  and  cultivation  of 
bacteria  from  fragments  of  dry  gum,  because  any  micro-organisms 
that  had  been  there  would,  during  the  process  of  drying,  have 
been  killed  plasmolytically.  Fresh  material  must  be  investigated, 
and  to  obtain  this  I  applied  to  Mr.  Maiden,  who  referred  me  to 
Prof.  Liversidge,  in  whose  paddock  at  Mittagong  he  had  seen  a 
specimen  of  Acacia  penninervis  bearing  <a  quantity  of  gum.  Prof. 
Liversidge  sent  me  a  gum-bearing  branch,  and  this  was  investi- 
gated. 

From  the  twigs  small  portions  containing  a  drop  of  gum  were 
cut;  from  these  the  gum  was  removed  and  the  part  sterilised  by 
rapidly  passing  it  through  a  bunsen  flame.     Smaller  parts  of  these 


Maiden,  Proc.  Roy.  Soc.  N.S.  Wales,  xxxv.,  171. 


BY    R.    GREIG    SMITH.  .   385 

portions  were  inserted  into  faintly  acid  saccharose-peptone  media"*^ 
and  incubated  at  30°.  From  the  tubes  of  media  that  became 
turbid,  tubes  of  saccharose-peptone  agar  were  infected  and  plates 
prepared  in  the  usual  manner.  The  media  infected  with  frag- 
ments of  heart-wood  were  sterile,  but  in  cases  where  the  bark 
had  been  taken,  two  kinds  of  colonies  developed.  One  of  these 
was  in  the  minority,  and  will  be  referred  to  again  in  dealing  with 
the  insoluble  gums.  The  prevailing  bacterium  was  stroked  upon 
saccharose  agar,  and  the  growth  was  found  to  contain  capsulated 
bacteria  together  with  slimy  bacterial  clumps.  A  bacterium  had 
therefore  been  isolated  which  promised  well  for  the  research. 

A  number  of  experimental  cultures  were  then  made  with  the 
object  of  determining  the  constituents  which  might  be  employed 
to  construct  a  prescription  for  the  manufacture  of  a  medium  that 
would  yield  a  maximum  amount  of  slime  and  at  the  same  time 
enable  one  to  trace  the  constituents  from  which  the  slime  is 
derived.  The  experiments,  however,  led  to  no  useful  result, 
partly  because  too  high  a  temperature  (30°)  was  employed  and 
partly  because  a  discrimination  was  not  made  between  the 
quantit}'^  of  bacteria  and  the  amount  of  slime.  The  question  of 
the  slime-forming  constituents  will  be  discussed  in  a  future  paper. 
On  a  peptone-saccharose  agar  medium,  however,  enough  slime 
was  obtained  to  enable  a  few  tests  to  be  made. 

The  slime,  which  adhered  to  the  surface  of  the  medium,  was 
moistened  with  water,  and  after  it  had  swelled  it  was  removed 
with  a  rubber  spade,  the  greatest  care  being  taken  to  avoid 
removing  portions  of  the  agar. 

The  slimy  emulsion  was  stirred  with  a  few  drops  of  dilute  hydro- 
chloric acid  and  precipitated  with  alcohol.  The  coagulum,  on 
treatment  with  water  and  then  alcohol,  produced  an  opalescent 
solution  from  which  the  slime  v/as  precipitated  by  potassium 
chloride.      The  watery    emulsion    was    found    to    be   free    from 

*''  An  improvement  upon  this  method  of  isolation  would  be  to  employ 
ordinary  gkicose  gelatine,  in  which  the  bacteria  grow  very  well.  Tubes  of 
molten  glucose  gelatine  should  be  infected  with  the  fragments  of  wood  and 
plates  formed  from  these  at  once  and  after  an  incubation  of  1,  2,  4,  8  and  24 
hours  at  .30^. 


386  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

saccharose  and  reducing  sugars,  and  was  used  for  the  test-tube 
reactions.  Flocculent  white  precipitates  were  obtained  with 
basic  and  ainmoniacal  lead  acetates.  Neutral  lead  acetate  gave 
no  reaction.  Neither  did  the  salts  of  silver  and  mercury.  The 
hydrates  of  calcium  and  barium  gave  no  precipitate.  Copper 
sulphate  followed  by  potassium  hydrate  gave  a  flocculent  pre- 
cipitate which  became  cohesive  on  boiling  and  did  not  darken  in 
colour.  When  tested  at  a  later  date  by  the  method  recommended 
by  Maben,*  borax  stiffened  the  mucilage,  while  basic  lead  acetate 
and  also  ferric  chloride  hardened  the  gum  acids  forming  cohesive 
masses.  Tannic  acid  had  no  effect  beyond  producing  an 
opalescence. 

The  natural  gum  of  Acacia  pe^minervis  is  not  entirely  soluble 
in  water;  the  major  portion  simply  swells  up  to  form  gelatinous 
lumps,  t     The  portion  that  does  dissolve  acts  towards  the  salts  of 

*  Maben,  Pharm.  Jour.  [3]  xx.,  719. 
t  ''Acacia  penniiierviA,  Sieb.,  '  B.  Fl.'  ii.,  362. 
A  "  blackwood  "  and  "  mountain  hickory."     Found  in  Tasmania,  Victoria, 
New  South  Wales  and  Queensland.     Sample  from  Quiedong,  near  Bombala, 
New  South  Wales. 

Arabin 70*3 

Metarabin 11  "42 

Moisture 1 6  "67 

Ash 0-66 

While  not  approaching  the  best  kinds  of  gum  arabic  in  lightness  of  colour, 
it  is  the  palest  and  cleanest  looking  of  the  coast  Wattle  gums  described  in 
this  paper.  Colour  pale  sherry  to  orange.  It  has  a  bright  fracture,  although 
a  sample  received  from  Brown's  Camp,  Delegate,  and  obtained  from  tarious 
trees  6  feet  2  inches,  6  feet  in  diameter  and  40  feet  to  60  feet  high,  has  a 
dull  fracture  like  A.  binervata.  It  is  interesting  to  note  that  in  the  Quiedong 
district  the  gum  was  obtained  in  abundance  from  shrubs  or  small  trees 
growing  in  low  ground,  while  in  the  Braidwood  district  no  gum  whatever 
could  be  found  on  trees  of  this  species,  where  it  grows  at  high  elevations,  and 
at  Delegate,  at  moderately  great  elevations,  only  very  small  quantities  of 
gum  could  be  obtained.  The  inference  is  that  this  species  yields  gum  plenti- 
fully from  shrubs  at  low  elevations,  but  little  or  none  from  trees  at  high 
ones.  The  same  thing  has  been  found  to  hold  good  to  some  extent  in  regard 
to  A.  dealhata.  These  observations  point  to  the  truth  of  the  oft-expressed 
surmise  that  gum  is  a  pathological  product  or  the  product  of  plants  in  uncon. 
genial  surroundings.  ' — Maiden,  Pharm.  -Jour.  [3]  xx.  (1890),  980-1. 


BY    R.    GREIG    SMITH.  387 

copper,  lead,  silver  and  mercury  and  to  the  hydrates  of  calcium 
and  barium  in  a  manner  precisely  similar  to  a  solution  of  com- 
mercial gum  arabic  and  to  the  bacterial  slime. 

The  next  step  was  the  testing  of  the  slime  for  pentosans.  A 
portion  was  distilled  with  hydrochloric  acid  of  sp.g.  1'06,  and  the 
distillate  on  treatment  with  a  hydrochloric  acid  solution  of 
phloroglucin  yielded  the  furfural  reaction.  Another  portion  was 
carefully  heated  with  strong  sulphuric  acid,  and  the  volatile 
products  gave  the  furfural  rose  colour  to  paper  moistened  with 
aniline  acetate.  These  reactions  were  also  obtained  with  portions 
of  the  natural  gum  of  Acacia  penninervis.  The  bacterial  slime, 
therefore,  contained  pentosans  as  evidenced  by  the  production  of 
furfural  on  treatment  with  acids. 

As  I  have  already  said,  the  cultures  upon  an  artificial  medium 
made  up  from  a  few  constituents  were  not  very  successful.  This 
medium  contained  saccharose,  peptone,  glycerine,  potassium  phos- 
phate and  chloride,  and  agar  in  proportions  that  experiment 
seemed  to  indicate  as  being  suitable.  Agar  must  be  employed, 
because,  although  on  all  gelatine  media  masses  of  slime  enclosing- 
bacteria  can  be  observed,  yet,  as  the  medium  is  slowly  liquefied 
by  the  bacterial  products,  the  slime  would  be  contaminated  with 
peptonised  gelatine.  In  the  course  of  experimenting  with  various 
media,  potato-extract  agar  gave  a  good  growth,  and  this  was 
increased  by  the  addition  of  glucose  or  saccharose.  But  even 
this  growth  was  not  what  would  be  called  rich  in  gum  or  slime. 

At  this  stage  several  observations  were  made.  After  scraping 
the  culture  from  the  surface  of  saccharose-potato  agar,  the  plates, 
which  had  been  incubated  at  30°,  were  allowed  to  stand  at  the 
laboratory  temperature  (15°)  for  several  days,  when  a  quantity  of 
slime  greater  than  the  original  crop  at  30°  was  obtained.  This 
appeared  to  indicate  that  a  lower  temperature  and  a  slower  growth 
was  accompanied  by  relatively  greater  slime  formation.  In 
separating  the  bacteria  from  the  tissues  of  a  specimen  of  gum- 


388  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

bearing  Acacia  binervata,  DC.,*  it  was  noted  that  subjacent  to 
the  hole  in  the  bark  through  which  a  large  globule  of  gum  had 
exuded  there  occurred  a  small  quantity  of  fluid  which  was  acid 
and  contained  a  tannin  body.  In  the  tissue  saturated  with  this 
fluid,  slime-forming  bacteria  identical  with  the  bacterium  from 
Acacia  penninervis  were  obtained  in  pure  culture.  The  occurrence 
of  tannin  at  the  site  of  production  of  the  gum  suggested  that  it 
might  influence  the  production  of  gum.  Accordingly  experiments 
were  made  with  it,  and  from  these  it  appeared  that  from  0*2  to 
0'4%  of  tannin  did  influence  the  formation  of  slime.  Smaller 
proportions  of  tannin  when  added  to  saccharose-potato  agar 
seemed  to  produce  a  relatively  greater  quantity  of  bacteria  and 
less  slime,  while  more  tannin  produced  less  bacteria  and  less  slime. 
With  regard  to  the  reaction  of  the  medium,  experiments  made 
with  varying  amounts  of  citric  acid  and  of  sodium   carbonate 

*  This  was  growing  in  a  garden  in  Macpherson-street,  Waverley,  Sydney, 
and  my  attention  was  drawn  to  it  by  Miss  S.  Hynes,  of  the  Botanic  Gardens. 

Maiden    (Pharm.  Jour.  [3]  xx.,   980)  gives  the  following  description  of 
the  gum : — 

"Acacia  binervata,  DC,  B.F.  ii.,  390. 

Arabin       76' 57 

Metarabin 4'24 

Ligneous  matter 1  '62 

Moisture -         ...     16-01 

Ash -771 


99-211 

"  A  '  black  wattle '  found  in  New  South  Wales  and  Queensland  ;  obtained 
from  old  trees,  Cambewarra,  New  South  Wales  ;  diameter  8-12  inches.  It 
is  obtained  in  pieces  from  the  size  of  a  pigeon's  to  that  of  a  hen's  egg,  and  is 
of  a  waxy  lustre.  The  freshly  exuded  gum  is  very  pale  brown;  old  gum  is 
often  nearly  black.  Being  in  comparatively  large  masses,  sorting  for  market 
would  be  easy.     It  has  a  dull  horny-looking  fracture. 

It  dissolves  fairly  well  in  water,  leaving  a  quantity  of  gum  in  the  form  of 
a  flocculent  deposit.  The  colour  of  the  dissolved  gum  is  rather  dark  owing 
to  the  presence  of  included  fragments  of  bark.  This  description  of  the  effect 
of  cold  water  will  apply  equally  well  to  A.  dealhata  and  A.  elata.  Like  other 
wattle  gums,  its  aqueous  solution  is  distinctly  acid  to  litmus  paper." 


BY    R.    GREIG    SMITH. 


389 


showed  that  acidity  or  alkalinity  of  the  medium  favoured  neither 
the  growth  of  bacteria  nor  the  production  of  slime. 

As  the  function  of  tannin  might  consist  in  slowing  the  growth 
of  the  bacteria,  glycerine  which  appeared  to  do  the  same  thing, 
was  also  employed  in  an  experiment  to  test  the  formation  of  slime. 

Several  plates  of  saccharose-potato  agar  which  contained 
0-06  grm.  tannic  acid  or  1  c.c.  glycerine  in  every  20  c.c.  of  medium 
were  prepared.  The  agar  surfaces  were  smeared  with  a  culture 
of  the  bacterium,  and  the  plates  were  incubated  at  22°  and  30'' 
for  three  days.  After  noting  the  appearances  of  the  cultures, 
the  slimes  were  removed  with  a  rubber  spade,  and  after  mixing 
with  water  slightly  acidified  with  hydrochloric  acid,  alcohol  was 
added.  The  compact  or  flocculent  precipitates  were,  after  wash- 
ing with  alcohol,  dried  at  100°  and  weighed.  The  results  are^ 
summarised  in  the  following  table  : — 


Addition  to  medium. 

Incubation 
temperature. 

Nature  of  growth. 

Order  as 

judged  by 

the  eye. 

Mgrms. 

crude  dry 

slime. 

Glycerine. 
Tannic  acid. 

22^ 
30^ 

Granular,  adherent, 
yellow  paste. 

Pale  buff  slime,  non- 
adherent. 

Buff  paste  non- 
adherent. 

2 

4 

1 

4 

58 

61 
83 

38 

The  results  show  that  it  might  be  possible  to  obtain  a  quantity 
of  slime  by  using  tannic  acid  and  by  using  the  lower  temperature. 
They  also  show  how  one  can  be  misled  by  appearances,  for  there 
appeared  to  be  much  more  growth  produced  on  the  glycerine 
medium  at  22°  than  at  30°,  while  in  reality  there  was  a  little  less. 

On  tannin  agar  the  production  of  slime  appears  to  have  finished 
in  from  four  to  five  days  at  22°  C,  and  the  medium  assumes  a 
blackish-green  colour.  If  the  plate  had  not  been  uniforml}^ 
smeared,  and  the  growth  in  consequence  had  formed  irregularly, 
it  is  noticed  that  the  culture  and  medium  are  buflf  coloured  at 
the  margin,  and  dirty  green  in  the  middle  where  the  growth  is 
26 


390  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

older.  The  change  from  buff  to  green  results  from  the  alteration 
in  the  reaction  of  the  medium  from  neutral  or  faintl}/  alkaline  to 
decidedly  alkaline,  as  is  shown  by  pressing  litmus  paper  against 
parts  of  the  under  side  of  the  plate — i.e.,  the  agar  in  contact  with 
the  bottom  of  the  petri-dish.  The  greenish,  mottled  slime  is 
converted  to  a  buff  colour  by  the  addition  of  a  few  drops  of  dilute 
hydrochloric  acid. 

When  the  culture  becomes  dark-coloured  there  is  not  so  much 
slime  obtained;  what  has  been  formed  appears  to  have  condensed 
and  to  have  become  less  soluble  in  water,  so  that  the  culture  forms 
a  suspension  with  water,  while  the  earlier  buff  cultures  form  a 
stiff  slime  under  similar  conditions. 

Further  experiments  with  the  tannin  medium  showed  that  the 
slime  was  formed  more  readily  by  growing  the  bacteria  at  15° — 
the  laboratory  temperature  at  the  time — than  at  22°.  The  most 
successful  method  consisted  in  growing  the  culture  at  15°  for 
three  days,  and  then  scraping  the  slime  from  the  plates.  In 
another  two  days  a  further  quantity  can  be  removed,  and  jDossibly 
still  another  in  two  days  later.  The  slime  is  acidulated  with  a 
few  drops  of  dilute  hydrochloric  acid  and  treated  with  alcohol, 
when  either  white  stringy  flakes  or  white  floccules  are  precipitated 
according  to  the  alcohol  which  has  been  added.  The  stringy, 
cohesive  flakes  are  changed  to  floccules  by  strong  alcohol.  The 
alcoholic  mother  liquor  is  coloured  a  bright  yellow  from  the 
lipochromes  of  the  bacterial  cells.  The  floccules  of  slime  and 
bacteria  are  white,  and  on  treatment  with  water  swell  up,  forming 
a  stiff  paste  like  that  made  from  flour.  The  opalescence  is  caused 
by  the  bacteria,  and  to  eliminate  them  the  paste  was  treated  with 
2  to  5  drops  of  dilute  sulphuric  acid,  and  heated  in  the  autoclave 
at  3  atmospheres'  pressure  for  15  minutes.  This  treatment  had 
been  found  very  useful  in  separating  bacteria  from  slime  on  a 
former  occasion.  By  this  treatment  a  faintly  opalescent,  gummy 
fluid,  which  could  easily  be  separated  by  filtration  through  paper 
from  the  precipitated  bacteria,  was  obtained.  The  faintly  opal- 
escent fluid  was  easily  clarified  with  aluminium  hydrate.  The 
difference  between  slime  and  gum  appears  to  be  caused  by  the 


BY    R.    GREIG    SMITH.  391 

presence  of  albuminoids  secreted  by  or  contained  in  the  bacteria. 
When  the  albuminoids  are  coagulated  by  treatment  in  the  auto- 
clave the  slime  is  altered  to  a  gum. 

Having  indicated  the  method  for  obtaining  a  solution  of  the 
gum  acids  free  from  dead  organisms,  a  word  may  be  said  about 
the  bacteria.  The  bacterium  from  Acacia  penninervis  was 
identical  with  that  separated  from  A.  binervata,  but  while  with 
the  former  another  bacterium  was  associated,  the  bacteria  in  the 
latter  were  in  pure  culture.  The  bacteria,  however,  were  of 
different  races — indicated  only  by  the  production  of  larger 
colonies  on  nutritive  media  and  more  slime  on  tannin-saccharose- 
potato  agar.  The  stronger  race  (from  Acacia  binervata)  was  used 
subsequently  for  the  production  of  the  artificial  gum. 

The  medium  upon  which  most  slime  was  obtained  consisted  of 

Potato  extract 1,000  c.c. 

Saccharose 50  grms. 

Agar 20     ,, 

Tannic  acid 3      ,, 

The  potatoes  were  washed,  pared,  eyed,  grated  and  finally 
strained  and  pressed  in  a  meat-press.  The  juice  was  then  allowed 
to  stand  overnight  in  a  flask  of  such  a  size  that  the  juice  filled 
the  neck.  The  darkening  of  the  fluid  by  contact  with  air  was 
thus  minimised.  In  the  morning  the  juice  was  siphoned  off  from 
the  starch  and  filtered.  An  equal  volume  of  water  was  added  to 
the  filtrate,  which  was  then  boiled  to  coagulate  the  albuminoids, 
which  were  removed  by  filtration  from  the  potato-extract.  To  the 
extract  thus  obtained,  the  sugar  was  added  and  the  solution 
heated  in  the  autoclave  to  three  atmospheres'  pressure"^  in  order  to 
kill  the  spores  of  Bac.  levaniformans,  which  are  generally  present  in 
commercial  sugar.  After  removal  from  the  autoclave,  the  chopped 
agar  was  added  and  the  solution  was  returned  to  the  autoclave 
and  heated  to  one  and  a  half  atmospheres'  pressure  to  bring  the 
agar  into  solution  The  tannin  was  then  added  and  the  medium 
was  steamed  for  an  hour,  after  which  it  was  cooled  to  50°  and 
poured  into  sterilised  (by  flaming)  large  damp-chambers  or  small 

*  This  would  be  unnecessary  if  dextrose  were  used. 


392  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

petri-dishes.  The  medium  in  the  larger  vessels  was  after  congela- 
tion infected  with  a  suspension  of  the  bacteria  in  10  c.c.  of  normal 
saline;  the  medium  in  the  petri-dishes,  by  smearing  with  a  loop 
of  the  culture.  The  culture  used  for  the  purposes  of  infection 
was  obtained  by  growing  the  bacteria  at  30°  in  potato-saccharose 
agar  prepared  in  the  same  way,  but  without  tannin.  The  incuba- 
tion at  30°  on  this  medium  gives  a  loose  yellow  growth  with  very 
little  slime;  the  culture  therefore  readily  becomes  distributed  in 
the  normal  saline.  The  plates  were  kept  at  the  laboratory  tem- 
perature (about  15°)  and  the  slime  was  removed  with  a  rubber 
spade  on  the  third,  fifth  and  seventh  days.  The  slime  is  easily 
removed,  and  there  is  no  danger  of  agar  being  taken  at  the  same 
time.  The  first  and  second  crops  were  preserved  by  the  addition 
of  a  few  drops  of  dilute  hydrochloric  acid  and  of  alcohol  until  all 
the  slime  was  obtained.  The  total  slime  was  then  treated  with 
an  excess  of  strong  spirit  and  worked  up  in  the  manner  already 
described.  The  difference  in  the  amount  of  slime  obtained  by 
adding  tannin  to  the  potato-saccharose  agar  is  ver}''  marked. 
Without  tannin  the  growth  is  bright  yellow  and  slightly  gummy, 
while  with  tannin  it  is  pale  buff,  thick  and  slimy. 

A  portion  of  the  gummy  solution  free  from  sugars  was  pre- 
cipitated with  alcohol  and  the  precipitate  treated  with  nitric  acid, 
sp.g.  1*1 '2.  A  white,  sandy  powder,  difficultly  soluble  in  cold 
water  but  readily  soluble  in  boiling  water,  was  obtained.  It  had 
an  indefinite  melting  point  over  210°  C,  and  had  the  same  crys- 
talline appearance,  viz.,  colourless  tables  with  a  straight  side,  as 
mucic  acid  obtained  from  commercial  gum  acacia.  The  mother 
liquor  contained  oxalic  acid.  The  oxidation  products  are  thus 
identical  with  those  of  gum  acacia. 

So  far  the  identity  of  the  bacterial  gum  with  the  natural  gum 
has  been  proved  by  the  test-tube  reactions,  the  presence  of  pen- 
tosans and  the  oxidation  products.  There  remained  the  optical 
activity  and  the  determination  of  the  constituents  of  the  gum 
acids.  The  former,  viz.,  the  optical  activit}',  is  of  little  con- 
sequence  in    view    of    the     researches    of    other    investigators. 


BY    R.    GREIG    SMITH.  393 

O'SuUivan*  has  shown  that  fragments  of  the  same  gum  contain 
gum  acids  which  rotate  the  ray  of  polarised  light  to  dififerent 
extents.  He  also  showed  that  diiferent  kinds  of  gum  of  the 
arabin  group  have  rotations  in  different  directions;  for  example, 
gum  arabic  is  Isevo-rotatory,  Gedda  gum  is  generally  dextro- 
rotatory, and  an  Australian  gum  was  optically  inactive.  It  is 
therefore  probable  that  the  gum  acids  of  the  natural  gum  and 
those  of  the  bacterial  gum  would  differ  in  their  optical  activitj^, 
since  the  conditions  under  which  the  gums  had  been  produced 
had  been  so  very  different.  This  proved  to  be  the  case.  I  pre- 
pared the  gum  acids  from  the  gum  of  A.  hinervata  and  from 
cultures  of  Bact.  acacice  and  submitted  them  to  Mr.  T.  U.  Walton, 
B.Sc,  of  the  Colonial  Sugar  Refining  Co.,  who  found  that  the 
natural  gum  acids  had  a  specific  rotation  of  [«]d  =  +0°-9  and 
the  bacterial  gum  acids  had  a  rotation  of  [a]^,  =  +  43°.  I  do  not 
lay  any  stress  upon  the  difference  in  the  optical  activities.  In 
the  future  I  shall  grow  the  bacterial  gum  in  different  ways  and 
from  different  materials,  to  see  how  the  optical  activity  is  thereb}'- 
influenced. 

A  portion  of  the  sugar-free  gum  acids  was  dissolved  in  15  c.c. 
of  water,  and  to  this  10  c.c.  of  5  %  sulphuric  acid  was  added,  thus 
making  a  2  %  solution.  This  was  heated  on  the  water  bath  for 
an  hour  to  hydrolyse  the  combined  arabinan.  Subsequent  work 
showed  that  this  heating  might  with  advantage  be  prolonged  for 
three  or  four  hours  to  hydrolyse  the  arabinan  more  completely. 
After  the  digestion  with  the  2  %  acid  the  solution  was  cooled 
and  the  unaltered  gum  acids  precipitated  with  alcohol.  The 
solution,  after  filtration,  was  distilled  until  most  of  the  alcohol 
had  been  removed;  the  residual  fluid  was,  after  the  addition  of 
about  50  c.c.  of  water,  neutralised  with  barium  carbonate  and 
filtered.  The  filtrate  was  boiled  down  to  one-half  and  clarified 
with  aluminium  hydrate.  The  clear  solution  was  used  for  the 
preparation  of  the  osazone. 


*  O'Sullivan,  .Journ.  Chem.  Soc.  1891,  1029. 


394  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

The  residual  gum  acids  which  did  not  hydrolyse  by  the  2  % 
acid  treatment  were  dissolved  in  25  c.c.  of  5  %  sulphuric  acid  and 
warmed  to  expel  the  traces  of  alcohol.  The  solution  was  then 
made  up  to  25  c.c.  with  water  and  boiled  under  a  reflux  condenser, 
at  first  gently  and  then  vigorously  after  the  foaming  had  ceased. 
The  boiling  was  continued  for  five  hours.  After  cooling,  the  acid 
solution  was  treated  with  alcohol  and  a  very  small  quantity  of 
unaltered  gum  removed.  The  alcoholic  solution  was  treated  as 
has  been  already  described,  and  the  final  solution  reserved  for 
the  preparation  of  the  osazone. 

The  solution  supposed  to  contain  arabinose  was  tested  with 
Fehling's  solution,  when  a  strong  reduction  was  obtained.  One- 
half  of  the  solution  was  set  aside  in  case  of  accidents.  The  other 
half  was  heated  on  the  water  bath  and  to  it  were  added  2  c.c.  of 
a  solution  containing  40  c.c.  phenylhydrazine,  40  c  c.  glacial 
acetic  acid  and  20  c.c.  water.  The  heating  was  continued  for  an 
hour  and  the  solution  was  cooled.  The  precipitated  osazone  was 
filtered  off,  dried  on  porcelain,  transferred  to  a  small  filter  and 
extracted  with  ether*  to  remove  a  black,  tarry  substance  which 
melted  at  about  98°.  This  was  always  found  in  the  osazones 
from  the  bacterial  as  well  as  the  natural  gums.  It  was  never 
obtained  in  working  with  pure  sugars  such  as  dextrose.  The 
precipitate  was  dried  at  100°  and  the  melting  point  determined. 
This  was  160°,  which,  together  w^ith  the  appearance  of  the  crystals, 
showed  it  to  be  the  osazone  of  arabinose. 

The  solution,  which  presumably  contained  galactose,  reduced 
Fehling's  solution.  One-half  was  treated  with  phenylhj^drazine- 
acetic  acid  and  the  osazone  extracted  with  ether.  The  dry  osazone 
melted  at  182-183°,  which  show^ed  it  to  be  a  mixture.  The 
mixed  osazone  was  boiled  with  10  c.c.  of  waterf  and  filtered 
through  a  hot  filter  which  had  been  moistened  with  boiling  water. 

"•'  A  preliminary  moistening  with  alcohol  is  sometimes  advantageous. 
t  In  some  cases  the  osazones  could  not  be  separated  by  water  alone,  but 
by  dissolving  everything  in  dilute  alcohol  and  slowly  boiling  off  the  alcohol, 
the  galactosazone  precipitated  out  from  the  hot  solution  and  was  separated 
by  filtering  through  a  hot,  wetted  filter. 


I 


BY    R.    GREIG    SMITH.  395 

The  filtrate  on  cooling  deposited  microscopic  crystals  which  dried 
as  an  Arabian-brown  skin  on  porcelain  and  which  melted  at 
157-159°,  showing  them  to  be  the  osazone  of  arabinose.  The 
portion  insoluble  in  water  melted  at  187°.  This  was  again  treated 
with  10  CO.  of  boiling  water.  The  insoluble  portion  dried  as  a 
yellow  powder  and  melted  at  192-193°,  the  melting  point  of 
the  osazone  of  galactose. 

The  bacterial  gum  acids  had  therefore  yielded  arabinose  and 
galactose  on  hydrolysis,  from  which  we  must  conclude  that  they 
contain  the  arabinan-galactan  complex  and  are  of  the  same  nature 
as  the  natural  gums  of  the  arabin  group. 

The  gum  which  was  found  upon  Acacia  hinervata  was  of  a  very 
pale  yellowish-brown  and  of  a  dark  brown  colour.  Portions  of 
the  same  mass  showed  both  colours.  It  occurred  in  hemispherical 
masses  and  in  tears  and  had  a  tough  gelatinous  consistency  which 
enabled  it  to  be  cut  without  fracture.  It  dissolved  readily  in 
water,  and  in  the  absence  of  particles  of  bark  it  formed  a  clear 
solution  with  an  acid  reaction.  The  acidity  of  100  grms.  of  gum 
to  litmus  paper  was  equal  to  3*27  c.c.  of  normal  acid. 

Although  there  could  be  no  doubt  that  the  gum  contained  the 
arabinan-galactan  complex,  yet  to  complete  the  identity  of  the 
bacterial  with  the  natural  gum  acids  the  proof  was  needed. 
Accordingly  a  portion  of  the  natural  gum  was  dissolved  in  water, 
acidified  with  hydrochloric  acid  and  treated  with  alcohol.  The 
gum  acids  were  hydrolysed  and  the  sugars  tested  by  means  of 
their  osazones  in  the  manner  already  indicated,  when  arabin- 
osazone  and  galactosazone  were  obtained.  Furthermore,  like  the 
bacterial  acids,  there  is  apparently  a  greater  proportion  of  arabinan 
than  galactan  in  the  complex. 

Summary. — A  bacterium  was  found  in  pure  culture  at  the 
place  from  which  the  natural  gum  was  exuding.  This  bacterium 
in  the  laboratory  formed  a  gum  which  behaved  to  reagents,  gave 
the  same  oxidation  products,  and  contained  the  same  constituents, 
viz.,  arabinan  and  galactan  as  the  natural  gum.  The  bacterium 
is  thus  the  producer  of  the  natural  gum. 


396  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIX    GROUP, 

The  bacterial  origin  of  that  variety  of  gum  acacia  exuding  from 
Acacia  binervata  having  been  proved,  it  is  a  just  assumption  that 
all  other  gums  of  the  arabin  group  are  likewise  bacterial  products, 
and  not  substances  formed  by  the  plants  in  a  pathological  con- 
dition. 

The  formation  of  gum  is,  therefore,  a  bacterial  disease,  for  the 
parasitic  bacteria  obtain  their  nourishment  from  the  plant  juices 
which  they  elaborate  into  gum,  ^vhich  is  not  required  by  the  host 
plant.  It  is  still  undecided  what  constituents  they  may  alter. 
Gum  can  be  formed  from  saccharose  and  from  dextrose,  but  I 
will  deal  with  this  side  of  the  subject  in  a  future  paper. 

The  bacteria  that  are  parasitic  in  plants  are  usually  found  in 
the  feebly  acid  or  neutral  juices  of  the  vessels,  and  probably 
the  gum  bacteria  are  located  in  the  sieve  tubes  of  the  soft  bast, 
as  Kraus  *  has  indicated,  although  the  gum  is  not  a  true  cell 
content  as  he  supposed. 

Bacterium  acacia,  n.sp. 

Shape,  etc. — The  bacteria  appear  as  short  rods  with  rounded 
ends,  and  occur  singly,  in  pairs,  and  in  groups,  sometimes  within 
a  well-defined  slime  case  (ascus).  On  nutrient  agar  the  rods 
measure  0'5  :  1  /x ;  in  bouillon  0*5  :  1-5-2  /x,  pairs  commonly 
occur  simulating  a  long  rod  ;  on  glycerine-saccharose  agar  the 
bacteria  vary  from  0-5  : 0'5-l'5  fx,  appearing  as  cocci  and  as  short 
rods;  on  saccharose-potato  agar  they  measure  0-6:l-2'5/x  and 
average  0*6  :  1'5  /z.     The  cells  stain  well  with  the  ordinary  stains; 

*  "Herr  G.  Kraus  has  determined  by  observations  on  the  exudation  of 
gum  from  Acacia  melanoxylon  that  it  is  formed  only  in  the  bark  and  not  in 
the  wood,  and  only  in  the  bast  layer,  never  in  the  parenchyma  nor  in  any 
more  external  portion  that  the  bast  fibres  have  no  share  in  its  formation  ; 
that  it  flows  from  the  cells  of  the  soft  bast,  and  especially  from  the  sieve 
tubes,  and  that  it  is  not  a  product  of  degradation  of  the  cellulose  but  is  a  true 
cell  content  flowing  out  unchanged  through  the  unchanged  cell  walls." 
Abstract  in  Journ.  Roy.  Micr.  Soc.  [ii.]  vi.  (1886),  90,  from  Ber.  Sitz. 
Naturf.  Gesell.  Halle,  1884,  pp.  19-20. 


BY    R.    GREIG    SMITH.  397 

the  Gram  stain  is  negative.  The  bacteria  are  actively  motile  ; 
the  flagella  vary  from  one,  terminal,  to  many,  peritrichous;  in 
the  latter  cases  up  to  six  have  been  observed,  but  there  may  be 
more. 

Temperature,  etc. — The  optimum  temperature  is  apparently 
about  22°  on  saccharose-potato  agar;  the  growth  is  most  bulky  at 
22°,  then  at  15°,  then  30°,  and  smallest  at  37°.  The  organism 
is  aerobic,  but  grows  scantily  under  anaerobic  conditions. 

Nutrient  agar  plate. — At  30°  the  colonies  are  circular,  white, 
slightly  raised,  and  moist  glistening.  When  magnified  they  are 
seen  to  have  a  smooth  circular  edge  and  contain  scattered 
granules.  The  deep  colonies  are  irregular,  oval  or  round,  and 
coarsely  granular.  The  colour  changes  from  white  to  primrose 
yellow. 

Glucose-gelathie  plate. — The  colonies  are  white,  circular  but 
sometimes  lacerate,  slightly  raised  and  glistening  like  drops  of 
gum.  The  colour  deepens  to  yellow.  When  magnified,  granules 
are  seen  scattered  throughout  the  colony,  and  as  growth  proceeds 
these  become  coarser  and  more  numerous.  The  deep  colonies  are 
at  first  dark,  rough  and  irregular,  but  become  rounded  or  moru- 
loid.  The  old  surface  colonies  are  raised,  slimy  and  streaky. 
The  medium  is  very  slowly  liquefied. 

N\Ltrient  gelatine  plate.  —As  on  glucose-gelatine,  but  the 
growth  is  not  so  good. 

Nutrient  agar  stroke. — A  white,  flat,  moist,  glistening  stroke  is 
first  formed;  this  changes  to  a  primrose  or  yellow,  dr}^,  terraced 
growth.     The  margin  may  be  straight  or  lobed. 

Saccharose-potato  agar  stroke.  —  The  temperature  of  incubation 
has  a  great  influence  upon  the  appearance  of  the  cultures.  In  3 
days  at  30°  the  growth  is  yellow-buff,  opaque  and  terraced.  At 
25°  the  stroke  is  transparent  at  the  margin  and  streaked  with 
white  in  the  centre,  while  the  growth  has  flowed  downwards  into 
the  condensed  water,  which  has  become  a  thick  slime.  At  22° 
the  opacity  and  growth  is  more  pronounced,  the  colour  is  pale 
buff.     The  growth  is  more  undulating  both  at  15°  and   at    22° 


398  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

than  at  30°  or  37'.     At  37°  the  growth  is  as  at  30°  but  more 
scanty. 

Glucose-gelatine  stroke. — The  growth  is  irregular,  spreading, 
sliiny,  and  becomes  depressed  in  places  owing  to  the  slow 
liquefaction  of  the  medium.  The  colour  changes  from  white  to 
canary-yellow. 

Xutrient  gelatine  stroke. — As  on  glucose-gelatine  but  poorer. 

Glucose-gelatine  stab. — A  strong  filiform  growth  with  a  trans- 
lucent, white,  flat,  slimy  nail-head.  The  nail-head  slowly  sinks, 
and  a  funicular,  then  stratiform,  liquefied  area  is  formed.  The 
medium  may  develop  a  few  gas  bubbles.  Saccharose  is  also 
fermented  with  an  evolution  of  gas.  Some  races  liquefy  the 
medium,  slowly,  and  an  air-bubble  is  produced  at  the  top  of  the 
filiform  stab.  Taken  as  a  whole  the  liquefaction  of  the  medium 
is  slow. 

Nutrient  gelatine  stab. — A  smaller  growth  than  glucose-gela- 
tine. 

Potato. — The  growth  is  at  first  dull  buff,  raised  and  irregular, 
then  becomes  glistening  and  deep  yellow. 

Bouillon. — The  medium  becomes  turbid,  and  forms  a  loose 
fiocculent  sediment,  and  a  thin  broken  surface  film  which  adheres 
to  the  sides  of  the  glass.  The  sediment  becomes  deep  yellow  and 
cohesive.  The  recently  isolated  races  produced  indol  together 
with  nitrite  in  from  10  to  15  days  at  SO''.  After  the  races  had 
been  cultivated  in  the.  laboratorj'  for  several  months  the  red 
nitroso-indol  reaction  was  in  some  cases  faint,  and  in  other  cases 
was  absent.  In  the  bouillon  ammonia  could  be  detected,  but 
neither  phenol  nor  sulphuretted  hydrogen. 

Milk. — The  medium  is  either  unaltered  or  made  slightly  ropy 
with  a  faint  acid  reaction. 

The  bacterium  has  characters  which  mark  it  as  being  new,  the 
most  important  of  these,  being  the  production  of  arabin.  I 
accordingly  name  the  organism  Bacterium  acacice  (Bacillus 
acacice  b}^  Migula's  s\'stem). 


BY    R.    GREIG    SMITH.  399 

II. — The  Insoluble  (Metarabin)   Wattle  Gums. 
(Bacterium  metarahinum,  n.sp.) 

While  the  natural  gum  of  Acacia  binervata  was,  in  the  absence 
of  particles  of  bark,  entirely  soluble  in  water,  the  gum  of  Acacia 
penninervis  was  only  partly  so.  The  bacteria  associated  with  the 
gum  of  the  former  were  entirely  JJact.  acacice,  while  from  the 
latter  this  organism  and  another  were  found.  As  separated  from 
my  culture  solutions  the  other  bacterium  was  in  the  minority, 
but  on  considering  the  matter  this  can  be  accounted  for  by  its 
method  of  growth.  Bad.  acacice  in  peptone  saccharose  fluid 
grows  as  single  cells,  while  the  other  bacterium  has  a  tendency 
to  grow  in  aggregates^*,  and  thus  in  plate  culture  the  aggregate 
comes  out  as  if  it  had  been  a  single  cell. 

I  did  not  think  that  the  plant  would  convert  soluble  gum  to 
the  insoluble  or  meta  modification,  and  expected  that  the 
insolubility  resulted  from  the  further  action  of  the  organism,  the 
action  of  tannin,  or  that  it  was  produced  by  quite  another 
bacterium.  The  first  is  unlikely,  the  second  is  quite  possible,  and 
the  last  is  most  probable.  Tannin  is  known  to  stiffen  the  gum, 
and  it  would  therefore  be  an  easy  thing  to  affirm  that  the 
insolubility  was  entirely  due  to  its  action.  But  the  gum  of 
Acacia  binervata  is  formed  in  the  presence  of  tannin  bodies,  and 
although  fragments  of  bark  are  imbedded  in  the  solid  exudate, 
yet,  when  these  are  picked  out,  the  gum  is  found  to  be  entirely 
soluble  in  w'ater.  There  is,  therefore,  strong  reason  to  believe 
that  tannin  has  nothing  to  do  with  the  production  of  the 
insoluble  portion  of  the  gums. 

The  second  bacterium  from  Acacia  pennhiervis  forms,  in  suit- 
able media,  slime  masses  similar  to  Bad.  acacice^  and  while  the 
latter  forms  gummy  colonies  on  gelatine  media,  the  colonies  of 
the  former  are  dry  and  cohesive,  so  that  the  colonies  are  removed 
from  the  medium  by  the  inoculating  needle  f:n  masse.     There  is 

On  this  account  the  bacterium  must  be  "  plated  "  several  times  before  a 
pure  culture  is  obtained. 


400  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

in  this  method  of  growth  a  strong  indication  that  the  slime  is 
more  insoluble  than  that  secreted  by  Bact.  acacice.  On  saccharose- 
potato  agar  with  or  without  tannin  Bact.  acacice  produces  at  15° 
a  gummy  slime  that  does  not  adhere  firmly  to  the  medium.  The 
other  bacterium  forms  a  thicker  slime,  and  the  portion  in  contact 
with  the  medium  adheres  firmly  and  cannot  be  entirely  removed 
with  a  rubber  spade.  When  the  two  slimes  are  carefully  removed, 
dried  in  the  steam  bath  at  100°  and  then  moistened  with  water, 
the  slime  of  Bact.  acacice  swells  up,  forming  a  diffusive  slime  very 
similar  to  its  original  condition,  while  the  slime  of  the  other  bac- 
terium smells  slightly  and  retains  the  shape  in  which  it  had 
dried.  There  is  thus  a  strong  probability  that  the  new  bacterium 
forms  the  insoluble  gum  (the  meta-arabin)  of  wattle  gum.  That 
the  slime  might  eventually  prove  to  be  of  the  arabinan-galactan 
class  was  indicated  by  the  great  similarity  of  the  growth  on 
tannin-saccharose-potato  agar  with  that  produced  by  Bact.  acacice. 

The  nature  of  the  slime  had  now  to  be  determined,  and  as  with 
Bact.  acacice,  large  plates  of  tannin-saccharose-potato  agar  were 
infected  with  a  suspension  of  the  bacteria  in  10  c.c.  of  normal 
saline,  and  these  were  kept  at  the  laboratory  temperature  (15°). 
The  slimes  wers  collected,  rendered  more  acid  with  a  few  drops 
of  dilute  hydrochloric  acid  and  treated  with  alcohol.  A  curdy 
precipitate  which  became  cohesive  and  an  opalescent  solution 
were  obtained.  The  precipitate  was  stirred  repeatedl}'  with 
alcohol  until  no  further  opalescence  was  produced.  The  pre- 
cipitate was  treated  with  water  and  then  with  alcohol,  when  a 
thin  starch  paste-like  suspension  was  produced.  This  was 
flocculated  with  barium  chloride  (potassium  chloride  did  not  act 
so  well  as  the  barium  salt),  and  a  contractile  curd  separated  from 
an  opalescent  solution.  The  opalescent  alcoholic  solution  was. 
added  to  the  opalescent  solution  previously  obtained  and  the  gum 
in  this  was  precipitated  by  the  addition  of  barium  hydrate  and 
reserved  for  future  examination. 

The  contractile  curd  was  treated  with  water,  with  which  it 
formed  a  thick  paste.  The  lumps  were  broken  up  by  passage 
through  a  wire-gauze  filter  and  the  whole  was  warmed  to  cause 


BY    R.    GREIG    SMITH.  401 

a  uniform  swelling.  It  had  an  acid  reaction  to  litmus  paper. 
By  heating  in  the  autoclave  for  ten  minutes  at  3-4  atmospheres  a 
slimy  sediment  (apparently  gummy)  and  an  almost  clear,  slightly  % 
acid,  supernatant  fluid  was  produced.  The  slimy  sediment  was 
mixed  with  water  treated  with  2-3  drops  dilute  sulphuric  acid 
and  again  heated  in  the  autoclave.  This  gave  a  curdy  precipi- 
tate and  supernatant  solution.  The  curdy  precipitate  was 
apparently  free  from  gum,  and  as  it  probably  consisted  of 
coagulated  bacteria  it  was  not  examined  further.  The  faintly 
acid  solution  from  the  first  autocla\'e  treatment  was  tested  with 
a  few  reagents.  Lead  acetate,  barium  hydrate  and  barium 
chloride  gave  no  precipitate.  Basic  lead  acetate  and  ammoniacal 
lead  acetate  gave  precipitates.  Fehling's  solution  gave  no  pre- 
cipitate and  no  reduction  on  heating.  Copper  sulphate  followed 
by  sodium  hydrate  gave  a  precipitate  which  coagulated  on  boiling 
without  change  of  colour.  Alcohol  gave  an  opalescent  solution 
which  flocculated  readily  with  potassium  chloride. 

The  gum  acids  in  the  mixed  solutions  from  the  autoclave  treat- 
ment were  precipitated  with  alcohol  and  potassium  chloride  and 
kept  over-night  in  contact  with  the  alcohol.  The  precipitate 
was  very  contractile,  and  on  treatment  with  water  it  partly 
dissolved  and  partly  swelled  up,  forming  a  practically  unfiltrable 
suspension.  A  portion  of  this  was  preserved;  it  had  not  dissolved 
at  the  time  of  reading  this  paper,  i.e.,  in  one  month;  the  gela- 
tinous lumps  were  still  evident.  We  have  in  this  insolubility  of 
the  gum  acids  a  condition  identical  with  what  occurs  on  treating 
the  semi-insoluble  wattle  gums  with  water,  and  confirms  the 
deduction  made  from  other  observations  that  this  bacterium  is 
responsible  for  the  production  of  the  metarabin  of  these  gums. 
The  gelatinous  lumps  of  gum  acids  were  insoluble  in  dilute  acid, 
but  readily  dissolved  in  dilute  sodium  hydrate,  from  which  dilute 
hydrochloric  acid  precipitated  the  gum  acids. 

The  acids  were  dissolved  in  dilute  sodium  hydrate,  neutralised 
with  sulphuric  acid,  and  enough  5  %  sulphuric  acid  was  added  to 
make  a  2  %  solution.  This  was  boiled  under  a  reflux  condenser 
for  two  and  a  half  hours.     A  white  precijDitate  which  contained 


402  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

no  o-um  acids  and  proved  to  be  barium  sulphate  was  filtered  from 
the  dilute  sulphuric  solution.  Alcohol  did  not  produce  a  pre- 
cipitate in  a  portion  of  the  filtrate,  so  that  the  gum  acids  appa- 
rently had  been  completely  hydrolysed  by  the  treatment.  The 
solution  was  neutralised  b}^  boiling  with  barium  carbonate, 
filtered  and  the  filtrate  clarified  with  aluminium  hydrate.  A 
portion  of  the  solution  was  treated  with  phenjdhydrazine  acetate 
solution,  and  a  mixed  osazone  was  obtained.  When  the  consti- 
tuents were  separated  in  the  manner  indicated  in  the  first  portion 
of  this  paper,  they  proved  to  be  arabinosazone  (m.p.  158-160°) 
and  galactosazone  (m.p.  192-193°).  The  gum  acids  therefore 
contained  the  arabinan-galactan  complex,  and  the  gum  belonged 
to  the  arabin  group. 

The  opalescent  alcoholic  solution  which  coagulated  on  the 
addition  of  barium  hydrate  was  washed  by  suspension  in  alcohol 
and  finally  treated  with  water,  in  which  it  formed  a  semi 
solution.  Dilute  sulphuric  acid  was  added  until  no  further 
white  precipitate  formed,  and  the  suspension  was  heated  in  the 
autoclave,  when  a  sediment  of  barium  sulphate  and  a  supernatant 
fluid  was  obtained.  From  this,  alcohol  threw  down  a  small 
quantity  of  a  contractile  precipitate  which  dissolved  easily  in 
water.  On  testing  the  solution,  which  was  acid  to  litmus  paper, 
precipitates  were  obtained  wdth  basic  lead  acetate,  ammoniacal 
lead  acetate  and  barium  hydrate.  Copper  sulphate  followed  by 
potassium  hj^drate  gave  a  precipitate  w^hich  contracted  but  did 
not  darken  on  boiling.  Barium  chloride,  sodium  h3^drate,  lead 
acetate  and  Fehling's  solution  gave  no  precipitates.  The  reaction 
with  barium  hydrate  suggests  the  possibility  that  the  compound 
may  be  a  pectin  substance.*     On  hydrolysis  with  sulphuric  acid 

*  The  pectin  substances  are  very  indefinite  and  yield  on  hydrolysis  various 
sugars.  Bourquelot  found  that  gentian  pectose  hydrolysed  to  arabinose  and 
doubtful  crystals  of  galactose.  Hebert  obtained  arabinose  from  the  pectic 
bodies  of  ripe  fruit  and  bulbs,  but  as  they  also  yielded  mucic  acid  on  oxida- 
tion, he  considered  that  gaiactan  had  been  present.  Bauer  hydrolysed  apple 
pectin  to  xylose  and  pear  pectose  to  galactose.  Herzfeld  concluded  that 
parapectic  acid  contained  arabinan  and  gaiactan. 


BY    R.    GREIG    SMITH.  403 

arabinose  and  galactose  were  obtained.  These  were  identified  as 
in  the  other  cases  by  the  isolation  of  the  osazones  and  the  deter- 
mination of  the  melting  points.  The  same  substance  was  not » 
obtained  from  the  natural  gum  of  Acacia  penninervis,  but  this 
may  be  accounted  for  by  the  small  quantity  of  the  gum  which  I 
had  at  my  disposal  at  that  time. 

The  growths  in  test-tubes  containing  saccharose-potato  agar 
with  and  without  tannin  apparently  show  that  tannin  has  little 
effect  in  increasing  the  quantity  of  slime.  The  difference,  however, 
is  more  marked  in  plate  cultures.  In  one  case  the  slime  scraped 
off  from  two  similar  large  plates  weighed  when  dry  : — tannin 
l'624grm.,  without  tannin  1*180  grm. 

In  purifying  the  gum  acids  it  was  noted  that  hydrochloric  acid 
prevents  the  precipitation  with  alcohol  much  more  than  in  the 
case  of  the  gum  acids  of  Bad.  acacice,  and  sometimes  a  strong- 
flocculating  agent,  i.e.,  barium  chloride,  has  to  be  employed. 
Again,  the  separation  of  the  bacteria  from  the  gum  acids  has  to 
be  very  carefully  done  if  one  would  obtain  the  acids  in  the  meta 
condition.  In  a  second  case,  when  a  few  drops  of  dilute  sulphuric 
acid  were  added  and  the  heating  in  the  autoclave  continued  for  a 
longer  period  (45  minutes)  the  gum  acids,  after  precipitation  with 
alcohol,  dissolved  readily  in  water.  The  insolubility  was,  how- 
ever, restored  by  heating  the  dried  gum  acids  at  100°  for  two 
hours.  After  this  treatment  they  did  not  dissolve  entirely  in 
water,  the  fioccules  being  visible  after  five  days.  The  gum  acids 
of  Bad.  acacice,  when  heated  at  100°  for  the  same  time,  dissolved 
in  water  as  readily  and  in  the  same  manner  as  commercial  gum 
acacia. 

The  natural  gum  of  Acacia  penninervis,  like  the  bacterial  gum, 
contained  the  arabinan-galactan  complex. 

Summary. — The  bacterium  which  I  have  named  Bacterium 
metarahinum  *  was  found  at  the  place  from  which  the  gum  was 
exuding.  It  produces  a  gum  which  is  tough  and  gelatinous,  as 
evidenced  by  the  consistency  of  the  colonies  on  nutritive  media. 

*  Bacillus  metarabinus,  by  Migula's  system  of  nomenclature. 


404  BACTERIAL    ORIGIN    OF    GUMS    OF    ARABIN    GROUP, 

The  gum  yields  gum  acids  which  partly  dissolve*  and  partly  swell 
up  with  water.  These  have  the  test  tube  reactions  of  gum 
acacia  and  contain  the  arabinan-galactan  complex.  The  gum 
acids  are  similar  to  those  of  the  natural  gum  of  Acacia  penni- 
7iervis,  and  there  can  be  no  doubt  that  the  natural  gum  is  of 
bacterial  origin.  Since  Bad.  acacice  was  found  with  Bad. 
metarahinum  the  gum  is  probably  produced  by  both  bacteria,  the 
former  producing  the  water-soluble  portion,  the  latter  the  water- 
insoluble  portion. 

Since  Bad.  acacice  was  found  in  the  only  two  instances  which 
were  examined,  it  is  possible  that  it  is  the  only  producer  of  all 
sorts  of  soluble  acacia  gums.  If  this  should  prove  to  be  the  case, 
the  varieties  of  the  gums  will  depend  upon  a  number  of  factors 
such  as  the  host,  plant,  the  juices  of  w^hich  undoubtedly  vary,  and 
thus  influence  the  relative  proportions  of  arabinan  and  galactan  in 
the  gum  complex.  The  situation  will  undoubtedly  influence  the 
susceptibility  of  the  tree  towards  the  microbe.  It  is  generally 
recognised  with  all  plant  diseases  that  unhealthy  surroundings 
have  much  to  do  with  the  invasion  by  bacteria  and  other  fungi. 
But  perhaps  the  most  important  factor  will  prove  to  be  the 
temperature.  The  slow  growth  at  the  higher  temperatures  (and 
the  appearances  of  the  growths  in  test  tube  cultures  indicated  a 
clearer  slime  at  comparatively  high  temperatures;  will,  I  think, 
explain  the  better  qualities  of  the  gums  produced  in  arid  regions. 
But  on  the  other  hand  it  is  possible,  and  analogy  supports  this 
contention,  that  there  is  a  class  of  bacteria  which  produce  the 
soluble  gums,  each  characteristic  gum  being  formed  by  a  different 
species.     Future  research  alone  will  settle  the  question. 

Since  the  arabin  and  the  metarabinf  gums  are  produced  by 
different  bacteria  it  is  extremely  probable  that  the  pararabin 
<yums  have  also  their  particular  microbe.     This  investigation  is 

""  The  partial  solution  may  be  accounted  for  by  the  treatment  with  acid  in 
the  autoclave  during  the  process  of  eliminating  the  bacteria. 

t  Metarabin  and  pararabin  are  insoluble  but  swell  up  with  water.  The 
former  is  soluble  in  dilute  alkalies,  the  latter  in  dilute  acids. 


BY    R.    GRBIG    SMITH.  405 

in    progress.     Gum   tragacanth  will   undoubtedly  be  shown  to 
have  a  bacterial  origin. 

The  various  gums  of  the  arabin  group  are  not  found  upon 
Acacias  only.  This,  however,  does  not  militate  against  the 
bacterial  origin  of  all  these  gums.  Thus  the  gums  exuded  by 
certain  Rosaceae*,  e.g.,  cherry,  peach,  plum,  are  very  similar  to 
wattle  gum,  and  are  probably  the  work  of  bacteria  such  as  Bad. 
acacice  and  metarabiuum.  These  fruit  tree  gums  are  being 
investigated. 

Bacterium  metarabinum,  n.sp. 

Shape,  etc. — The  organism  is  a  stout,  motile,  short  rod,  with 
rounded  ends.  The  dried  and  imbedded  bacteria  when  taken 
from  cultures  on  saccharose-potato  agar  appear  as  oval  cells, 
measuring  0-8  :  1-2  /z,  and  as  rods  measuring  1  :  2  /^t.  On  the  oval 
cells  the  flagella  are  generally  terminal,  but  as  the  cell  grows 
longer  many  peritrichous  flagella  can  be  seen,  up  to  seven  have 
been  counted;  but  as  the  growths  on  agar  are  cohesive  many  more 
may  occur  on  the  cells.  Staining  by  Gram's  method  is  negative, 
and  spores  are  not  formed. 

Temperature,  etc.  — The  optimum  temperature  is  about  30°.  At 
37°  the  growth  is  less  than  at  22°  or  15°.  The  bacterium  is 
aerobic,  but  grows  slightly  under  anaerobic  conditions. 

Nutrient  agar  plate. — The  colonies  are  dirty-white,  raised  and 
rough.  When  magnified  they  appear  rounded,  irregular  or 
lobular,  with  a  smooth  edge  and  convoluted,  puckered  or  crinkled 
centre.  The  deep  colonies  appear  moruloid  or  very  irregular,  as 
if  several  colonies  had  fused  together  at  different  parts.  As  the 
colony  increases  in  size  the  colour  deepens  to  a  yellowish-bnff. 

Glucose-gelatine  plate. — The  colonies  are  ivory-white,  dry  and 
puckered,   and  have  a  very  cohesive   consistency;  it  is   almost 

*  The  slime-flux  or  mucilage-flux  and  the  gum-flux  of  certain  trees  have 
been  ascribed  to  the  action  of  certain  Ascomycetes  by  Ludwig  and  by 
Beijerinck.  In  view  of  my  research  it  is  possible  that  in  some  cases  at  least 
these  ascomycetes  have  appeared  in  the  flux  at  a  period  subsequent  to  its 
formation  by  bacteria. 
27 


406  BACTERIAL    ORIGIN    OF    GUxMS    OF    ARABIX    GROUP, 

impossible  to  pick  up  a  fragment  of  the  young  colony  with  the 
needle,  the  whole  colony  comes  away.  Microscopically,  the  colonies 
are  dark,  very  coarsely  granular  and  crinkled.  The  margin  is 
indefinite  on  account  of  the  consumption  of  the  medium.  The 
deep  colonies  are  granular  and  very  irregular.  The  colour  deepens 
to  a  yellowish  tint  and  the  medium  is  slowly  liquefied. 

Nutrient  gelatine  plate.  —  As  on  glucose-gelatine,  but  the 
growth  is  poorer. 

Nutrient  agar  stroke. — A  narrow,  slightly  raised,  rough,  dry, 
glistening,  lobular,  yellowish-white  growth  forms  at  30°;  it 
adheres  firmly  to  the  medium.  The  condensed  water  carries  a 
film  from  which  the  rough  growth  spreads  upwards. 

Glucose-gelatine  stroke. — A  translucent  white,  very  wrinkled, 
rough,  dry,  dull  growth,  with  an  irregular  margin  is  formed  at 
22°.  A  pit  forms  at  the  base  of  the  stroke  owing  to  the  con- 
sumption of  the  gelatine  at  the  place  of  strongest  growth.  The 
medium  slowly  liquefies  and  gravitates,  producing  an  appearance 
like  the  clear  condensed  water  of  an  agar  tube.  The  culture 
coheres  as  a  wrinkled  skin.  On  gelatine  the  colour  is  always 
white  or  very  pale  yellow;  while  on  agar  it  changes  from  white 
to  yellow  or  bufi". 

Glucose-gelatine  stab. — The  stab  becomes  filiform-tuberculate. 
The  nail-head  is  sunken  and  a  rough  growth  lines  the  surface  of 
the  depression.  Below  the  consumed  medium  the  gelatine  may 
be  slightly  and  locally  liquefied.  Gas  bubbles  are  formed  in  the 
medium;  the  gas  formation  is  more  pronounced  in  saccharose- 
(^elatine,  on  plates  of  saccharose-potato  agar,  and  in  saccharose- 
potato  extract. 

Nutrient  gelatine  stab. — As  in  glucose-gelatine  but  poorer. 

Saccharose-potato  agar  stroke. — The  appearance  of  the  culture 
depends  upon  the  incubation  temperature.  At  30°  in  24  hours 
the  growth  is  luxuriant  and  has  spread  over  the  greater  portion 
of  the  slope.  It  is  undulating,  much  raised,  glistening,  granular 
and  apparently  very  slimy.  The  colour  is  the  same  as  the 
medium,  viz.,  buff  or  very  light  oak.  The  condensed  water  has 
been  absorbed.     In  48  hours  the  growth  has  spread  entirely  over 


BY    R.    GREIG    SMITH.  407 

the  agar  surface.  At  37°  in  24  hours  the  growth  is  broad,  raised, 
_^  glistening,  with  a  translucent  margin  and  opaque  centre.  The 
culture  is  apparently  gummy.  The  condensed  water  has  a  film. 
In  48  hours  the  growth  by  reflected  light  appears  homogeneous 
and  the  colour  of  the  medium.  At  22°  in  24  hours  the  growth 
is  broad,  whitish,  very  rough  and  undulating.  The  margin  is 
more  glistening  than  the  centre.  The  condensed  water  carries  a 
film.  In  48  hours  the  growth  is  mottled  and  pale  buff".  At  15° 
the  growth  is  as  at  22°,  but  less  luxuriant.  On  this  medium  at 
22°  the  growth  is  very  like  that  of  Bad.  acacice^  but  differs  in 
having  a  lumpy  (gelatinous)  consistency,  while  that  of  Bact. 
acacice  is  homogeneous.  At  the  higher  temperature  the  differ- 
ences are  more  pronounced. 

Potato. — A  dry,  pale  buff  scattered  growth  spreads  irregularly 
over  the  medium.  It  is  raised  and  becomes  undulatino-.  A 
glistening  appearance  begins  at  the  margin  and  spreads  inwards 
as  the  colour  deepens  to  orange-yellow. 

Bouillon. — The  medium  becomes  turbid   and  forms  a  strong 

o 

surface  ring  and  a  floating  puckered  film.  The  indol  reaction 
was  obtained,  but  nitrite  is  not  produced  as  with  Bact.  acacice. 
In  nitrate-bouillon  the  nitrate  is  reduced  to  nitrite. 

Milk. — The  medium  slowly  coagulates  and  the  reaction  is 
faintly  acid. 


40S 


REVISION  OF   THE   AUSTRALIAN   CURCULIONID^ 

BELONGING  TO  THE  SUBFAMILY 

CR  YPTORHYNCHIDES. 

By  Arthur  M.  Lea. 

Part  V. 

This  part  deals  with  the  genus  Cryptorhynchus  and  some  of 
the  allied  genera;  of  these  34  genera  and  93  species  have  been 
described  and  tabulations  of  the  genera  and  species  have  been 
prepared,  but  as  the  whole  is  too  length}'-  for  publication  in  one 
part  only  a  portion  of  the  genera  and  species  are  now  described, 
and  the  tabulation  of  the  genera  is  withheld  till  the  whole  of  the 
section  is  completed. 

Genus  Cryptorhynchus,  Illiger. 

Illig.  Magaz.  vi.  p.  330;  Schonherr,  Gen.  Sp.  Cure.  iv.  p.  47, 
No.  304;  Lacordaire,  Gen.  Col.  viii.  p.  121. 

Head  of  moderate  size,  convex,  not  concealed.  Eyes  moderately 
large,  ovate,  separation  less  than  width  of  rostrum,  moderately 
coarsely  faceted.  Rostrum  moderately  long  and  not  very  thin, 
distinctly  curved.  Scape  inserted  nearer  apex  than  base  of  ros- 
trum, slightly  shorter  than  funicle;  two  basal  joints  of  the  latter 
moderately  long;  club  elliptic-ovate.  Prothorax  transverse,  sides 
rounded,  base  feebly  bisinuate,  constriction  feeble;  ocular  lobes 
obtuse.  Scutellum  distinct.  Elytra  oblong-cordate,  wider  than 
prothorax,  each  rounded  at  base.  Pectoral  canal  deep  and  rather 
narrow,  terminated  between  intermediate  coxae.  Mesosternal 
receptacle  feebly  raised,  Y-shaped,  walls  thin;  feebly  cavernous. 
Metasternum  not  much  shorter  than  the  following  segment ; 
episterna  distinct  and  rather  wide.  Basal  segment  of  abdomen 
shorter    than    the    two    following  combined,   intercoxal  process 


BY    ARTHUR    M.    LEA.  409 

rather  narrow;  211(1  just  perceptibly  longer  than  3id  or  4th  and 
(like  them)  drawn  slightly  backwards  at  the  sides.  Legs  rather 
short  and  stout;  femora  very  feebly  grooved,  dentate,  posterior 
not  passing  elytra;  tibiae  feebly  compressed,  straight  except  at 
extreme  base;  tarsi  slightly  shorter  than  tibiae,  1st  and  4th  joints 
equal,  3rd  wide  and  deeply  bilobed.  Oblong-elliptic,  convex, 
squamose,  winged. 

The  above  diagnosis  is  drawn  up  from  European  specimens  of 
the  typical  C.  Lapathi,  and  with  which  the  two  species  described 
below  are  decidedly  congeneric.  Of  the  sixteen  species  referred 
to  Cryptorhynchus  and  standing*  under  that  name  in  Masters' 
^Catalogue'  I  am  unacquainted  with  australis,  corosus,  teter,  longi- 
manus,  rncestas  and  solidus,  in  all  six  species.  Some  of  these  six 
species  may  belong  to  Cryptorhy7ichus,  but  the  genus  of  each  one 
of  them  is  very  doubtful.  Of  the  others  stigmaticus  is  a  true 
Cryptorhynchus  and  is  redescribed  below;  Uthodermus  and  suc- 
cisits  belong  to  Poropterus;  ephippiger  to  Camptorrhinus^  alhicollis 
to  Metyrios,  cariosus  to  Exithius,  femoralis  to  Amydala,  sirius  and 
antares  to  a  new  genus,  and  infulatus  also  to  a  new  genus. 

Elytra  with  six  spots  at  summit  of  posterior  declivity stigmaticus,  Fasc. 

Elytra  f asciate  at  summit  of  declivity vems,  n.  sp. 

Cryptorhynchus  stigmaticus,  Pasc;  Mast.  Cat.  No.  5550. 

Brownish-black,  antennae  dull  red.  Eather  sparsel}'-  clothed 
(except  on  scutellum,  where  the  clothing  is  dense  and  rather  pale) 
with  small  scales  varying  from  a  dingy  whitish-grey  to  brown, 
each  elytron  with  three  small  spots  about  summit  of  posterior 
declivity,  the  median  ones  slightly  anterior  to  the  two  lateral  ones 
and  which  are  on  the  3rd  and  5th  interstices.  Under  surface 
with  not  very  sparse  whitish  scales;  legs  with  thin  whitish  scales, 
denser  at  apex  of  tibise  than  elsewhere.  Head  with  sparse  scales, 
thin  in  front,  rounded  behind;  sides  (behind  antennse)  and  base 
of  rostrum  with  thin  scales. 


*  Except  O.  solidus,  accidentally  omitted. 


410  REVISION    OF   THE    AUSTRALIAN    CURCULIONIDiE, 

Head  with  not  very  dense  punctures.  Rostrum  the  length  of 
prothorax,  sides  scarcely  visibly  incurved  to  middle;  basal  third 
rather  strongly  punctate,  sides  elsewhere  moderately  strongly 
punctate.  Scape  inserted  slightly  nearer  apex  than  base  of 
rostrum:  1st  joint  of  funicle  stouter  and  slight^  shorter  than  2nd. 
Prothorax  rather  strongly  transverse,  apex  less  than  half  the 
width  of  base;  with  a  dense  slightly  shining  seta  bearing  granules, 
becomingveryindistinct  anteriorly;  with  a  narrow  and  distinct  but 
not  quite  continuous  median  carina.  Elytra  slightly  wider  than 
prothorax;  with  series  of  large  punctures,  each  of  which  is  deep 
only  in  its  middle;  interstices  rather  densely  granulate,  the 
alternate  ones  scarcely  visibly  raised  above  but  with  denser 
granules  than  the  others.  Under  surface  indistinctly  punctate. 
Femora  thin  and  very  feebly  dentate.  Length  9,  rostrum  3; 
width  4|;  variation  in  length  8-9  mm. 

Hah. — "  Queensland  "  (Pascoe),  Rockhampton  (Macleay  Mus.), 
Upper  Endeavour  River  (Mr.  C.  French). 

Cryptorhynchus  verus,  n.sp. 

Brownish-black,  antennae  dull  red.  Moderately  densel}'-  clothed 
with  scales  mostly  of  an  ochreous-fawn,  but  varying  from  white 
to  brown;  scutellum  with  dense  whitish  scales;  elytra  with  an 
irregular  whitish  fascia  at  summit  of  posterior  declivity.  Under 
surface  with  rounded  scales  varying  from  white  to  fawn;  legs 
with  whitish  scales.  Head,  base  and  sides  of  rostrum  with 
ochreous-fawn  scales. 

Head  with  dense  concealed  punctures.  Rostrum  rather  wide, 
feebly  curved  and  scarcel}^  the  length  of  prothorax;  basal  third 
coarsely  punctate,  elsewhere  shining  and  (except  at  sides)  very 
lightly  punctate.  Scape  inserted  slightly  nearer  apex  than  base 
of  rostrum;  1st  joint  of  funicle  slightly  longer  than  2nd.  Pro- 
thorax lightly  transverse,  apex  more  than  half  the  width  of  base; 
granules  rather  less  numerous  and  the  setae  less  distinct  than  in 
C.  stigmaticus;  median  carina  continuous  to  base  and  almost  to 
apex.  Elytra  slightly  wider  than  prothorax;  with  series  of  rather 
large  punctures;  alternate  interstices  feebly  raised  and  all  with 


BY    ARTHUR    M.    LEA.  411 

series  of  very  obtuse  granules.  TJndar  surface  with  more  distinct 
punctures  than  in  C  stigmaticus;  the  femora  stouter  but  just  as 
feebly  dentate.  Length  8J,  rostrum  2|;  width  4;  variation  in 
length  8-9  mm. 

Zrrt6.— Australia  (Herr  J.  Faust);  N.W.  Australia  (Mr.  G. 
Masters) — Q.  :  Cairns  (Macleay  Mus.). 

Close  to  the  preceding  species,  but  considerably  narrower,  the 
rostrum  shorter  and  wider  and  the  two  basal  joints  of  the  funicle 
differently  proportioned.  The  granules  also  are  different, 
especially  on  the  elytra.  The  clothing  is  much  the  same  except 
that  it  is  denser  and  that  the  elytra  are  supplied  with  a  fascia 
instead  of  six  spots  at  the  summit  of  the  posterior  declivity. 

Genus  Tyrt^osus,  Pascoe. 

Trans.  Ent.  Soc.  1870,  p.  479. 

Head  convex,  ocular  fovea  indistinct.  Eyes  subtriangular, 
rather  large,  coarsely  faceted.  Rostrum  rather  long  and  thin,  but 
sometimes  comparatively  short  and  stout,  distinctly  curved. 
Antennce  moderately  stout;  scape  usually  inserted  nearer  apex 
than  base  of  rostrum,  shorter  than  funicle;  1st  joint  of  funicle 
stout  but  moderately  long;  club  ovate.  Prothorax  subconical, 
constriction  feeble,  sides  rounded,  base  sinuate,  ocular  lobes  pro- 
minent. Scutellum  subtriangular  or  subquadrate.  Elytra  more 
than  twice  the  length  of  prothorax  and  usually  distinctly  wider 
at  base,  cordate  or  oblong-cordate.  Pectoral  canal  deep  and 
narrow,  terminated  between  four  anterior  coxae.  Mesosternal 
receptacle  moderately  raised,  either  semicircular  or  shaped  like  a 
short  Y,  rather  narrow  at  base,  emargination  usually  distinctly 
transverse;  cavernous.  Metasternum  subequal  in  length  to  the 
following  segment;  episterna  rather  wide.  Abdomen  with  straight 
distinct  sutures ;  1st  segment  large,  intercoxal  process  rather 
narrow,  3rd  and  4th  conjoined  as  long  as  or  slightly  longer  than 
2nd  or  5th.  Legs  moderately  long;  femora  rather  stout,  dentate 
(but  occasionally  edentate),  with  a  more  or  less  distinct  lateral 
groove  to  receive  tibiae,  posterior  not  extending  to  apex  of  abdo- 
men; tibiae  short,  grooved   their  entire  length,  the  grooves  less 


412  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

distinct  on  the  anterior  pair,  usually  slightl}'  curved;  tarsi  rather 
short,  claw-joint  elongate.  Elliptic  or  elliptic-ovate,  convex, 
striate,  moderately  squamose,  winged. 

This  genus  is  very  close  to  Cryptorhynchus;  compared  with  the 
typical  C.  Lajmthi  it  differs  in  having  the  mesosternal  receptacle 
raised  in  front  and  much  more  cavernous,  the  prothorax  more 
decidedly  bisinuate  at  the  base,  the  tibiae  sulcate-striate  or  punc- 
tate and  the  clothing  never  fasciculate.  The  two  genera,  how- 
ever, are  remarkably  close,  but  it  appears  to  be  as  well  not  to 
degrade  Tyrtceosus  into  a  section  of  Cryptorhynchus,  as  the  latter 
is  in  anything  but  a  satisfactory  state. 

The  majority  of  the  species  occur  in  Australia  or  Tasmania, 
but  several  are  recorded  from  New  Guinea.  All  are  arboreal  and 
very  homogeneous  in  their  general  appearance.  The  sexual  charac- 
ters are  either  very  slight  or  else  are  so  strong^  marked  as  to 
cause  the  sexes  to  appear  as  distinct  species.  I  believe,  however, 
that  they  are  very  slight.  The  ciliation  of  the  ocular  lobes  in  all 
the  species  is  dense,  verj'  short  and  silvery,  and  appears  as  a 
white  rim  when  the  head  is  removed;  with  head  in  position  it  is 
traceable  only  with  extreme  difficult}^  It  has  not  been  made 
use  of  here. 

Femora  edentate. 

Elytral  interstices  impunctate.. inerm'u,  n.sp. 

Ely tral  interstices  densely  punctate crassirostris,  n,  sp. 

Elytral  interstices  divided  into  squares  each  con- 
taining a  puncture quadratoUneatns,n. sip. 

Anterior  femora  edentate urens,  n. sp. 

Femora  dentate. 

Punctures   of   metasternal   episterna    continuous 
throughout. 
Sutural  interstices  each  with  a  row  of  rather 

large  punctures i)nitator,  n.  sp. 

Interstices  regular. 

Each  of  3rd  and  4th  abdominal  segments  Nvith 
with  one  row  of  punctures  in  middle. 
Kostrum  with  strong  punctures  at  base  only. 
Funicle  with  1st  joint  slightly  longer  than 

2nd incallidux,  Pasc. 


BY    ARTHUR    M.    LEA.  413 

Funicle   with  1st  joint  slightly  shorter 

than  2nd castor,  n.sp. 

Rostrum  with  distinct  punctures  throughout    doloms,  n.sp. 
Each  of  the  3rd  and  4th  abdominal  segments 
with  two  rows  of  punctures  or  densely 
punctate. 

Prothorax  very  distinctly  carinate cai'inatus,  n. sp. 

Prothorax  feebly  carinate. 
Elytra  slightly  and  not  suddenly  wider 

than  prothorax ichthyosomus,  n.sp. 

Elytra  distinctly  and  suddenly  wider. 
Elytra  densely  clothed  with  brick-red 

scales laterarius,  n.sp. 

Elytra  much  less  densely  clothed pollux,  n.sT^. 

Punctures  of  metasternal  episterna  not  continuous 
throughout. 
Each  of  the  .3rd  and  4th  abdominal  segments 
with  one  row  of  punctures  in  middle. 
Prothorax    comparatively   small   and   feebly 

transverse microthorax,  Pasc. 

Prothorax  larger  and  more  distinctly  trans- 
verse. 

Antennse  almost  black corpulentus,  n.  sp. 

Antennae  red. 

Head  impressed  behind  each  eye vetustus,  Pasc. 

Head  not  so  impressed. 
Each  of  the  .3rd  and  4th  abdominal 
segments  with  irregular  punctures 

and  elongate  scales ustulatus,  Pasc. 

These  segments  with  regular  punctures 

and  rounded  scales ccmulus,  n.  sp. 

Each  of  the  3rd  and  4th  abdominal  segments 
with   two    rows   of  punctures   or   densely 
punctate. 
Funicle  with  1  st  joint  slightly  shorter  than  2nd     cinerascens,  n.  sp. 
Funicle  with  1st  joint  the  length  of  or  slightly 
longer  than  2nd. 
Eyes  separated  almost  the  width  of  rostrum 
at  base. 
Elytra  (viewed  in  profile)  perfectly  flat  at 

base ceqmis,  n.  sp. 

Elytra  distinctly  convex  from  base hiseriatus,  n.sp. 


414  REVISION    OF    THE    AUSTRALIAN    CURCULlONIDiE, 

Eyes  separated  considerably  less  than  width 
of  rostrum  at  base. 

.  Femora  feebly  dentate assimilis,  n.s^. 

Femora  strongly  dentate. 

Elytral  interstices  with  punctures  along 

middle imrdalU,  Pasc. 

Elytral  interstices  without  such  punc- 
tures      religiosns,  n.sp. 


TvRTiEOSUS  MiCROTHORAX,  Pasc.  I  l.c.  No.  555 


Black;  antenn£e  and  claws  dull  piceous-red.  Rather  unevenly 
clothed  with  scales  of  varying  shades  of  brown  and  yellow;  vary- 
ing in  shape  from  short,  quadrate  or  spatulate,  to  elongate  seta?. 
Head  with  pale  brown  scales  at  base  of  rostrum  and  behind  eyes; 
prothorax  with  pale  brown  setose  scales  and  a  few  shorter  and 
almost  white  at  sides;  disc  with  three  feeble  longitudinal  stripes 
of  setose  scales ;  scutellum  with  brownish  scales ;  elytra  with 
rather  small  scales,  a  large  very  irregular  patch  of  pale  yellowish 
scales  near  the  sides  and  extending  obliquely  backwards  to  near 
suture,  then  irregularly  to  apex,  pale  brown  ones  at  base,  along 
suture  and  sides,  a  few  small  patches  of  dark  brown  ones  scattered 
about,  short  pale  setse  at  apex  and  sides;  sterna  and  legs  with 
long  pale  setose  scales;  metasternal  episterna  with  embedded 
scales  at  each  end;  abdomen  with  pale  and  rather  setose  scales 
on  1st  segment,  2nd  with  shorter  sooty-brown  scales  and  a  few 
pale  ones  scattered  about,  3rd  and  4th  each  with  one  distinct 
row  of  sooty  scales  and  a  few  smaller  and  paler  ones  scattered 
about,  their  sides  with  paler  and  denser  scales. 

Head  densely  punctate;  a  shining  distinct  carina  from  base  to 
ocular  fovea  (which  is  partially  concealed);  a  depression  on  each 
side  behind  eyes.  Rostrum  shining;  almost  parallel  to  near  apex 
where  it  is  distinctly  widened;  coarsely  punctate  at  base,  the 
punctures  decreasing  in  size  and  number  to  apex.  Funicle  with 
lst-2nd  joints  equal,  moderately  elongate,  the  length  of  4th-6th 
combined,  6th-7th  globular.  Prothorax  transverse,  base  bisinuate;. 
a  shining  median  carina  continuous  from  base  to  apex,  but  partially 
concealed   by  scales;  densely  punctate   and   with  small  shining. 


BY    ARTHUR    M.    LEA.  415- 

oblong  or  rounded  granules.  Elytra  cordate,  much  wider  than^ 
prothorax,  and  about  four  times  its  length ;  base  trisinuate  ; 
shoulders  rounded;  punctures  almost  concealed  by  scales  and 
apparently  very  shallow;  interstices  very  narrow,  shining,  raised 
and  keel-like,  ten  on  each  elytron,  the  first  appearing  as  a  row  of 
elongate  granules;  apex  with  small  shining  granules.  Mesosternal 
receptacle  slightly  angular.  Metasternum  rather  coarsely  punctate;; 
episterna  with  moderately  large  punctures  at  their  bases  and 
fewer  and  smaller  at  their  apices.  Abdomen  with  moderately 
dense  punctures  on  1st  and  5th  segments,  fewer  on  2nd,  and 
forming  a  transverse  row  across  each  of  the  3rd  and  4th,  these 
closely  joined  together  but  sutures  distinct.  Femora  moderately 
thick,  posterior  almost  reaching  apical  segment,  teeth  in  all  dis- 
tinct and  rather  sharp;  tibiae  short,  grooved,  base  strongly  curved, 
the  posterior  widening  to  apex,  apex  itself  obliquely  truncate  and 
very  thin.     Length  8,  rostrum  24;  width  4jmm. 

//a6.— S.  Queensland  (Mr.  G.  Masters). 

This  is  a  rather  prettily  marked  species  with  very  narrow 
elytral  interstices.     The  carina  on  the  head  is  very  distinct. 

Tyrt.eosus  vetustus,  Pasc;  I.e.  No.  5560. 

Black;  antennae  and  claws  dull  piceous-red.  Base  of  rostrum,, 
between  and  behind  eyes  with  short  pale  scales;  prothorax  with 
dull  brown  scales  embedded  in  punctures  and  seldom  rising 
above  the  general  level,  the  sides  with  larger  pale  and  broader 
scales;  scutellum  with  pale  scales;  elytra  with  two  transverse 
fascia3  of  short  pale  scales  interrupted  towards  and  very  irregular 
about  suture:  one  near  apex  and  one  near  base;  duller,  smaller 
and  scarcely  visible  scales  scattered  elsewhere;  metasternal  epi- 
sterna with  embedded  scales  at  each  end;  sterna  and  abdomen 
with  large  embedded  scales,  pale  but  becoming  darker  on  abdomen, 
3rd  and  4th  segments  each  with  one  distinct  row  of  dull  scales 
and  a  much  feebler  scarcely  traceable  row. 

Head  densely  punctate,  ocular  fovea  indistinct;  eyes  large,  pro- 
minent, subcontiguous.  Rostrum  feebly  shining,  parallel  almost 
to  apex;  base  coarsely  punctate,  the  punctures  gradually  decreas- 


416  REVISION    OF    THE    AUSTRi^LIAN    CUKCULIONIDA:, 

ing  in  size  to  apex.  Scape  inserted  slightly  nearer  apex  than 
base  of  rostrum;  funicle  rather  thick,  joints  lst-2nd  briefly  obtri- 
-angular,  3rd-7th  transverse,  their  sutures  indistinct.  Prothorax 
transverse;  with  dense  round  punctures;  a  shining,  slightly  inter- 
rupted median  carina  continued  from  base  to  apex.  Elytra 
oblong-cordate,  wider  than  prothorax  and  about  thrice  its  length; 
punctate-striate,  punctures  subquadrate  and  rather  shallow:  inter- 
stices narrow,  raised,  and  appearing  as  rows  of  oblong  granules, 
terminating  at  apex  in  bead-like  granules.  Metasternwrn  as  in 
preceding.  Abdomen  with  1st,  2nd,  and  5th  segments  rather 
densely  punctate;  the  3rd  and  4th  with  regular  punctures,  their 
sutures  wide.  Femora  more  slender  than  in  preceding,  the  teeth 
(except  posterior)  smaller  but  sufficient!}'  distinct ;  posterior 
reaching  middle  of  apical  segment;  tibiae  short,  strongly  curved 
at  base,  the  posterior  slightly  wider  at  their  apices  than  the 
anterior.     Length  7,  rostrum  1|;  width  3|  (vix)  mm. 

Hah. — Queensland,  Victoria,  S.  Australia. 

In  this  species  the  eyes  are  perhaps  closer  together,  larger  and 
more  coarsely  faceted  than  in  any  other  of  the  genus.  One  of 
my  specimens  was  received  from  Herr  J.  Faust  and  marked  as 
having  been  so  named  by  Mr.  Pascoe. 

TYRTiEOSUS    RELIGIOSUS,  n.sp. 

Black;  antennae  dull  reddish-piceous,  claws  dull  red.  Head 
and  rostrum  with  pale  scales,  more  distinct  about  eyes  than  else- 
w^here;  prothorax  with  pale  ochreous  setose  scales  rising  above 
punctures,  more  regular,  shorter  and  slightly  paler  on  sides  than 
on  disc;  scutellum  v/ith  very  small  pale  scales;  elytra  with  short 
ochreous  scales,  unevenly  distributed  and  denser  towards  apex 
and  base  than  elsewhere,  seen  from  above  with  semi-nude  spaces 
forming  an  irregular  Maltese  cross;  metasternal  episterna  with 
embedded  scales  at  each  end;  sterna  and  two  basal  segments  of 
abdomen  with  moderately  elongate  pale  scales,  3rd  and  4tli  each 
with  two  or  more  irregular  rows  of  sooty  and  pale  scales  inter- 
mingled together. 


BY    ARTHUR    M.    LEA.  417 

Head  punctate  and  granulate;  ocular  fovea  concealed;  eyes 
rather  large.  Hostrura  subparallel,  moderately  shining,  coarsely 
punctate,  the  punctures  smaller  and  more  elongate  at  apex  than 
base.  Scape  inserted  about  two-iifths  from  apex  of  rostrum;  two 
basal  joints  of  funicle  equal,  moderately  elongate,  the  rest  trans- 
verse. Prothorax  slightly  transverse,  sides  rounded,  increasing 
to  base;  a  shining  median  carina  continuous  throughout;  with 
deep  round  punctures,  the  surface  scarcely  cut  up  into  granules.. 
Elytra  slightly  wider  than  prothorax  and  about  four  times  its 
length;  punctate-striate,  punctures  separated  by  granules,  con- 
cealed or  not;  interstices  narrow,  raised,  continuous  throughout 
or  cut  up  into  elongated  granules;  minute  granules  about  suture 
and  apex.  Mesosternal  receptacle  and  metasternum  as  in  preced- 
ing, punctures  of  episterna  almost  continuous.  Abdomen  densely 
punctate.  Posterior  femora  extending  to  apical  segment;  tibiae 
short,  curved  at  extreme  base,  rather  thin,  grooved,  jDosterior 
widening  to  apex.     Length  9,  rostrum  2J;  width  4|  mm. 

Hah. — Q. :  Cooktown  (Herr  J.  Faust). 

May  be  distinguished  from  microthorax  by  its  noncarinate  head 
and  from  the  preceding  by  its  less  parallel  outline. 

Tyrt^osus  pardalis,  Pasc;  Lc.  No.  5558. 

Piceous-black;  elytra  dark  reddish-brown;  legs,  rostrum  and 
antenna?  dull  piceous-red,  claws  dull  red.  Head  about  eyes  and 
base  of  rostrum  with  short  ochreous  scales;  prothorax  with  dull 
ochreous,  scarcely  visible  scales  embedded  in  punctures  and  not 
rising  to  general  level,  except  in  three  scarcely  traceable  lines  at 
the  sides;  near  apex  the  scales  are  larger,  paler  and  much  more 
distinct;  elytra  with  ochreous  scales  very  irregularly  distributed 
and  forming  small  patches  between  the  interstices;  metasternal 
episterna  with  scales  at  each  end ;  under  surface  uniformly 
clothed  with  short  spatulate  pale  yellowish  scales;  two  distinct 
rows  on  each  of  the  3rd  and  4th  abdominal  segments. 

Head  densely  punctate,  ocular  fovea  concealed.  Rostrum 
shining,  parallel  almost  to  apex;  base  densely  punctate.  Scape 
inserted  slightly  in  advance  of  middle;  1st  joint  of  funicle  slightly 


418  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

longer  than  2nd,  3rcl-7th  transverse.  Prothorax  scarcel}^  wider 
than  long,  apex  narrow,  sides  rounded  and  feebly  increasing  to 
base  ;  median  carina  shining,  not  continued  to  apex  ;  densely 
punctate,  punctures  deep  and  round.  Scutellum  almost  circular. 
Elytra  suboblong-cordate,  slightly  wider  than  prothorax  and  not 
thrice  its  length ;  shoulders  feebly  rounded  ;  punctate-striate, 
punctures  partially  concealed  ;  interstices  moderately  narrow, 
raised,  interrupted  throughout  and  appearing  on  each  elytron  as 
nine  rows  of  elongate  punctured  granules.  Under  surface  densely 
punctate;  metasternal  episterna  with  a  shining  impunctate  space 
near  apex;  intermediate  segments  rather  narrow,  their  sutures 
wide.  Vo^teviov  femora  reaching  apical  segment:  tibiae  grooved, 
base  not  very  strongly  curved,  the  posterior  wider  than  the  four 
anterior;  claw-joint  and  claws  smaller  than  usual.  Length  5f, 
rostrum  If;  width  2f  mm. 
Hah. — N.  Queensland. 

Tyrt^osus  biseriatus,  n.sp. 

Piceous-black  ;  legs,  rostrum  and  elytral  interstices  paler, 
antennse  pale  red.  Head  with  ochreous  and  brown  scales  inter, 
mingled;  prothorax  with  three  moderate!}'  distinct  longitudinal 
stripes  of  ochreous  scales,  sides  with  larger  and  more  regular 
scales;  scutellum  densely  squamose,  the  scales  very  small  and 
whitish  ;  elytra  with  ochreous  scales  giving  them  a  spotted 
appearance;  metasternal  episterna  with  embedded  scales  at  each 
end;  scales  of  under  surface  uniform  in  colour,  each  of  the  3rd 
and  4th  abdominal  segments  with  two  distinct  rows  of  obtrian- 
gular  scales. 

Head  punctate  and  granulate;  ocular  fovea  invisible;  eyes 
rather  widely  separated.  Rostrum  parallel,  highly  polished,  a 
feeble  carina  extending  almost  its  entire  length,  elsewhere  rather 
coarsely  punctate.  Funicle  with  1st  joint  obtriangular,  slightly 
longer  than  2nd,  the  others  transverse  and  feebly  widening  to 
apex;  club  large.  Protliorax  feebly  transverse;  densely  granulate, 
the  granules  subopaque;  with  a  feeble  uninterrupted  median  carina. 
Elytra  wider  than  prothorax  and   scarcely   thrice    its    length ; 


BY    ARTHUR    M.    LEA.  419 

punctures  marked  by  scales,  interstices  feebly  raised,  scarcely 
distinguishable.  Under  surface  moderately  densely  punctate; 
abdominal  segments  with  rather  narrow  sutures,  the  second  with 
about  three  rows  of  moderately  regular  punctures,  each  of  the 
3rd  and  4th  with  two  distinct  and  very  regular  rows.  Posterior 
femora  scarcely  extending  to  middle  of  4th  segment,  the  teeth  in 
all  rather  small;  tibiae  short,  grooved,  equal,  parallel  except  at 
base.     Length  4,  rostrum  1;  width  2  mm. 

ZTaS.— N.S.W.:  Gosford. 

On  a  first  glance  the  elytral  interstices  appear  to  be  wider  than 
the  punctures,  but  this  is  not  the  case,  although  they  are  wider 
than  usual;  from  the  following  species  (which  it  closely  resembles) 
its  larger  size,  more  convex  form,  and  shining  rostrum  will 
sufficiently  distinguish  it. 

Tyrt.eosus  usTULATUs,  Pasc. ;  Ix.  No.  5559. 

Piceous-black;  antennae  and  tarsi  dark  reddish-brown.  Ros- 
trum with  setose  scales  throughout;  prothorax  with  three  feeble 
longitudinal  stripes  of  pale  setose  scales;  elytra  with  griseous 
scales  forming  very  ill-defined  fascitis  and  variable  in  pattern, 
sparser  at  base  and  behind  the  middle  than  elsewhere;  meta- 
sternal  episterna  with  embedded  scales  at  each  end,  each  of  the 
3rd  and  4th  abdominal  segments  with  elongate  scales  forming  at 
least  two  very  irregular  rows. 

Head  granulate  and  punctate;  ocular  fovea  invisible;  eyes 
rather  widely  separated.  Rostrum  opaque,  parallel  to  near  apex, 
coarsely  punctate  throughout.  Prothorax  feebly  transverse,  sides 
feebly  rounded;  with  deep  round  punctures  giving  the  surface  a 
granulated  appearance;  median  carina  scarcely  traceable.  Scii- 
tellum  oblong,  smooth,  slightly  raised.  Elytra  about  once  and 
one-fourth  the  width  of  prothorax  and  not  thrice  its  length; 
slightly  the  widest  behind  middle;  shoulders  rounded;  striate- 
punctate,  punctures  rather  large,  sufficiently  distinct;  striae 
apparently  wider  than  interstices,  these  raised  and  appearing  as 
rows  of  oblong  granules.  Mesosternal  rece'ptade  rather  wide. 
Under  surface  coarsely  punctate,  punctures  and  scales  of  3rd  and 


420  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

4th  abdominal  segments  very  irregular.  Femora  rather  thin, 
feebly  dentate,  the  posterior  extending  to  middle  of  apical  seg- 
ment; tibia?  grooved,  equal.     Length  4,  rostrum  f;  width  If  mm. 

Hah. — Tasmania. 

The  rostrum  is  coarsely  punctate  throughout;  the  femora  are 
long  and  rather  thin,  the  anterior  pair  being  very  feebly  dentate. 

Tyrt.eosus  iEMULUS,  n.sp. 

Differs  from  the  preceding  species  in  having  elytral  scales  very 
much  sparser  and  slightly  more  elongate  (and  under  a  microscope 
seen  to  be  very  different  in  shape);  the  scales  on  the  intermediate 
segments  almost  circular  and  forming  one  distinct  and  very  regular 
row ;  rostrum  slightly  shorter,  much  less  coarsely  and  more 
regularly  punctate,  less  densely  squamose  and  with  a  shining 
carina;  the  elytral  punctures  are  coarser;  the  femora  somewhat 
stouter,  and  the  posterior  pair  do  not  reach  the  apical  segment. 
Length  4,  rostrum  f ;  width  1|  mm. 

^«6._N.S.W.  :  Galston,  Forest  Reefs  (Dumbrell  k  Lea). 

This  species  very  strongly  resembles  ustulatuSj  but  the  differ- 
ences noted  above  appear  to  me  to  be  incompatible  with  specific 
identity.  I  have  five  specimens  agreeing  in  all  particulars  except 
size,  and  that  variable  to  but  a  very  slight  extent  (not  half  a  milli- 
metre between  the  two  extremes).  Of  ustulatus  I  have  under 
examination  two  specimens  kindly  lent  to  me  by  Mr.  Masters. 

Tyrt^osus  dolosus,  n.sp.  (Faust  MS.). 

Black,  elytral  interstices  tinged  with  red;  antennse  dull  reddish- 
piceous;  claws  dull  red.  Head  with  pale  ochreous-brown  scales, 
densest  about  eyes  and  base  of  rostrum;  prothorax  with  three 
feeble  longitudinal  rows  of  ochreous  scales  on  disc,  disc  with  sooty 
black  scales  not  rising  to  general  level  and  scarcely  visible,  the 
sides  with  sooty  and  white  scales  and  large  pale  ochreous  ones 
near  the  apex;  scutellum  densely  squamose,  the  scales  pale;  elytra 
with  sooty  scarcely  visible  scales  and  others  of  an  ochreous-red 
colour  and  irregularly  distributed;  the  darker  spaces  appearing 
as  a  feeble  Maltese  cross;  metasternal  episterna  with  scales  con- 


BY    ARTHUR    M.    LEA.  421 

tiiiuous  throughout;  sterna  and  abdomen  with  short  uniformly 
whitish  scales;  each  of  the  3rd  and  4th  abdominal  segments  with 
a  distinct  row  of  scales  as  the  rest  of  the  abdomen,  and  a  few 
feeble  ones  about  suture. 

Head  densely  punctate  and  very  feebly  carinate  in  front;  ocular 
fovea  concealed ;  eyes  large,  subapproximate.  Rostrum  sub- 
parallel,  densely  punctate,  punctures  naked  towards  apex,  squa- 
mose  towards  basC;  a  feeble  carina  extending  from  base  almost  to 
apex.  Antennae  inserted  slightly  in  advance  of  the  middle;  scape 
short:  basal  joint  of  funicle  obtriangular,  longer  than  2nd,  3rd- 
7th  transverse.  Prothorax  transverse;  densely  punctate,  punc- 
tures round  and  deep;  a  narrow  median  somewhat  irregular 
carina  extending  from  base  to  apex.  Sc^itellum  oblong.  Elytra 
distinctly  wider  than  prothorax  and  not  thrice  its  length ; 
shoulders  slightly  produced,  rounded,  granulate;  punctures  large, 
shallow,  separated  by  feeble  transverse  ridges;  ten  slightly  raised 
and  rather  wavy  interstices  on  each  elytron,  becoming  granulate 
towards  apex.  Mesosternal  receptacle  semicircular.  Metasteriial 
episterna  punctate  throughout,  the  punctures  forming  double  rows 
at  base  and  apex.  Ahdoinen  punctate,  each  of  the  3rd  and  4th 
segments  with  one  distinct  row.  Femora  rather  thick,  posterior 
extending  to  middle  of  apical  segment;  tibiae  short,  grooved,  all 
(but  especially  the  posterior)  dilating  towards  apex.  Length  1\, 
rostrum  2  (vix);  width  4  mm. 
['      Hab.—l^.Q.:  Cooktown  (Herr  J.  Faust). 

Very  strongly  resembles  T.  religiosus,  but  besides  the  carinate 
head  it  is  of  a  shorter  and  broader  build,  with  shorter  and  broader 
rostrum  and  punctures  of  metasternal  episterna  different.  A 
specimen  from  the  Northern  Territory  (probably  Palmerston) 
appears  to  belong  to  this  species,  but  differs  in  being  considerably 
smaller  (5  mm.),  the  clothing  rather  denser  and  paler  and  the 
Maltese  cross  even  less  distinctly  defined. 

Tyrt^osus  incallidus,  Pasc;  I.e.  No.  5555. 

Dark  reddish-brown;  prothorax  and  under  surface  darker, 
antennae  dull  red.      Head  and  base  of  rostrum  with  pale  uniformly 

28 


422  REVISION    OF    THE    AUSTRALIAN    CURCULIONID^, 

distributed  scales;  prothorax  densely  clothed  with  rather  elongate 
pale  scales  all  converging  towards  the  middle;  elytra  with  four 
feeble  much  interrupted  fasciae  of  pale  scales,  one  at  base,  one 
before  and  one  behind  middle,  and  one  at  apex;  metasternal 
episterna  with  scales  continuous  throughout;  under  surface  with 
moderately  large  and  rather  sparse  pale  scales;  3rd  and  4th 
abdominal  segments  each  with  one  distinct  row,  a  few  small  scales 
in  the  prosternal  portion  of  the  pectoral  canal. 

Head  punctate  and  granulate ;  ocular  fovea  shallow ;  eyes 
moderately  separated.  Rostrum  long,  thin,  rounded,  extreme 
base  coarsely  punctate,  elsewhere  highly  polished  and  finely 
punctate.  Antennse  slender;  scape  curved  and  inserted  slightly 
nearer  base  than  apex;  two  basal  joints  of  funicle  elongate,  5th- 
7th  transverse;  club  rather  small.  Prothorax  distinctly  trans- 
verse, sides  feebly  rounded,  apex  not  much  narrower  than  base, 
base  bisinuate;  densely  punctate,  the  punctures  almost  concealed. 
Scutellum  transverse.  Elytra  not  much  wider  than  prothorax 
and  about  thrice  its  length,  subparallel  to  near  apex;  shoulders 
feebly  rounded;  punctures  subquadrate,  separated  by  transverse 
ridges  almost  level  with  interstices;  these  feebly  raised,  rather 
broad,  in  parts  appearing  as  elongated  granules  and  granulate 
towards  apex.  Mesosternal  receptacle  longer  than  wide.  Meta- 
sternal episterna  with  one  continuous  row  of  punctures.  Abdomen 
moderately  densely  punctate,  2nd  segment  in  about  three  rows, 
each  of  the  3rd  and  4th  with  one  distinct  and  regular  row. 
Femora  rather  stout,  teeth  distinct ;  posterior  reaching  apical 
segment;  tibiae  grooved,  anterior  feebly  bisinuate,  four  posterior 
almost  parallel  (except  at  base).  Length  5  (vix),  rostrum  If; 
width  2^-  mm. 

Hab. — S.  Queensland. 

Tyrt^osus  ichthyosomus,  n.sp. 

Dark  reddish-brown;  prothorax,  under  surface  and  rostrum 
darker  ;  antennae  dull  red.  Densely  clothed  with  short  pale 
scales  almost  entirely  concealing  derm,  those  on  prothorax  larger 
.than  on  elytra  and  all  converging  towards  the  middle,  scutellar 


BY    ARTHUR    M.    LEA.  423 

scales  very  minute;  each  of  the  3rd  and  4th  abdominal  segments 
with  two  irregular  rows  in  the  middle  and  more  towards  sides; 
metasternal  episterna  with  scales  continuous  throughout. 

Head  densely  punctate;  ocular  fovea  feeble.  Rostrum  long, 
thin,  densely  punctate  at  base,  feebly  elsewhere,  highly  polished, 
slightly  increasing  in  width  almost  from  base  to  apex.  Antennae 
slender;  scape  inserted  in  exact  middle  of  rostrum;  two  basal 
joints  of  funicle  elongate,  subcylindrical,  as  long  as  those  follow- 
ing combined,  7th  transverse.  Prothorax  transverse;  punctures 
and  median  carina  almost  concealed.  S cutellum  covdia.tQ.  EJytra 
not  much  wider  than  prothorax  and  about  thrice  its  length, 
parallel-sided  to  near  apex;  each  with  about  eight  rows  of  punc- 
tures marked  by  depressions,  but  the  punctures  themselves  con- 
cealed; interstices  apparently  rather  broad,  feebly  raised  and 
punctate.  Mesosternal  receptacle  longer  than  wide.  Metasternal 
episterna  with  punctures  continuous  throughout  and  forming  two 
rows  at  base  and  apex.  Abdomen  moderately  densely  punctate, 
the  3rd  and  4th  segments  each  with  two  very  irregular  rows,  the 
sides  more  densely  punctate.  Femora  almost  linear,  the  grooving 
and  teeth  very  slight,  posterior  not  reaching  apical  segment ; 
tibiae  thin,  anterior  feebly  bisinuate  and  very  feebly  grooved,  the 
four  posterior  almost  straight,  somewhat  wider  and  with  more 
distinct  grooves.     Length  6^,  rostrum  1|;  width  3  mm. 

//a6.— N.S.W.:  Tenterfield  (Mr.  G.  Masters)— Queensland 
(Mr.  C.  French). 

An    uniformly  clothed   and   subparallel    species,   with    longer 

k mesosternal  receptacle,  legs  and  rostrum  than  is  usual. 
TYRT.E0SUS  CARINATUS,  n.sp. 
.  Dull  brownish-black;  legs  dark  reddish-brown,  antenna3  dull 
bed.  Sparsely  clothed  with  dingy  setose  scales ;  metasternal 
episterna  with  setose  scales  continuous  throughout;  each  of  the 
3rd  and  4th  abdominal  segments  with  long  setose  scales  not 
formed  in  rows. 

Head  rather  densely  punctate,  a  distinct  carina  continued  from 
summit  to  about  the  middle  of  rostrum,  from  its  termination  an 


424  REVISION    OF    THE    AUSTRALIAN    CURCULIONID^, 

impunctate  line  to  apex;  eyes  large,  rather  distant.  Rostrum 
parallel-sided,  shining;  densely  punctate  at  base  and  sides,  feebly 
so  elsewhere.  Scape  inserted  much  closer  to  apex  than  base  of 
rostrum;  two  basal  joints  of  funicle  rather  stout,  subequal,  3rd-7th 
feebly  transverse;  club  acuminate.  Prothorax  feebly  transverse, 
apex  rather  narrow,  sides  rounded,  feebly  decreasing  to  base; 
median  carina  very  distinct,  continued  from  base  to  apex;  densely 
punctate,  punctures  deep  and  round.  Scutelluni  small,  subtrian- 
gular.  Elytra  wider  than  prothorax  and  not  thrice  its  lengthy 
elongate-cordate,  base  trisinuate,  shoulders  somewhat  prominent, 
rounded;  interstices  slightly  raised,  rather  broad  and  punctate, 
scarcely  granulate  at  apex.  Mesosternal  recpptacle  feebly  trian- 
gular. Metasternal  episterna  each  with  a  narrow  row  of  punc- 
tures, continuous  throughout.  Abdomen  densely  punctate,  3rd 
and  4th  segments  with  dense  and  very  irregular  punctures. 
Femora  rather  slender,  teeth  small  but  distinct;  posterior  extend- 
ing beyond  middle  of  apical  segment;  tibiae  feebly  grooved,  thin, 
parallel  except  at  base.     Length  4J,  rostrum  4;  width  2  mm. 

^«5._N.S.W.  :  Glen  Innes. 

A  small  species  which  should  be  easily  recognised  on  account 
of  its  distinct  carina  continued  from  head  on  to  rostrum  and 
which  is  not  at  all  interrupted  at  the  position  of  the  ocular  fovea. 

Tyrt^osus  castor,  n.sp. 

Piceous-black ;  antennae  and  claws  dull  red.  Moderately 
densely  clothed  with  ochreous-yellow  scales  mixed  with  paler  and 
sooty  scales;  head  and  base  of  rostrum  densely  squamose;  a  dis- 
tinct median  stripe  on  prothorax  and  a  much  more  feeble  one  on 
each  side;  each  puncture  bearing  a  scale  varying  from  white  to 
ochreous;  elytra  more  densely  squamose  towards  apex  than  else- 
where; usually  with  a  small  whitish  macula  on  4th  interstice 
before  the  middle.  Under  surface  with  paler  and  smaller  scales 
than  above,  a  continuous  row  on  each  of  the  metasternal  episterna 
and  one  row  on  each  of  the  3rd  and  4th  abdominal  segments. 

Head  punctate;  ocular  fovea  absent,  its  place  being  occupied 
by  squamose  punctures;  eyes  rather  large,  depressed  and  distant. 


BY    ARTHUR    M.    LEA.  425 

Rostrum  shining,  sides  slightlj''  incurved  to  middle,  base  and  sides 
punctate,  a  shining  impunctate  line  on  basal  third.  Scape 
inserted  almost  in  exact  middle  of  rostrum;  1st  and  2nd  joints 
of  funicle  moderately  long,  subequal,  as  long  as  3rd-7th  combined. 
Prothorax  feebly  transverse,  sides  rounded,  feebly  decreasing  to 
base;  median  carina  very  feeble;  densely  punctate,  punctures 
round  and  moderately  deep.  Scutellum  oblong.  Elytra  slightly 
granulate,  sides  decreasing  to  apex  from  about  the  middle;  inter- 
stices feebly  elevated,  somewhat  flattened,  as  wide  as  punctures 
at  base  and  wider  at  apex,  with  depressed  granules.  Mesosternal 
receptacle  semicircular.  Metasternum  more  densely  punctate  in 
middle  than  sides;  each  episternumwithonedistinctand continuous 
row  of  punctures.  Abdomen  (except  apical  segment)  rather 
sparsely  punctate,  the  3rd  and  4th  each  with  one  regular  row  of 
rather  shallow  punctures.  Femora  slightly  thickened,  posterior 
scarcely  reaching  apical  segment,' tooth  in  all  rather  small  but 
distinct;  tibiae  grooved,  subequal.  Length  G  (vix),  rostrum  14; 
width  3  mm. 

Hah. — Q. :  Endeavour  River,  Cairns  (Macleay  Mus.),  Somerset 
(Mr.  C.  French). 

This  species  must  be  close  to  T.  concretus^  also  from  North 
(Queensland;  the  following  passages  in  Mr.  Pascoe's  description, 
however,  would  certainly  not  apply  to  the  present  insect,  "Elytra 
interstitiis  valde  elevatis;  femora  .  .  .  mutica." 
"  The  scales  on  the  prothorax  are  embedded  in  the  punctures, 
both  being  of  rather  unusual  size."  He  also  says : — "Tibia:'  anticse 
sulcatse,"  thereby  at  least  implying  that  the  anterior  tibiae  are 
more  noticeably  grooved  than  the  four  posterior;  in  the  jDresent 
insect  the  reverse  is  the  case. 

TvRTiEosus  POLLUX,  n.sp. 

Piceous-black;  elytra  feebly  tinged  with  red,  rostrum  piceous- 
red,  antennae  and  claws  paler.  Moderately  squamose,  scales 
obscure  ochreous  and  sooty;  metasternal  episterna  with  scales 
continuous  throughout ;  abdomen  (especially  apical  segment) 
rather  densely  squamose,  each  of  the  intermediate  segments  with 
two  distinct  rows. 


426  REVISION    OP    THE    AUSTRALIAN    CURCULIONIDiE, 

Head  densely  punctate ;  ocular  fovea  moderately  distinct. 
Rostrum  feebly  shining,  subparallel,  coarsely  punctate  except  in 
middle.  Scape  inserted  distinctl}^  closer  to  apex  than  base;  basal 
joint  of  funicle  subquadrate,  slightly  shorter  than  2nd,  3rd-7th 
transverse.  Prothorax  transverse,  sides  rounded,  very  feebl}'- 
decreasing  to  base;  median  carina  very  feeble  and  appearing  wavy 
through  interruptions  by  punctures,  these  dense  and  round. 
Scutellum  oblong.  Elytra  noticeably  v/ider  than  prothorax  and 
about  thrice  its  length,  subparallel  to  near  apex ;  interstices 
feebly  elevated,  somewhat  flattened,  slightly  wider  than  punctures. 
Mesosternal  receptacle  semicircular,  thinner  than  in  preceding 
species.  Metasiermal  episterna  with  punctures  continuous 
throughout,  and  forming  double  rows  at  base  and  apex.  Abdomen 
rather  densely  punctate,  3rd  and  4th  segments  each  with  two 
very  distinct  and  regular  rows.  Legs  rather  long ;  posterior 
femora  not  passing  middle  of  4th  abdominal  segment,  all  with 
very  feeble  teeth;  tibi?e  shorter  than  usual,  subparallel.  Length 
6,  rostrum  If;  width  2|  mm. 

Rah. — N.S.W.  :  Clarence  River. 

Very  close  to  the  preceding,  but  besides  colour  and  clothing 
differs  in  being  narrower,  in  the  puncturation  and  shape  of  ros- 
trum, in  antennal  insertion,  shape  and  size  of  funicular  joints, 
shape  of  mesosternal  receptacle  and  punctures  of  metasternum 
and  abdominal  segments. 

TYRT.E0SUS    ASSIMILIS,  n.sp. 

Blackish-brown,  elytra  and  legs  somewhat  paler,  antennae  of  a 
rather  pale  red.  Prothorax  with  a  sooty  scale  in  each  puncture, 
but  in  addition  a  few  larger  and  paler  ones;  elytra  with  dense 
pale  ochreous  and  whitish  scales  formed  into  very  feeble  spots 
and  fasciae,  but  a  moderately  distinct  median  fascia  of  dark  scales. 
Under  surface  with  subelongate  whitish  scales,  variegated  with 
darker  embedded  ones.  Legs,  head  and  base  of  rostrum  with 
obscure  whitish  scales. 

Head  coarsely  punctate  between  and  behind  eyes,  elsewhere 
finely  punctate;  with  a  short  median  ridge  separating  a  shallow 


I 


BY    ARTHUR    M.    LEA.  427 

impression  behind  each  e3^e,  these  not  very  close  together.  Ros- 
trum i-ather  short  and  wide,  sides  feebly  incurved  to  middle; 
basal  half  coarsely  punctate  and  with  a  median  ridge,  apical  half 
shining  and  impunctate  along  middle,  but  with  strong  punctures 
on  the  sides.  Scape  inserted  two-fifths  from  apex  of  rostrum, 
and  shorter  than  funicle;  1st  joint  of  funicle  much  stouter,  but 
scarcely  longer  than  2nd,  the  others  transverse.  Prolhorax 
moderately  transverse,  sides  rather  strongly  rounded;  densely 
and  coarsely  punctate;  median  carina  feeble  and  continuous. 
Elytra  elongate,  wider  than  and  almost  thrice  the  length  of  pro- 
thorax;  with  series  of  large  punctures;  interstices  regular  and 
narrower  than  punctures.  Metasternal  episterna  with  large  but 
not  continuous  punctures;  3rd  and  4th  abdominal  segments  each 
with  two  irregular  rows.  Femora  feebly  dentate;  tibiae  curved  at 
base  only.     Length  4,  rostrum  4;  width  2  mm. 

Hah. — N.W.  Australia  (type  in  Macleay  Mus.). 

Very  close  to  j^f^'^'dalis,  but  the  metasternum  with  three  very 
large  punctures  on  each  side  close  to  the  episternum. 

Tyrt^osus  imitator,  n.sp. 

Dark  reddish -brown,  elytra  and  antennae  paler.  Rather 
sparsely  clothed  with  ochreous  scales,  thin  and  subsetose  on  pro- 
thorax,  under  surface  and  legs,  but  moderately  stout  on  elytra. 
Head  and  sides  of  base  of  rostrum  feebly  clothed. 

Head  rather  coarsely  punctate,  depressed  behind  each  eye;  with 
a  short  median  carina;  eyes  large  and  subapproximate.  Rostrum 
moderately  long,  sides  incurved  to  middle,  basal  half  strongly 
punctate  and  with  three  feeble  ridges,  elsewhere  shining  and  with 
smaller  but  very  distinct  punctures.  Scape  inserted  one-third 
from  apex  of  rostrum;  2nd  joint  of  funicle  slightly  longer  than 
1st,  none  of  the  others  transverse.  Prothorax  small,  densely  and 
coarsely  punctate;  with  a  rather  wide  median  carina.  Elytra 
wider  than  and  more  than  thrice  the  length  of  prothorax;  punc- 
tate-striate,  striae  deep  and  punctures  shallow  (larger  on  sides 
than  elsewhere);  the  five  sutural  interstices  each  with  a  strong 
row  of  punctures  along  its  middle,  the  others  with  punctures  on 


428  REVISIOX    OF    THE    AUSTRALIAN    CURCULIONID^, 

sides.  Under  surface  (including  metasternal  episterna  and  3rd 
and  4th  abdominal  segments)  densely  punctate.  Femora  feebly 
dentate;  posterior  tibiae  very  wide  at  apex.  Length  10,  rostrum 
2|-;  width  5.^  mm. 

Hah. — E.  Australia  (Mr.  Horace  W.  Brown). 

In  appearance  remarkably  close  to  microthorax,  but  the  punc- 
tures very  different;  those  of  the  ely  tral  interstices  should  prevent 
the  species  from  being  readily  confounded  with  any  other  of  the 
genus.  The  specimen  described  is  either  from  Rockhampton,  Q., 
or  Orange,  N.S.W. 

Tyrt.eosus  laterarius,  n.sp. 

Dark  reddish-brown,  antennae  and  claw-joints  paler.  Prothorax 
moderately  clothed  with  elongate  ochreous  scales;  elytra  very 
densely  clothed  with  brick-red  or  ochreous  scales  intermingled 
with  obscure  patches  of  paler  and  darker  scales,  a  moderately 
distinct  subtriangular  patch  on  each  side  about  the  middle. 
Under  surface,  legs,  head  and  base  of  rostrum  moderatel}'  densely 
clothed  with  ochreous  scales. 

Head  with  dense  concealed  punctures;  eyes  rather  distant. 
Rostrum  scarcely  the  length  of  prothorax,  sides  lightly  incurved 
to  middle ;  basal  half  coarsely  punctate  but  without  median 
ridge,  apical  half  shining  and  moderately  punctate.  Scape 
inserted  two-fifths  from  apex  of  rostrum,  the  length  of  funicle; 
1st  joint  of  funicle  stouter  and  slightly  longer  than  2nd,  the  4th- 
7th  transverse.  Prothorax  moderateh^  transverse,  sides  lightly 
rounded ;  with  ver}^  dense  and  moderately  large  punctures ; 
median  carina  indistinct.  Elytra  elongate,  considerably  wider 
than  and  almost  thrice  the  length  of  prothorax;  with  series  of 
large  concealed  punctures;  interstices  apparently  wider  but  in 
reality  narrower  than  punctures.  Under  surface  (including  3rd 
and  4th  abdominal  segments  and  metasternal  episterna)  densely 
punctate.  Femora  feebly  dentate  ;  tibia?  straight  except  at 
extreme  base.     Length  5,  rostrum  1;  width  2 J  mm. 

Ilah. — N.S.W.  :  Blue  Mountains  (Macleay  Mus.). 


BY    ARTHUR    M.     LEA.  429 

The  clothing  of  the  elytra  is  unusually  dense  and  is  of  a  brick- 
red  colour. 

Tyrt^osus  inermis,  n.sp. 

Dark  reddish-brown,  antennae  and  claw-joints  paler.  Densely 
and  almost  uniformly  (except  that  they  are  paler  on  the  sterna 
and  absent  on  the  apical  half  of  rostrum)  clothed  with  soft  sooty 
scales,  feebly  variegated  with  sooty  white  ones. 

Head  with  dense  concealed  punctures;  eyes  large,  but  not  very 
close  together.  Rostrum  short,  wide  and  almost  straight,  sides 
very  feebly  incurved  to  middle;  basal  two-fifths  coarsely  punctate, 
elsewhere  shining  and  (except  along  middle)  very  distinctly  punc- 
tate. Scape  inserted  almost  in  exact  middle  of  rostrum;  1st  joint 
of  funicle  stouter  but  no  longer  than  2nd,  the  others  transverse. 
Frothorax  strongly  transverse,  sides  strongly  rounded;  with  dense, 
round,  clearly-cut  but  concealed  punctures;  median  carina  absent. 
Elytra  slightly  and  not  suddenly  wider  than  prothorax  and  about 
twice  and  one-half  as  long;  striate-punctate,  punctures  large  but 
more  or  less  concealed  ;  interstices  convex,  the  width  of  but 
apparently  much  wider  than  striae.  Punctures  of  metasternal 
episterna  continuous  throughout;  3rd  and  4th  abdominal  segments 
each  with  one  row.  Feinora  edentate.  Length  4,  rostrum  |; 
width  2.  mm. 

Hab. — INorth  Queensland  (Macleay  Mus.). 

The  clothing,  except  on  the  tarsi,  is  nowhere  setose  in  cha- 
racter. The  scape  is  inserted  almost  in  the  middle  of  the  rostrum 
and  the  prothoracic  carina  is  entirely  absent. 

Tyrt^osus  corpulentus,  n.sp.  * 

Brownish-black,  antennae  not  much  paler,  elytra  reddish-brown. 
Prothorax  with  a  mouse-coloured  scale  in  each  puncture,  and  with 
a  few  small  spots  of  dingy  whitish  scales;  elytra  with  mouse- 
coloured  scales  and  with  two  fasciae  of  white  (bordered  or  inter- 
mingled with  pale  ochreous)  scales.  Under  surface  with  large 
soft  white  scales  intermingled  on  the  apical  segments  with  darker 
ones;  legs  mostly  with  white  scales.  Head  and  basal  half  of 
rostrum  not  very  densely  squamose. 


430  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

Head  with  rather  coarse  punctures;  shallow!}^  depressed  behind 
eyes,  which  are  large  and  subapproximate.  Rostrum  scarcely 
the  length  of  prothorax,  sides  feebly  incurved  to  middle;  distinctly 
punctate  throughout,  but  punctures  much  coarser  on  basal  than 
on  apical  half;  with  a  feeble  median  ridge  behind  antennae.  Scape 
inserted  two-fifths  from  apex  of  rostrum  and  shorter  than  funicle; 
1st  joint  of  funicle  slightly  longer  than  2nd,  4th-7th  transverse. 
Prothorax  moderately  transverse,  sides  strongly  rounded;  with 
dense  round  clearly-cut  punctures;  with  a  continuous  median 
carina,  which,  however,  becomes  rather  feeble  anteriorly.  Elytra 
scarcely  wider  than  and  not  thrice  the  length  of  prothorax,  with 
series  of  large,  round,  regular  punctures;  interstices  (except  the 
suture)  narrower  than  punctures  and  regular  throughout.  Punc- 
tures of  metasternal  episterna  not  continuous  throughout;  those  of 
3rd  and  4th  abdominal  segments  somewhat  irregular  at  sides  but 
in  only  one  row  on  each  across  middle.  Femora  acutely,  the 
four  posterior  strongly  dentate.  Length  9,  rostrum  2;  width 
5  mm. 

Hah. — Q. :  Cooktown  (Mr.  C.  French). 

Very  close  to  vetustus,  but  distinguished  by  the  larger  size  and 
the  almost  jet-black  antennae.  It  appears  also  to  be  close  to 
lateralis^  but  the  elytral  punctures  and  clothing  do  not  agree  with 
Mr.  Pascoe's  description.  The  elytral  fasciae,  though  very  dis- 
tinct, are  not  sharply  defined  except  on  the  sides  :  the  first  com- 
mences on  the  side  at  the  basal  third,  but  at  the  4th  interstice 
divides  into  two,  of  which  one  part  touches  the  scutellum,  the 
other  the  suture  before  the  middle;  the  enclosed  space  is  some- 
what diamond-shaped,  but  contains  a  few  pale  scales;  the  posterior 
fascia  is  at  the  apical  fourth  and  scarcely  touches  the  suture. 

Tyrt^osus  cinerascens,  n.sp. 

Black,  elytra  almost  black,  antennae  dull  red.  Upper  surface 
(including  head  and  base  of  rostrum)  moderately  densely  clothed 
with  whitish  scales  intermingled  with  mouse- coloured  ones,  the 
whole  having  a  speckled  appearance.  Under  surface  and  legs 
with  dense  whitish  scales. 


BY    ARTHUR    M.    LEA.  431 

Head  coarsely  punctate;  eyes  moderately  large  and  scarcely 
approximate.  Rostrum  moderately  long,  sides  incurved  to  middle, 
but  slightly  swollen  at  insertion  of  antennae;  distinctly  punc- 
tate throughout  but  coarsely  so  on  basal  half;  without  median 
ridge.  Scape  inserted  two-fifths  from  apex  of  rostrum  and  shorter 
than  funicle;  2nd  joint  of  funicle  slightly  longer  than  1st,  5th-7th 
transverse.  Prothorax  rather  strongly  transverse,  sides  strongly 
rounded;  with  dense  round  punctures;  with  a  continuous  but 
rather  feeble  median  carina.  Elytra  not  much  wider  than  pro- 
thorax  and  about  twice  and  one-half  as  long;  sculpture  as  in  the 
preceding  species.  Punctures  of  under  surjace  dense,  but  on  the 
metasternal  episterna  not  quite  continuous.  Femora  strongly 
and  acutely  dentate.     Length  8J,  rostrum  2;  width  4|  mm. 

ZTaJ.— N.S.W.  :  Blue  Mountains  (Rev.  T.  Blackburn),  Went- 
worth  Falls  (Mr.  A.  Simson)— Tas. :  Hobart  (Mr.  F.  A.  Rodway). 

On  one  of  the  specimens  under  examination  the  dark  scales 
form  four  very  feeble  elytral  fasciae. 

Tyrt^osus  urens,  n.sp. 

Black ;  tarsi  and  antennae  dull  red.  Moderately  densely 
clothed  with  blackish  scales;  prothorax  with  several  indistinct 
patches  of  pale  scales;  elytra  with  distinct  but  irregular  and  not 
sharply  defined  fasciae;  scutellum  with  dense  pale  scales.  Under 
surface,  legs,  head  and  rostrum  obscurely  clothed. 

Head  densely  punctate  ;  eyes  moderately  widely  separated. 
Rostrum  rather  long  and  thin,  sides  very  feebly  incurved  to 
middle;  with  strong  punctures  throughout,  but  smaller  in  front 
of  antennae.  Scape  inserted  one-fourth  from  apex  of  rostrum  and 
the  length  of  funicle;  1st  joint  of  funicle  stouter  and  longer  than 
2nd,  the  others  transverse.  Prothorax  feebly  transverse,  sides 
moderately  rounded;  densety  and  coarsely  punctate,  median  carina 
traceable  but  nowhere  distinct.  Elytra  narrow,  not  much  wider 
than  and  about  twice  and  one-half  the  length  of  prothorax ; 
striate-punctate,  punctures  large  and  distinct;  interstices  densely 
punctate  and  the  width  of  striae.  Under  surjace  (including  3rd 
and  4th  abdominal  segments)  densely  punctate;  metasternal  epi- 


432  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

Sterna  less  densely  and  nob  continuously  punctate.  Legs  long 
and  thin;  anterior  femora  edentate,  the  others  feebly  dentate. 
Length  4,  rostrum  1  J;  width  IJ  mm. 

I/ab.—Tsis.:  Hobart  (Mr.  H.  H.  D.  Griffith)— N.S  W. :  Moon- 
bar  (Mr.  R.  Helms). 

A  small  species  with  much  thinner  legs  than  usual.  The  speci- 
men from  M  r.  Helms  is  marked  as  having  been  taken  at  an 
elevation  between  3,000  and  3,500  feet. 

Tyrt^osus  .equus,  n.sp. 

Black;  antennse  (club  excepted)  and  tarsi  dull  red.  Clothing 
as  in  the  preceding  species. 

Head  and  eyes  as  in  the  preceding  species.  Rostrum  moder- 
ately long,  but  not  very  thin;  basal  third  coarsely  punctate  and 
with  a  feeble  median  ridge;  elsewhere  shining  and  lighth*  punc- 
tate. Scape  inserted  two-fifths  from  apex  of  rostrum  and  slightly 
shorter  than  funicle;  1st  joint  of  funicle  much  stouter  but  no 
longer  than  2nd,  the  others  transverse.  Prothorax  feebly  trans- 
verse, sides  strongly  rounded,  densely  and  coarsely  punctate,  with 
a  moderately  distinct  and  continuous  median  carina.  Elytra  as 
in  the  preceding  species,  except  that  they  are  rather  wider, 
Punctures  of  under  surface  as  in  the  preceding  species.  Femora 
feebly  dentate.     Length  5,  rostrum  1 ;  width  2  mm. 

Hah. — N.S.  Wales  (Macleay  Mus.)— Tas.:   Huon  River  (Lea). 

Close  to  the  preceding  species,  but  larger  and  all  the  femora^ 
dentate;  it  is  also  close  to  pollux,  but  the  punctures  of  the  meta- 
sternal  episterna  are  continuous  throughout. 

Tyrt^osus  crassirostris,  n.sp. 

Blackish-brown  ;  elytra,  legs  and  antennae  paler.  Upper 
surface  with  indistinct  sooty  scales,  with  small  patches  of  obscure 
whitish  scales  scattered  about  and  forming  \evy  feeble  fascia3  on 
the  elytra  and  three  feeble  stripes  on  the  prothorax;  scutellum 
with  dense  pale  ochreous  scales.  Under  surface,  legs,  head  and 
rostrum  with  rather  thin  and  not  very  dense  whitish  scales. 


BY    ARTHUR    M.    LEA.  433 

Head  with  moderately  large  punctures  in  front,  becoming 
smaller  behind;  a  shallow  impression  between  eyes;  eyes  widely 
separated  and  much  smallf^r  than  usual.  Rostrum  slightly  shorter 
than  prothorax,  rather  wide  and  thick,  sides  increasing  in  width 
to  apex  and  not  incurved  to  middle;  with  coarse  punctures  in 
strong  series  throughout.  KScape  inserted  two-fifths  from  apex  of 
rostrum  and  shorter  than  funicle;  1st  joint  of  funicle  stouter  but 
scarcely  longer  than  2nd,  the  others  transverse.  Prothorax 
strongly  transverse,  sides  strongly  rounded;  densely  and  moder- 
ately strongly  punctate,  the  punctures  in  places  confluent;  median 
carina  distinct  and  continuous.  Elytra  elongate-cordate,  wider 
than  and  about  twice  and  one-half  the  length  of  prothorax ; 
punctate-striate,  striae  deep  and  punctures  indistinct;  interstices 
densely  punctate  and  wider  than  striae.  Under  surface  (including 
3rd  and  ith  abdominal  segments)  with  dense  (but  on  the  meta- 
sternal  episterna  not  continuous)  punctures.  Femora  edentate. 
Length  6,  rostrum  IJ;  width  3  mm. 

Hab.—Q. :  Somerset  (Mr.  C.  French). 

The  rostrum  and  eyes  in  this  and  the  following  species  are  very 
different  from  those  of  any  other  here  described.  In  both  also  the 
femora  are  feebly  ridged  beneath,  but  even  on  the  posterior  pair 
these  ridges  do  not  become  dentate. 

Tyrt.^osus  quadratolineatus,  n.sp. 

Dark  reddish-brown,  antennae  and  claw-joints  paler.  Clothing 
as  in  the  preceding  species,  but  rather  sparser. 

Head  and  eyes  as  in  the  preceding  species.  Rostrum  as  in  the 
preceding  species,  except  that  the  punctures  though  everywhere 
coarse  are  not  in  distinct  series.  Antennae  paler,  but  otherwise 
as  in  the  preceding.  Prothorax  feebly  transverse,  sides  moder- 
ately rounded  ;  densely  and  coarsely  punctate,  median  carina 
feeble  but  continuous.  Elytra  narrow,  scarcely  wider  than  and 
not  much  more  than  twice  the  length  of  prothorax;  striate-punc- 
tate,  punctures  indistinct  but  not  small;  interstices  the  width  of 
striae,  flat  and  all  (but  the  lateral  ones  less  noticeably  so)  in  series 
of  small  squares,  each  of  which  contains  a  rather  large  and  more 


434  REVISION   OP    THE    AUSTRALIAN    CURCULIONID.E, 

or  less  central  puncture.  Abdomen  with  two  rows  of  punctures; 
those  on  metasternal  episterna  not  continuous.  Femora  edentate. 
Length  5,  rostrum  1  J;  width  2J  mm. 

Hab.—l^.W.  Australia  (Macleay  Mus.). 

The  elytral  interstices  are  very  peculiar;  each  appears  (especially 
towards  the  base)  to  be  divided  into  series  of  squares  each  of 
which  contains  a  more  or  less  central  puncture.  It  is  this  char- 
acter which  renders  it  so  distinct  from  the  preceding  species, 
which  otherwise  it  strongly  resembles. 

H  Y  P  E  R  I  0  s  o  M  A,  n.g. 

Head  rather  large  and  feebly  convex,  not  concealed.  Fyes 
large,  ovate,  not  very  distant,  coarsely  faceted.  Rostrum  moder- 
ately long  and  not  very  thin,  curved.  Scape  inserted  nearer  apex 
than  base  of  rostrum  and  the  length  of  funicle;  two  basal  joints 
of  the  latter  moderately  long;  club  elliptic.  Prothorax  transverse, 
sides  rounded,  base  bisinuate,  constriction  feeble,  ocular  lobes 
obtuse.  SciUelluM  distinct.  Elytra  oblong-subcordate,  base 
trisinuate  and  wider  than  prothorax.  Pectoral  canal  terminated 
between  intermediate  coxse.  Mesosternal  receptacle  slightly  raised, 
crescent-shaped,  emargination  briefly  U-shaped;  cavernous.  Meta- 
sterniim  shorter  than  the  following  segment;  episterna  distinct. 
Basal  segment  of  abdomen  as  long  as  the  two  following  combined, 
its  apex  incurved;  2nd-4th  with  their  sides  drawn  slightly  back- 
wards, 2nd  slightly  longer  than  3rd  and  4th  combined  and  dis- 
tinctly longer  than  5th.  Legs  rather  short;  femora  stout,  dentate, 
not  grooved,  posterior  not  passing  elytra;  tibise  compressed  and 
distinctly  curved;  tarsi  normal.  Subelliptic,  convex,  squamose, 
winged. 

This  genus  is  not  a  very  distinct  one,  but  the  combination  of 
characters  is  such  as  to  forbid  the  species  described  below  (and 
which  in  shape  and  to  a  certain  extent  in  clothing  resembles  the 
European  Hypera  ])unctata)  being  placed  in  any  other  genus. 
From  Cryptorliynchus  it  is  distinguished  by  the  base  of  the  elytra, 
2nd  abdominal  segment  and  legs. 


BY    ARTHUR    M.    LEA.  435 

Hyperiosoma  falcatum,  n.sp. 

Dark  reddish-brown;  antennse  (club  excepted)  paler.  Densely 
clothed  with  soft  muddy-brown  scales,  each  elytron  with  a  large 
triangle  of  dingy  whitish  scales,  its  base  on  the  sides  and  its  apex 
touching  the  3rd  interstice.  Under  surface,  legs,  head  and 
rostrum  with  paler  scales  than  on  pro  thorax,  and  with  rather 
numerous  whitish  scales  scattered  about. 

Head  with  dense  concealed  punctures.  Rostrum  the  length  of 
prothorax  and  almost  parallel-sided;  basal  half  with  strong  con- 
cealed punctures;  apical  half  shining  and  lightly  punctate.  Scape 
inserted  two-fifths  from  apex  of  rostrum ;  1st  joint  of  funicle 
stouter  and  slightly  longer  than  2nd,  7th  longer  than  6th.  Fro- 
thorax  moderately  transverse,  convex,  apex  more  than  half  the 
width  of  base  and  scarcely  produced;  with  dense,  round,  regular, 
concealed  punctures.  Elytra  about  once  and  one-third  the  width 
and  twice  and  one-half  the  length  of  prothorax,  sides  parallel  to 
apical  third;  with  series  of  large  and  round  but  almost  entirely 
concealed  punctures.  Under  surface  with  dense  concealed  punc- 
tures. Femora  stout,  teeth  of  anterior  pair  rather  small,  of  inter- 
mediate large  and  of  posterior  very  large,  posterior  extending 
almost  to  apex  of  abdomen;  tibiae  equally  and  rather  strongly 
curved  and  diminishing  in  width  to  apex.  Length  5|,  rostrum 
1^;  width  3  mm. 

Hah. — Q. :  Cooktown  (Herr  J.  Faust). 

The  elytral  triangles  of  pale  scales  are  very  distinct;  the  teeth 
of  the  posterior  femora  are  very  large  and  triangular. 

SCLEROPOIDES,  n.g. 

Head  moderately  large  and  convex,  not  concealed.  Eyes  of 
moderate  size,  distant,  coarsely  faceted.  Ghib  ovate.  Mesosternal 
receptacle  distinctly  and  rather  suddenly  raised,  sides  incurved  to 
base,  emargination  briefly  Y-shaped;  cavernous.  Femora  edentate 
and  not  very  stout;  tibiae  compressed  and  straight  except  at  base. 
Other  characters  as  in  the  preceding  genus. 

Elytral  triangles  of  white  scales  not  conjoined sqiiamicollis. 

These  triangles  conjoined trianguliferus. 


436  REVISION    OF    THE    AUSTRALIAN    CURCULIONID.E, 

SCLEROPOIDES    SQUAMICOLLIS,    n.sp. 

X.  Black;  antennae  and  tarsi  dull  red.  Densel}^  clothed  with 
large  soft  scales  of  an  almost  uniform  and  rather  dingy  shade  of 
brown;  each  elytron  with  an  obscure  triangle  of  pale  scales  about 
the  middle  but  not  extending  to  suture.  Legs  with  somewhat 
paler  scales  than  above,  the  sterna  and  abdomen  with  still  paler 
scales;  pectoral  canal  distinctly  squamose.  Head  and  rostrum 
(except  near  apex  where  the  clothing  is  feeble)  densely  squamose. 

Head  with  dense  concealed  punctures.  Rostrum  slightly  longer 
than  prothorax,  sides  feebly  incurved  to  middle;  densely  punctate, 
punctures  concealed  behind  antennae,  but  three  narrow  costae 
traceable.  Scape  inserted  at  one-third  from  apex  of  rostrum;  1  st 
joint  of  funicle  slightly  longer  than  2nd,  2nd  almost  the  length 
of  3rd  and  4th  combined,  7th  strongly  transverse.  Prothorax 
lighth^  transverse,  sides  rather  strongly  incurved  near  apex,  base 
rather  strongly  bisinnate;  punctures  entirely  concealed.  Elytra 
almost  thrice  the  length  of  prothorax  and  at  base  not  much  (but 
rather  suddenly)  wider,  sides  subparallel  to  near  apex;  seriate- 
punctate,  punctures  concealed;  interstices  convex,  the  alternate 
ones  distinctly  raised.  Under  surface  with  dense,  rather  strong, 
and  almost  uniform  but  concealed  punctures.  Legs  densely 
punctate;  femora  linear,  the  posterior  not  extending  to  apical 
segment.     Length  7,  rostrum  2;  width  3|-  mm. 

Hah.—^.^   Wales  (Sydney  Mus.). 

The  scales  on  the  prothorax  are  much  larger  than  elsewhere, 
are  rounded  and  thick  and  entirely  conceal  the  sculpture.  The 
outlines  of  this  and  the  following  species  are  very  similar  to  those 
of  Sclerojms  sticticus,  Faust,  from  Borneo. 

SCLEROPOIDES    TRIANGULIFERUS,  n.Sp. 

^  (?).  Black,  antennae  and  tarsi  darker.  Densely  clothed  with 
large  soft  scales  of  a  rather  dingy  shade  of  brown;  each  elytron 
with  a  large  triangle  of  white  scales  which  are  conjoined  at 
suture.  Under  surface  and  legs  with  rather  pale  scales;  pectoral 
canal  squamose.  Head  and  basal  third  of  rostrum  densely 
clothed. 


BY    ARTHUR    M.    LEA.  437 

Head  with  dense  round  concealed  punctures.  Rostrum  slightly 
longer  than  prothorax  and  rather  strongly  curved,  sides  very 
feebly  incurved  to  middle;  sides  at  base  rather  strongly  punctate, 
elsewhere  shining  and  with  moderately  small  and  rather  sparse 
punctures.  Scape  inserted  two-fifths  from  apex  of  rostrum;  1st 
joint  of  funicle  distinctly  longer  than  2nd,  2nd  the  length  of  3rd 
and  4th  combined,  7th  moderately  transverse.  Prothorax  rather 
widely  transverse,  sides  rather  strongly  incurved  near  apex,  base 
moderately  bisinuate;  with  dense,  round,  deep,  partially  concealed 
punctures.  Elytra  oblong-cordate,  about  thrice  the  length  of 
prothorax,  and  at  base  suddenly  wider,  sides  parallel  to  near 
apex;  with  series  of  rather  large  but  almost  entirely  concealed 
punctures;  interstices  convex,  wider  than  punctures,  the  alternate 
ones  rather  feebly  raised.  Under  surface  and  legs  as  in  the  pre- 
ceding species.     Length  6J,  rostrum  2;  width  3  mm. 

Hah. — N.8.W. :  Clarence  River. 

The  clothing  of  the  elytra  is  divided  into  four  distinct  sections  : 
a  basal  portion  triangularly  advanced  on  suture  and  of  a  brown 
shade,  a  larger  and  almost  circular  apical  patch  of  similar  scales, 
with  two  whitish  triangles  (each  of  which  is  as  large  as  the  basal 
portion)  filling  the  intervening  space. 

BOTHYNACRU  M,*  n.g. 

Head  of  moderate  size,  convex,  not  concealed.  Eyes  large, 
ovate,  lateral,  rather  coarsely  faceted.  Rostrum,  long,  thin,  and 
curved.  Antennce  thin;  scape  inserted  rather  close  to  apex  of 
rostrum,  the  length  of  funicle;  club  elliptic,  joints  oblique.  Pro- 
thorax transverse,  sides  strongly  rounded,  base  feebly  bisinuate, 
constriction  feeble,  ocular  lobes  rounded  but  rather  prominent. 
Scutellum  distinct.  Elytra  oblong-cordate,  considerably  wider 
than  prothorax,  base  almost  truncate.  Pectoral  canal  deep  and 
narrow,  terminated  between  intermediate  coxae.  Mesosternal 
recei^tacle  feebly  raised,  U-shaped,  but  base  somewhat  thickened; 
slightly  cavernous.     Metasternum  shorter  than  the  following  seg- 

*  ^oOvvos,  fovea;   aKpov,  apex. 
29 


438  REVISION    OF    THE    AUSTRALIAN    CURCULIONID^, 

ment;  episterna  distinct.  Abdomen  with  the  1st  segment  as  long 
as  2nd  and  3rd  combined,  intercoxal  process  narrow;  2nd-4th  with 
the  sides  drawn  slightly  backwards,  2nd  slightly  shorter  than  5th 
and  about  once  and  one-half  the  length  of  each  of  the  following. 
Legs  moderately  long;  femora  scarcely  grooved  but  each  with  a 
thin  ridge  terminating  in  a  tooth,  posterior  passing  elytra;  tibiae 
thin,  flat  and  curved;  tarsi  moderately  long,  1st  joint  slightly 
longer  than  4th.     Elliptic,  convex,  non-tuberculate,  winged. 

Most  of  the  characters  described  denote  an  approach  to  Tyr- 
tceosus;  the  legs  are  much  as  in  Pe^HssojJs  (except  that  the  tibiae 
are  thinner);  and  the  rostrum  and  antennae  are  suggestive  of 
affinity  with  Pezichus.  The  species  described  below  is  remarkable 
for  the  large  round  fovea  on  the  apical  segment  of  the  ^. 

BOTHYNACRUM    STOREOIDES,   n.sp. 

^.  Reddish-castaneous  and  lightly  shining.  Sparsely  clothed 
with  subochreous  scales;  elytra  with  a  very  distinct  but  small 
transverse  patch  of  white  scales  at  summit  of  posterior  declivity 
and  a  longitudinal  patch  on  each  side  at  apex.  Under  surface 
and  legs  with  white  scales;  head  and  basal  two-thirds  of  rostrum 
with  whitish  scales. 

Head  with  small  punctures.  Rostrum  slightly  longer  than 
prothorax,  thin  and  parallel-sided;  basal  half  with  concealed  but 
rather  small  punctures,  and  with  a  moderatel}'  distinct  median 
carina  and  a  much  feebler  one  on  each  side;  apical  half  shining 
and  lightly  punctate.  Scape  inserted  one-fifth  from  apex  of 
rostrum;  two  basal  joints  of  funicle  subequal,  their  combined 
length  slightly  less  than  that  of  the  others,  6th  and  7th  trans- 
verse. Prothorax  moderately  transverse,  with  dense  and  moder- 
ately strong  (but  slightly  obscured)  punctures ;  with  a  very 
narrow  and  almost  continuous  median  carina.  Elytra  about 
once  and  one-third  the  width  and  twice  and  one-half  the  length 
of  prothorax;  striate-punctate,  punctures  large  but  shallow  at 
each  end;  interstices  convex  and  regular,  narrower  or  the  width 
of    punctures,    but    posteriorly    becoming    considerably    wider. 


BY    ARTHUR    M.    LEA.  439 

Under  surface  with  rather  sparse  but  regularly  distributed  punc- 
tures, each  of  which  contains  a  scale.  Apical  segment  with  a 
large  round  fovea  occupying  more  than  half  the  surface.  Femora 
rather  feebly  but  distinctly  dentate,  posterior  just  passing  el3'tra. 
Length  5  J,  rostrum  1  J;  width  2 J  mm. 

9.  Differs  in  being  without  the  large  apical  fovea  of  the 
abdomen;  the  rostrum  squamose  at  extreme  base  only,  and  the 
scape  inserted  at  its  apical  two-fifths. 

Hah. — Q. :  Endeavour  River  (Macleay  Mus.). 

A  very  distinct  species  in  general  appearance,  and  in  clothing 
(except  of  the  under  surface)  resembling  many  of  the  species  of 
Storeus.  The  prothorax  at  first  sight  appears  to  be  almost 
regularly  granulate,  but  this  is  entirely  due  to  the  clothing  con- 
cealing portions  of  the  interstices  between  the  punctures. 

Genus  E  n  t  e  l  e  s,  Schonherr. 

Spec.  Cure.  iv.  (1),  p.  269,  ISTo.  321;  Lacordaire,  Gen.  Col.  vii. 
p.  128. 

Head  convex,  not  concealed,  ocular  fovea  distinct.  Eyes  very 
large,  finely  faceted,  subcontiguous.  Rostrum  slender,  longer 
than  prothorax,  strongly  curved.  Antennae  slender ;  scape 
inserted  nearer  apex  than  base  of  rostrum;  2nd  joint  of  funicle 
elongate  ;  club  elongate.  Prothorax  subconical,  ocular  lobes 
almost  rectangular,  constriction  very  feeble.  Scutellum  distinct 
bat  small.  Elytra  subcordate,  more  than  twice  the  length  of 
prothorax.  Pectoral  canal  narrow,  moderately  deep,  just  passing 
intermediate  coxse.  Mesosteimal  receptacle  not  raised,  sides  equal 
throughout  and  rather  thin,  emargination  elongate  U-shaped ; 
open.  Metasternum  about  half  the  length  of  the  following  seg- 
ment; episterna  not  very  wide.  Basal  segment  of  abdomen  large, 
intercoxal  process  wide  and  truncate,  2nd  two-thirds  the  length 
of  1st;  3rd  and  4th  conjointly  slightly  longer  than  2nd  or  5th. 
Legs  long:  femora  thickened,  dentate,  posterior  passing  elytra; 
tibiae  thin,  compressed;  tarsi  elongate,  1st  and  4th  joints  long 


440  REVISION'    OF    THE    AUSTRALIAN    CURCULIONID^., 

and  subequal,  3rd  short,  deeply  bilobed.      Elliptic,  convex  above 
and  below,  non-tuberculate,  winged. 
A  very  distinct  genus. 

Prothoiax  with  two  transverse  lines  of  scales,  elytra  with 

three Vigorm,  Gyll. 

Prothorax  with  one  transverse  line  of  scales,  elytra  with  two    vicinus,  Faust. 

Enteles  Yigorsii,  Gyll.;  I.e.  No.  5568. 

Black,  shining;  antennae,  claws  and  tibial  hooks  piceous-red. 
Elongate,  setose,  adpressed  scales  (in  ^  dark,  in  9  pale  strami- 
neous), forming  two  lines  across  prothorax,  one  almost  at  extreme 
apex  (usually  but  not  always  interrupted  in  middle)  and  one  at 
extreme  base,  the  two  conjoined  at  sides;  elytra  with  three  similar 
lines,  one  slightly  before  middle,  curved,  and  extending  to  meta- 
sternum;  one  after  the  middle,  straight  and  reaching  2nd  abdo- 
minal segment;  the  third  semicircular,  its  base  and  sides  touching 
the  middle  line,  the  enclosed  spaces  with  a  few  feeble  spots; 
between  base  and  first  line  there  are  numerous  spots,  and  the 
suture  is  bare;  between  1st  and  2nd  lines  there  are  usually  no 
spots,  but  the  suture  is  clothed.  The  under  surface  has  similar 
scales  to  the  upper;  they  narrowly  margin  apex  of  prosternum, 
sides  of  pectoral  canal,  mesosternal  receptacle,  margins  of  1st 
abdominal  segment  (except  intercoxal  process)  and  sides  of  the 
other  segments;  a  few  scales  scattered  about,  especially  on  the 
two  basal  and  the  apical  segment  of  abdomen;  legs  with  scattered 
scales,  causing  them  to  appear  grey  to  the  naked  eye.  Sides  of 
eyes  and  of  rostrum  at  base  with  scales.  Ciliation  short,  dense, 
silvery;   visible  with  head  in  position. 

Head  feebly  and  minutely  punctate;  ocular  fovea  deep,  distinct, 
in  9  open  at  apex.  Rostrum  shorter  and  more  noticeably  curved 
in  (J  than  9?  finely  punctate  throughout,  the  punctures  very 
feeble  in  ^^  S  ^^'ith  ^  shallow  squamose  groove  on  each  side  from 
eye  almost  to  insertion  of  antenme,  a  narrow  costa  separating  it 
from  scrobe;  in  $  the  groove  is  almost  absent.  Scape  inserted 
about  two-fifths  from  apex  of  rostrum,  in  (J  slightly  passing,  in 
9  scarcely  reaching  apex;  1st  joint  of  funicle  short,  obtriangular, 


BY    ARTHUR    M.    LEA.  441 

2nd-6th  subcyliiidric,  7th  transverse,  2nd  as  long  as  3rd-4th. 
Prothorax  slightly  transverse,  convex,  sides  slightly  incurved 
towards  apex,  slightly  rounded  towards  base,  base  bisinuate; 
scutellar  lobe  with  a  very  feeble  longitudinal  ridge,  a  still 
more  feeble  one  at  apex ;  with  minute  and  very  minute 
punctures  throughout.  Scutelhmi  small,  shining,  transverse, 
within  a  depression.  Elytra  closely  applied  to  prothorax,  base 
trisinuate,  shoulders  feebly  clasping  prothorax,  widest  slightly 
behind  base;  very  feebly  striate,  the  striae  almost  invisible  near 
apex;  with  rather  small,  distinct  punctures  at  base  and  sides, 
rather  distant  and  only  in  striae.  Urider  surface  scarcely  visibly 
punctate;  intercoxal  process  with  an  oblique  groove  on  each  side. 
Legs  finely  punctate;  claw-joint  setose.  Length  8,  rostrum  3^; 
width  4  mm. 

Hah. — Queensland;  New  South  Wales  (Tweed  and  Richmond 
Rivers). 

An  elegant  and  very  abundant  species;  the  derm  beneath  the 
lines  of  scales  is  feebly  grooved,  so  that  the  scales  do  not  rise 
above  the  general  surface,  a  character  also  possessed  by  the  fol- 
lowing species. 

Enteles  vicintus,  Faust,  Stett.  Ent.  Zeit.  xlix.  1888,  p.  309. 

Black,  feebly  shining;  antennae  and  claws  piceous-red.  With 
similar  but  much  sparser  scales  than  on  preceding;  base  of  pro- 
thorax very  feebly  margined,  two  lines  on  elytra — one  about  one- 
sixth  from  base  on  the  disc  only  and  not  extending  to  suture  by 
more  than  one  interstice;  one  about  one-fourth  from  apex  partly 
continued  on  to  sides  and  interrupted  at  suture,  usually  without 
scales  elsewhere,  but  a  few  spots  occasionally  in  the  vicinity  of 
suture  about  the  middle.  Under  surface  and  legs  with  feeble 
scattered  scales.     Ciliation  duller,  but  otherwise  as  in  preceding. 

Rather  broader  and  less  convex  than,  but  very  similar  to,  the 
preceding.  Head  densely  and  not  very  minutely  punctate;  ocular 
fovea  deep,  distinct,  in  ^  separated  from  eyes  by  a  flat,  narrow, 
punctate  space,  in  9  rising  gradually  to  eyes.  Rostrum  rather 
coarsely    and    densely    punctate    in     ^J,    feebly    in    9  '■>     lateral 


442  REVISION    OF    THE    AUSTRALIAN    CURCULIONIDiE. 

grooves  as  in  preceding.  Scape  as  in  preceding;  funicle  a  little 
stouter,  2nd  joint  not  much  longer  than  1st,  and  not  the  length 
of  3rd-4th,  6th-7th  transverse.  Prothorax  scarcely  if  at  all 
transverse,  convex;  sides  slightly  incurved  towards  apex,  slightly 
rounded  towards  base;  base  bisinuate;  a  scarcely  traceable  median 
line;  with  small  punctures  on  disc,  becoming  very  minute  on  sides, 
apex  with  rather  strong  punctures,  each  carrying  a  small  seta. 
Elytra  much  as  in  the  preceding,  but  each  with  ten  distinct  rows 
of  distant  rounded  punctures,  visible  but  much  smaller  near  apex, 
those  at  the  base  (before  first  line  of  scales)  strong  and  apparently 
each  with  a  small  granule.  Groove  of  inter coxal  process  longer 
and  shallower  than  in  preceding.  Tihice  stouter  than  in  preced- 
ing, the  posterior  with  a  narrowly  separated  squamose  ridge  on 
each  side  at  apex,  more  pronounced  in  ^  than  5.  Length  10, 
rostrum  4;  width  5  mm. 

Hah. — North  Queensland. 

It  has  been  suggested  to  me  that  this  species  is  but  a  variety 
of  the  preceding,  but  this  is  incorrect.  It  is  a  rather  gloomy- 
looking  insect,  without  the  polished  derm  and  beautiful  markings 
of  E.  Vigorsii. 


443 


NOTES  ON  PROSOBRANCHIATA. 
No.  l—LOTORIUM. 

By  H.  Leighton  Kesteven. 

(Communicated  by  jL>er??^^ss^07^  of  the  Trustees  oj  the  Australian 

Museum.) 

Part  i. — Synonymy. 

Family  LOTORIID^,  Harris. 

(Harris,  Cat.  Tert.  Moll.  Brit.  Mus.  Pt.  i.,  p.  185,  1897.) 

Genus  L  o  t  o  r  i  u  m,  Montfort. 

Buccinum,  Tournefort,   1742.     Lotorium  nodiferum^  Lamarck 

{fide  Bayle,  Journ.  de  Conch.,  1880,  p.  241). 
Buccinum-tritonis,  Klein,  Tent.  Meth.  Ostrac.  1753,  p.  43,  pi.  7, 

f.  117.     B.  penatum,  &lc.  =  Lotorium.  tritonis,  Linn. 
ArgohuGcinu,m,  Klein,  op.  cit.  p.  44,  pi.  7,  f.  128.     A.  /asciatum 

=  Gyrineum  vexillum,  Sowb. 
Lagena,  Klein,  op.  cit.  p.  49,  pi.  3,  f.  61.     L.  toroides  =  Lotorium 

clandestinum,  Chemnitz. 
Simjndum^  Klein,  op.  cit.  p.  50,  pi.  3,  f.  62,  63.      S.  forosum  = 

Lotorium  costatum,  Born. 
Gictturiiiiwi,  Klein,  op.  cit.  p.  51,  pi.  3,  f.  64.      G.  ranula  — 

Lotorium  tuberosum,  Lamarck. 
*Epidromus,  Klein,  op.  cit.  p.  52.     B.  buccijium-sulcatum,  &c.  = 

Colubraria  7naculosa,  Chemnitz. 
*Murex,  Linnaeus,  Syst.  Nat.  1767,  Tom.  i.,  Pars  2,  p.  1213. 
Tritonium,  Bolten,  Mus.  Bolt.  1798,  p.  125;  ed.  ii.  p.  88,  1819. 

Not  Tritonium,  O.   F.   Miiller,  Prodr.  Zool.   Dan.,  p.  243, 

1776. 


444  NOTES    ON    PROSOBRANCHIATA, 

Cymatium,  Bolten,  op.  cit. 
Cahestana,  Bolten,  op.  cit. 

Tritojiium,    Link,     Beschr.     Rostock.    Samml.    p.   121,    1807. 
Lotoriutn  tritonis,  Linn. 
*  Persona,  Montfort,  Conch.  Syst.  ii.  1808,  p.  633.     Type  P.  anus, 
Linn.  =  Distortrix  anus,  Linn. 
Aquillus,  Montfort,  op.  cit.  p.  579.     Type  A.  cutaceus,  Linn.  = 
Lotorium  cutaceum,  Linn,     ^ot  Aqidlus,  Brisson,  Ornithol. 
i.,  p.  419  (AvEs). 
^Apollon,  Montfort,  op.  cit.  p.  571.     Type  A.  gyrinus,  Linn.  = 
GyrineuTYi  gyrinum,  Linn. 
Lotorium,  Montfort,  op.  cit.  p.  583.     Type  L.  lotor  =  Lotorium 
lotorium,  Linn. 
,  Triton,  Montfort,  op.  cit.  p.  587.     Type  T.  tritonis,  Linn.  = 
Lotorium  tritonis,  Linn.  Not  Triton,  Linn.,  1768(Crustacea), 
nor  of  Laurenti,  1768  (Batrachia). 
Monoplex,   Perry,   Conchology,    1811,   pi.  iii.     M.    cornutus  = 
Lotorium  exaratum,  Reeve. 
*Biplex,   Perry,   op.   cit.    pi.  iv.      B.   rosa  =  Gyrineiun   hufonia, 

Gmelin. 
Septa,    Perry,   op.    cit.    pi.   xiv.     S.   ;ja7*^i7iso?iia?ia  —  Lotorium 

fusiforme,  Kiener. 
Lampusia,  Schumacher,  Essai  Nouv.  Syst.  Habit.  Testaces,  1817, 
pp.  72,  250.     Ij.  jnleare,  Linn.  =  Lotoritun  pileare,  Linn. 
^Coluhraria,  Schum.,  op.  cit.  pp.  76,  251.      C.  granulata  =  Colu- 
braria  maculosa,  Gmelin. 

*Gyrina,  Schum.,  op.  cit.  pp.  77,  253.      G.  maculata==Gyrineum 
giganteum,  Lamk. 
Ranularia,  Schum.,  op.  cit.  p.  253.     {Ranula,  p.  77)  R.  lahiata 

—  Lotorium  pyrum,  Linn. 
Luterium,  Herrmannsen,  Indicis  Gen.  Malac.  i.  1846,  j^p.  625, 

632.     Emend,  for  Lotorium. 
Cumia,  Bivon,  Caratt.  Nuov.  Gen.  Conchiglie,  1838. 


BY  H.  LEIGHTON  KESTEVEN.  445 

Chat'onia,  Gistel,  Naturg.  des  Tierreichs,  1848,  p.  107. 
Linatella,  Gray,  Guide  Moll.  Brit.  Mus.  1857,  p.  39.     Type  L. 

cinc/ulata  =  Lotoriinn  cingulatum,  Lamk. 
Neptunella  (Gray),  H.  &  A.  Adams,  "Genera"  ii.,  p.  654, 1858. 
*Priene,  H.  &  A.  Adams,  op.  cit.  p.  654.     P.  rudis,  Broderip. 
1  Trachytriton,  Meek,  Smithsonian  Miscell.  Coll.  vii,  1864;  Smith, 
Check  List  Tert.  Foss.,  pp.  22,  37.     T.  vinculum,  Hall  k 
Meek. 
"^Tritonopsis,  Conrad,  Am.   Journ.   Conch,  i.   1865,  p.  20.      T. 
suhalveatum  =  Cymia  woodii,  Dall  (Jide  Dall). 

%  Personella,  Conrad,  op.  cit.  p.  21.     P.  sejjtemdentata,  Gabb. 
1  Ranellina,  Conrad,  op.  cit.  p.  21.     R.  maclurii,  Conrad. 
*Buccitriton,  Conrad,  op.  cit.  p.  21.     B.  alius  =  Nassa  {fide  Dall). 
ISassia,  Bellardi,  "  T.  Moll.  Terreni  Terz.  del  Piedmont  e  della 

Ligura,"  Mem.   Reale  Accad.  Sci.  Torino  xxvii.   (ser.  ii.) 

1873,  p.  219.      First  sp.  Triton  apenniniciom,  Sassi. 
"^Aspella,  Morch,  Malak.  Blatt.  xxiv.  1877,  p.  24.    Asj)ella  ancepti, 

Lamk. 
"i  Plesiotritoii,  Fischer,  Man.  de  Conch.  1884,  p.  654. 
"I  Hilda,  Hoernes  et  Auinger,  1884.     Die  Gasterop.  der  Meeres- 

Ablagerungen,  p.  182,  pi.  xxii.  f.  17-20.     Abhand.  derk.  k. 

Geologischen    Reich.    Band    xii.       Type    Triton    (Hilda) 

transsylvanicum,  H.  &  A. 

The  Generic  Name. 

The  names  in  the  above  synonymy  marked  with  an  asterisk 
are  included,  because  they  have  been  associated  with  "  Tritoyi,'* 
more  or  less  erroneously,  by  Tryon,  Fischer  and  others.  Some 
are  good  genera,  others  synonyms  of  other  groups.  Klein's  pre- 
Linnsean  names  acquired  a  right  to  quotation  when  endorsed  by 
later  writers. 

Murex,  Linn.,  being  retained  for  another  distinct  genus,  the 
first  names  we  need  consider  are  those  of  Bolten.  I  have  been 
unable  to  consult  his  work,  but  they  seem  to  be  nomina  nuda,  and, 


446  NOTES    ON    PROSOBRANCHIATA, 

therefore  unquotable.  Fischer  (n,  206)  reviewed  the  '  Museum 
Boltenianum,'  and  came  to  this  conclusion;  and  Dall's  sentence, 
*'  There  is  no  good  reason  why  we  should  not  adopt  the  name 
proposed  by  Bolten,  given  a  scientific  standing  by  Link,  and 
adopted  by  Cuvier  "  (7,  225);  and  his  repeated  rejection  of  Bolten's 
noinina  nuda  points  to  his  being  of  the  same  opinion.  (The  italics 
are  mine.)  The  name  to  which  the  above  sentence  refers  is 
Tritonium.  He  has  since  regarded  the  name  as  preoccupied  at 
the  time  of  its  proposal,  and  inadmissible  on  that  account  (8,  4:16). 

With  reference  to  Aquillus,  I  cannot  do  better  than  quote 
Harris  (15, 186): — "Commencing  with  Aquillus,  the  etymology 
of  the  word  is  uncertain,  and  in  any  case  is  hybrid.  When  it  is 
emended  in  the  manner  suggested  by  Agassiz  (2,  p.31,  Moll. p.  7)  and 
others,  we  have  Aquilus  or  Aquila,  which  is  anticipated  by  the 
well  known  Aquila,  Brisson,  in  ornithology,  and  by  several  other 
authors  prior  to  the  appearance  of  Montfort's  work.  To  prevent 
difficulty,  therefore,  it  is  not  advisable  to  select  Aquillus,  the 
more  so  that  Montfort  suggested  another  name  at  the  same  time, 
in  the  same  work  which  will  do  very  well." 

The  next  name  on  my  list,  which  is  arranged  chronologically, 
is  Lotorium,  which  is  not  preoccupied,  is  proposed  in  a  thoroughly 
scientific  manner,  and  for  which  a  type  (Murex  lotorium,  Linn.) 
is  named,  described  and  figured.  Triton  is  preoccupied;  Septa 
and  Laiyipusia  were  proposed  subsequent  to  Lotorium. 

I  am  of  the  opinion  that  Harris  is  right  in  maintaining  that 
Lotorium  should  be  accepted.  With  this  conclusion  both  Messrs. 
E.  A.  Smith  {fide  Harris,  I.e.)  and  C.  Hedley  agree. 

Until  a  generic  nonien  nudum  is  absolutely  defined  there  will 
be  an  element  of  uncertainty  in  this  synonymy.  Verrill  (38,  54) 
says  Bolten  worked  in  a  rational  manner,  and  that  "  he  gave  no 
diagnoses,  but  he  cited  well  known  and  figured  species  as  types, 
so  that  his  meaning  is  clear."  If  this  is  so,  and  it  constitutes  a 
generic  description,  Montfort's  name  must  give  place  to  that  of 
Bolten.  It  is  a  point  which  can,  it  seems,  be  only  settled  finally 
by  a  consultation  of  a  few  malacologists  of  experience,  and  a 
careful  consideration  of   the   consequences,  rather  than  a  rigid 


BY    H.    LEIGHTON    KESTEVEN.  447 

application  of  the  rules  of  nomenclature.  The  following  extract 
from  Dall's  Report  to  the  American  Association  for  the  Advance- 
ment of  Science  (1877  Meeting)  "On  Nomenclature  in  Zoology 
and  Botany"  (p.  45,  §  Ixii.)  makes  a  fitting  conclusion  to  a  dis- 
cussion on  this  synonymy  : — 

"The   following  kinds   of   works   are   entitled   to   citation  in 

bibliography,  but  not  in  synonymy: — 1 2 3.  Works 

not  published. 

"  It  may  seem  superfluous  to  object  to  works  of  the  third 
category.  But  besides  several  MSS.  preserved  in  museum  libraries 
and  sometimes  quoted,  though  never  printed,  there  are  a  few 
works  that  have  been  printed  but  never  published.  This  is  the 
case  with  a  Museum  Catalogue  prepared  by  Link  about  1806. 

"  It  was  printed  and  contained  a  host  of  new  names.  But 
whether  the  author  was  ashamed  of  his  work,  or  the  authorities 
of  the  University  declined  to  be  sponsors  for  the  innovations,  the 
work  was  never  offered  for  sale,  distributed,  or  advertised  by  the 
author. 

"Only  one  copy  is  definitely  known  to  have  escaped  from  the 
University  cellars,  and  it  has  been  stated  that  the  remainder,  or 
most  of  them,  were  destroyed  by  fire.  Yet  in  1851,  the  solitary 
copy  having  been  discovered,  one  or  two  authors  called  attention 
to  it,  and  demanded  that  these  names  should  take  precedence  of 
those  of  Lamarck  and  others,  which  had  been  in  use  for  nearly 
half  a  century.  A  few  writers  have  adopted  this  suggestion,  and 
in  one  branch  of  science  at  least,  deplorable  confusion  has  resulted. 

"  The  auctioneer's  catalogue  of  Bolten's  collection  printed  in 
1798,  but  fortunately  containing  no  diagnoses,  and  of  which  only 
one  or  two  copies  are  known,  falls  nearly  in  the  same  category.  A 
reprint  was  made  in  1819,  but  is  also  one  of  the  rarest  books." 

The  Family  Name. 

The  consideration  of  what  name  should  be  applied  to  the 
family  has  been  complicated  by  the  action  of  Dall  and  Simpson. 
They  have,  without  giving  any  reason,  divided  the  group  known 
of  old  as  Triton  into  four  genera,  and,  while  admitting  Lotoriiim^ 


448  NOTES    ON    PROSOBRANCHIATA, 

they  make  Septa  the  type  genus  of  the  family  (Septid?e)  [8,416, 
et  seq.]. 

Perry  enumerated  six  species  of  the  genus  Septa;  they  are,  in 
the  order  he  gave  them  : — 

Septa  parkinsoniana  =  Triton  fusi for  mis,  Kiener. 
,,     scarlatina  =        ,,     r^ibecida,  Linn. 

,,     sp>engleri  —        ,,     spengleri,  Chemn. 

,,      rubicunda         =        .,     nodiferus,  Lamk. 
,,      rubecula  =       ,,     pilearis,  Linn. 

,,     triangularis     =       ,,     costatus,  Born. 

Lotoriimi  parkinsoniarium  is  not  related  to  the  tritonis-group 
(vide  post,  p.  475),  but  makes  with  some  of  the  Australian  Tertiary 
species  a  rather  distinct  section,  which  is,  however,  connected 
with  L.  waterhousei.  Thus  the  first  three  species  are  all  referable 
to  the  so-called  genus  Laynpusia,  which  is,  therefore,  a  synonym 
of  Septa.  These  again  are  generically  inseparable  from  Lotoriam. 
Therefore  it  is  only  by  admitting  Lotoriium  as  the  type  of  the 
family,  and  removing  Perry's  first  three  species  thereto,  that  the 
name  Sep)ta  can  be  applied  to  the  tritonis-gvou^.  It  is  obvious 
that  they  cannot  be  referable  to  Lampusia  or  Ranularia,  both  of 
these  being  later  names.  If,  therefore.  Septan  are  typical  forms 
(and  parkinsonianum  is  the  type  thereof),  Lotorium  must  be 
regarded  as  a  synonym,  which  has  been  shown  to  be  impossible. 
Should  m}'-  "lines  of  generic  similarity"  not  be  considered 
sufficient!}^  complete,  nor  the  other  evidence  conclusive,  then  the 
name  Septa  must  be  applied  to  the  group  formed  by  pjarkinsoni- 
anwn  and  the  Australian  fossils,  this  section  being  more  distinct 
than  any  of  the  others.  ■ 

The  tritonis-gvoviY)  cannot  be  retained  as  the  typical  section  of 
the  genus;  it  has  not  yet  received  a  name  which  can  be  used. 
Unless  we  are  to  regard  L.  parkiyisonianum  as  the  type  of  the 
genus,  the  name  Septidm  cannot  be  adopted;  under  these  circum- 
stances it  seems  far  more  reasonable  to  accept  Harris's  term 
Lotoriidce. 


BY  H.  LEIGHTON  KESTEVEN.  44& 

This  is  not  only  correct  in  point  of  nomenclature,  but  it  will  be 
found  more  in  accordance  with  a  natural  arrangement  of  the 
groups  if  it  is  advantageous  to  split  the  genus  up  into  systematic 
divisions. 

Lotorinin  {scmsu  stricto)  will  embrace  all  those  typical  forms 
which  have  been  placed  by  Tryon  under  Simjndum,  Cymatium^ 
and  Gutturtiium.  The  sections  will  then  include  a  few  forms 
which  cannot  be  regarded  as  in  any  way  typical. 

The  conclusions  of  this  Part  are  that  the  following  should  be 
adopted  : — 

Family  LOTORIID^,  Harris. 

Genus  Lotorium,  Montfort. 


Part  ii. — Arrangexment  of  the  Species. 

"The  original  group  has  been  considerably  divided;  in  fact, 
Bolten,  Montfort,  and  others  began  the  work  of  division.  The 
whole  matter  is  worth  an  exhaustive  discussion.  ..."  (Dall, 
'  Blake  Mollusca,'  p.  225,  1889). 

In  the  following  pages  I  discuss  this  subject  at  some  length, 
and  have  come  to  the  conclusion  that  the  whole  of  the  species 
included  by  Tryon  in  Triton  {sensu  stricto),  Shn^ulum,  Cymatiiim 
and  Gutturnium  form  one  natural  genus.  From  a  study  of 
figures  and  descriptions,  and  of  one  species  {P.  scaber,  King),  I 
feel  inclined  to  regard  Priene,  H.  &  A.  Adams,  as  a  good  genus. 
I  am  unable  to  express  any  opinion  as  to  the  value  of  the  various 
fossil  groups  proposed  by  Fischer,  Conrad,  Gabb  and  others. 
Epidromus  ( =  Cohihraria)  has  rightly  been  treated  as  a  distinct 
genus  by  most  late  w^riters.  I  agree  with  Dr.  Dall  that  Fischer 
(12,  655)  incorrectly  referred  the  Apollon  group  of  Gyi^ineum 
and  Aspella  to  "  Triton  "  as  subgenera. 

In  this  essay  I  have  used  conchological  and  embryonic  characters 
only.  I  have,  however,  also  studied  the  matter  from  an  anato- 
mical standpoint,  and,  although  my  investigations  here  have  been 
by  no  means  extensive,  I  think  it  may  safely  be  said  that  investi- 
gations in  this  line  will  have  no  important  modifying  effect  on 


450  NOTES    ON    PROSOBRANCHIATA, 

the  conclusions  arrived  at.  So  far  as  our  knowledge  at  present 
goes  the  species  of  the  "  tritonis ''-group  (post,  p.  474)  have  a 
dentition  somewhat  different  from  that  of  other  groups,  and 
the  so-called  subgenus  Guttumium  has  been  stated  by 
Fischer  (12, 655)  to  have  an  operculum  distinct  from  that  of 
the  rest.  I  would,  however,  draw  attention  to  the  fact  that  only 
about  half-a-dozen  radulse  have  been  figured,  and  of  these  few 
that  of  L.  femorale,  Linn.,  is  intermediate  between  those  of  L. 
tritonis,  Linn.,  as  the  one  extreme,  and  L.  cutaceum,  Linn.,  as  the 
other.  Hitherto  there  have  been  known  three  apparently  distinct 
types  of  protoconchs — those  typified  by  L.  riitihim,  Menke 
(PL  xvii.,  fig.  21),  L.  cormUum,  Perry  (PI.  xvii.,  fig.  10),  and  L. 
woodsi,  Tate  (PL  xvii.,  fig.  1).  These  are  later  shown  to  be 
extreme  forms  of  one  generic  type.  It  is  more  than  probable  that 
when  the  subject  is  systematically  investigated  the  two  types  of 
dentition  will  suffer  the  same  fate,  and  all  the  intermediate  forms 
will  be  forthcoming. 

The  fact  that  Guttumium  has  an  operculum  differing  from  that 
of  other  groups  is  not  necessarily  of  much  importance;  since  there 
is  much  variation  in  the  form  of  the  protoconch  and  dentition 
(and  these  variations  of  little  note),  a  certain  amount  of  variety 
in  the  operculum  is  also  to  be  expected. 

As  already  stated,  my  anatomical  investigations  have  not  been 
extensive;  the  cause  of  this  has  been  the  difficulty  of  obtaining 
material.  Having  in  view  the  proposal  to  use  four  generic  names 
for  this  group,  I  have  thought  it  as  well  to  publish  this  essay  in 
its  present  incomplete  form,  and  to  follow  it  with  an  anatomical 
supplement  as  soon  as  sufficient  material  can  be  obtained. 

Concho  logical  Evidence  (Recent  Species). 

The  genus  has  hitherto  been  divided  on  the  form  of  the  shell 
only;  with  what  diversity  of  opinion  is  shown  in  the  next  part 
of  my  paper.  In  the  present  part  I  propose  to  use  that  evidence 
with,  I  submit,  rather  conclusive  results,  to  point  out  the  use- 
lessness  of  such  divisions  as  nomenclatural  entities.  For  want  of 
better  I  have  had  to  use  such  phrases  as  "nearest  ally"  and  "  nearly 


BY    H.    LEIGHTON    KESTEVEX.  451 

related "  in  this  discussion;  but  it  is  not  argued  that  any  one 
species  is  more  nearly  related  to  any  one  other  than  to  the  rest. 
Such  a  line  of  argument,  were  it  substantiated,  would  prove 
disastrous  to  my  contention,  for  community  of  relationship  pre- 
supposes community  of  descent.  Were  it  possible  to  prove  a 
common  progenitor  for  any  group,  distinct  from  that  of  another, 
it  would  certainly  be  advisable  to  separate  such,  at  least  sub- 
generically.  I  would  suggest  that  the  groups  are  to  be  accounted 
for  by  variation  along  similar  lines  from  the  original  parent. 

The  four  genera  of  Dall  and  Simpson  are  Sejyta^  Ranularia, 
Lampusia  and  Lotorium.  Being  full  genera  it  is  important  that 
they  should  be  discussed.  The  descriptions  given  with  the 
rehabilitation  of  each  of  these  are  not  sufficient  to  define  them; 
there  is,  however,  another  way  of  determining  their  value.  On 
comparison  with  sections  previously  proposed  they  appear  to  equal 
those  of  Try  on. 

Septa  can  be  no  other  than  Triton  (sens.  sir.).  Two  out  of  the 
three  species  enumerated  under  Lampusia  occur  in  Simpulum. 
Lotorium  evidently  equals  Cymatium.  The  species  named  under 
Ranularia  is  placed  by  Tryon  in  his  subgenus  Gutturnium.  That 
this  is  an  unsatisfactory  way  of  deciding  the  matter  must  be 
admitted,  but  it  receives  support  from  the  fact  that  there  are 
only  four  divisions  in  each  work  (though  Tryon  divides  his  sub- 
genera into  groups),  and  that  Septa  could  not  have  been  applied 
to  the  tritonis-gvoM'^  had  not  these  authors  accepted  Tryon's 
classification  of  L.  parkinsoniamcm,  Perry  ( =  X.  fusiforme, 
Kiener).  If,  however,  this  deduction  is  incorrect,  and  Dall  and 
Simpson's  genera  do  not  equal  Tryon's  subgenera,  the  following- 
statements  will  not  be  invalidated.  It  is  submitted  that  they 
serve  to  prove  that  the  group  known  of  old  as  Triton  is  a  natural 
genus  and  cannot  be  divided.  The  specific  nomenclature  here 
employed  is  that  of  Tryon's  "  explanation  of  plates  "  as  a  read}'- 
means  of  intimating  to  my  readers  the  particular  forms  referred 
to;  not  that  I  entirely  disagree  with  his  synonymy.  The  values 
of  a  few  specific  names  are  dealt  with  in  the  concluding  i^art  of 
the  paper. 


■452  NOTES    ON    PROSOBRANCHIATA, 

The  species  which  are  referred  to  Lotoriuniy  Dall  and  Simpson 
( =-  Cymatium,  Tryon)  ?ire—/emorale,  Linn.  ;  tigrinus,  Brod.  ; 
lotorium,  Linn.,  pyrum,  Linn.;  dunkeri,  Lischke;  and  miinsteri, 
Anton.  The  difference  between  L.  pyrum  and  L  cynocephalum 
is  if  avi^'thing,  less  than  that  between  the  former  and  L.  lotorium. 
It  would,  therefore,  be  unreasonable  to  separate  generically 
pyrtcm  and  cynocephalum.  By  the  same  argument  we  may  include 
first  L.  sarcofiioma,  then  L.  trilineatum,  Reeve;  thence  all  the  rest 
of  the  species  included  in  Tryon's  Gutturnium. 

A  series  of  species  generically  similar  in  every  way,  but  with 
gradually  shortening  canals,  is  formed  by — sarcostoma,  tripus, 
aegrotus,  vespaceus,  thersites,  crispus  and  quoyi.  It  would  be 
impossible  to  place  any  one  in  this  series  in  one  genus  and  either 
of  those  next  to  it  in  another.  They  are  all  included  in  Tryon's 
Gutturnhim.  The  same  remarks  maybe  made  of  the  series,  with 
gradually  elongating  canals,  formed  by  doliarium,  spengleri, 
waterhouseij  tranquebaricus,  pilearis,  mundum,  gemmatus  and 
orientoJis.  This  series  is  placed  by  Tryon  in  Simpidum.  The 
resemblance  between  ye^nmatus  and  quoyi  is  very  much  greater 
than  between  quoyi  and  sarcostoma.  Again,  crispus  is  more 
nearly  related  to  orientalis  than  orientalis  is  to  spengleri.  Another 
line  of  generic  similarity  is — waterhousei,  chemnitzi,  succinctus, 
cingulatus,  connecting  completely  the  tun-shaped  species  of 
Tryon's  group  Linatella  with  Simpulum.  Yet  another  perfect 
series  is  presented  by — verracosum,  quoyi,  intermedius,  rmindum, 
vespaceus,  eraratus,  chlorostoma,  thersites,  orientalis  and  crispus. 
This  is  not  merely  a  series  but  a  complete  ring,  for  crisjnis  is 
nearly  related  to  verrucosum.  Of  this  series  Nos.  1,  2,  5,  6  and  8 
are,  according  to  Tryon's  classification,  referable  to  Ranularia. 
Nos.  3,  4.  7,  9  and  10  to  Lampusia.  Lotorium  tritonis  is  con- 
nected, though  not  so  perfectly,  to  the  typical  group  by — nodiferus, 
hassi,  suhdistortus,  tumidosus,  philomelce,*  oligostirus,  eburneum, 
quoyi,  etc. 

*  Watson,  Chall.  Eep.  Zool.  xv.,  p.  391,  pi.  xiv.,  f.  10  (1885). 


BY  H.  LEIGHTON  KESTEVEN. 


453 


Nos.  4  and  6  of  this  series  are  Australian  Tertiary  species 
(34,  122,  126). 

An  attempt  has  been  made  in  the  accompanying  tables  to  show 
at  a  glance  the  intricate  relationship  existing  between  the  various 
groups. 


lotorium 

I 
j)yium 


cynocephalus 

I 
sarcostoma 

I 
exaratus 

I 


I 
vespaceus 


gemmatus 

I 
mundum 

pilearis 

tranquebaricus 

waterhousei 


gemmatus 

I 
intermedius 

I 

quoyi 

I 
verrucosum 


spengleri 

I 
doliarium 


I 
chemnitzi 

I 
succinctus 

cingulatus 

I 
etc. 


I 
chlorostoma 

thersites 

I 
orientalis 

I 
crispus 

I 
quoyi 

i 
verrucosum 

eburneum 

oligostirus 

I 
philomelae 

I 
tumulosus 

! 
subdistortus 


I 

nodiferus 

I 
tritonis 


dunkeri 

I 
tuberosus 

I 
encausticus 

I 
caudatus 

I 
exaratus 


lotorium 

I 
pyrum 

dunkeri 

I 
tuberosus 

I 
encausticus 

i 
caudatus 

I 
exaratus 

I 
vespaceus 

I 
gemmatus 

I 
intermedius 

I 
quoyi 

i 
crispus 

orientalis 

thersites 

I 
chlorostomus 

I 
exaratus 

I 
sarcostoma 

cynocephalus 

I 
pyrum 

I 
lotorium 


These  tables,  judged  by  the  form  of   the  shell,  are,  I  submit, 
correct,  and  serve   to  prove  the   impossibility  of    dividing  the 
genus  into  four. 
30 


454  NOTES    ON    PROSOBRANCHIATA, 

Australian  Fossil  Species. 

That  their  relationships  are  to  recent  species  occurring  only  in 
southern  waters  is  one  of  the  facts  which  strike  the  student  of 
Australian  fossil  Lotoriid^.  L.  quoyi,  Reeve,  exists  fossil  as  well 
as  recent.  L.  philomelce  is  so  like  L.  oligostirum,  Tate,  that  it  might 
be  as  aptly  termed  a  "living  fossil"  as  Trigonia.  Other  recent 
allies  of  these  are  L.  verrucosum,  ehurneuin,  and  nodocostatum  from 
Southern  Australia.  L.  parkinsonianu7)i  is  the  recent  represen- 
tative of  L.  radiale,  abbotti,  textile,  woodsi,  and  tortirostris.  This 
group  is  more  distinct  than  any  I  have  studied.  L.  tortirostris 
bears  a  resemblance  to  L.  waterliousei;  but,  to  institute  a  com- 
parison, this  resemblance  is  not  so  great  as  that  existing  between 
L.  subdistortum  and  bassi.  A  s  already  stated,  if  it  is  advantageous 
to  name  the  groups.  Septa  must  be  reserved  for  this  one.  The 
species  tumulosum  is  nearly  allied  to  subdistortum,  whilst  bassi  is 
represented  hj  ovoideum,  and  an  undetermined  species  intermediate 
between  them.  Lotorium  jjrotensum  is  possibly  an  archaic  form, 
and  resembles  somewhat  the  Fusid^e  from  which  the  genus  is 
apparently  descended.  Lotorium  cyphus  seems  to  stand  alone; 
it  might  be  considered  as  shadowing  forth  Distortio,  and  bears  a 
striking  resemblance  to  Cassidaria  gradrata,  Tate  (35,  169);  this 
resemblance  is,  however,  dispelled  on  comparing  young  specimens. 
The  existence  of  such  widely  different  species  as  protensum  and 
cyphus  in  an  early  Tertiary  stratum  points  to  a  much  earlier 
horizon  for  the  genus  than  the  text-books  admit;  the  wide  dis- 
tribution of  the  genus  in  Tertiary  times  is  also  evidence  of  this. 

The  parkinsonianum-grouip  is  of   particular   interest  from  a| 
zoogeographic  and  palseontological  point  of  view.     The  type  is  toj 
be  regarded  as  the  only  surviving  species  of  an  otherwise  extinct 
Antarctic  group).     There  are  in  Southern  Australian  beds  six  01 
nine  species;  in  Patagonian  beds  one  {L.  bicegoi);  and  the  New^ 
Zealand  Tertiary  species,  L.  minimum,  is  probably  referable  here 
also.     We  have  here,  then,  another  link  in  the  already  strong 
chain  of  faunal  relationship  shown  to  exist  between  these  three 
countries. 


BY  H.  LEIGHTON  KESTEVEN.  455 

To  the  palaeontologist  this  group  should  be  of  particular 
interest  as  throwing  light  on  the  vexed  question  of  the  age  of 
the  deposits  in  which  they  are  found. 

A.  E.  Ortmann  (25,  303),  compares  the  Patagonian  with  Euro- 
pean fossils,  and  then  compares  the  former  with  Australian  and 
New  Zealand  fossils,  and  says  : — 

"  The  result  of  the  foregoing  consideration  is  :  We  regard  the 
Patagonian  beds  as  of  Lower  Miocene  age;  contemporaneous  dejjosits 
are  found  in  the  southern  hemisphere^  not  only  in  Chili  (within 
the  Navidad  series),  hut  also  in  Nevj  Zealand  ( Pareora  beds  of 
Hutton)  and  Australia.'^  The  italics  are  in  the  original.  The 
Australian  beds  referred  to,  are  apparently  those  from  which  the 
fossils  under  discussion  are  derived. 

As  a  support  to  the  "Lyellian  percentage"  method  of  estimating 
the  age  of  a  deposit,  the  broader  comparison  of  the  genera  con- 
tained therein,  with  those  from  deposits  of  determined  age, 
and  with  recent  representatives,  should  give  more  satisfactory 
results  than  would  a  comparison  of  the  species  in  detail.  For 
this  purpose  the  genus  Lotorium,  being  well  represented,  is  of 
especial  value  to  the  Australian  palaeontologist.  Thus,  if  we 
compare  this  genus  as  it  occurs  in  the  lower  Australian  strata 
with  European  Miocene  representatives,  we  are  presented  with 
two  entirely  different  types  of  the  genus.  The  predominating 
feature  of  the  Australian  section — that  of  the  extinct  Antarctic 
group — finds  expression  in  only  one  European  fossil  (Z.  tarbel- 
lianum).  Again,  if  the  two  groups  be  compared  with  the  recent 
representatives,  it  will  be  seen  that  the  European  section  has  the 
general  facies  of  the  recent  species,  whilst  the  Australian  fossils 
can,  with  one  exception,  be  onl}-  compared  inter  se.  L.  quoyi  and 
the  Australian  fossils  possibly  referable  to  its  group  are  from 
more  recent  deposits  than  the  parkinsonianu7n-gYO\x^.  These 
facts,  namely,  that  the  predominating  feature  of  the  Australian 
grouj?  is  that  of  an  extinct  section,  and  that  the  European  group 
has  the  general  facies  of  the  recent  species,  assuredly  point  to  the 
greater  antiquity  of  the  Australian  fossils. 


456 


NOTES    ON    PROSOBRANCHIATA, 


A  comparison  of  the  species  in  detail  would  show  a  single 
related  species  in  all  four  localities,  and,  therefore,  give  an  erroneous 
impression  that  the  lieds  were  of  similar  age. 

The  Evidence  of  Literature. 

A  detailed  examination  of  literature  has  resulted  in  the  dis- 
covery of  greater  support  for  the  arguments  I  have  advanced 
than  I  expected  when  writing  the  last  part  of  this  paper.  I  have 
found  that  the  majority  of  the  species  spoken  of  as  being  referable 
to  or  connecting  two  of  the  divisions  proposed  by  Tryon  and 
others  have  been,  by  different  writers,  placed  in  both  sections. 

Tryon  (37,  9)  says  : — "  The  species  of  Triton  being  numerous 
several  attempts  have  been  made  to  separate  them  into  generic 
or  subgeneric  groups,  the  most  successful  being  the  arrangement 
of  Messrs.  H.  and  A.  Adams  (1)."  Previous  arrangements  never 
having  attained  any  degree  of  acceptance,  I  will  date  my  com- 
parisons from  this  one.  Subsequent  classifications  are  those  of 
Kobelt  (20),  Gray  (13),  Fischer  (12),  Chenu  (5),  Tryon  (36),  Melvill 
and  Standen  (24),  and  Dall  and  Simpson  (s).  Kobelt's  arrange- 
ment being  the  same  as  Tryon's,  and  Chenu's  the  same  as  that  of 
H.  and  A.  Adams,  they  are  not  included  in  the  following  table  : — 


1 

Gray.            Tryon. 

H.  &A. 

Adams. 

J"--- 1  ^ijr,t™ 

Dall  and 
Simpson. 

Triton,  s.st.i  Triton,  s.st. 

Tritonium,  s.st. 

Triton,  &.&i.\             ? 

Septa 

1 
Lampusia    \  Simjmlum 

Simpuhim 

Simpulum    |  ?  Simpubnn 

LAilPUSIA 

Aqniiius      Cabestana 

Cabestana 

AquilluS   \1  Aqmllus,i=.Bi. 

Do. 

?  Monopiex  \  Linatella 

Wanting 

Linatella  i?    Do. 

Do. 

Lotoriuni     j  Cymatlum 

Cymatiuin 

Lotorium :  ?  Lotorium 

Lotorium 

Ranularia 

Guttnrnium 

Gutturnium 

Bamdaria 

?     Do. 

Ranularia 

Wanting 

Cumia 

Wanting 

Wanting 

Wanting 

Wanting 

Generic  names  in  small  capitals,  subgeneric  in  italics,  group  names  in 
black  type. 


BY    H.    LEIQHTON    KESTEVEN.  457 

From  this  table  it  will  be  seen  that  though  there  is  a  general 
agreement  as  to  the  number  of  sections,  in  no  one  point  do  any 
five  of  these  writers  agree  as  to  the  values  of  the  sections. 

Cymatium  is  regarded  by  the  brothers  Adams,  Kobelt,  and 
Tryon  as  a  subgenus;  Fischer  deemed  it  a  section  of  Simpulum; 
Melvill  and  Standen  apparently  treat  it  as  a  section  of  Guttiirnium; 
whilst  Dall  and  Simpson  consider  it  a  full  genus.  Such  a  diversity 
of  opinion  can  only  be  attributed  to  the  imperfection  of  this  line 
of  classification. 

The  arrangement  of  Melvill  and  Standen  is  given  in  such  a 
haphazard  way  that  it  is  possible  I  have  not  rightly  interpreted 
it.  They  regard  Aquillus  as  the  name  which  should  be  accepted 
for  the  genus  as  a  whole,  and  enumerate  five  species  in  the  follow- 
ing order :  — 

Aquillus  cheniiiitzi,  Gray  {=A.  tranquebaricus,  Lamk.) 
,,         [Lotorium)  lotoriuvi,  Linn. 
,,        {Simjndum)  aquatills,  Reeve. 
,,         {Lotorium)  retusus,  Lamk. 
,,        {Lotoriuin)  tripus,  Lamk. 

Two  facts,  however,  are  plain — (1)  Lotorium,  according  to  these 
writers,  includes  forms  referred  by  Tryon  to  Cymatium  and 
Gutturnium;  and  (2)  Si7nj)ulum  of  Tryon  is  divided  into  two. 

I  now  proceed  to  discuss  the  "  shuttle-cock  "  fate  of  some  of  the 
species.  The  first  species,  instanced  as  connecting  Cymatium  and 
Gutturnium  (L.  cynocephalum)  has  been  referred:  by  Kobelt,  to 
Cymatium;  by  Tryon,  to  Gutturnium;  and  by  Dall  and  Simpson, 
to  Simpulum.  Ij.  pyrum  is  referred  by  the  brothers  Adams  to 
Gutturnium,  and  by  Tryon  to  Cymatium.  Kobelt  has  placed  L. 
dunkeri  in  Gutturnium,  whilst  Tryon  assigns  it  a  place  in 
Cymatium.  I  have  spoken  of  the  "  ^^^.02/^  "-group  as  connecting 
Simjndum  and  Gutturnium.  Watson  compared  his  species  L. 
philomelce  to  L.  quoyi,  and  yet  placed  it  in  Simpulum,  from  which 
it  may  be  concluded  that  he  regarded  the  latter  as  referable  to 
that  section  also.  Dall  (7,226)  has  referred  L.  olearium,  Linn 
(Tryon  Vj  =  L.  costatum,  Born,  to  Ranularia. 


458 


NOTES    ON    PROSOBRANCHIATA, 


Believing  that  it  is  the  best  form  in  which  to  put  matter  of 
this  kind,  I  have  tabulated  these  differences  of  opinion. 


Species. 

Sections  to  wh 

KOBELT. 

ich  they  have  been  referred  by 
Tryox.        H.&  A.  Adams.  : 

1         L.  lignarium,  Brod. 

Simpulum 

Linatella 

Simpulum 

L.  pfeifferiamtm,  Reeve 

Simpulum 

Gutturnium 

Simpulum 

L.  c y HOC ejih alum,  Lamk. 

Cymatium 

Gutturnium 

Gutturnium 

L.  dunkeri,  Lischke 

Gutturnium 

Cymatium 

not  mentioned 

L.  pyrum,  Linn. 

Cymatium 

Cymatium 

Gutturnium 

The  Apices. 

Becent  species. 

It  is  here  shown  that  the  apices  do  not  present  a  feature  on 
which  the  species  of  the  genus  can  be  grouped.  Twenty-six 
protoconchs,  including  nine  of  fossil  species,  are  figured  and 
described ;  and  descriptions  of  a  few  others  are  culled  from 
various  sources.  The  species  are  from  all  the  groups  except 
Lotorium.  I  have  found,  as  might  have  been  anticipated,  that 
species  conchologically  very  similar  have  similar  apices.  It  has 
not,  however,  followed  that  species  conchologically  dissimilar  have 
dissimilar  apices.  L.  succinctum,  Lamarck,  has  an  apex  exactly 
the  same  as  that  of  L.  exaratum,  Reeve;  and  the  apex  of  L. 
pileare,  Linn.,  differs  very  slightly  from  that  of  ves^Mceum,  Reeve. 
The  protoconchs  of  the  recent  species  are  all  of  one  type,  differing 
from  one  another  in  the  number  and  convexity  of  the  whorls  and 
in  colour.  They  consist  of  a  thin  coating  of  lime  inside  a  corneous 
original.  The  lime  is  apparently  not  generally  deposited  until 
the  mollusc  starts  the  adult  structure.  Protoconchs  of  L. 
spengleri,  Chemn.,  and  L.  exaratinn,  Reeve  (?),  to  which  no  adult 
structure  was  attached,  were  not  acted  upon  in  an}'-  way  when 
immersed  in  pure  hydrochloric  acid.  That  the  corneous  coating 
is  only  outside,  not  inside  as  stated  by  Reeve  (30),  is  proved  by 


BY    H.    LEIGHTOX    KESTEVEN.  459 

the  fact  that  the  apices  of  adult  specimens  from  which  the  external 
coating  had  been  removed,  were  entirely  demolished  by  the  same 
treatment.  In  none  of  the  species  studied  has  it  been  found 
possible  to  define  a  nepionic  stage;  in  every  instance  the  sculpture 
of  the  postembrj^onic  structure,  which  immediately  adjoins  the 
protoconch,  is,  in  miniature,  that  of  the  adult;  nor  is  there  in  any 
of  the  species  a  varix  formed  of  embryonic  structure.  In  one  or 
two  instances,  however,  the  adult  structure  begins  with  a  small 
varix.  It  appears  that  a  suggestion  made  by  the  writer  in  a 
previous  paper  (19)  applies  particularly  to  the  species  of  Lotoi'ium, 
and  that  none  of  them  have  left  a  conchological  record  of  the 
nepionic  stage.  It  might  have  been  inferred  that  the  protoconch 
had  been  deposited  inside  an  original  corneous  one.  Judging  by 
those  species  which  I  have  been  able  to  study,  the  whole  shell  is 
cast  inside  its  epidermal  coating.  After  a  rest-period  the  epidermis 
grows  very  quickly  and  extends  beyond  the  shell  (the  writer  has 
seen  as  much  as  half  an  inch  overhanging,  with  the  merest 
"  stiffening "  of  lime  inside  it),  and  inside  this  the  shell  is 
deposited.  It  may  be  worthy  of  note  here  that  in  Lotoriiim,  at 
least,  a  varix  is  indicative  of  a  "  period  of  growth  "  of  the  mollusc 
rather  than  of  a  "period  of  rest"  of  the  shell.  After  the 
formation  of  a  varix  [the  gutter  which  Reeve  describes  (I.e.)], 
and  while  the  shell  anterior  to  the  preceding  varix  is  yet  thin, 
the  mollusc  does  not  fill  the  cavity  nearly  so  much  as  when  this 
part  of  the  shell  is  thickened.  The  shell  is  enlarged  by  periods, 
faster  than  the  mollusc  grows,  which  while  growing  is  employed 
strengthening  its  shell,  not  enlarging  it.  Harris  (15,  p.  xii.)  speaks 
of  variation  in  the  size  of  apices  of  the  same  species.  Among 
fossils  the  variation  is  plainly  noticeable,  but  as  far  as  my 
experience  goes,  it  is  practically  non-existent  among  recent  species. 
The  following  descriptions,  like  the  figures  on  the  plates,  have 
been  arranged  to  show  the  perfect  graduation  from  the  tall  apex 
of  L.  exaratum  to  the  very  flat  one  of  quoyi.  The  letter  A 
indicates  that  species  so  marked  have  been  referred  by  Tryon  to 
Shnpulum;  B  indicates  Gutturnium.  It  is  worthy  of  note  that 
the  Jii' St  and  last  of  the  series  are  marked  with  a  B.     Contour  is 


460  NOTES    ON    PROSOBKANCHIATA, 

not  described,  the  figures  being  considered  sufficient  description 
thereof. 

LoTORiUxM  coRNUTUM,  Perry,  sp.  B. 

(Plate  xvii.,  fig.  10.) 

Triton  exaratus,  Reeve;  Tryon,  Man.  Conch. iii.  p.  22,  pi.  12, fig.  104. 

Apex  of  four  and  one-half  whorls,  brown,  semitransparent, 
smooth,  shining,  covered  with  a  thin  brown  epidermis.  I  have  in 
the  paper  referred  to  above  figured  this  apex  under  the  name  of 
I'ritoniwm  olearium,  Linn. 

LoTORiUM  cosTATUM,  Born,  sp.  A. 

I'riton  succinctus,  Lamarck;  Tryon,  op.  cit.  p.  11,  pi.  6,  f.  37. 

I  have  not  thought  it  necessary  to  figure  this  apex;  it  is  exactly 
similar  to  that  of  the  preceding  species. 

LoTORiUM  VESPACEUM,  Lamk.,  sp.  B. 

(Plate  xvii.,  fig.  11.) 

Triton  vespaceus,  Lamk.;  Tryon,  op.  cit.  p.  22,  pi.  12,  f.  94. 

Apex  of  four  and  one-half  whorls,  fairly  solid,  opaque,  dark 
brown  at  the  sutures  with  a  lighter  band  medially;  covered  with 
a  light  brown  epidermis. 

LoTORiUM  CHLOROSTOMUM,  Lamk.,  sp.  A. 

(Plate  xvii.,  fig.  12.) 

Triton  chlorosiomus,  Lamk.;  Tryon,  op.  cit.  p.  13,  pi.  7,  f.  47. 

Apex  of  four  and  one-half  whorls,  semitransparent,  smooth, 
shining,  covered  with  a  thin  light  brown  epidermis. 

LoTORiUM  AQUATiLE,  Reeve,  sp.  A. 

(Plate  xvii.,  fig.  13.) 

Triton  pilearis,  Linn.;  Tryon,  op.  cit.  p.  12,  pi.  6,  f.  31. 

Apex  of  four  and  one-half  whorls,  rather  solid,  semitransparent, 
light  brown,  smooth,  shining,  covered  with  a  thick  brown  epi- 
dermis. 


BY    H.    LEIGHTON    KESTEVEN.  461 

LoTORiUM  siNENSE,  Reeve,  sp.  B. 

(Plate  xvii.,  fig.  14.) 

Triton  sinensis,  Reeve;  Tryon,  op.  cit.  p.  20,  pi.  11,  f.  85. 

Apex  of  about  four  whorls,  very  slightly  transparent,  light 
brown,  smooth,  shining,  covered  with  a  rather  thick  epidermis. 
The  tip  of  the  specimen  figured  is  broken  off,  but  there  is  enough 
to  show  its  position  in  the  series. 

LoTORiUM  GEMMATUM,  Reeve,  sp.  A. 

(Plate  xvii.,  fig.  15.) 

Triton  gemmatus,  Reeve;  Tryon,  op.  cit.  p.  13,  pi.  7,  f.  41. 

Apex  of  four  and  one-half  whorls,  semitransparent,  white, 
smooth,  shining,  covered  with  a  thin  light  brown  epidermis. 

LoTORiUM  ELONGATUM,  Reeve,  sp.  B. 

(Plate  xvii.,  fig.  16.) 

Triton  elongatus,  Reeve;  Tryon,  op.  cit.  p.  22,  pi.  12,  f.  96. 

Apex  shorter  and  narrower  than  that  of  L.  vespaceum,  but 
resembling  it  in  other  respects. 

/         LoTORiUM  TUBEROSUM,  Lamk.,  sp.  B. 

(Plate  xvii.,  fig.  17.) 

Triton  tuberositas,  Lamk.;  Tryon,  op.  cit.  p.  23,  pi.  13,  f.  111. 

Apex  of  four  whorls,  semitransparent,  light  brown  above,  dark 
brown  at  the  anterior  suture,  smooth,  shining,  covered  with  a  thin 
brown  epidermis. 

LoTORiUM  THERSiTES,  Reeve,  sp.   B. 

(Plate  xvii.,  fig.  18.) 

Triton  thersites,  Reeve;  Tryon,  op.  cit.  p.  32,  pi.  12,  ff.  99,  100. 

Apex  of  three  whorls,  semitransparent,  light  brown,  smooth, 
shining,  covered  with  a  thin  light  brown  epidermis. 


462  NOTES    ON    PROSOBRANCHIATA, 

LoTORiUM  SPENGLERI,  Chemn.,  sp.  A. 
(Plate  xvii.,  fig.  19.) 
Triton  sjjengleri,  Chemn.;  Tryon,  op.  cit.  p.  16,  pi.  9,  f.  61. 

The  apex  of  this  species  has  been  described  in  a  previous  paper 
(l9,  p.  713).  A  figure  is  given  here  to  complete  the  series.  The 
apex  of  L.  tvaterhousei,  Angas,  is  almost  exactly  similar  to  this. 

LoTORiUM  STRANGEi,  Ad.  and  Ang.,  sp.  A. 
(Plate  xvii.,  fig.  20.) 
Triton  strangei,  Ad.  and  Ang.;  Tryon,  op.  cit.  p.  17,  pi.  9,  f.  67. 

Apex  of  three  and  one-half  whorls,  fairly  solid,  opaque,  shining, 
smooth,  dark  brown,  covered  with  a  thick  dark  brown  epidermis, 
which  under  a  lens  shows  four  spiral  threads,  a  few  transverse 
ones  equally  thick,  and  close  set  growth-striae. 

LoTORiUM  RUTiLUM,  Menke,  sp.  A. 
(Plate  xvii.,  fig.  21.) 
Tritonium  rutilvm,  Menke,  Moll.  Nov.  Holl.  Spec.  1843,  p.  25. 

Apex  of  three  and  one-half  whorls,  rather  solid,  opaque,  polished, 
smooth,  light  brown;  covered  with  a  thick  dark  brown  epidermis, 
which  is,  under  a  lens,  ornamented  with  four  spiral  threads  and 
close-set  growth-stride. 

LOTORIUM  LABIOSUM,   Wood,  Sp.    A. 

(Plate  xvii.,  fig.  22.) 
Triton  labiosus,  Wood;  Tryon,  op.  cit.  p.  17,  pi.  9,  f.  65. 

Apex  of  three  whorls,  semitransparent,  smooth,  shining;  the 
epidermis  was  cleaned  off  the  specimens  studied. 

LoTORiUM  VERRUCOSUM,  Reeve,  sp.  B. 
(Plate  xvii.,  fig.  23.) 
Ti'iton  verrucosus^  Reeve;  Tryon,  I.e.,  p.  24,  pi.  13,  f.  117. 

Apex  of  one  and  one-half  whorls,  white,  semitransparent,  smooth, 
shining.     No  epidermis  on  the  specimens  studied. 


I 


BY    H.    LEIGHTON    KESTEVEN.  463 

LoTORiUM  EBURNEUM,  Reeve,  sp.  B. 
(Plate  xvii.,  fig.  24.) 
Triton  eburneiis,  Reeve;  Tryon,  op.  cit.  p.  24,  pi.  13,  f.  115. 

Apex  of  one  and  one-half  whorls,  light  brown,  semitransparent, 
smooth,  shining;  covered  with  a  thin  light  brown  epidermis. 

LoTORiUM  QUOYi,  Reeve,  sp.  B. 
(Plate  xvii.,  fig.  25.) 
Tritoji  quoyi,  Reeve;  Tryon,  op.  cit.  p.  24,  pi.  13,  f.  116. 

Apex  of  one  and  one-half  whorls,  dark  brown,  smooth,  shining. 
No  epidermis  on  the  specimens  examined. 

LoTORiUM  NODOCOSTATUM,  Tate  and  May,  sp. 

(Text  fig.  1.) 

Lcunpusia  nodocostata,  Tate  and  May,  Trans.  Roy.  Soc.  S.  Aust., 
xxiv.,  1900,  p.  90;  Proc.  Linn.  Soc.  N.S.  Wales,  xxvi.,  1901, 
p.  355,  pi.  xxiii.,  f.  2. 

Apex  of  three  and  one-half  whorls,  perforate,  turbinate,  white, 
semitransparent,  polished.  The  sculpture  consists  of  six  faint 
revolving  lines,  of  which  that  on  the  periphery 
is  the  most  pronounced,  and  of  oblique  growth- 
stride,  some  of  which  are  stronger  than  others. 
Dimensions:  length  5,  breadth  3-3  mm. 

The  large  size  of  this  protoconch  marks  it 
as  unique  among  recent  species.  Notwith- 
standing its  entirely  normal  plan  of  coiling, 
it  appears  to  me  to  be  a  direct  link  between  the 
mammillate,  excentric  apices  of  some  fossils, 
and  the  normal  apices  of  the  recent  species. 

To  Mr.  C.  Hedley  I  am  indebted  for  the  p^™^  1 

opportunity  of    figuring  and    describing    this     L.  nodocostatum, 
apex,  from  material  dredged  by  him  from   a         'I&te  &  May. 
depth  of  one  hundred   fathoms    16   miles   east  of  Wollongong, 
N.S.W.     The  species  has  been  identified  by  comparison  with  a 
cotype  kindly  lent  me  by  Mr.  W.  L.  May. 


464  NOTES    ON    PROSOBRANCHIATA, 

LOTORIUM    RUBICUNDUM,  PeiTJ,  sp. 

(Text  fig.  2.) 

Triton  nodiferunij  Lamk.;  Tryon,  op.  cit.  p.  10,  pi.  3,  f.  17. 

Apex  of  three  and  one-half  whorls,  semitransparent,  delicate 
pink,  smooth,  shining. 

The  large  size  and  bright  colour  of  this  apex  give  it  a  different 
character  from  any  of  the  others.  Although  semitransparent,  it 
has  an  appearance  of  strength  which 
others  lack.  The  epidermal  original  is 
cast  very  early  in  the  life  of  the  indivi- 
dual; a  specimen  of  onty  four  adult 
whorls,  obtained  alive,  shows  no  sign  of 
it.  That  it  had  an  original  corneous 
mould  may  be  generalh'  proved  bj'- 
Fig.  2.  breaking  off  the  tip  of  the  protoconch, 

L.( Septa) nodiferum^'LQ.mk.  ^hen  the  remains  will  be  seen  attached 
to  the  base  of  the  piece  broken  off,  or  to  the  top  of  the  part 
remaining. 

The  following  descriptions  of  the  apices  of  L.  parkinsonianum 
and  L.  philoynelce  are  reprinted  that  they  may  be  compared  with 
those  of  L.  tortirostris  and  L.  oligostirum  on  pp.  469,  470. 

LoTORiUM  PARKINSONIANUM,  Perry,  sp. 

Triton  fusiformis,  Kiener ;  Tryon,  op.  cit.  p.  11,  pi.  4,  f.  22; 
Kesteven,  Proc.  Linn.  Soc*  N.S.  Wales,  xxvi.,  1901  (1902), 
p.  712,  pi.  xxxv.,  ff.  3,  4,  5. 

"  Protoconch  ovoid,  umbilicate,  thin,  semi-pellicid,  shining, 
corneous,  light  brown,  variously  marked  with  spots  or  stripes  of 
darker  colour;  consisting  of  about  three  whorls,  covered  with  a 
very  thin  light  brown  epidermis.  Transversely  sculptured  with 
very  fine  growth-lines;  the  epidermis  supplies  four  ciliated  ridges 
which  encircle  the  last  whorl,  the  top  one  of  which  is  continued 
on  the  earlier  whorls.     .     .     ." 


BY    H.    LEIGHTON    KESTEVEN.  465 

LOTORIUM   PHILOMEL.E,   Watson,  Sp. 

Trito7i  (Simpuhnn)  philomelce,  Watson,  ChalL  Rep.  Zool.  xv.,  1885, 
p.  391,  pi.  xiv.,  f.  10. 

"  Apex  consists  of  four  polished  but  spirally  threaded,  white, 
turbinated  whorls,  of  which  the  first  is  extremely  small  and  some- 
what immersed." 

LOTORIUM    PHARCIDUM,  Dall,  Sp. 

Lampusia     pharcida,    Dall,    "Blake    Mollusca,"    pt.  ii.,    p.  227, 
pl.xxxvi.,  f.  1. 

"  A  four-whorled  subcylindrical  nucleus;  nucleus  smooth  with 
rounded  subequal  whorls  and  a  rather  blunt  apex."  This  pro- 
toconch  somewhat  resembles  that  of  L.  vespaceum,  but  is  not  so 
tapering. 

Fossil  Species. 

One  of  the  most  noticeable  features  of  these  fossils  was  the 
large  number  of  specimens  which  had  their  apices  complete  (over 
70  per  cent.).  Among  recent  species  I  had,  like  Mr.  Baker  (3), 
found  that  only  about  five  per  cent,  retained  their  apices.  Talking 
over  this  matter  with  Mr.  C.  Hedle}',  he  suggested  that  it  was  in  a 
great  measure  due  to  their  having  been  inhabitants  of  deep  and 
still  waters.  Though  the  greater  strength  of  the  fossil  apices  has 
doubtless  been  a  contributing  cause,  an  inquiry  into  the  habitats 
of  living  allies  has  provided  support  for  Mr.  Hedley's  sugges- 
tion. He  has  (17)  in  dealing  with  tlie  "Thetis"  Pelecypoda 
instanced  several  cases  of  fossil  species  or  their  near  allies  occurring 
off  the  coast  of  New  South  Wales  in  deep  water.  My  own  investi- 
gations have  disclosed  the  fact  that  many  of  the  shells  collected 
by  the  "Challenger"  in  deep  waters  of  high  southern  latitudes 
have  near  allies  in  the  fossil  beds  from  which  the  species  under 
discussion  were  obtained.  It  is  thus  seen  that  many  of  the 
nearest  living  allies  of  these  fossils  are  inhabitants  of  deep  waters, 
and  it  might  reasonably  be  inferred  that  they  themselves  occupied 
similiar   localities.     This   inference    cannot,   however,   safely  be 


466  NOTES    ON    PROSOBRANCHIATA, 

pressed  too  far,  for  Tate  (35)  has  shown  that  man}'-  of  the  species 
now  inhabiting  the  tidal  zone  also  occur  in  these  beds.  The 
tabulation  of  these  relationships  would  be  productive  of  interest- 
ing results. 

The  protoconchs  of  Lotoriurti  p^'otensutn  and  ahhotti  are  similar 
to  those  of  the  Fusion,  the  elongate,  fusiforme  shape  of  the  latter 
also  recalling  that  family.  It  is  from  this  group  that  I  would 
suggest  the  genus  Lotoriurti  is  descended. 

A  prominent  character  of  all  the  apices  examined  is  their  spiral 
sculpture.  This,  it  is  interesting  to  find,  exists  in  two  recent 
species  {L.  nodocostatum  and  L.  philomelce)*  and  is  sometimes 
found  on  the  corneous  originals  of  other  species.  On  a  superficial, 
or  first  examination,  the  student  is  apparently  presented  with  two 
or  three  different  types  of  apex.  A  closer  study  discloses  the 
fact  that  they  are  morphologically  only  varieties  of  one  form. 
The  difference  lies  in  the  size  of  the  nucleus,  and  the  extent  of 
its  depression  or  elevation.  A  study  of  the  apices  in  profile  gives 
the  first  impression,  whilst  this  view  studied  in  conjunction  with 
the  view  from  above,  proves  the  correctness  of  my  conclusion. 
The  following  are  the  apices  studied;  like  those  of  the  recent 
species  they  have  been  arranged  in  a  graduation;  that  from  the 
very  eccentric  protoconch  of  L.  looodsi  to  the  normal  ones  of 
oligostirum  and  tortirostris.  These  two  being  normal  they  have 
not  been  figured  from  above  : — 


'&' 


LoTORiUM  wooDsi,  Tate,  sp. 

(Plate  xvii.,  fig.  1.) 

Triton  tvoodsi,  Tate,  Proc.  Roy.  Soc.  S.  Aust.,  x.,  1886-7  (1888), 
p.  119,  pl.v,  f.4. 

Apex  of  two  whorls;  the  first  half  whorl  perpendicular  to  the 
plane  of  coiling  of  the  shell;  second  half  normal,  but  slightly 
overlapping  the  second  whorl;  first  whorl  smooth,  dull;  second 
whorl  normal,  polished,  faintly  biangled,  sculptured  by  two  revolv- 

*  Vide  also  P.  [  =  L.]frateradum. 


BY  H.  LEIGHTON  KESTEVEN.  467 

inar  lirse  and  close-set  transverse  strise.     Towards  the  end  of  the 

o 

embryonic  structure  there  are  faint  indications  of  three  extra 
spiral  lirse,  which  are  suddenly  developed  on  the  adult  structure. 
The  sculpture  of  the  anterior  whorl  of  this  and  all  other  species 
studied  (with  one  exception  ■?)  fades  away  as  the  nucleus  is 
approached. 

LOTORIUM  RADIALE,  Tate,  Sp. 

(Plate  xvii.,  fig.  2.) 
Triton  radialis,  Tate,  op.  cit.  p.  118,  pi.  v.,  f.  8. 

Apex  of  a  little  more  than  one  and  one-half  whorls,  polished; 
nucleus  mammillate,  eccentric,  placed  rather  to  one  side  of  the 
centre,  in  some  instances  slightly  overlapping  the  succeeding 
whorl;  the  rest  of  the  protoconch  normal,  gradually  acquiring  a 
sculpture  consisting  of  three  spiral  threads,  the  uppermost  median, 
and  faint  transverse  growth-stri?e.  There  is  also  a  very  faint 
indication  of  a  fourth  spiral  thread  half-way  between  the  upper- 
most of  the  three  mentioned  and  the  posterior  suture. 

Harris  (15,  p.  187,  pi.  vi.,  f,  6)  describes  this  apex  as  consisting  of 
two  and  one-half  whorls,  whilst  his  figures  show  the  number  I 
have  found.  The  term  "  lateral  "  applied  by  Tate  and  Harris  to 
the  nucleus  of  this  and  other  protoconchs  of  this  series  is  some- 
what misleading.  The  whole  apex  is  truly  spiral,  but  the  nucleus 
being  somewhat  mammillate  and  large  relatively  to  the  rest  of  the 
protoconch  is  placed  eccentrically.  This  apex  is  defined  by  a 
complete  change  in  the  sculpture  of  the  shell. 

LoTORiUM  TUMULOSUM,  Tate,  sp. 

(Plate  xvii.,  fig.  3.) 

Triton  tumulosus,  Tate,  op.  cit.  p.  122,  pi.  v.,  f.  2. 

"Apex  of  two  polished  whorls;  the  anterior  one  is  high, 
bicarinated,  and  transversely  striated;  the  posterior  one,  encircling 
a  shallow  concavity,  at  first  suddenly  narrowed,  then  somewhat 
depressedly  dilated,  and  ending  in  a  blunt  appressed  point." 


468  NOTES    ON    PROSOBRANCHIATA, 

The  above  is  Tate's  description  of  this  apex,  and  well  describes 
the  specimens  I  have  been  enabled  to  examine,  except  that  in 
some  instances  two  small  spiral  threads  may  be  seen  below  the 
anterior  carination.  Though  all  the  specimens  examined  exhibit 
the  same  characters  the  abrupt  termination  of  the  sculpture  and 
*'  suddenly  narrowed  "  posterior  whorl  present  all  the  features  of 
a  caducous  cast  and  subsequent  shedding  of  the  nuclear  whorls. 
At  the  beginning  of  the  adult  structure  the  lower  threads  become 
suddenly  strengthened,  and  all  become  somewhat  nodulose. 

Among  recent  species,  L.  suhdistortum^  Lamk.,  drops  its  pro- 
toconch  at  a  very  early  stage.  Dr.  Yerco,  who  has  sent  me 
specimens  of  only  two  and  one-half  whorls,  says  he  has  never 
taken  a  specimen  with  its  protoconch  attached. 

LoTORiUM  ABBOTTi,  Tenison-Woods,  sp. 

(Plate  xvii.,  fig.  4.) 

Triton  abbotti,  Ten.-Woods,  Proc.Roy.  Soc.  Tas.  1875  (1876),  p.  24, 
pi.  i.,  f.  8. 

Apex  of  a  little  more  than  two  whorls;  nucleus  very  small, 
central,  rapidly  enlarging;  posterior  whorl  nearly  as  large  as  the 
anterior,  first  whorl  and  one-half  smooth,  polished;  thence  the 
sculpture  which  ornaments  the  adult  shell  is  gradually  developed. 

Tate  describes  (I.e.,  p.  117)  four  and  one-half  whorls  to  this 
apex;  it  is  only  very  faintly  defined  (at  the  dotted  line  on  my 
figure),  and  it  is,  therefore,  probable  that  he  reckoned  from  the  first 
varix. 

The  specimens  from  which  this  apex  is  described  are  from 
Schnapper  Point,  but  they  have  been  carefully  compared  with 
specimens  from  the  type  locality,  and  there  can  be  no  doubt  as 
to  their  identity. 

LOTORIUM  PROTENSUM,  Tate,  Sp. 

(Plate  xvii.,  fig.  5.) 
Triton  protensus,  Tate,  op.  cit.  p.  124,  pi.  v.,  f.  10. 

Apex  of  a  little  less  than  two  whorls,  polished;  nucleus  small, 
placed  a  little  to  one  side  of  the  centre,  rapidly  enlarging;  anterior 


BY    H.    LEIGHTON    KESTEVEN.  469 

whorl  sculptured  with  two  or  three  spiral  threads,  and  very  fine 
transverse  striae.  This  apex  is  absolutely  undefined.  I  agree 
with  Tate  in  considering  its  termination  to  be  a  little  anterior 
to  the  first  appearance  of  sculpture. 

LoTORiuM  CYPRUS,  Tate,  sp. 
(Plate  xvii.,  fig.  6.) 
Triton  cyphus,  Tate,  op.  cit.  p.  119,  pi.  v.,  f.  11. 

Apex  of  one  and  one-half  whorls,  polished;  nucleus  slightly 
mammillate,  but  placed  centrally;  the  anterior  whorl  sculptured  by 
a  distinct  median  and  two  postero-median  spiral  threads,  and  by 
transverse  growth-striae. 

This  protoconch  is  defined  by  the  acquisition  of  the  numerous 
spiral  lirse  with  which  the  adult  shell  is  ornamented. 

LOTORIUM   ANNECTANS,  Tate,  Sp. 

Triton  annectans,  Tate,  op.  cit.  p.  121,  pi.  v.,  f.  3. 

"  The  protoconch  of  this  species  is  similar  to  that  of  L.  torti- 
rostris  in  being  somewhat  elevated,  roundly  turbinate,  and  spirally 
striated;  it  is  composed  of  three  and  one-half  to  four  whorls,  and 
the  striae,  which  are  four  in  number,  equidistant  and  very  pro- 
nounced, decrease  in  size  on  being  traced  backwards;  the  top  of 
the  larval  shell  is  extremely  minute  and  central "  (Harris,  I.e., 
p.  192,  pi.  vi.,  f.  7). 

LoTORiuM  OLiGOSTiRUM,  Tate,  sp. 
(Plate  xvii.,  fig.  7.) 
Triton  oligostirus,  Tate,  op.  cit.  p.  126,  pi.  vi.,  f.  7. 

Apex  normal,  of  three  polished  whorls,  nucleus  slightly  de- 
pressed, first  two  whorls  smooth,  thence  an  ornamentation  of  four 
spiral  threads  and  oblique  growth-striae  is  gradually  developed. 
At  the  termination  of  the  embryonic  shell  the  structure  changes 
completely. 

Compare  with  this  Watson's  description  of  the  apex  of  Lotorium 
philomelce.  .^^^^'^^''^''7^^ 

.  •  -  "^'  >• 


470  NOTES    ON    PROSOBRANCHIATA, 

LoTORiuM  TORTiROSTRis,  Tate,  sp. 
(Plato  xvii.,  fig.  8.) 
Triton  tortirostris,  Tate,  op.  cit.  p.  123,  pi.  v.,  f.  7. 

Apex  normal,  of  three  whorls,  polished,  nucleus  very  slightly 
depressed,  the  anterior  whorl  ornamented  by  a  median  spiral 
ridge,  and  a  very  slight  thread  midway  between  this  and  the 
posterior  suture,  transversely  obliquely  striate,  defined  by  the 
acquisition  of  the  adult  sculpture. 

Compare  with  this  the  description  of  the  protoconch  of  L. 
parkinsonianum,  Perry. 

LOTORIUM  TEXTILE,  Tate,  sp. 
(Plate  xvii.,  fig.  9.) 
Tritort  textilis,  Tate,  op.  cit.  p.  120,  pi.  v.,  f.  12. 

"  Apex  acute  of  three  rounded  Urate  whorls,  ending  in  an  acute 
upward-curved  point." 

The  above  is  Tate's  description;  none  of  the  specimens  I  have 
examined  possess  perfect  protoconchs.  The  sculpture  on  the 
anterior  whorl  of  the  specimen  figured  consists  of  six  revolving 
threads  and  rather  distant  fine  growth-strise.  This  apex  is  not 
clearly  defined;  its  termination  seems  to  be  indicated  by  a  slight 
transverse  swelling  (an  aberrant  varix  ?),  and  the  ending  of  the 
polished  surface.  Judging  from  the  material  to  hand  it  seems 
probable  that  Tate  counted  from  the  first  varix. 

The  apex  has  rightly  been  considered  a  feature  of  systematic 
importance  by  most  recent  writers.  Unless,  as  is  evident  we 
should  do,  we  regard  all  the  apices  here  discussed  as  varieties  of 
one  generic  type,  the  division  of  this  group  will  present  some 
extraordinary  anomalies.  If  the  group  is  to  be  split  up  into 
various  genera  it  will  be  impossible  to  disregard  the  form  of  the 
apices,  now  that  we  know  so  much  about  them  ;  and  we  shall 
have  such  dissimilar  species  as  L.  costatum,  Born,  and  L.  cornutum, 
Perry,  in  the  same  genus,  and  species  so  absolutely  alike  as  L. 


BY  H.  LEIGHTON  KESTEVEN.  471 

tortirostris  and  L.  radiale  in  different  genera.  Considered  as 
varieties  of  one  type,  they  may,  for  the  convenience  of  the 
monographer,  be  disregarded. 

'*A  number  of  species  have  a  world-wide  distribution,  which 
is  doubtless  due  to  their  free-swimming  or  pelagic  larvse  "  (Tryon, 
op.  cit.  p.  7).  I  am  inclined  to  carry  this  statement  further,  and 
to  say  that  the  distribution  of  a  species  is  largely  decided  by  the 
size  of  its  protoconch.  Most  of  the  species  with  many-whorled 
protoconchs  have  a  very  wide  distribution,  whilst  those  with 
small  protoconchs  are  rather  local.  There  are  exceptions  to  this. 
L.  cornutum,  Perry,  with  a  many-whorled  apex  ranges  over  a 
small  area,  whilst  if  Tryon  is  correct  in  regarding  L.  loroisii, 
Petit,  as  a  synonym  of  L.  labiosum,  we  are  presented  with  a  species 
having  a  small  protoconch  and  a  practically  world-wide  distribu- 
tion. 

Conclusions  o^  Part  ii. 

My  conclusions  are  that  the  subgeneric  names  heretofore  used 
under  "  Triton  "  are  redundant  and  altogether  useless.  To  quote 
them  conveys  no  more  than  would  the  generic  name  alone.  It 
is  impossible  to  treat  them  as  full  genera;  to  do  so  would  create 
confusion.  References  to  several  species  would  have  to  be  sought 
under  two  or  three  genera.  After  eliminating  L.  clandestinum, 
all  the  species  enumerated  by  Tryon  under  Simpulum,  Cymatium 
and  Guturnium  form  one  section,  which  it  is  impossible  to  divide 
into  groups,  if  such  groups  are  to  be  regarded  as  nomenclatural 
entities.  If  from  Tryon's  "  Triton  s.st"  L.  suhdistortum  and  L. 
parkinsonianum  are  withdrawn,  and  L.  ovoideum  added,  we  have 
a  second  section ;  a  third,  as  already  stated,  is  formed  by  L. 
jyarkinsonianum  and  some  of  the  Australian  Tertiary  species. 
Should  it  be  decided  advisable  to  give  the  sections  quotable 
names,  the  above  are  the  three  which  must  be  first  recognised; 
any  further  subdivision  is  to  be  deprecated. 

I  am  of  the  opinion  that  none  of  the  sections  are  sufficiently 
distinct  to  rank  as  subgenera,  and  submit  the  following  arrange- 
ment of  the  species  of  the  genus. 


472  NOTES    ON    PROSOBRANCHIATA, 

In  this  arrangement  I  have  included  only  those  fossil  species 
of  which  I  have  been  enabled  to  examine  actual  specimens.  I 
note,  however,  that  Lotorium  apenninicum,  Sassi  (the  type  of 
Bellardi's  subgenus  Sassia),  might,  judging  by  Hoernes  and 
Auigner's  figures  (is),  be  included  in  my  "quoyi'^-group.  If  this 
is  so,  I  cannot  concede  that  the  subgeneric  name  is  a  useful  one : — 

Genus  Lotorium. 

Typical  group. 

L.  lotorium,  Linn.  L.  fetnorale,  Linn. 

L.  tigrinurtiy  Brod.  L.  grandimaculatuin,  Reeve. 

{L.  miinsteri,  Anton?) 

Group  of  L.  cosiatum. 
L.  costatimi,  Born.  L.  spengleri,  Chemn. 

L.  waterhousei,  Angas.  Z.  chemnitziy  Gray. 

L.  barthelemyi,  Bern.  L.  cutaceum,  Linn. 

L.  brasilianum,  Gould.  Z.  tranquebaricum,  Lamk. 

Z.  chlorostomum,  Lamk.  Z.  doliarium,  Linn. 

(Z.  africanum,  A.  Ad.;  L.  fossatum,  Gould?) 

Group  of  Z.  cingulatum. 

L.  cingulatum,  Lamk.  Z.  voigtii,  Anton. 

Z.  poulsfinii,  Morch. 

Group  of  Z.  j^iZeare. 

Z.  pileare,  Linn.  Z.  krebsii,  Morch. 

Z.  aquatile,  Reeve.  Z.  intermedium,  Reeve. 

Z.  tnundum,  Reeve.  Z.  gemmatum,  Reeve. 

L.Jicoides,  Reeve.  Z.  rubeculum,  Linn. 

Z.  durbanense,  Smith.  Z.  beccai'ii,  Tap.-Can. 

Z.  veliei,  Calkins.  Z.  lineatum,  Brod. 

Group  of  Z.  labiosum. 

Z.  labiosum.  Wood.  Z.  strangei,  Ad.  &  Ang. 

Z.  rutilum,  Menke.  Z.  loroisii,  Petit. 

Z.  orientale,  Nevill.  Z.  crispum.  Reeve. 


BY    H.    LEIGHTON    KESTEVEN.  473 

Group  of  L.  cynocephalum. 
L.  cynocephalum,  Lamk.  L.  pyrum,  Linn. 

L.  moritinctum,  Reeve.  L.  sarcostomum,  Reeve. 

Group  of  L.  tuberosum. 
L.  tuberosum,  Lamk.  L.  encausticum,,  Reeve. 

L.  retusum,,  Lamk.  Z.  clavator,  Lamk. 

L.  dunkerij  Lischke.  L.  mauritianum,  Tap. -Can. 

Group  of  L.  irilineatum. 
L.  trilineatiun,  Reeve.  L.  testudinarium,  Ad.  &  Reeve. 

L.  sitiense,  Reeve.  L.  aegrotum,  Reeve. 

L.  gallinago,  Reeve.  L.  moniliferum,  Ad.  &  Rve. 

L.  exile,  Reeve.  L.  caudatum,,  Gmel. 

L.  cormitum,  Perry.  L.  'pyrulum,  Ad.  k,  Rve. 

L.  pachycheilos,  Tap. -Can.  L.  tripus,  Lamk. 

L.  pfeifferianum,  Reeve. 

Group  of  L.  yibbosum, 
L.  gibbosum,  Brod.  L.  jjharciduin,  Dall. 

L.  Icebecckei,  Lischke. 
Group  of  L.  vespaceum. 
L.  vespaceum,  Lamk.  L.  elongatum,  Reeve. 

L.  thersites.  Reeve.  L.  gracile.  Reeve. 

L.  amictum.  Reeve.  L.  tenuiliratum,  Lischke. 

Group  of  L.  quoyi. 
L.  quoyi.  Reeve.  L.  eburneutn,  Reeve. 

L.  verrucosum.  Reeve.  Z.  nodocostatum,  Tate  &  May. 

L.  pliilomelce,  Watson.  L.  oligostirum,  Tate. 

Group  of  L.  co7ivolutum. 
L.  convolutum,  Brod.  L.  scalariforme,  Brod.^ 

Group  of  L.  parkinsonianum. 
L.  parkinsonianum,  Perry.  Z.  radiale,  Tate. 

Z.  abbotti,  T.-Woods.  Z.  woodsi,  Tate. 

Z.  textile,  Tate.  Z.  tortirostris,  Tate. 

*  These  two  species  will  probably  prove  to  be  Trophon. 


474  NOTES    ON    PROSOBRANCHIATA, 

Group  of  L.  tritonis. 

L.  tritonisj  Linn.  L.  ovoideum,  Tate. 

L.  rubicundum,  Perry.  L.  variegatum,  Lamk. 

L.  saulice,  Reeve.  Z.  australe,  Lamk. 

L.  bassi,  An  gas. 

The  following  species  are  not  conveniently  referable  to  any  of 
the  above  groups  : — 

L.  subdistorlicm,  Lamk.  L.  tumulosum,  Tate. 

L.  cyphus,  Tate.  L.  protensum,  Tate. 

L.  clandestinum,  Lamk.f  L.  fraterculum,  Dkr. 

Species  incertse  sedis. 

L.  gramdatum,  Dunker.  L.  contabulatum,  Anton. 

L.  birostoinum,  A.  Ad.  L.  tringa,  A.  Ad. 

L.  papillosum,  A.  Ad.  L.  nodoliratum,  A.  Ad. 

L.  dorsuosujii,  A.  Ad. 

This  arrangement,  which  is  similar  to  that  adopted  by  Pilsbry 
for  the  species  of  the  various  genera  of  Helices  (28),  and  Simpson 
for  the  Naiades  (31),  has  advantages  possessed  by  no  previous 
arrangement  of  the  genus.  None  of  the  sectional  names  are 
([uotable,  and  are,  therefore,  not  additions  to  an  already  over- 
burdened nomenclature.  As  the}'-  consist  of  essentially  similar 
species  a  reference  to  one  of  these  groups  should  at  once  convey 
to  the  reader  a  tangible  type;  they  should,  on  that  account,  be 
of  service  to  the  specialist  and  to  the  general  conchologist  when 
describing  new  forms.  A  monograph  arranged  in  this  way  should 
greatl}'-  facilitate  identifications.  Since  I  have  worked  to  some 
extent  from  figures  and  descriptions  only,  some  of  m}'-  groups 
may  contain  species  which  break  their  uniformity. 

Miscellaneous  Notes. 

Wliile  at  work  on  this  paper  I  have  made  miscellaneous  notes 
of  interest,  and  arrived  at  conclusions  at  variance  with  Tryon's 

t  Vide  post,  p.  479. 


BY    H.    LEIGHTON    KESTEVEN.  475 

specific  nomenclature.     I  have  thought  it  would  be  useful  to 
collect  these  together. 

LOTORIUM  WATERHOUSEI,  Ad.   (t  Ang. 

This  species,  rightly  treated  as  distinct  by  Tryon,  has  been 
relegated  to  the  synonymy  of  L.  spengleri,  Chemnitz,  by  Pritchard 
and  Gatliff  (29,  264).  The  revolving  ribs  of  spengleri  are  broader 
than  their  interstices;  those  of  luaterhousei  are  only  half  as  broad, 
and  duplicate.  The  transverse  lirse  of  the  former  are  very  much 
coarser  than  those  of  the  latter.  The  varices  of  L.  waterhousei 
are  filled  up  inside,  whereas  in  L.  sj^engleri  the  porcellanous 
interior  may  be  seen  in  every  varix.  The  epidermis  of  L. 
spengleri  is  quite  smooth;  that  of  ivaterhousei  is,  as  its  authors 
say,  "squamato-pilose."  In  Port  Jackson,  where  L.  spengleri  is 
to  be  found  on  most  of  the  rocks,  L.  waterhousei  never  occurs. 

LoTORiUM  GRACiLE,  Reeve. 

Both  Watson  (39,  394)  and  Dall  (7,  227)  disagree  with  Tryon 
that  this  is  a  synonym  of  L.  vespaceum. 

LoTORiuM  PARKiNSONiANUM,  Perry. 

I  have  said  of  this  species  that  it  is  not  related  to  the  ^Hritonis '' 
group.  In  arriving  at  this  conclusion  I  have  been  influenced  by 
the  evolution  of  the  species  as  evidenced  by  its  stages  of  growth. 
It  is  one  of  the  few  species  in  which  a  neanic  stage  is  plainly 
definable.  I  regard  it  as  being  a  very  recent  species  evolved 
from  L.  to7'tirostris,  Tate.  My  reasons  for  so  regarding  it  are — 
(1)  it  has  the  contour  of  this  species;  (2)  had  it  existed  in  Tertiary 
times  it  is  reasonable  to  suppose  it  would  have  been  found  in 
these  beds^  especially  as  (3)  it  is  common  on  the  coasts  of  South- 
Eastern  Australia;  (4)  in  the  neanic  stage  it  is  ornamented  with 
the  complex  sculpture  of  this  form  (vide  19,  pi.  36,  ff.  3,  4);  (5) 
some  specimens  of  L.  tortirostris  show  a  tendency  to  lose  their 
complex  sculpture  on  the  ultimate  whorls.  On  the  other  hand, 
there  is  undoubtedly  an  underlying,  but  indefinable,  resemblance 
to  the  "  tritonis  "-group.      This,  it  seems,  is  to  be  attributed  to  a 


476  NOTES    ON    PROSOBRANCHIATA, 

**  convergence  of  development "  rather  than  a  real  affinity;  for  if 
the  species  of  this  group  have  been  evolved  from  ancestors  orna- 
mented with  a  complex  sculpture,  it  was  at  an  earlier  period  than 
that  at  which  L.  parkinsonianum  was  evolved,  since  none  of  them 
show  any  sign  thereof  on  their  early  whorls.  Further,  two  species 
belonging  typically  to  the  tritonis-grow^  occur  in  Tertiary  strata 
— L.  ovoideum,  Tate,  from  Australian  beds,  and  L.  nodi/erum, 
Lamk.,  which  has  been  recorded  from  Mediterranean  beds  by 
Hoernes  and  Auinger  (is). 

LoTORiUM  PiLEARE,  Linn. 

For  this  species  Linnaeus  quoted  figures  of  two  species,  and 
described  a  third.  Hanley  (14)  says  his  type,  on  which  his 
description  was  founded,  was  the  Triton  corrugatus  of  Lamarck, 
and  refers  to  Reeve's  figure  in  the  '  Conchologica  Iconica'  (pi.  v., 
f.  15).  The  first  of  these  three  species  to  be  recognised  as  distinct 
was  Lotorium  costatum  by  Born  in  1780.  Lamarck,  in  dealing 
with  the  species,  unfortunately  bestowed  a  new  name  on  Linnaeus' 
type,  and  retained  the  name  pilearis  for  the  species,  which  has 
since  been  known  under  that  name. 

The  principles  of  nomenclature,  however,  require  that  in 
eliminating  from  an  heterogeneous  group,  the  name  originally 
bestowed  thereon  shall  be  retained  for  the  type  when  that  t3'pe 
is  available. 

The  synonymy  of  these  two  species  will,  therefore,  stand  thus  : — 

Lotorium  pileare,  Linn.  H 

'i 

Mur ex  pilearis,  Linn.,  Syst.  Nat.  1767,  p.  1217,  sp.  534. 
Triton  corrugatus,  Lamarck,  An.  s.  Vert,  vii.,  1822,  p.  181;  Reeve, 
Conch.  Icon.  ii.  ''Triton,''  pi.  v.,  f.  15,  1844. 

Lotorium  aquatile,  Reeve. 

Triton  pilearis,  Lamarck,  An.  s.  Vert.  vii.  1822,  p.  182  (non  Linn.). 
Triton  aquatilis,  Reeve,  Conch.  Icon.  ii.  "  Triton,''  pi.  vii.,  f.  24, 

April  1844;  P.Z.S.  1844  (December),  p.  114. 
Triton  vestitus,^A\x\^^,  Zool.  Voy.  Sulphur,  "Mollusca,"  Oct.  1844; 

P.Z.S.  1844  (July),  p.  21. 


BY    H.    LEIGHTON    KESTEVEN.  477 

T.  martinianum,  D'Orb.,  Moll.  Cuba,  ii.,  p.  162,  1847. 

T.  hrasilianum,  Gould,  Proc.  Bost.  Soc.  Nat.  Hist,  iii.,  p.  142,  1849. 

T.  americanum,  D'Orb.,  Moll.  Cuba,  ii.,  p.  163,  pi.  23,  f.  22,  1853  (?) 

(Jide  Morch,  Malac.  Blatt.  xxiv.,  p.  28,  1877). 
T.  intermedius,  Pease,  Amer.  Jour.  Conch,  v.,  1869,  p.  84. 
T.  veliei,  Calkins,  Proc.  Davenport  Acad.   Sci.  ii.,  p.  235,  pi.  viii., 

ff.  1-2,  1878  (ex  type,  Dall). 
In  giving  the  above  synonymy  of  L.  aquaiile,  I  have  accepted 
the  opinions  of  and  quoted  from  Dall  (7,  227)  and  Tryon  (36.  12). 
Not  having  specimens  of  some  of  them  I  am  unable  to  judge  for 
myself.  The  inclusion  of  T.  americanum,  D'Orb.,  seems  doubtful, 
since  Watson  (39,  390)  regarded  it  as  a  synonym  of  L.  costatum, 
Born.  I  have  recovered  the  exact  date  of  the  publication  of  the 
Mollusca  of  the  Sulphur  from  Pace's  admirable  work  on  the 
Columhellidce  (26).  In  arriving  at  the  above  conclusions  I  have 
had  the  advantage  of  consulting  with  Mr.  C.  Hedley,  and  they 
may,  therefore,  be  regarded  as  a  joint  opinion. 

LoTORiuM  QUOYi,  Reeve. 

I  cannot  agree  with  Pritchard  and  Gatliff  {op.  cit.)  that  this  is 
only  a  form  of  L.  verrucosum,  Reeve;  and  that  the  name  should 
be  relegated  to  the  synonymy  thereof.  Than  that  species  L. 
quoyi  is  smaller,  has  flatter  whorls,  is  more  regular  in  growth, 
has  a  slightly  smaller  and  higher  protoconch.  The  body  whorl 
of  quoyi  is  sculptured  by  revolving  and  transverse  riblets  of  equal 
strength,  seven  of  the  former  and  nineteen  of  the  latter.  Z. 
verrucosum  has  six  of  the  former  and  eleven  of  the  latter,  these 
being  twice  as  strong  as  revolving  riblets.  This  sculpture  gives 
L.  quoyi  a  beaded  appearance  which  the  other  species  does  not 
possess.  They  were  always  treated  as  distinct  species  by  Prof. 
R.  Tate,  whose  acquaintance  with  them  was  almost  life-long. 

LoTORiuM  RUTiLUM,  Menke. 

This  species,  considered  by  Reeve  to  be  synonymous  with  L. 
labiosum,  Wood,  has  lately  been  identified  by  Mr.  C.  Hedley 
from  the  type  locality.  He  finds  that  it  is  a  valid  species,  and 
has  in  preparation  a  note  on  the  subject. 


478  NOTES    ON    PROSOBRANCHIATA, 

LoTORiUM  ELONGATUM,  Reeve. 

Treated  by  Tryon  as  a  synonym  of  L.  vespaceum.  It  is  more 
elongate,  not  so  heavil}^  transversely,  but  more  heavily  spirally 
sculptured,  and  has  a  smaller  protoconch.  Watson  {oj).  cit.)  has 
already  maintained  the  validity  of  this  species. 

LoTORiuM  THERSITES,  Reeve. 

Tryon  regards  this  as  the  mature  condition  of  L.  vespaceimi,  sl 
conclusion  with  which,  after  studying  a  fairly  large  series,  I 
cannot  agree.  Whilst  being  slightly  shorter,  it  is  decidedly 
broader,  has  a  shorter  canal,  is  somewhat  distorted  (vespaceum  is 
quite  regular),  is  more  prominently  angled  at  the  periphery,  has 
more  varices,  fewer  and  stronger  intervarical  ribs,  and  is  more 
delicately  sculptured.  The  intervarical  ribs  do  not  continue  on 
the  body  whorl  so  far  towards  the  canal  as  Reeve's  figure  shows. 
The  revolving  lirse,  which  are  smaller,  more  finely  beaded,  and 
equal,  incline  to  fade  away  between  the  transverse  ribs,  on  the 
crests  of  which  and  on  the  varices  they  are  strongest.  The  inter- 
stitial striae  are  also  finely  beaded.  The  protoconch  of  L.  vesjmceum 
is  the  larger  and  set  on  straight,  whilst  that  of  the  species  under 
discussion  is  set  obliquely. 

LoTORiUM  MORiTiNCTUM,  Reeve. 

This  was  quoted  by  Dunker  (op.  cit.),  and  more  recently  by 
Stearns  and  Pilsbry  (op.  cit.).  Martens  also  used  the  name  (07;. 
cit.). 

LoTORiuM  SAULiii:,  Reeve. 

Quoted  by  Stearns  and  Pilsbry  (op.  cit.).  With  reference  to 
Lotorium  australe,  Lamk.,  Pritchard  and  Gatlifi"  (o^:>.  cit.)  write: 
"In  our  opinion  it  is  doubtful  that  T.  nodifer,  Lamk.,  and  T. 
saulice,  Reeve,  are  identical,  and  we  cannot  concede  that  T. 
australis  is  also  identical.'' 

Lotorium  strangei,  Ad.  &  Ang. 

This  species,  confounded  by  Tryon  with  L.  labiosum,  Wood, 
and  by  Pritchard  and  Gatlifi"  (op.  cit.)  treated  as  the  young  of  L. 


BY    H.    LEIGHTON    KESTEVEN. 


47^ 


spengleri,  Chemnitz,  has  rightly  been  maintained  a  good  species 
by  Smith  (32).     Its  affinities  are  with  L.  labiosum. 


"Triton  speciosus,"  Angas. 

(Text  fig.  3.) 

This  species  is,  as  Tryon  suspected, 
a  Troplion.  I  give  figures  of  its  oper- 
culum and  dentition. 

LOTORIUM  CLANDESTINUM,   Chcmn. 

This  species  would  be  best  trans- 
ferred to  Apollon. 

LoTORiUM  cosTATUM,  Born. 
The  complicated  synonymy  of   this 
species    is    dealt  with    at    length   by 
Watson  (39).     Some  of  the  synonyms 


Fig.  3. 
Tritoniiim  speciosum,  Angas. 


might  advantageously  be  retained  for  varieties. 

LOTORIUM  NODOCOSTATUM,  Tate  ife  May. 

(Text  fig.  4.) 

From  the  material  above  referred  to  I  have 
drawn  the  accompanying  sketch.  From  this 
it  will  be  seen  that  New  South  Wales  specimens 
'f/^  have  a  longer  canal  than  the  southern  form, 
unless  the  description  and  figure  of  the  species 
were  drawn  from  a  mutilated  specimen.  The 
co-type  lent  me  by  Mr.  May  has  the  canal 
broken  short  oif,  and  is,  therefore,  of  no 
assistance  on  this  point. 

The  length  of  canal  w^ould  seem  to  exclude 
the  species   from  the  group  in  which  I  have 
placed  it,  but  the  tout  ensemble  is  that  of  this  group. 


Fig.  4. 

L.  nodocostatum, 
Tate  &  May. 


480  NOTES    ON    PROSOBRANCHIATA, 

LoTORiUM  FRATERCULUM,  Dunker. 
(Text  fig.  5.) 
Triton  f rater  cuius,  Dkr.,  Malac.  Blatt.  1871,  p.  166. 
Sipho(1)  mimeticus,  Tate,  Trans.  Roy.  Soc.  S.  Aust.,  1893,  Pt.  i., 

p.  189,  pi.  i.,  f.  10. 
Triton  mimeticus,  Verco,  loc.  cit.,  1895,  p.  88,  pi.  2,  ff.  4,  4a. 
Triton  [Argohuccinum)  mimeticus,  Verco,  loc.  cit.,  1896,  p.  221, 
pi.  5,  ff.  6,  6a,  6b. 

Some  little  time  ago  Mr.  Hedley  drew  my  attention  to  the  fact 

that  the  description  given  by  Dunker  under  the  above  name  could 

not   apply  to  Lotorium   bassi,   Angas,   as 

stated  bySchmeltz,"^  Tryon  and  others.     It 

subsequently    occurred    to    me    that    the 

diagnosis  described  Tate's  Sipko  (?)  minieti- 

cus.     Thanks  to  the  courtesy  of  Dr.  Verco, 

who  has  very  kindly  sent  me  specimens 

^    .    ,       ,       -Tk     1        to  study,  I  am  enabled  to  oive  the  follow- 
L.  Jraterculum,  Dunker.    .  •"  ° 

ing  note  on  the  subject : — 

The  specimens  do  not  harmonise  perfectly  with  Dunker's 
description,  his  statement  that  his  specimens  had  a  short  canal, 
in  particular  disagreeing.  The  description  is,  however,  fairly 
detailed,  and,  but  for  this,  applies  to  the  examples  before  me,  the 
measurements  and  proportions  fitting  exactly.  Mr.  Hedley  agrees 
with  me  that,  on  the  whole,  the  species  had  best  be  known  under 
Dunker's  name.  It  is  a  true  Lotorium,  though  it  cannot  be 
referred  to  any  of  my  groups,  and  has  a  faint  resemblance  to  the 
parkinsonianum-gvow'^,  but  from  these  its  rather  long  canal 
separates  it. 

This  species  has  a  most  striking  resemblance  to  Stre2jtosip>hou 
porphyrostoma,  Ad.  &  Rve.f  The  apex  also  bears  no  little 
resemblance  to  that  of  the  family  to  which  Streptosiphon  belongs. 
The  dentition,  however,  shows  the  species  to  belong  to  Lotorium. 

*  Mus.  God.  Cat.  v.,  1874,  p.  139. 
t  Reeve,  Conch.  Icon.  iv.  {Faciolaria),  pi.  v.,  f.  11,  1847. 


BY    H.    LEIGHTON    KESTEVEN.  481 

LoTORiUM  BARTHELEMYi,  Bemardi. 

This  unlocalised  species  is,  as  stated  by  Pritchard  and  Gatliff 
(o^j.  cit.),  a  synonym  of  L.  spengleri,  Chemn. 

The  following  names  proposed  by  Perry  in  1811  (27)  have  lately 
been  reinstated  by  Mr.  C.  Hedley  (le) : — 

Perry's  specific  name.  Specific  synonym  in  use  heretofore. 

Lotoriutn  cornutum.  Lotorium  exarcUum,  Reeve. 
,,       parkinsonianum  ,,        fusiforme,  Kiener. 

,,        ruhicundum  ,,        nodiferurn,  Lamk. 

In  the  same  paper  Mr.  Hedley  says  "  Monoplex  formosus,  Perry, 
is  recognised  by  Deshayes  as  a  synonym  of  Triton  clavator, 
Lamarck,  1822  (  —  Murex  clavator,  1817).  Triton  sinensis,  Reeve, 
seems  to  me  identical."  At  my  request  he  has  gone  over  this 
ground  again  with  me,  and,  as  far  as  our  material  allows  us  to 
judge,  we  have  come  to  the  following  conclusions  : — 

(1)  Triton  clavator,  Reeve,  is  not  clavator,  Chemn.,  1795, 
Dillwyn  (9),  Lamarck  (21),  and  should  be  quoted  as  Lotorium 
formosum,  Perry. 

(2)  Lotorium  clavator,  Chemn.,  (4)  (a  binonial  name),  is  probably 
the  correct  name  for  the  form  hitherto  known  as  Triton  sarcostoma^ 
Reeve. 

(3)  Lotorium  sine^ise,  Reeve  (30),  is  a  valid  species. 

In  conclusion  I  have  to  thank  Mr.  C.  Hedley  for  a  great  deal 
of  assistance  rendered  me  in  my  search  through  literature;  and 
the  Curator,  Mr.  R.  Etheridge,  Junr.,  through  whose  kindness  I 
have  been  enabled  to  study  Australian  Museum  material. 


Works  Eeferred  to. 

1. — Adams,  H.  &  A. — Genera  of  Recent  Shells,  Vol.  i. 

2. — Agassiz,  L. — Nomenclator  Zoologicus  (Index  Univers.),  p.  51. 

3.— Baker.— Ann.  New  York  Acad.  1897,  Vol.  ix. 

4.— Chemnitz.— Conch.  Cab.  xi.,  1795,  p.  110,  pi.  190,  figs.  1825-26. 

5.— Chenu. — Manuel  de  Conchyliologie. 


482  NOTES    ON    PROSOBRANCHIATA, 

6.— Dall,  W.  H.— 'Tertiary  Mollusca  of    Florida.'     Wagner  Free   Inst. 

Science,  Vol.  iii. 
7. 'Blake  Mollusca,' Pt.  ii.      Bull.  Mus.  Comp.   Zool.  Harv.  Coll. 

Vol.xvii.,  1889. 
8. Dall  &  Simpson. — 'Mollusca  of    Porto  Kica.'     Bull.  U.S.  Fish  Com- 
mission, 1900,  Vol.  i.,  pp.  416-417. 
9.— DiLLWYN— Descrip.  Cat.  of  Recent  Shells,  1817,  Vol.  ii.,  p.  701. 
10.— DuNKER.— Index  Mollusc.  Maris  Japonici,  1882. 
11. —Fischer. — Journ.  de  Conchyliologie,  1858,  p.  206. 

12. Manuel  de  Conchyliologie,  1887. 

13.— Gray,  J.  E.— Proc.  Zool.  Soc,  London,  xv.,  1847,  p.  133. 

14. — Hanley. — Ipsa  Linnsei  Conchylia,  1855,  p.  287. 

15. Harris,  G.  F.— Cat.  Tertiary  Moll,  in  Brit.  Mus.    Pt.  i.    Australasia, 

1897,  p.  186. 
16.— Hedley,  C— Proc.  Linn.  Soc.  N.S.  Wales,  1902,  xxvii.,  pp.  24-28. 
17. 'Thetis  Pelecypoda.'     Mem.  Aust.  Mus.  iv.,  1902,  pt.  5, 

p.  288. 
18.— HoERNES  &  AuiNGER. — Abhaud.  K.  K.  Geologischen  Eeich.     Bandii., 

1884,  p.  178,  Taf.  xxi. 
19. — Kesteven,  H.  Leighton— Proc.    Linn.   Soc.   N.  S.  Wales,  1901,  xxvi. 

(1902). 
20.— KoBELT.— Jahrb.  Malak.  Gesell.  1878,  pp.  241-250,  360-369. 
21. — Lamarck. — An.  s.  Vert.,  Deshayes'  edit.,  ix.,  1845,  p.  635. 
22.  — Martens     in    Mobius,    '  Meeresf auna   der  Insel    Mauritius   und    der 

Seychellen. ' 
23. — Melvill  &  Standen. — Journal  of  Conchology,  1895,  p.  110. 

24. Oj).  cit.,  1898,  Vol.  ix.,  pp.  43-44. 

25. — Ortmann,  a.  E. — Reports  of  Princeton  Univ.  Expeditions  to  Patagonia, 

Vol.  iv.  p.  ii. 
26. — Pace. — Proc.  Malac.  Soc,  Vol.  v.,  pt.  2,  1902,  p.  119  (sub  voce  Colum- 

bella  pavonia,  Hinds). 
27. — Perry. — Conchology,  1811. 

28. — PiLSBRY. — Manual  of  Conchology,  Vol.  ix.  (2nd  ser.),  1894. 
29. — Pritchard  &  Gatliff. — '  Cat.  Marine  Shells  of  Victoria.    Pt.  i.'     Proc. 

Roy.  Soc.  Vic.  1897. 
30. — Reeve. — Conch.  Icon.,  ii,,  Triton. 

31. — Simpson.— 'Synopsis  of  the  Naiades.'   Proc.  U.S.  Nat.  Mus,,  Vol.  xxii. 
32.— Smith,  E.  A.— Proc.  Zool.  Soc.  London,  1878,  p,  816,  pi.  50,  f.  16. 
33. — Stearns  &  Pilsbry. — Cat.  Marine  Moll,  of  Japan,  1895. 
34.— Tate.— Proc.  Roy.  Soc.  South  Australia,  1886-7  (1S88). 
35. Oj).  cit.  for  1888,  p.  245. 


BY  H.  LEIGHTON  KESTEVEN.  483 

36. — Tryon. — Structural  and  Systematic  Conchology. 

37. Manual  of  Conchology,  Vol.  iii. 

38. — Vekrill,  A.  E. — '  Eevision  of   the  Pectinidse.'       Trans.   Connecticut 

Acad.,  X.,  pt.i.,  1899. 
39. — Watson — Challenger  Rep.  Zool.,  xv..  Gasteropoda. 


EXPLANATION  OF  PLATE  XVII. 

LOTOEIUM   APICES. 

Fig.  1. — Lotoriinn  woodsi,  Tate, 

Fig.  2. —  ,,  radiale,  Tate. 

Fig.  3. —  ,,  tumulosum,  Tate. 

Fig.  4. —  ,,  ahbotti,  Tenison- Woods. 

Fig.  5. —  ,,  protensum,  Tate. 

Fig.  6. —  ,,  cyphus,  Tate. 

Fig.  7. —  ,,  oUgosfirum,  Tate. 

Fig.  8. —  ,,  tortirostris ,  Tate. 

Fig.  9.—  ,,  textile,  Tate. 

Fig.  10. —  ,,  cornutum,  Perry. 

Fig.  11. —  ,,  vespaceum,  Lamarck. 

Fig.  12. —  ,,  chlorostomum,  LiSiniSiYck. 

Fig.  13. —  ,,  aquatile,  Pteeve. 

Fig.  14. —  ,,  sinens e,  BiBewe. 

Fig.  15. —  ,,  gemmatum,  Eeeve. 

Fig.  16. —  ,,  elongatum,  Reeve. 

Fig.  17. —  ,,  tuherosiim,  Lamarck. 

Fig.18. —  ,,  thersites,  Reeve. 

Fig.  19. —  ,,  spengleri,  Chemnitz. 

Fig.  20. —  ,.  strangei,  Adams  &  Angas. 

Fig.  21. —  ,,  rutilum,  Menke. 

Fig. 22. —  ,,  lahioswn,  Wood. 

Fig.  23. —  ,,  verrucosum.  Reeve. 

Fig. 24. —  ,,  ehurneum,  Reeve, 

Fig.  25, —  ,,  quoyi,  Reeve. 


484 


DESCRIPTIONS  OF  SOME  NEW  ARANEID^  OF  NEW 
SOUTH  WALES.     No.  10. 

By  W.  J.  Rainbow,  F.L.S.,  F.E.S. 

(Entomologist  to  the  Australian  Museum.) 

Plate  xviii. 

The  present  paper  contains  descriptions  of  three  apparently- 
new  species,  viz.,  one  Storena,  one  Araneus,  and  one  Stephanopsis. 
A  species  of  Celcenia,  probably  distincta,  O.  P.  Camb.,  is  also 
described. 

Family  ZODARIID^. 

Subfamily  ZODARIIN.aE. 

Simon  divides  this  subfamily  into  five  sections,  viz.,  Hermippese, 
Storene{«,  Trygetinse,  Suffuciese,  and  Zodariese.  The  first  of  these 
embraces  only  one  species — Hermippus  loricahis,  E.  Sim. — and 
this  is  only  known  from  Central  Africa;  the  second  includes  four 
genera  :  Lachea,  Thor. ;  Storenosoma,  Hogg ;  Storena,  Walck. ; 
and  Lufica,  G.  Marx;*  this  is  the  only  group  in  the  Zodariinse 
that  is  represented  in  Australia,  and  this,  again,  by  two  genera 
— Storenosoma  and  Storena.  The  latter,  which  is  one  of  thej 
most  polymorphous,  has  a  wide  range,  occurring,  according  to| 
Simon,  in  the  "  Regio  mediterranea  occid.  et  orient.;  Afric.  trop.; 
Indo-China;  Malaisia;  N.  Guinea,  N.  Hollandia,  N.  Caledonia  etj 
Polynesia;  Amer.  septentr.  sequator.  et  australis." 

Storena  appears  to  be  more  numerous  in  species  in  Australia] 
than  elsewhere,  fifteen    having    been    previously  recorded,  and' 
all  from  the  mainland.     S.Jormosa,  L.  Koch,  which  was  originally 
recorded  from  Bowen,  and  which  is  not  uncommon  around  Sydney, 
occurs  also  in  Tasmania. 

*  Simon,  Hist.  Nat.  des  Araignees,  2nd  ed.,  pt.  ii.,  1893,  p.  423,  et  seq. 


BY    W.    J.    RAINBOW.  485 

The  eyes  of  the  species  are  very  varied,  no  two  forms  being 
exactly  alike,  whilst  in  some  instances  the  differences  are  striking. 
The  form  described  hereunder  is  no  exception  to  the  rule;  it  is 
the  first,  however,  that  I  have  examined  in  which  the  eyes  are  of 
equal  size.  The  legs  are  also  very  variable  in  respect  of  their 
relative  lengths.  They  are  as  follows  :  (a)  4,  1,  2,  3;  (b)  4,  1,  3,  2; 
or  (c)  4,  2,  3,  1.     S.  albomaculata  conforms  to  the  first  of  these. 

A  number  of  Australian  species  were  described  by  L.  Koch 
under  the  generic  name  Habronestes,  L.  Koch.  This  has  been 
sunk  by  Simon  as  a  synonym. 

Genus  S  t  o  r  e  n  a,  Walck. 

Storena  albomaculata,  sp.nov. 

(Plate  xviii.,  figs.  1,  la.) 

9-  Cephalothorax  3  7  mm.  long,  2*9  mm.  broad;  abdomen  5 -6 mm. 
long,  3-5  mm.  broad. 

Ce2jhalothorax  obovate,  shining,  black,  relieved  by  patches  of 
white,  scale-like  hairs,  and  a  margin  or  belt  of  the  same;  there 
are  also  a  few  long  black  hairs  scattered  over  the  surface.  Pars 
cephalica  strongly  arched,  somewhat  obtusely  truncated  in  front, 
black,  relieved  on  each  side  (near  the  margin)  by  a  large  and 
prominent  patch  of  white  hairs;  normal  groove  separating  cephalic 
from  thoracic  segment  faintly  visible.  Clypeus  broad,  fringed 
with  a  few  rather  long  black  bristles.  Pars  thoracica  broad, 
strongly  arched,  black,  with  lateral  and  posterior  patches  of  white 
hairs ;  median  and  longitudinal  groove  distinct,  lateral  radial 
grooves  indistinct;  sides  and  posterior  angle  declivitous.  Mar- 
ginal  band  broad,  thickly  clothed  with  short  white  hairs. 

Eyes  prominent,  of  equal  size,  and  arranged  in  two  procurved 
rows  of  four  each,  of  which  the  posterior  is  the  most  strongly 
bent;  those  comprising  the  posterior  row  are  closely  set,  and 
nearly  touch  each  other,  whilst  the  four  constituting  the  posterior 
series  are  equidistant,  each  eye  being  separated  from  its  neigh- 
bour by  a  space  equal  to  about  once  its  own  diameter. 

Legs  moderately  long  and  tapering,  robust,  black,  with  patches 
of  white  hairs,  and  throwing  metallic  scintillations  at  all  the 
32 


4:SQ  DESCRIPTIONS    OF    SOME    NEW    ARANEID^E    OF    N.S.^Y., 

joints;  each  ambulatory  limb  is  armed  with  a  few  long  but  not 
strong  spines,  and  clothed  with  short  black  hairs.  Relative 
lengths :  4,  1,  2,  3. 

Falpi  long,  tapering,  black,  armed  with  long  but  not  strong 
spines,  and  clothed  with  short  black  hairs. 

Falces  black,  shining,  arched,  apices  divergent,  furnished  with 
long  black  bristles. 

Maxillce  strong,  arched,  shining,  apices  obtuse  and  divergent, 
outer  angles  and  upper  surface  black,  inner  margins  golden  brown 
at  base,  pale  yellowish  at  tips;  inner  angles  fringed  with  long- 
white  hairs. 

Labium  broader  than  long,  arched,  rounded  off  at  apex, 
shining,  the  base  black,  and  the  apex  golden  brown. 

Sternum  oblong-oval,  black,  glossy,  moderately  convex,  hairy. 
Abdomen  ovate,  convex,  barely  overhanging  base  of  cephalo- 
thorax,    pubescent,     black-brown     with    metallic    scintillations, 
ornamented  with  large  and  small  median  and  lateral  oval,  round, 
and  reniform  patches  of  white  hairs,  distributed  as  illustrated  in 
the  figure;  sides  speckled  with  white;  ventral  surface  dark  brown. 
Epigyne  as  in  figure. 
Loc. — Burwood. 

This  species,  by  its  scintillations,  recalls  the  Rev.  0.  P.  Cam- 
bridge's spider,  S.  scintillaiis,  but  differs  therefrom  not  only  in 
size  and  ornamentation,  but  also  by  the  fact  that  the  eyes  of 
the  posterior  row  are  equally  as  large  as  those  of  the  anterior 
row,  whilst  in  scintillans  the  anterior  eyes  are  much  smaller  than 
their  posterior  neighbours. 

Family  ARGIOPID^. 

Subfamily  ARGIOPIN-ffi. 

Genus  A  r  a  n  e  u  s,  Walck. 

Araneus  liberalis,  sp.nov. 

(Plate  xviii.,  figs.  2,  2a.) 

9.  Cephalothorax  2-G  mm.  long,  18  mm.  broad;  abdomen 
4  4  mm.  long,  4  mm.  broad. 


BY    W.    J.    RAINBOW.  487 

Cephalothorax  obovate,  longer  than  broad,  yellowLsli-brown, 
smootli,  shining.  Pars  ce2:)halica  narrow  in  front,  but  becoming 
gradually  wider  towards  the  segmental  groove,  strongly  arched, 
sparingly  furnished  with  pale  yellowish  pubescence.  Clypetis 
broad,  sloping  inwards,  sparingly  clothed  with  fine  and  moderately 
long  hairs.  Pars  thoracica  sparingly  pubescent,  broad,  strongly 
arched,  median  longitudinal  depression  deep,  radial  grooves 
distinct.      Marginal  band  broad. 

Eyes  black;  the  four  comprising  the  median  group  are  of  equal 
size,  and  elevated  upon  a  tubercular  eminence,  and  form  a 
trapezium;  of  these  the  anterior  pair  are  separated  from  each 
other  by  a  space  equal  to  twice  their  individual  diameter,  whilst 
those  of  the  second  row  are  less  than  once  their  individual 
diameter  apart,  and  the  two  rows  are  separated  from  each  other 
by  a  space  equal  to  rather  more  than  the  diameter  of  one  eye; 
the  lateral  eyes  are  minute,  contiguous,  and  elevated  obliquely  upon 
small  tubercles;  of  each  of  these  pairs  the  anterior  eye  is  slightl}' 
larger  than  the  posterior. 

Legs  long,  tapering,  concolorous  with  cephalothorax,  sparingly 
clothed  with  yellowish  hairs,  and  armed  with  long  black  spines. 
Relative  lengths:  1,  2,  4,  3. 

Pal'pi  long,  tapering,  similar  in  colour,  clothing  and  armature 
to  legs. 

Falces  long,  concolorous  also,  arched,  smooth,  shining,  apices 
divergent,  sparingly  pubescent;  the  lower  margin  of  each  falx  is 
furnished  with  a  row  of  four  teeth,  and  the  upper  with  a  row  of 
three;  fangs  reddish-brown. 

Maxillce  pale  yellowish,  glabrous,  convex,  inclining  inwards, 
apices  truncated,  inner  margins  thickly  fringed  with  long  yellowish 
hairs,  surface  sparingly  hairy. 

Labium  concolorous,  short,  broad,  arched,  glabrous,  convex, 
obtusely  pointed. 

Sternum  concolorous  also,  shield-shaped,  convex,  glabrous, 
sparingly  hair3\ 

Abdomen  broadly  ovate,  overhanging  base  of  cephalothorax, 
arched,  chrome-yellow,  faintly  tinged  with  green  down  the  middle; 


488  DESCRIPTIONS    OF    SOME    NEW    ARANEID^    OF    N.S.W., 

upon  the  superior  surface  there  is  a  faintly  visible  leaf -like  design, 
and  an  ornamental  scheme  of  tracery  commencing  near  the  centre, 
and  continuing  to  posterior  extremity ;  the  sides  are  chrome- 
yellow,  relieved  by  a  series  of  faint  markings  as  indicated  in  the 
figure;  ventral  surface  yellowish-green  at  the  middle,  and  yellowish- 
brown  laterally. 

Ejngyne  as  in  figure. 

Hah. — Prospect. 

Genus  C  e  l  ^  n  i  a,  Thor. 

Cel.enia  distincta,  0.  P.  Cambr. 

(=  Thlaoso77ia  distinction,  O.  P.  Cambr.) 

(Plate  xviii.,  figs.  4,  4a,  46,  4c,  id,  ie.) 

In  1870  the  Rev.  O.  P.  Cambridge,  F.Z.S.,  described  and 
figured  a  spider  under  the  name  of  Thlaosoma  distinctum*  which, 
judging  by  his  figure  (fig.  36),  appears  to  bear  a  superficial 
appearance  to  the  one  herein  described,  and  which  is  now 
included  in  the  national  collection  at  the  Australian  Museum. 
The  specimen  upon  which  Cambridge  founded  his  species  was 
an  old  and  dry  example;  moreover  it  was  a  single  specimen,  and 
vaguely  labelled  "  New  Holland."  It  is,  or  was,  in  the  Hope 
Collection,  University  Museum,  Oxford. 

Although  Cambridge's  figure  (fig.  36  above  referred  to)  appears, 
as  I  have  pointed  out,  to  bear  a  superficial  appearance  to  my 
specimen,  his  description  and  supplementary  figure  (fig.  37)  do  not 
agree  in  several  important  details.  If,  to  the  possibility  that  the 
Hope  specimen  was  an  immature  example — which  Cambridge 
was  unable  to  determine — it  be  borne  in  mind  that  the  type  was 
also  dry,  it  will  be  readily  conceived  that  differences,  both  in 
appearance  and  coloration,  from  a  fresh  example  would  only  be 
natural.  Nevertheless  there  are  points  mentioned  in  Cambridge's 
description  which  one  would  hardly  expect  to  undergo  change, 
however  old  or  dry  the  specimen  should  be. 

It  is  impossible,  in  the  absence  of  the  type,  to  say  positively 
whether  the  example  in  my  possession  is  distinct  or  not,  and, 
*  Cambridge,  Journ.  Linn.  Soc.  Zool.,  x.,  1870,  pp.  274-5,  pi.  ix.,  figs.  36-38. 


BY    W.    J.    RAINBOW.  489 

tlierefore,  I  prefer  for  the  present  to  regard  it  as  synonj^mous 
with  Cambridge's  C.  distincta,  and  to  further  elucidate  the  matter 
by  description  and  figures. 

9.  Cephalothorax  2*4  mm.  long,  2-8  mm.  broad;  abdomen  5  mm. 
long,  6*6  mm   broad,  8-2  mm.  high. 

Ceph'dothorax  broader  than  long,  rounded  laterally,  granulated, 
and  marked  with  two  lateral,  parallel,  longitudinal  grooves; 
yellowish-brown,  densely  clothed  with  short  coarse  hairs,  and 
having,  in  addition,  two  prominent  brushes  of  long,  thick,  coarse 
hairs  running  from  the  anterior  region  of  the  cephalic  segment 
to  the  posterior  angle  of  the  thoracic  segment,  and  there,  com- 
mencing one  on  each  side  of  the  caput,  incline  inwards  until  near 
the  centre,  where  they  meet,  and  from  whence  they  again 
gradually  widen  out,  and  do  not  terminate  until  the  posterior 
angle  is  attained.  Pars  cejyhalica  arched,  ascending,  and  termina- 
ting in  front  with  an  obtuse  projection  or  eminence,  upon  which 
the  four  larger  eyes  are  elevated;  at  the  anterior  extremity  of  the 
caput  there  are  two  small  projecting  tubercles  (fig.  4:d).  Clypeus 
broad,  deep,  sloping  inwards.  Pars  thoracica  broad,  arched; 
radial  grooves  indistinct. 

Ut/es  distributed  in  three  series  of  2,  4,  2;  of  these  the  median 
group  of  four  are  much  the  largest,  and  arranged  in  a  somewhat 
trapezoidal  form,  the  anterior  being  separated  from  each  other 
by  a  space  equal  to  about  three  times  their  individual  diameter, 
and  the  posterior  pair  by  about  two  diameters;  the  lateral  eyes 
are  seated  well  back,  and  each  pair  is  elevated  upon  a  prominent 
tubercle;  they  are  minute  and  nearly  contiguous;  the  lower  one 
is  somewhat  the  smaller. 

Legs  laterigrade,  yellowish-brown,  clothed  with  short  hairs, 
and  armed  with  spines;  the  first  and  second  pairs  are  much  the 
longest,  and  are  folded  when  at  rest ;  the  femora  are  rough, 
granulated,  long,  hairy,  and  each  has  on  the  underside  a  deep 
groove  for  the  reception  of  the  lower  joints  which  fit  therein  when 
folded  like  the  blade  of  a  pocket-knife;  the  edges  of  the  grooves 
are  armed  with  numerous  short,  sharp,  strong,  tooth-like  spines; 
the  patellae  are  long,  attenuated,  elbowed  at  the  base;  the  patella 


490  DESCRIPTIONS    OP    SOME    NEW    ARANKIDiE    OF    N.S.W., 

and  tibia  of  each  of  the  first  and  second  pairs  are  smooth,  and 
are  (together)  as  long  as  the  femur;  the  tibise  are  very  attenuated; 
the  metatarsi  and  tarsi  are  short,  armed  with  small  strong  spines, 
and  clothed  rather  thickly  with  short  scale-like  hairs;  the  second 
and  third  pairs  of  legs  are  short,  and  thickly  clothed  with  short 
stiff  hairs.     Relative  lengths  :   1  =  2,  4,  3. 

Palpi  short,  concolorous,  hairy. 

Falces  normal. 

Maxilloi  short,  broad,  arched,  apices  divergent. 

Labium  short,  broad,  arched,  apex  rather  more  obtuse  than  in 
C.  excavata,  L.  Koch. 

Abdomen  overhanging  base  of  cephalothorax,  yellow-brown, 
clothed  with  short  hairs,  broader  than  long,  and  higher  than 
broad,  deeply  sculptured,  granulated,  and  surmounted  at  the 
summit  with  two  lateral  lobes,  from  each  of  which  a  tapering  and 
somewhat  acutely  pointed  process  extends;  in  front  there  are  two 
prominent  and  well-defined  ridges,  uneven  in  outline,  the  space 
})etween  which  is  rather  flat,  and  punctuated  with  six  deep 
depressions  or  pits;  the  anterior  extremity  or  angle  is  strongly 
recurved;  the  areas  to  the  right  and  left  of  the  ridges  are  rough, 
sculptured,  and  pitted;  sides  uneven  and  wrinkled;  behind  the 
contour  is  somewhat  sinuous  in  outline,  the  surface  rough  and 
uneven,  and  punctuated  with  deep  pits. 

Epigyne  :  two  deep  circular  pits  as  in  figure. 

Log. — Prospect,  near  Sydney. 

Obs. — According  to  Cambridge  his  specimen  had  "  a  very 
distinct  pattern  on  the  upper  or  forward  side  of  the  abdomen, 
formed  by  lines  of  pure  white,  longish,  scaly  hairs,  enclosing 
various  irregularly  triangular  spaces  of  deep  chocolate-brown,  on 
a  paler  yellowish-brown  ground;  three  of  these  patches  or  spaces 
are  nearly  contiguous  in  the  median  line,  a  large  one  in  front, 
and  two  smaller  ones  immediately  behind;  some  small  tufts  of 
white  hairs  also  occupy  tlie  sides,  hinder  portion,  and  other  parts 
of  the  abdomen."* 

*  Loc.  cii.,  p.  274. 


BY    W.    J.    RAINBOW.  491 

Family  THOMISID^. 

Subfamily  STEPHANOPSIN^. 

Genus  S  t  e  p  h  A  N  o  P  s  i  s,  O.  P.  Cambr. 

Stephanopsis  obtusiprons,  sp.nov. 

(Plate  xviii.,  figs   3,  3a,  ?)b.) 

9.  Cephalothorax  4'1  mm.  long.  3-5  mm.  broad ;  abdomen 
5-9  mm.  long,  5-3  mm.  broad. 

Cephahtho7'ax  longer  than  broad,  yellowish-brown,  coarsely 
rugose.  Pars  cephalica  sloping  gentl}'-  forwards  and  upwards 
until  near  the  ocular  eminence,  where  it  makes  a  slight  descent, 
and  from  whence  it  rises  almost  perpendicularly,  thereb}'-  causing 
a  deep  and  sharpl}'-  defined  groove  at  the  base  of  the  latter;  sides 
gently  rounded ;  surface  coarsely  granulated  and  rugose ;  the 
ocular  tubercular  eminence  is  not  so  high  as  in  altifrons  (O.  P. 
Cambr.)  or  aspera  (mihi);  the  summit  is  rounded  off  and  obtuse, 
and  the  sides  and  front  are  somewhat  precipitous.  Pcwa  tlioracica 
broad,  gently  arched,  coarsel}'  rugose  and  granulated ;  radial 
grooves  unevenly  directed. 

Eyes  arranged  in  a  ring  or  corona  around  the  ocular  tubercular 
eminence;  the  four  seated  at  the  rear  form  a  slightly  curved 
series — the  curvature  directed  downwards;  they  are  of  equal  size 
and  equidistant;  in  front  the  two  lower  eyes  are  minute,  and  are 
separated  from  each  other  by  a  space  equal  to  rather  more  than  once 
their  individual  diameter,  whilst  the  upper  pair  are  not  only  widely 
separated  from  each  other,  but  are  much  the  largest  of  the  group. 

Legs  yellowish-brown,  coarsely  granulated  and  rugose;  each  is 
armed  with  short  strong  spines,  and  thickly  clothed  with  short 
fine  hairs.  Relative  lengths:  1,  2,  4,  3;  the  first  and  second 
pairs  are  much  the  longestand  strongest,  and  of  nearly  equal  length. 

Palpi  short,  similar  in  colour,  clothing  and  armature  to  legs. 

Falces  concolorous,  hairy,  apices  divergent. 

Maxillce  concolorous  also,  glabrous,  punctated,  apices  inclining 
inwards,  thickly  fringed  with  long  yellowish  hairs,  surfaces 
furnished  sparingly  with  long  hairs. 

Labium  dark  brown,  broad,  slightly  arched,  apex  narrow  and 
obtusely  pointed;  a  few  hairs  scattered  over  its  surface, 


492  DESCRIPTIONS    OP    SOME    NEW    ARANEIDiE    OF    N.S.W. 

Sternum  normal,  yellowish,  surrounded  with  a  fringe  of  rather 
long  coarse  hairs;  the  surface  very  finely  granulated,  and  closely 
clothed  with  short  adpressed  hairs. 

Abdomen  somewhat  obovate,  slightly  overhanging  base  of 
cephalothorax,  moderately  arched,  yellowish-brown  with  dark 
brown  markings,  the  surface  being  finely  tuberculated  and  granu- 
lated; anterior  extremity  sinuous;  at  about  one-third  its  length 
from  anterior  extremity  there  are  two  large  but  shallow  discular 
depressions,  and  these  are  separated  from  each  other  by  a  space 
equal  to  about  once  their  individual  diameter;  at  about  the  centre 
there  are  two  others,  equally  as  large  but  wider  apart,  being 
separated  from  each  other  by  a  space  equal  to  twice  their  indi- 
vidual diameter;  sides  yellowish,  marked  with  dark  wavy  granu 
lated  lines;  ventral  surface  much  smoother,  dark  yellowish-grey, 
and  marked  at  the  middle  with  two  lines  of  punctures  rather 
widely  separated;  these  commence  just  below  the  epigastric  fold 
and  run  in  a  tolerably  straight  line  for  two-thirds  their  distance, 
and  then  converge  inwards  towards  the  spinnerets,  but  do  not 
meet;  those  near  the  spinnerets  are  much  the  closest  together;  in 
addition  to  these  there  are  a  few  smaller  lateral  punctures 
scattered  promiscuously. 

Ejngyne  as  in  figure. 

Hah. — Jervis  Bay. 


EXPLANATION  OF  PLATE  XVIIL 

Fig.  1  . — Storena  alhomaculatn  (  $  ). 

Fig.  la.—      ,,  ,,  epigyne. 

Fig.  2  . — Amneu>:  Uheralu  (  $  ). 

Fig.  2a.—       ,,  „  epigyne. 

Fig.  3  . — Steplutnoima  olduHfrom^  (  $  ). 

Fig.  3a. —  ,,  ,,  abdomen,  profile. 

Fig.  36.—  ,,  ,,  epigyne. 

Fig.  4  .—Celi'nia  (li^tincta  (  $  ). 

Fig.  4a. —      ,,  ,,  abdomen  from  the  rear. 

Fig.  4h.—      ,,  „  „        profile. 

Fig.  4c. —      ,,  ,,  femur  of  1st  and  2nd  legs. 

Fig.  4(Z. —      ,,  ,,  caput,  profile. 

Fig.  4e..—      ,,  „  epigyne. 


[Printed  off  December  Utb,  1902.] 


P.L.S.IN.S.W,  1902. 


-V.di. 


v.d 


vot 


eoc.v 


irvt 


eoo  p 


G.J.  del. 


Pi. XIII. 


-e^oc,.  c 


RLS.N.5.W.  1902. 


PI.  XIV. 


G.A.W.deL 


OGYRIS 


P.L.SNSW.   1902. 


PI  XV. 


/^ 


ELAE0CARPU5    sp. 


P.L.S.N.S.W. 


PI. XVI. 


ARDISIA      RACEMOSA,  RI.B. 


RL.S.N.5.W.  1902. 


PI.  XVII. 


p.L.S.N.S.W.   1902 


tV.  J.  R.,  del.         4a. 

CEL/ENIA  DlSTINCTA,C<9/7?6r 


PL  XVIII 


■la  STORENA  ALBOMACULATA.  2-2a  ARANEUS  LIBERALI5. 

3-3a-3b  5TEPHAN0PSI5  0BTU5IFR0MS. 


OF    THE  '      -• *=— "'^ 

LINNEAN    SOCIETY 

OF 


SUPPLEMENT  TO  PROCEEDINGS,   1902. 


CENSUS  MUSCORUM  AUSTRALIENSIUM. 

A  CLASSIFIED  Catalogue  of  the  Frondose  Mosses  of  Australia 
AND  Tasmania,  collated  from  available  Publications  and 
Herbaria  Records,  by  the  Rev.  W.  Walter  Watts  and 
Thomas  Whitelegge,  F.L.S. 

Part  I. 

(MS.  received,  May  8th,  1902.) 


Introduction. 

By  the  Rev.  W.  Walter  Watts. 

In  1899  Mr.  Whitelegge  and  I  prepared  a  Catalogue  of  the 
Mosses  of  N.S.W.  My  colleague  had  applied  himself,  with 
characteristic  ardour,  to  the  study  of  the  Mosses  of  the  State, 
especially  in  the  neighbourhoods  of  Sydney,  Moss  Yale,  Gosford 
and  the  Blue  Mountains,  first  in  1884-85,  and  again,  at  the  urgent 
request  of  Dr,  Brotherus,  in  1890-91,  securing  altogether  nearly 
100  new  species.  In  1892  he  had  compiled  a  list,  revised  by 
Dr.  Brotherus,  of  some  300  New  South  Wales  species.     This  work 


2  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

had  not  been  printed  when,  about  1898,  I  first  made  Mr. 
Whitelegge's  acquaintance.  It  was  embodied,  under  a  different 
classification,  in  our  conjoint  Catalogue  which,  containing  upwards 
of  500  species,  was  accepted  for  publication  in  March,  1900,  by 
this  Society. 

My  own  ambition  had  been  to  prepare,  in  the  course  of  a  few 
years,  a  Catalogue  of  the  Mosses  of  Australia  and  Tasmania, 
towards  which  I  had  been  steadily  accumulating  material.  It 
soon  became  evident,  however,  that  it  would  be  impossible  to  get 
such  a  w^ork  printed  by  the  Society  in  addition  to  the  New  South 
Wales  Catalogue;  and  the  Council  kindl}'  allowed  us  to  withdraw 
the  earlier  work  and  substitute  a  Census  of  Australian  Mosses, 
so  far,  at  any  rate,  as  the  species  usually  called  Acrocarps  were 
concerned. 

My  colleague's  pre-occupations  in  the  Australian  Museum 
having  prevented  him  from  taking  an  active  part  in  this  larger 
work,  the  responsibility  for  the  collation  and  classification  of  the 
material  falls  upon  my  shoulders  alone. 

It  ca.nnot,  unfortunately,  be  claimed  that  there  are  no  synonyms, 
nor  nomina  nuda,  in  this  Census.  The  inaccessibility  of  speci- 
mens, and  even,  in  some  cases,  of  descriptions,  the  differing 
principles  of  determination  ado^Dted  by  specialists,  and  the  large 
number  of  new  species  of  which  we  know  nothing  except  the 
names,  make  an  unchallengeable  list  of  Australian  Mosses  imprac- 
ticable at  the  present  stage. 

To  illustrate  one  of  our  many  perplexities.  Mitten  and  Wilson 
introduced  an  excessive  number  of  European  and  Antarctic  names 
into  our  Moss  Flora.  Dr.  Carl  Mueller,  on  the  other  hand, 
regarded  all  our  species  as  endemic,  so  much  so  that,  with  him, 
our  Ficnaria  hygrometrica  was  F.  sphaerocarpa,  our  forms  of 
Bryum  argenteum  became  Br.  Bateanum  and  Br.  HamjyeanKm,  and 
even  our  Ceratodon  purimreus  w^as  only  recognised  by  him  as  var. 
australis.  Dr.  Brotherus,  in  his  '  Bryales,'  now  in  course  of 
publication,  goes  a  long  wa}'^  in  the  direction  of  Dr.  Mueller's 
views,  but  by  no  means  regards  all  our  species  as  endemic. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  3 

The  introduction  of  so  many  European  names  by  the  earlier 
bryologists,  and  of  so  many  new  species  by  the  late  Dr.  C. 
Mueller  (a  number  of  which  were  due  to  the  erroneous  presuppo- 
sition above  mentioned),  has  created  a  series  of  difficult  problems 
for  Australian  students.  It  will  be  long  years  before  our  Moss 
Flora  is  placed  upon  a  satisfactory  footing.  It  might  be  contended 
that  this  Census  should  have  awaited  a  complete  investigation. 
We  entirely  dissent  from  that  view.  Knowing,  as  we  do,  the 
enormous  task  before  the  bryological  student  in  this  countr}^  we 
have  endeavoured  to  lay,  in  some  manner,  a  foundation  for  the 
labours  of  future  builders;  and,  in  this  Census,  we  submit  the 
result  of  much  patient  toil,  in  the  hope  that  it  will  be  of  service 
to  our  fellow-bryologists. 

It  need  scarcely  be  said  that  we  do  not  profess  to  give  an 
exhaustive  list  of  localities  and  collectors.  Even  if  the  informa- 
tion had  been  available,  space  would  have  failed  us.  It  is  to  be 
hoped  that  students  throughout  the  Commonwealth  will  develop 
the  bryology  of  the  separate  States,  and  build  up  detailed  accounts 
of  the  Moss  flora  of  different  districts. 

To  the  works  named  in  the  appended  list  we  desire  to 
acknowledge  our  great  indebtedness, — to  Jaeger  and  Sauerbeck's 
'  Adumbratio,'  Paris'  'Index  Bryologicus,'  Mueller's  'Genera  Mus- 
corum,'  Brotherus's  '  Bryales,'  and  Warnstorf's  '  Sphagnacese ' 
especially.  Dr.  Brotherus's  kindness  and  his  great  services  in 
the  study  and  determination  of  plants,  as  well  as  in  his  descrip- 
tion of  new  species,  deserve  the  sincerest  gratitude  of  Australian 
bryologists. 

We  desire  very  heartily  to  express  our  obligations  for  the 
assistance  received  from  Mr.  Luehmann,  Director  of  the  IS'ational 
Herbarium,  Melbourne ;  from  the  publications  of  Mr.  F.  M. 
Bailey,  Colonial  Botanist  of  Queensland;  from  Mr.  R.  A.  Bas tow's 
Tasmanian  records;  from  Mr.  F.  M.  Reader,  of  Warracknabeal, 
Victoria;  Mr.  C.  T.  Musson,  of  the  Hawkesbury  Agricultural 
College;  Mr.  J.  H.  Maiden,  Director  of  the  National  Herbarium, 
Sydney  (per  Mr.  Wm.  Forsyth,  Superintendent  of  the  Centennial 
Park);  Mr.  R.  T.  Baker,  Curator  of  the  Technological  Museum, 


4  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

Sydney;  Mr.  Wm.  Bauerlen,  of  the  same  institution;  Mr.  Jas. 
Stirling,  late  Government  Geologist  of  Victoria;  Mr.  W.  A. 
Weymouth,  and  others.  Mr.  Reader  was  the  first  to  respond  to 
our  application  for  assistance  by  sending  a  list  of  all  the  records 
known  to  him,  thus  rendering  valuable  service  from  the  Victarian 
side.  Mr.  Luehmann  very  kindly  supplied  a  list  of  the  records 
in  the  Melbourne  Herbarium,  which,  though  containing  no  details 
of  localities,  etc.,  gave  the  States  in  which  the  species  had  been 
collected,  confirming  many  of  the  records  in  our  possession  and 
adding  others.  At  the  eleventh  hour  a  valuable  list  arrived 
from  Mr.  Stirling,  who  was  absent  in  London  when  our  applica- 
tion for  assistance  reached  Melbourne.  Mr.  Stirling  has  "for 
many  years  collected  specimens  of  the  plants  growing  all  over  the 
Australian  Alps,  from  Mt.  Baw  Baw  at  the  heads  of  the  Yarra, 
to  Mt.  Kosciusko,"  and  during  the  years  1880-88  he  "made 
several  collections  of  the  cryptogamic  florula  of  the  highest 
elevations,"  being  accompanied  on  one  of  these  excursions  to  Mt. 
Kosciusko  by  Mr.  D.  Sullivan,  of  Moysten,  since  deceased. 
Acknowledgment  should  also  be  made  of  the  assistance  received 
from  the  Librarian  and  Assistant  Librarian  of  the  Public  Library, 
Sydney,  by  whom  some  valuable  Papers,  not  previously  known  to 
us,  were  brought  within  our  reach. 

Systems  of  classification  in  bryology  have  varied  greatly.  Dr. 
Mueller  adhered  throughout  to  his  principle  of  "  centralisation," 
grouping  the  mosses  of  the  world  under  large  and  bold  generali- 
sations. A  principle  of  "decentralisation,"  represented  in  Jaeger 
and  Sauerbeck's  'Adumbratio,'  has  reigned  for  many  years  among 
Continental  bryologists,  including  Dr.  Brotherus.  Mitten 
followed  a  scheme  of  his  own,  a  scheme  which,  through  the 
influence  of  his  Catalogue  of  Australian  Mosses,  has  been  in  force, 
for  the  most  part,  in  this  country.  Dr.  Brotherus,  in  his 
'  Bryales  ' — a  master  work.  Part  i.  of  which  has  been  received  — 
departs  greatly  from  Jaeger's  system,  and  follows  an  independent 
course  that  brings  him,  to  some  extent,  into  line  with  Mitten. 

We  have  adopted  the  following  scheme,  based  upon  many  con- 
siderations, but  in  large  part  upon  Mr.  H.  N.  Dixon's  invaluable 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGC4E.  0 

work    on    the    British    Mosses,   with    whioli    Dr.    Brotherus,   in 
*  Bryales,'  is  in  considerable  agreement : — 

Order  MUSCI. 

Suborder  1.  SPHAGNALES. 

Suborder  2.  ANDRE.EALES. 

Suborder  3.  BRYALES. 
Group  A.  NEMATODONTES  (Polytrichaceffi). 
Group  B.  ARTHRODONTES. 

Subgroup  i.  Acrocarpi. 

Subgroup  ii.   Pleurocarpi. 

Among  the  Acrocarpi  we  have  followed,  without  distinct 
headings,  the  Order  Haplolepidei  {Fissidentacece,  Archidiacece, 
Ditrichacece,  Dicranacece,  Leucohryacece,  C alymperaceoi^  Pottiacecc, 
Enca^yptacice,  Grimmiacece),  and  Diplolepidei  [Orthotrichacece, 
Sjjlachnacece,  Funariaceoi,  Bartramiacece,  Bi'yacece,  JIniacece). 
Among  the  Haplolepidei  we  have  placed  first,  w4th  Dr.  C.  Mueller, 
the  distichophyllous  Fissidentacece.  The  terms,  Haplolepidei  and 
Diplolepidei,  scarcely  apply  to  the  non-peristomate  members  of 
the  included  families,  but  they  form  a  valuable  practical  guide  to 
the  arrangement  of  the  Acrocarps.  The  Pleurocarps  are  all 
Diplolepidei. 

The  Mosses  of  Lord  Howe  and  Norfolk  Islands  are  excluded 
from  this  Census  on  zoogeographic  srrounds. 

We  have  refrained  from  stating  the  fruiting  season  of  the 
different  species,  there  being  considerable  variation  north  and 
south,  east  and  west.  July  to  December  may  be  regarded  in 
general  as  the  best  time  for  collecting;  but  allowance  must  be 
made  for  altitude  and  latitude.  The  Fissidentes  may  be  found 
in  full  fruit  as  early  as  April  and  May,  and  there  is  no  time  of 
the  year  when  fruiting  specimens  of  many  of  our  mosses  may  not 
be  collected,  especially  in  damp  gullies  and  about  the  shady  banks 
of  creeks. 


6  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

COXTRACTIONS    AND    REFERENCES. 

Bail.  Syn. — Bailey,  F.  M.     Synopsis  of  Queensland   Flora  '83. 

Suppls.  '86,  'SS,  and  '90. 
Bail.  Bot.  Bull.— Bailey,  F.  M.     Botanical  Bulletins,  '91-'96. 
Bastow,  M.  Tas. — Bastow,  R.  A.     Mosses  of  Tasmania.     Papers 

and  Froc.  B.  Soc.  Tasm.  '86. 
Bot.  Centr. — Botanisches  Centralblatt. 
Bot.  Zeit. — Botanisches  Zeitung. 
Broth.  A.M.- — Brotherus,  Dr.  Y.  F.,  of  Helsingfors.    Some  New 

Sp.  of  Australian  Mosses,  1890-1900  (5  Parts).     The  sp.  are 

numbered  consecutively. 
Broth.  Andreteales,  and  Bryales — Brotherus,  Dr.  V.  F.,  on  the 

Andreseales,  '01,  and  Bryales  (Part  1,  '01). 
CM.,  Syn. — Mueller,  the  late  Dr.  Carl.     Synopsis  Muscorum, 

'49-'51. 
C.  M. ,  Symb. — Mueller,  the  late  Dr.  Carl.    Symbolse  ad  Bryologiam 

Australia),  Hediuigia  '97-8;  numbered  consecutively. 
CM.,  Gen.  M. — Mueller,  the  late  Dr.  Carl.     Genera  Muscorum 

Frondosorum,  1901  (posthumous).   Sometimes  later  informa- 
tion will  be  found  in  Symb. 
F.v.M.  Austr.  M. — Mueller,  The  late  Baron  F.  von.     Analytical 

Drawings  of  Australian  Mosses,  '64. 
F.v.  M.  Frag.  Phyt. — Mueller,  the  late  Baron  F.  von.    Fragmenta 

Phytographi^e  Australise  (Vol.  xi.,  with  Suppl.  '78-'81). 
Fl.  Ant. — Hooker,  J.  D.     Flora  Antarctica,  '44-7. 
Fl.  N.Z.— Hooker,  J.  D.     Flora  Novte  Zealandi?e,  '54  (cir.). 
Fl.  Tas.— Hooker,  J.  D.     Flora  Tasmania*,  '60. 
H.f.W.— Hooker  fil.  (J.  D.  Hooker)  et  Wilson,  W. 
Hdbk.  N.Z.  Fl.— Hooker,  J.  D.      Handbook  of  the  KZ.  Flora. 
Hpe.  List. — Musci  frondosi  Australia?,  continentalis  prsesertim  e 

Bar.  de   Mueller  collectionibus.  Dr.   E.   Hampe  enumerati, 

1880. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  7 

Jaeg.  Ad. — Jaeger  und  Sauerbeck.  Adumbratio  Florae  Mus- 
corum  '70-78. 

Jour.  L.  Soc. — Journal  of  the  Linnean  Society. 
Linn. — Linnsea. 

Lond.  J.  Bot. — London  Journal  of  Botany. 

Mitt.  Cat. — Mitten,  Wm.  Catalogue  of  Australian  Mosses,  '82-3. 
Proc.  JR.  Soc.   Victoria. 

Mitt.  Jour.  L.  Soc.  '60— Paper  read  Mch.  '59,  published  '60,  on 
"  Some  New  Sp.  of  Musci  from  N.Z.  and  other  parts  of  the 
S.  Hemisphere."  (Wilson  had  the  use  of  the  MS.  in  pre 
paring  the  Moss  section  of  Fl.  Tas.) 

Par.  Index — Paris,  General.  Index  Bryologicus,  '93-'98.  Suppl.  i., 
1900. 

Melb.  Census— List  from  Mr.  Luehmann,  1902,  of  the  Mosses 
recorded  in  the  National  Herbarium,  Melbourne. 

KS.W.— New  South  Wales. 

Q. — -Queensland. 

Rev.  Bry. — Revue  Bryologique,  edited  by  M.  Husnot. 

S.  A. — South  Australia. 

Stirling  Crypt.- — Stirling,  James,  Govt.  Geologist  of  Victoria. 
The  Cryptogamia  of  the  Australian  Alps,  Part  i.  Froc.  R. 
Soc.  Victoria,  '85-'86. 

Sullivan,  Paper — Paper  read  before  the  Field  Naturalists'  Club 
of  Victoria,  Aug.  '87 — Victorian  Naturalist,  '87-8. 

Tas. — Tasmania. 

Tate,  List — List  of  Mosses,  etc.,  collected  by  F.v.  M.  in  extra- 
tropical  S.A.  in  the  early  days  of  colonisation,  communicated 
to  the  Royal  Soc.  of  Vic.  by  the  late  Prof.  Tate,  '82. 

Vic. — Victoria. 

W.  A.— West  AustraHa. 

Warnst.  Sphag. — Warnstorf,  Dr.  C,  of  Neuruppin.  On  the 
Sphagnaceae,  1901. 


8  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

Weymouth,  M.  Tas. — Weymouth,  W.  A.,  of  Hobart.  Some 
additions  to  the  Moss  Fl.  of  Tasmania,  '93-'95.  Pajyers  and 
Proc.  R.  Soc.  Tcif^m. 

Order  MUSCI. 
Suborder  SPHAGNALES. 
Tribe  i.  Sphagnaceae. 
Sphagnum  (Dill.),  Ehrh. 

1.  S.  ANTARCTicuM,  Mitt.,  Jour.  L.  Soc.  '60;  S.  comimctum 
var.  Fl.  Ant.  (fid.  Mitt.). 

Tas.,  Longley,  Huon  Rd.  '89  and  '92,  Snake  Plains,  Mt.  Welling- 
ton '94 :  Weymouth.  Long  Bay  Channel :  A.  J.  Taylor. — 
Vic.  :  Miss  Campbell.— N.S.W.,  Mt.  Kosciusko,  4000',  and  Mt. 
Aberdeen  {S.  cristatum,  Hpe.,  Linn.  '74,  fid.  Warnst.). 

2.  S.  AUSTRALE,  Mitt.,  ibid.;  S.  compactum  var.  ovatum,  Fl. 

Ant.  (fid.  Mitt.). 
Tas.,   Huon   R.  :   Oldfield.   Rivulet,   nr.    Cumming's    Head,   W. 
Mts. :  Archer  {S.  confertum,  Mitt.,  Jour.  L.   Soc.  '60).   Longley, 
Huon  Rd. :  Weymouth  '89.  Mt.  Wellington :  Bastow  et  al. 

3.  S.  Brotherusii,  Warnst.,  Bot.  Centr.  1900. 

N.S.W.,  Richmond  R.,  East  Ballina,  Sept.  '98,  and  Byron  Bay, 
Aug.  '99  :  Watts. 

4.  S.  centrale,  Jens.,Sv.  Vet.  Akad.  Handl.   (An  European 

sp.).     S.  subbicolor,  Hpe. 
N.S.W.,  S.  Coast,  Kiama:  Gilbert  Weir,  '96  (det.  Warnst.). 

(a)    var.     flavo-glaucescens,     Russ.,    /.     brachyclada, 
Warnst. 
Tas.  :  teste  Par.  Index  Suppl.  i. 

5.  S.  COMOSUM,  CM.,  Flora  '87;  Warnst.,  Hedw.  '9L 

N.S.W.,  Waterloo  Swamps,  nr.  Sydney;  Whitelegge,  Oct.-Nov. 

'83. 

S.  compactum  vars.     See  Nos.  2  and  \. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  d 

Sp  H  A  G  N  u  M  {continited).  [Sphagnaceo'. 

S.  conjertum,  Mitt.  See  No,  2  [CM.,  in  Gen.  M.,  retains 
S.  covfertum,  and  says  it  has  the  appearance  of  S. 
rigidum  var.  compactum.  Name  not  in  Warnst. 
Sphag.] 

«S'.  contortum^  Schultz,  vars.  Fl.  Tas.     See  No.  8. 

»S'.  cristatum^  Hpe.,  Linn.  74.  See  No.  1  [CM.,  in 
Gen.  M.,  retains  S.  cristatum  for  the  Mt.  Kosciusko 
and  Mt.  Aberdeen  Mosses.] 

6.  S.  CUSPIDATUM  (Ehrh.)  Russ.  et  Warnst. 

CosMOPOL.— N.S.W.,  Blue  Mts.,  Nellie's  Glen:  Whitelegge,  Oct. 
'91. — Q.,  Maroochie  :  herb.  Bailey  (Suppl.  '86) 
S.  cymhifolioides,  CM.     See  No.  8. 

7.  S.  CYMBIFOLIUM  (Ehrh.),  Warnst.;  *S'.   Wilco:cii,C.M.  {fid. 

Warnst.) 

CosMOPOL.  (Warnst.). — Widespread  in  Tas.  (Gunn.  Oldfield,  J.  D. 
Hooker,  et  al.),  and  in  Australia  (Brisbane  R.  :  Bailey.  Blue 
Mts.,  Lawson  :  Whitelegge.  Gippsland :  F.v.M.  Austral.  Alps: 
Stirling)  [CM.  gives  Europe  and  N.  Amer.  only]. 

(a)  var.  carneo-pallidum,  Warnst.,  in  sched.  '98. 
N.S.W.,  Richmond  R.,  Park  Gray's  Isl.  :  Watts,   96. 

(b)  var.  GLAUCESCENS  f.  brachyclada,  Warnst.,  in  sched. 
'98. 

N.S.W.,  Richmond  R.,  Park  Gray's  Isl.  :  Watts,  '96. 

8.  S.  CYMBOPHYLLUM,  F.V.M. ,  Secoud  Gen    Report  '54;  S. 

cymhifolioides,  CM..  Bot.  Zeit.  '51  (?),  '62  (teste  Par.). 

Prob.  S.  conto7'tuin  vars.  Fl.  Tas. 
Tas.,  Cheshunt :  Archer.     And  prob.   Mossman,  Gunn,  Oldfield, 
et  al.  on  Mt.  Wellington. — Vic,  Victorian  Ranges,  Gippsland, 
Mt.  Aberdeen  :  F.v.M.— N.S.W.,  nr.  Sydney  :  Kayser  coll.— Q., 
Nerang  Creek  :  herb.  Bailey. 


10  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Sphagnum  {continued).  [^Sphagnacece. 

9.  S.  DREPANOCLADUM,  Wamst.,  Bot.  Centr.  1900 
N.S.W.,  Richmond  R.,  Shaw's  Bay,  on  wet  face  of  cliff:  Watts, 
April  '99ff. 

10.  S.  DUBiosuM,  Warnst.,  Hedw.  '91. 
S.A.  :  teste  Warnst ,  Sphag.  1900. 

11.  S.  GRANDIFOLIUM,  Warnst ,  Bot.  Centr.  1900. 
N.S.W.,  N.  Coast,  Byron  Bay,  etc.  :  Watts,  Aug.  '99ff. 

12.  S.  LANCiFOLiuM,  CM.  ct  Wamst.,  Hedw.  '97. 
N.S.W.,    Sydney,   prob.     Botany :    Whitelegge,    Nov.    '83.       In 
Warnst.  Sphag.  given  for  S.A.,  evidently  in  error. 

13.  S.  LATicoMA,  CM.  (teste  F.  M.  Reader). 
Vic,  Blacks'  Spur :  Miss  Campbell  (Mrs.  Martin). 

14.  S.  MACROCEPHALUM,   Wamst.,  Hcdw.  '93;  Weymouth, 

M.  Tas.  '93  et  '95. 
Tas.,  Lake  Bellenger  Track,  Zeehan  Railway,  W.   Coast;  Wey- 
mouth, Feb.  '91. 

15.  S.  MAXIMUM,  Warnst.,  Hedw.  '91. 

Tas.,  loc.  unknown.  Mitt,  found  it  among  packing  material  for 
Tasmanian  apples. 

16.  S.  MOLLicuLUM,  Mitt.,  Jour.  L.  Soc.  '60  et  Fl.  Tas.;  S. 
Jlossmanniamcin,  CM.  (teste  Par.  Index). 

Tas.,  Cheshunt,  Little  Bridge's  Head  Creek  :  Archer. — N.S.W., 
nr.  Sydney,  Waterloo  Swamps:  Whitelegge  '83.  "Tas.,  N.S.W., 
and  elsewhere  in  Australia"  (CM.  in  Gen.  M.). 

17.  S.  MooREi.  Warnst.,  Allgem.  Bot  Zeitschr. in  Karlsruhe, 

'95. 
Tas.,  Kelly's  Basin  and  Gordon  R.,  Macquarie  Harbour,  on  damp 
ground:  T.  B.  Moore  '93  (herb.  Weymouth). 

S.  Mossmannianum,  CM.     See  No.  16. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  11 

Sphagnum  {continued.)  [Sphagnacece. 

18.  S.  Naumanni,  cm.,  Gen.  M.,  p.  102. 
Q.  :  teste  Melb.  Census  and  CM.  loc.  cit. 

19.  S.  Novo-Zelandicum,  Mitt.,  Jour.  L.  Soc.  '60;  Warnst., 

Hedw.  '91. 
Tas.  :  Archer  (teste  Mitt.  Cat.) — N.S.W.,  nr.  Sydne}^  Maroubra  : 
Whitelegge,  Sept.  '91  [CM.  and  Warnst.  Sphag.  give  N.Z.  only]. 

»S'.  pachycladiwi,  CM.,  given  by  F.v.M.,  Frag.  Phyt.  xi., 
SuppL,  for  N.S.W. — a  doubtful  species. 

20.  S.  PSEUDO-RUFESCENS,  Warnst.,  Hedw.  '93;  Weymouth, 

M.  Tas.  '93-95. 
Tas.,  in  bog,  top  of  Mt.  Wellington:  Weymouth  '88,  Watts  '95. 

21.  S.   ScoRTECHiNii,  CM.,  in  litt.;  descr.  by  Warnst.  in 

Hedw.  '97. 
Q. :  Rev.  B.  Scortechini. 

22.  S.  SERRATiFOLiUM,  Wamst.,  Bot.  Centr.  1900. 
N.S.W.,  K  Coast.,  Byron  Bay:  Watts,  Aug.  '99. 

23.  S.  SERRULATUM,  Wamst.,  Hedw.  '93;  Weymouth,  M. 

Tas.  '95. 
Tas.,  in  ditch,  Zeehan  Railway,  W.  Coast:  Weymouth,  Feb.  '91. 

24.  S.  SUBCONTORTUM,  Hpo.,  Linn.  '76;  Warnst.,  Bot.  Centr. 

1900.       aS'.  suhsecundum  var.  in  Jaeg.  Ad. 
N.S.W.,  N.  Coast,  Mt.  Warning:  Guilfoyle  '71. 

25.  S.  suBMOLLicuLUM,  Warnst.,  Hedw.  '97. 

Tas.,  Kelly's  Basin  '93 :  J.  B.  Moore.  Port  Esperance:  Weymouth, 
June  '92. 

26.  S.  suBSECUNDUM(Nees),Limpr.  var.  macrophylla,  CM. 
Vic,  Blacks'  Spur:  Miss  Campbell  (Mrs.  Martin). 

[The  species  itself  is  recorded  for  Tas.  in  Par.  Index,  and  for 
Vic.  in  Melb.  Census,  possibly  because  Schultz's  S.  contortumi  was 
taken  as  a  var.  of  this  sp.  Neither  Warnst.  (Sphag.)  nor  CM. 
(Gen.  M.)  gives  Tas.  or  Australia.] 


12  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

S  p  H  A  G  N  u  M  {continued).  [Sphaifnacece. 

27.  S.  SuLLivANi,  CM.,  Gen.  M.  p.  103. 
Vic,  Grampians,  Mt.  William,  Creek :  Sullivan  '75. 

28.  S.  TRicHOPHYLLUM,  Warnst.,  Hedw.  1900. 
Tas.,  Mt.  Wellington:  Bastow,  Dec.  '87. 

29.  S.  Wattsii,  Warnst.,  Bot.  Centr.  '98. 

N.S.W.,  Richmond  R.,  common  on  heaths;  East  Ballina.  Heath 
8wamp:  Watts,  May  '96.  Pimlico:  Mrs.  A.  Heugh,  Oct.  '96. 
Byron  Bay,  etc. :  Watts  '96  ff.,  often  in  full  fruit. 

30.  S.   Whiteleggei,  CM.,  Flora '87;  S.  leionotum,  CM. 

(fid.  Par.) 
N.S.W.,  Blue  Mts.,  Nellie's  Glen:  Whitelegge,  '84.  Nr.  Sydney: 
Kayser  coll.  Braid  wood  District:  Biiuerlen. 

S.  Wilcoxii,  CM.,  Flora  '87  and  Gen.  M.  (collected  by 
Mr.  Wilcox  on  the  Clarence  R.,  N.S.W.)    See  No.  7. 

Suborder  ANDRE.EALES. 

Tribe  ii.  Andreaeaceae. 
A  N  D  R  E  ^  A,  Ehrh. 

31.  A.  ACUMINATA,  Mitt.,  Jour.  L.  Soc.  '60  et  Fl.  Tas. 
Tas.,  on  rocks,  Cheshunt:  Archer.  Mt.  Wellington:  Weymouth, 
Bastow,  Watts  et  al. 

32.  A.  AMBLYOPHYLLA,  CM.,  in  Broth.  A.M.  '95,  No.  53; 

Symb.,  No.  99. 
N.S.W.,  Blue  Mts.,  3500':  Whitelegge.— Tas.,  on  rocks,  the  Glen, 
Knock  Lofty,  Hobart :  Weymouth.     Also  R.  A.  Bastow  and  A. 
J.  Taylor  in  herb.  Weymouth.  Top  of  Mt.  Wellington :  Weymouth, 
Watts. 

33.  A.  asperula.  Mitt.,  Jour.  L.  Soc.  '60;  A.  MueUeri,  Sond. 

(fid.  Mitt.) 
Vic,   Australian   Alps:    F.v.M.— N.S.W. :    teste    F.v.M.,    Frag. 
Phyt.  xi.,  Suppl. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  13 

A  N  D  R  E  .E  A  {continued).  [Andreceacece. 

34.  A.  ATTENUATA,  CM.,  ^jmh.,  No.  105. 

Tas..  on  rocks,  Mt.  Wellington:  Weymouth,  Sept.  '91  (ster.). 

35.  A.  australis,  F.v.M.  in  sched.;  Mitt.,  Hook.  Jour.  Bot. 

'56;  CM.,  Gen.  M.,  p.  5. 
Vic,  Mun3'ang  Mts.,  Australian  Alps:  F.v.M. 

36.  A.  ERUBESCENS,  CM.,  Sjmb.,  No.  90. 

Tas.,  top  of  Mt.  Wellington,  4500'.  Weymouth,  Jan.  '89. 

(a)  var.  nigrita,  CM.,  ibid.;  A.  suhuli/olia,  C.^l.,  herb. 
Burchard,  '90,  and  Gen.  M. 
Tas.,  top  of  Mt.  Wellington:  Weymouth,  Jan.  '89  (ster.) 

37.  A.  ExiMiA,  CM.,  Symb.,  No.  103. 
Tas.,  Mt.  Wellington:  Weymouth,  Jan.  '^S. 

38.  A.  JULiCAULis,  CM.,  Symb.,  No.  89. 

Tas.,  Mt.  Wellington,  4000':  Weymouth,  Sept.  '91  (ster.). 

39.  A.  MONTANA,  Mitt.,  Jour.  L.  Soc.  '60  and  Fl.  Tas. 
Tas.,  W.  Mts.,  Cumming's  Head,  on  rocks  in  rivulet:  Archer. 

A.  Mueller l^  Sond.     See  No.  33. 

40.  A.  nitida,  H.f.W.,  Lond.  J.  Bot.  '44. 

Tas.,  on  stones  in  rivulet,  nr.  Cumming's  Head:  Archer. 

41.  A.  PETROPHILA,  Ehrh. 

Tas.,   the  Falls,  Cheshunt:  Archer.  Lake  Echo:  J.   D.  Hooker. 
Mt.  Wellington:  Gunn,  Weymouth  et  al. 

A.  pseudo-suhulata.     See  Note  after  No.  43. 

A.  subulata.     See  No.  42. 

42.  A.  subulatissima,  CM.,  Symb.,  No.   100;  A.  suhidatay 

Harv.,  of  earlier  bryologists. 
Tas.,  Recherche  Bay:  Oldfield.   On  rocks,  Jackey's  Plain  Creek, 
and    rivulet    nr.   Cumming's   Head  :    Archer.     Mt.   Wellington, 
frequent :  Bastow,  Weymouth  et  al.   Millhouse's  Falls:  Weymouth. 
A.   suhulifolia,  CM.     See  No.  36  var. 


14  CENSUS    MUSCORUM  AUSTRALIENSIUM, 

A  N  D  R  E  ^  A  {continued).  [Andreceacece. 

43.  A.  TENERA,  CM.,  Symb.,  No.  104. 

N.S.W.  Blue  Mts.,  Nellie's  Glen:  Whitelegge,  Oct.  '91.  (Par. 
Index  gives  Vic. — apparently  in  error.) 

[Note. — CM.,  in  Gen.  M.,  questions  Mitt.'s  determ.  of  A. 
siihulata,  A.  nitida,  and  A.  jjetrophila  for  Tas.  Broth.,  in 
Bryales,  keeps  the  last  two,  but  accepts  CM.'s  A.  suhulafissima 
for  A.  subidata  in  Tas.  In  Bot.  Zeit.  '64,  CM.  gives  A.  pseudo- 
suhulata  for  Tas.,  but  in  Gen.  M.  he  limits  it  to  "das  Inselmeer 
des  Feuerlandes."] 

Suborder  BRYALES. 

Group  A    NEIVIATODONTES. 

Tribe  iii.  Polytrichaceae. 
B  u  X  B  A  u  M  I  a.  Hall. 

44.  B.  TASMANiCA,  Mitt.,  Jour.  L.  Soc.  '60  and  Fl.  Tas. 
Tas.,  nr.  Cheshunt:  Archer. 

D  A  ws  0  N  I  A,  R.Br. 

45.  D.  APPRESSA,  Hpe.,  Linn.  '60;  F.v.M.,  Aust.  M.  pi.  10. 
S.A.,  Onkaparinga  Valley,  St.  Vincent's  Gulf,  in  fissures  of  rocks  : 
F.v.M. 

46.  D.  INTERMEDIA,  CM.,  Symb.,  no.  12. 

Yic. — N.S.W. — Q.  C?)  First  recorded,  apparently,  by  Luehmann 
for  Fernshaw,  Upper  Yarra,  in  '81  (c.fr.).  Also  recorded  from 
Wilson's  Promontory,  Rocky  Cape,  nr.  Sydney,  Clarence  R.  (A. 
Rudder,  '82),  Richmond  R.  (Night  Cap  Mts.:  De  Camara  Alston- 
ville  Cutting,  Pearce's  Creek,  etc. :  Watts). 

[A  specimen  determ.  as  D.  polytrichoides,  coll.  by  C  T.  Musson  on 
the  N.  Pine  R.,  Q.,  appears  to  me  to  be  D.  intermedia — W.W.  W.] 

47.  D.  LONGiSETA,  Hpe.,  Linn.  '60;  F.v.M.,  Aust.  M.,  pi.  9. 
]^.S.\V. — Q.  First  record,  Parramatta:  Woolls,  or  F.v.M.  Parra- 
matta  Park:  Whitelegge,  Dec.   '84   (c.fr.).    Canterbury,  face  of 


13Y  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  15 

D  A  w  s  0  N  1  A  (continued).  [Polytrichacecb. 

cutting,  '96,  and  at  Emu,  foot  of  Blue  Mts.,  on  ground  and  moist 
banks,  Nov.  1900:  Watts  (c.fr.  plentiful).  Nr.  Barber's  Creek: 
Forsyth,  '99.  Nr.  Brisbane:  C.  T.  Musson,  Bailey,  and  J.  H. 
Simmonds. 

[Victorian  records  must  be  changed,  apparently,  to  D.  Victorue. 
—  VV.W.W.] 

48.  D.  POLYTRICHOIDES,  R.Br. 

Tas. — ViG. — N.S.W. — Q.  Our  commonest  Dawsonia,  collected 
first,  apparentl}^,  by  R.  Brown,  nr.  Sj^dney.  Austral.  Alps,  up  to 
4000'  (teste  Stirling). 

(a)  var  minor,  CM. 
N.S.W.,  S.   Coast,  mountain  nr.   Kangaroo  Valley:  Whitelegge, 
Dec.  '86  (c.fr.).     N.   Coast,  Richmond  R.,  Alstonville  Cutting  : 
Watts,  Jan.  '98  (c.fr.). 

49.  D.  superba,  Grev.  '47;  D.  longifolia,  Gunn  in  sched. 
Tas.— Vic.— N.S.W.— Q.      Shady   banks,    etc.    (also    in    N.Z.) 
frequent.    Very  fine  specimens  from  the  Upper  Yarra  (Luehmann) 
and   from  the  Head  of  Wilson's  Creek,  Richmond  R.   (Watts). 
Austral.  Alps,  up  to  4000'  (teste  Stirling). 

50.  D.  Victoria,  CM.,  Symb.,  No.  11. 

Vic.,  Doncaster,  nr.  Melbourne:  Reader,  July  '84  (c.fr.).  Other 
re'cords  in  herbaria,  possibly,  under  D.  longiseta. 

A  T  R  I  c  H  u  M,  P.  Beauv.     (Sect,  of  Cailierinea^  CM.) 

A.  angustatum,  Brid.,  var.  jjolysetum.     See  No.  52. 

51.  A.  LEPTOCYLiNDRicuM,  CM.,  Symb.,  No.  18. 

Vic,  Genoa  R.  '85:  Bauerlen. — N.S.W. ,  Delegate,  '85:  Biiuerlen. 
Katoomba  Falls:  Schauinsland,  '96-7  (also  in  N.Z.). 
A,  ligulatum,  Mitt.     See  No.  52. 

52.  A.  MuELLERi,  CM.  et  Hpe.,  Linn.  '53,  Catheriyim;  A. 

ligulatum,  Mitt.  '56  (fid.  Mitt.  '60);  A.  angustatum  var. 
poly se turn,  Fl.  Tas. 


16  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

A  T  R  I  c  H  u  M  (continued).  [Polytrichacece. 

Vic,  not  uncommon,  Bunyip  Creek,  Dandenong  Ranges,  Apollo 
Bay,  Sealer's  Cove,  Brodribb  R.,  Tarwin  R.,  Austral.  Alps,  etc.  : 
F.V.M.,  Biiuerlen,  Miss  Campbell,  Stirling  et  al.  Healesville  : 
Schauinsland,  '96-7. — Tas.,  Mt.  Wellington,  etc. :  Archer,  Bastow, 
Weymouth,  Watts  et  al. — N.S.W.,  Parramatta:  Woolls.  Bolara 
Mts.,  Clyde  R. :  Bauerlen,  Aug.  '98  (herb.  Tech.  Mus.  Syd.). 

53.  A.  PusiLLUM,  CM.,  Symb.,  No.  17. 
Tas.,  Marydale:  Weymouth,  Dec.  '90. 

54.  A.  siDEROLOMA,  CM.,  Symb,,  No.  16. 
Yic,  Gippsland,  Moe  R. :  Luehmann,  '81. 

PsiLOPiLUM,   Brid.     (Sect,  of  Polytrichum,  H.f.W.;  Sect,  of 
Catharlnea,  CM.) 

55.  Ps.  AUSTRALE,  H.f.W.,  Fl.  N.Z. 

Tas.,  W.  Mts.,  on  earth  and  stones  in  rivulet  behind  Cumming's 
Head:  Archer.  Mt.  Wellington:  J.  D.  Hooker  et  al.— N.S.W. 
Mt.  Kosciusko:  Forsyth,  Jan.  '99. 

56.  Ps.  CRISPULUM,  H.f.W.,  Fl.  N.Z.,  and  Fl.  Tas. 
Tas.,  W.  Mts.:  Archer. 

57.  Ps.  PiRiFORME,  Hpe.,  Linn.  '71-3,  Catherinea;  Mitt.  Cat. 
N.S.W.,  Blue  Mts.:  Mrs.  Calvert.— Vic,  Munyang  Mts.:  F.v.M. 

O  L  I  G  0  T  R  I  c  H  u  M,  Lam.  et  DeCand.   (Sect,  of  Catherinea,  CM.) 

58.  O.  MiNUTUM,  CM.,  Symb.,  no.  14. 
Tas.,  Mt.  Wellington:  Kayser  in  herb.  Melb. 

P  0  L  Y  T  R I  c  H  A  D  E  L  p  H  u  Sj  CM.     (Scct.  of  Catherinea,  CM.  et 
Hpe.) 

59.  P.  Arnoldi,  Hpe.,  Linn.  '74. 

Vic,  Australian  Alps,  Mt.  Arnold,  4000',  also  Upper  Yarra  and 
G(  ml  burn  Rs. :  F.v.M. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  17 

POLYTRICHADELPHUS  {continued).  [Polytrichacece. 

60.  P.  AUSTRALAsicus,  Hpe.,  Linn.  '76. 

Q.,  "subtropical  East  Australia":   Eaves. — N.S.W.,  Richmond 
R. :  Capt.  Stackhouse. 

P.  innovans,  CM.,  Bot.  Zeit.  '51.     See  No.  62. 

61.  P.  LAGENACEUS,  CM.,  Sjmb.,  No.  19. 
Tas.,  Mar^^dale  :  Weymouth,  Dec.  '91. 

62.  P.  Magellanicus  (L.),  Mitt.,  Jour.  L.  Soc.  '60. 

Tas.,  nr.  Cheshunt :  Archer.  Mt.  Wellington,  frequent. — Vic.  to 
N.S.W.,  in  alpine  regions,  Bonang  :  Biiuerlen,  Feb.  '99  (ster.). 

[C.M.'s  P.  hmovans  represents  this  Moss,  which  he,  in  Gen.  M., 
distinguishes  from  the  Magellan  sp.  His  name  may  yet  displace 
P.  Magellanicus  for  Tas.] 

63.  P.  PROLiFiCANs,  CM.,  Symb.,  No.  20. 

Tas.,  Mt.  Wellington  :  J.  and  B.  Gulliver,  herb.  Melb. 

P  0  G  o  X  A  T  u  M,    P.    Beauv.     [PolytricMim-Catharinella   mostly 
in  CM.) 

P.  aloides,  Hedw.     Recorded  for  Vic.  in  Melb.  Census. 
Confirmation  needed. 

64.  P.    ALPINUM  (L.),  Roehl.  ;    P.    j^seudo-alpinum,     CM., 

Bot.  Zeit.  '55  and  Gen.  M.;  P,  austro-al2?inum,  F.v.M. 
Vic,  Cobberas  Mts.,  6000':  F.v.M.  Austral.  Alps  :  Stirling.— 
Tas.,  W.  Mts  ,  the  Falls  and  rivulet  behind  Cumming's  Head  : 
Archer. — N.S.W.,  Mt.  Kosciusko:  Biiuerlen,  Feb.  '90 ;  Forsyth, 
Jan.  '99. 

65.  P.  AUSTRALASicuM,  Hpe.  et  CM., Linn.  '53,  Polytrichum. 
Vic,  Gippsland:  F.v.M.  Bonang:  Biiuerlen,  Feb.  '99.— N.S.W., 
shady  banks  of  cuttings  throughout  coastal  districts;  fine  speci- 
mens on  the  Northern  Rivers.  Blue  Mts.:  Watts. — Tas.,  St. 
Crispin's  Track,  Mt.  Wellington,  '88,  and  Castra  Rd.,  Leven,  '92: 
Weymouth.  .  ^ 


LI  B  R  A  R  Y   i 


18  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

PoGONATUM  {continited).  [PoJytrichacece. 

P.  austro-aJjnnum,  F.v.  M.     See  No.  64. 
P.  Baileyi,  Broth.     See  No.  67. 

Q6.     P.  BKACHYPODiuM,  CM.,  Sj'inb.,  No.  28;  PsUopilumiw 
sched. 
N.S.W.,  Fitzroy  Falls  on  logs  in  swamp:  Whitelegge,  Nov.  '84 
(c.fr.).  Cambewarra:  C.  Harris,  Sept.  '85.   Blue  Mts.,  Katoomba  : 
Schauinsland,  '96-7. 

67.  P.  Camar^,  cm.,  Sjmib.,  No.  26;  Pog.  Baileyi,  Broth. 
N.S.W.,  Clarence  R. :  Wilcox,  Nov.  75.  Richmond  R.,  Night  Cap 
Mts.:  De  Camara,  herb.  Melb.  '81.  Blue  Mts.,  Nellie's  Glen: 
Whitelegge,  Oct.  '84  (c.fr.).— Q.:  F.  M.  Bailey,  herb.  Broth.  '91. 
Eudlo:  Field  Naturalists. 

68.  P.  GiPPSLANDLE,  CM.,  Symb.,  No.  25;  P.  Tysdaleanum, 

Gen.  M.  p.  177. 
Vic  ,  Gippsland,  Pyers  R. :   Henry  Tysdale,  herb.  Melb.  '81. 

69.  P.  GuLLiVERi,  Hpo.,  Linn.   '76,  Polytrichnm  {Cathari- 

nella). 

Tas.,  Mt.  Wellington:  J.  and  B.  Gulliver.— Vic. :  teste  F.v.M., 
Frag.  Phyt.  xi.  Suppl.  (prob.  in  error). 

70.  P.   nanocarpum,   cm.,  Sj^mb.,   No.    22,    Polytrichiim 
[Aloidella). 

Vic,  Gippsland,  Walhalla:  Tysdale,  herb.   Melb.  '84  (Tisdall  in 
Gen.  M.). 

P.  2^s''-'^(^o-alpinum,  CM.  See  No.  64.  Jaeg.  Ad. 
follows  CM.  in  naming  the  Australian  specimens 
P.  pseudo-alpinum.  Dr.  Brotherus  returns  P. 
alpinum,  and  we  have  followed  him. 

P.  suhulatam,  Menz.  (a  N.Z.  sp.).  In  herb.  Melb. 
recorded  for  Vic.  as  *' (?)  P.  Camarce.^^  In  Bastow, 
M.  Tas.,  a  syn.  of  P.  tortile. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  19 

PoGONATUM  {coittinued).  [Polytrichacece. 

71.  P.  TORTILE,  Swartz,  Polytrichum  (a  W.  Ind.  sp.). 

Tas.,  Mt.  Wellington:  teste  Bastow,  M.  Tas. 

P.  Tysdaleanitm,  CM.  This  name  occurs  in  Gen.  M. 
p.  177,  but  in  Symb.  it  is  evidently  changed  to  P. 
Gijypslandice  (q.v.)  to  avoid  confusion  with  Polytri- 
chum Tysdalei. 

Polytrichum,  Dill. 

P.  Antillarum,  Rich.  Records  in  Melb.  Census  for  Tas., 
Vic.  and  N.S.AV.,  but  the  occurrence  of  this  Central 
and  South  American  sp.  requires  confirmation. 

72.  P.  Beccarii,  cm.,  Symb.,  No.  36. 
Tas.,  Mt.  Wellington:  Beccari,  Feb.  78. 

73.  P.  BRACHYPELMA,  CM.,  Symb  ,  No.  39. 

N.S.W.,  nr.  Sydney:  Kayser  coll.,  herb.  Geheeb  '72.  Blue  Mts., 
Lawson,  swampy  ground :  Whitelegge  '84. 

P.  brachypellina,  CM.  Name  in  F.v.M.,  Frag.  Phyt.  xi. 
for  N.S.  W.     Apparently  an  error  for  P.  hrachypelma. 

74.  P.  CATARACTARUM,  CM.,  Symb.,  No.  40. 

N.S.W.,  Fitzroy  Falls,  swampy  ground:  Whitelegge,  Nov.  '84. 

75.  P.  COMMUNE,  L. 

CosMOPOL.  In  Fl.  Tas.  recorded  for  Tas.,  Brown's  R.,  Cheshunt, 
Mt.  Wellington  (Oldfield,  Archer,  Gunn),  and  a  var.  in  bogs, 
Huon  Rd. :  Oldfield.  In  Melb.  Census  for  Tas.,  Vic,  and  N.S.W 
Austral.  Alps,  at  5000' :  Stirling. 

(a)  var.  perigoniale  (Rich.),  Br.  Eur. 
Vic,  Cobberas  Mts. :  F.v.M. 

76.  P.  CROCEUM,  Hpe.,  Linn.  '52. 
Tas.  :  teste  Par.  Index. 


20  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

PoLYTRiCHUM  (contimied).  [Polytrichacece. 

77.  P.  CYPELLOMiTRiUM,  CM.,  Symb.,  No.  32. 

N.S.W.,  Fitzroy  Falls,  on  logs  in  swamp,  Nov.  '84,  and  Kangaroo 
Valley,  Dec.  '85:  Whitelegge  (c.fr.). 

78.  P.  DENSiFOLiUM,  Hpe.,  Linn.  '60;  P.  Nov(e-Hollandice,y 
Jaeg. 

Vic,  Mt.  Wellington,  subalpine:  F.v.M. 

79.  P.  JUNiPERiNUM,  Hedvv. 

CosMOPOL. — Tab.,  common  on  Mt.  Wellington,  etc. :  Gunn,  Lyall, 
J.  D.  Hooker,  Mossman,  Archer,  Bastow,  Weymouth,  Watts  et  al. 
Vic,  Austral.  Alps:  Stirling  et  al.  "Common  on  light  gravelly 
soils,  especially  about  the  roots  of  trees.  Ascends  all  the  summits 
of  our  highest  mts."  (D.Sullivan).  Nr.  Melbourne:  Adamson. — 
N.S.W.,  Mt.  Kosciusko:  Forsyth,  Jan.  '99.— S.A.,  Mt.  Lofty 
Range:  F.v.M. 

[CM.  evidently  limits  this  sp.  to  the  N.  Hemisphere,  and 
includes  the  Tasm.  and  Austr.  specimens  under  his  P.  hycopo- 
dioides,  etc.] 

80.  P.  LONGiPiLUM,  CM.,  Symb.,  No.  34. 

Vic,  Upper  Ovens  R. :  McCann,  herb.  Melb.  '82.  Grampians 
and  Fowler's  Bay:  herb.  Melb.  '81.  Nr.  Melbourne,  StudleyPark: 
Reader,  Aug.  '83.  Daylesford:  Wallace,  '87. 

8L     P.  LYcoPODioiDES,  CM.,  Symb.,  No.  41. 
Tas.,  loc.  not  specified:  herb.  Melb.  '8L 

82.  P.  NODicoMA,  CM.,  Symb.,  No.  37. 
Vic,  Oakleigh:  Reader,  Sept.  '86  (m.pl.  only). 

P.  Novce-Hollandice,  Jaeg.     See  No.  78. 

83.  P.  OBLiQuiROSTRE,  CM.,  Symb.,  No.  29. 
Vic,  Mt.  William:  Sullivan,  Oct.  '78. 

P.  jnliferum,  Schreb.   "Pac.  Austral." :  teste  Par.  Index. 
Record  doubtful. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  21 

PoLYTRiCHUM  {continued).  \Polytrichacece. 

84.  P.  RECURViPiLUM,  CM.,  Syiiib.,  No.  31. 
N.S.W.,  Braidwood  District,  3400':  Biiuerlen,  Nov.  '84. 

85.  P.  RYPAROMiTRiuM,  CM.,  Syiiib.,  No.  33. 

N.S  W.,  Liverpool,  on  banks,  left-hand  side  of  railway  station: 
AVhitelegge,  Nov.  '84  (c.fr.). 

S^.     P.  SuLLivANi,  Hpe.,  Linn.  '76. 
Vic,  Grampians,  between  Mts.  Ararat  and  William  (parce  statu 
juniore):  Sullivan.     "Gullies  and  crevices  of  rocks,  Grampians 
and  Pyrenees"  (SulL).  Dandenong  Ranges :  herb.  Melb.  — N.S. W., 
Mt.  Kosciusko:  Biiuerlen,  '89. 

87.  P.  Tasmanle,  CM.,  Symb.,  No.  30. 
Tas.,  Marydale:  Weymouth,  Dec.  '90. 

88.  P.  Tysdalei,  cm.,  Symb.,  No.  38;  P.   Tisdallii,  CM., 

Gen.  M. 
Vic,  Gippsland:  H.  Tysdale,  herb.  Melb.  '84. 

Group  B.  ARTHRODONTES. 

Subgroup  AcROCARPi. 

Tribe  iv.  Fissidentaceae. 
F  I  s  s  I  D  e  N  s,  Hedw. 

F.  adiantoides,  Hedw.  Recorded  for  Tas.  in  Bastow, 
M.  Tas.  (Gunn  et  Archer  lect.)  and  in  Par.  Index,  but 
the  specimens  of  Gunn  and  Archer  were  named  with 
uncertainty  (see  Fl.  Tas.),  and  Mitt,,  in  Cat.,  gives 
F.  adiantoides,  Wils.  (non  Hedw.).  Stirling  records 
this  sp.  for  the  Austral.  Alps  (confirmation  needed). 
Broth,  and  CM.  restrict  F.  adiantoides  to  the  N. 
Hemisphere. 

F.  alternifolius.  Given  for  Vic.  in  Melb.  Census.  Par- 
ticulars needed. 


22  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

FissiDENS  {continued).  [Fissidentacece. 

89.  F.  AMPLiRETis  (CM.,  Gen.  M.,  p.  73,  Conomitruim). 
N.S.W.,  nr.  83'dney:  Whitelegge,  '84. 

90.  F.  APPLANATUS,  CM.,  Geii.  M.,  p.  62;  F.v.M.,  Fra,g. 
Phyt.  xi.  Suppl.  (CM.  repeats  under  a  different  sec- 
tion, p.  67,  op.  cit.). 

N.8.W.,  Richmond  R. :  teste  CM.  in  loc.  cit. 

91.  F.  ARBOREUS,  Broth.  A.M.  '90,  No.  7. 

Q.,  Pimpama,  on  trees:  C  J.  Wild,  Aug.  '87. — N.S.W.,  Richmond 
R.,  common  on  trees,  saplings  and  roots;  occasionally  on  rocks: 
Watts,  '96  ff.  (c.fr.). 

F.  Archeri,  F.v.M.  Name  in  Mitt.  Cat.  Recorded  in 
Melb.  Census  for  Vic.  Confirmation  needed.  Not 
in  CM.,  Gen.  M. 

F.  asplenioides  (Sw.),  Hedw.     See  No.  113. 

F.  assimilis,  CM.     See  No.  124. 

92.  F.  AUSTRALiENSis,  Jaeg.,  Fissid.   Enumerat. ;   Conomi-' 

trium  perjmsilhom,  Hpe.,  Linn.  '55  and  '60,  Jaeg.  Ad., 
CM.,  Gen.  M. 

Vic,  Sealer's  Cove:  F.v.M.  (Aust.  M.,  pi.  16).  Australian  Alps: 
Stirling.— N.S.W.,  Valley  of  Waters,  Blue  Mts.,  and  Taylor's  Bay, 
Port  Jackson:  Forsyth,  Aug.  '99. 

F.  bartra7niocarpus,  CM.     See  No.  131. 

93.  F.  BASiLARis,  Hpe.  et  CM ,  Linn.  '53. 

S.A.,  Barossa  Range,  assoc.  with  F.  pungens:  F.v.M. — Vic,  R. 
Yarra:  teste  Melb.  herb,  (a  doubtful  record). 

94.  F.  BREviFOLius,  H.f.W.,  Fl.  N.Z.,  Fl.  Tas. 

Tas.,  Johnny's  Creek,  Derwent  Valley,  and  Kangaroo  Point : 
Weymouth. — Vic:  teste  Par.  Index,  Suppl.  and  Melb.  Census. 
"  E.  Australia":  teste  Broth.  Bryales. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  23 

F  I  s  s  I  D  E  N  s  (continued).  \^Fissidentacece. 

(a)  var.  floribundus,  H.f.W.,  Fl.  Tas. 
Tas.,  shady  banks  by  the  Derwent,  New  Norfolk:  Oldfield.    (F. 
scalaris,  Mitt.,  according  to  Bastow,  M.  Tas.) 

F.  bryoides  var.  C(espita7is.     See  No.  98. 

95.  F.  CALODiCTYON,  Broth.,  A.M.  '90,  No.  6. 

Q.,  Ashgrove,  nr.  Brisbane:  C.  J.  Wild,  May  '88.— N.S.W., 
Richmond  R  ,  at  many  places:  Watts,  96  if. 

96.  F.  COARCTATUS  (CM.,  Gen.  M.,  p.  76,  Conomitrium). 
N.S.W.,   nr.   Sydney:    Whitelegge,   '84.     Cook's    R.,   April   '96, 
Parsley   Bay   '99,   Richmond   R.,   frequently   from   '96:    Watts. 
National  Park  '98,  Bot.  Gdns.,  Syd.  '99 :  Forsyth.  Manning  R.  : 
E.  Cheel,  herb.  Syd. 

97.  F.  coMMUTATUs,  CM.,  Gen.  M.,  p.  64. 
Australia,  prob.  Q.,  but  reference  in  loc.  cit.  not  clear. 

98.  F.  CuRNOWii,  Mitt.,  Jour.  L.  Soc.  '85;  F.  bryoides  var. 
ccespitaois,  Schimp. 

N.S.W.:  teste  Par.  Index. 

99.  F.  cuspiDiCALYX,  CM.,  Gen.  M.,  p.  56. 
Tas.,  Pedder's  Gully:  Weymouth  (?) 

100.  F.  dealbatus,  H.f.W.,  Fl.  N.Z. 

Tas.,  on  wet  rocks:  Archer  et  al. — N.S.W.,  Richmond  R.,  Ballina, 
on  swampy  ground,  Aug.  '96,  Wilson's  Creek,  Aug,  '98:  Watts 
(det.  Broth.). 

101.  F.  DENSiFOLius,  Broth.,  Bail.  Bot.  Bull.,  Dec.  91;  CM., 

Gen.  M.,  p.  59. 
Q.,  Mt.  Perry:  J.  Keys. 

102.  F.  DiETRicHi^,  CM.,  Linn.  '71-2. 

Q.,  Brisbane  R. :  Mrs.  Dietrich,  '64.— N.S.W.,  Richmond  R.,  many 
places,  especially  on  rocks  in  creeks,  also  on  wet  sea  cliffs  of 
Skinner's  Head,  and  on  bricks  of  footpath,  W.  Gollan's,  Wood- 
burn:  Watts,  96ff. 


24  CENSUS  MUSCORUM  AUSTRALIENSIUM, 

F  I  s  s  I  D  E  N  s  {continued).  [Fissidentacece. 

(a)  f.  minor,  Broth,  (det.) 
N.8.W.,  alluvial  soil,  bank  of  Nepean  R.,  at  Emu  Plains:  Watts, 
Oct.  '99. 

103.  F.   ELAMELLOSUS,  CM.  et  Hpe.,  Linn.   '56  ;   F.v.M., 
Aust.  M.,  pi.  20. 

Vic,  Yarra  R. :   F.v.M.  (mixed  with  F.  pungens  and  F.  hasilaris). 

104.  F.  EQUITANS,  CM.,  Gen.  M  ,  p.  75,   Conomitrium. 
N.S.W.:  teste  CM.,  loc.  cit. 

105.  F.  HOMOMALLULUs,  CM.  in  sched.  (not  in  Gen.  M.) 
N.S.W.,   Lilyvale:    Whitelegge,   Sept.   '91.     Richmond   R.   and 
Brunswick   R.,  at  several    places,  '96  ff,  and  under   rocky  cliff, 
Cook's  R.  '96:  Watts  (det.  Broth.) 

106.  F.  HYOPHiLUs,  Mitt.  Cat. 

Q.,  between  the  Burnett  and  Brisbane  Rs. :   F.v.M. 

F.  incurvo-hryoides.     See  No.  131. 

F.  incurvus,  Schw.  vars.  Records  in  Fl  Tas.  (St. 
Patrick's  R.  and  South  Port :  Gunn,  Oldfield),  and 
in  Melb.  Census  for  Tas.  and  Q.;  but  see  No.  131. 

107.  F.  iNTEGERRiMus,  Mitt.,  Jour.  L.  Soc.  '60. 
Tas.,  nr.  Cheshunt:  Archer. 

F.  intermedins,  CM.  This  name  (already  occupied  by 
CM.)  was  given  to  a  sp.  coll.  by  Whitelegge  at 
Mosman's  Bay  in  '84.  Apparently  the  sp.nov.  has 
been  dropped,  since  it  does  not  find  a  place,  so  far  as 
we  can  see,  in  Gen.  M. 

108.  F.  Kerianus,  cm.  in  Broth.  A.M.  '95,  No.  63. 
Q.,  Mulgrave  R.,  Ker  Range,  5200':  Bailey. 

109.  F.  LEPTOCLADUS,  CM.,  Gen.  M.,  p.  59. 
Tas.  :  teste  C  M.  loc.  cit. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  25 

FissiDENS  (continued).  [Fissiclentace(e. 

110.  F.  LEPTOLOMA,  CM.,  Gen.  M.,  p.  59. 

N.S.W.,  nr.  Sydney,  Eandwick  Rd. :  Whitelegge,  June  '84  (c.fr.). 
Richmond  R.,  Tintenbar  '96,  Brunswick  R.,  top  of  Myocum, 
about  300',  Sept.  '97  :  Watts. 

111.  F.  LEPTOPELMA,  CM.,  Gen.  M.,  p.  66;  F.v.M.,  Frag. 
Phyt.  xi.,  Suppl. 

N.S.VV.:  teste  CM.  loc.  cit. 

112.  F.  LEucoNEURUs,  CM.,  Gen.  M.,  p.  63. 
Q. :  teste  CM.,  loc.  cit. 

113.  F.  LiGULATUS,  H.f.W.,  Fl.   N.Z.;  F.  asple7iioides  (fid. 

Jaeg.  Ad.). 
Tas.:  teste  Bastow,  M.  Tas.  'S6,  and  Jaeg.  Ad.— Vic,  Austral. 
Alps:  Stirling. — N.S.W.,  Richmond  R.,  many  places,  on  wet 
rocks  by  creeks,  also  on  sea-clifF,  Ballina:  Watts,  '96  ff.  (c.fr.  Aug., 
rare).  Nepean  R.,  creeklet  at  foot  of  Blue  Mts. :  Watts,  Oct.  "99 
(det.  Broth.). 

114.  F.  liliputano-bryoides,  CM.,  Gen.  M.,  p.  56. 

Q.,  Victoria  Park:  H.  Tryon.  ("Vic."  in  Melb.  Census  — evidently 
in  error). 

115.  F.  liliputano-incurvus,  CM.,  Gen.  M.,  p.  56. 

Q.,  Beenleigh:  C  J.  Wild.— N.S. W.,  nr.  Sydney,  Lane  Cove, 
National  Park,  Bot.  Gdns.  Syd.,  also  Jamieson  Valle}',  Kiama, 
and  Shoalhaven  R. :  Forsyth,  July  '98-9.  Lily  vale:  Hamilton. 
On  upturned  roots  of  trees,  on  rocks  and  ground,  at  several  jDlaces 
on  the  Richmond  R. :   Watts,  '99  ff. 

116.  F.  linearis,  Brid. 

Australia:  herb.  Desfont. — Vic,  Gippland,  Moe  R. :  Luehmann, 
'81. 

117.  F.  LuEHMANNiANus,  CM., Gen.  M.,  p.  75,  Conomitrium.. 
Vic,  Moe  R. :  Luehmann,  '81,  mixed  with  F.  linearis. 


26  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

FissiDENS  (continued).  [Fissideniacece. 

118.  R  MACERATUs,  Mitt,  Cat.  '83. 
Q.,  Brisbane  R. :  Bailey. 

119.  F.  MACRODUS,  Hpe.,  Linn.  '60;  F.v.M.,  Aust.  M.,  pi.  19. 
Vic,  Yarra  R.,  on  grassy  ground:  F.v.M.  Banks  of  Yarra,  nr. 
Melbourne:  F.  M.  Reader,  Sept.  '84.— N.S.W.,  Barber's  Creek, 
in  gullies,  Sept.  '99,  and  Warrumbungle  Ranges,  Oct.  '99 :  Forsyth. 

(a)  f.  lamina  vera  sublimbata  (Broth,  det.). 
N.8.W.,  Shoalhaven  R.:  Forsyth,  Sept.  '99. 

F.  macropus,  Hpe.,  Par.  Index;  evidently  err.  for  F, 

macrodus. 
F.  microhlastus,  Broth,  in  sched.     See  No.  121. 

120.  F.  MiCROLECYTHTS,  CM.,  Gen.  M.,  p.  59. 
Q.,  Trinity  Bay:  teste  CM.,  loc.  cit. 

121.  F.  MoNTECOLLi,  Broth.,  Watts;  F,  microhlastus^  Broth. 

in  sched. 
N.S.W.,  Brunswick  R.,  top  of  Montecollum,  about  500':  Watts, 
Sept.  '97  (c.fr.). 

122.  F.  NANOPYXis,  CM.,  Gen.  M.,  p.  59. 
Vic:  teste  CM.,  loc.  cit. 

123.  F.  Nov.E-HoLLANDL?:,  Hsch.;  Conomitriicm,  CM. 
N.S.W.:  Sieber. 

124.  F.  Nov.E  Valesle,  Watts;  F.  assimilis,  CM.,  Gen.  M., 

p.  62  (cf.  Conomitr.  assimile,  Hpe.  '77,  a  Brazilian  sp.). 
N.S.W.,  "von  Sydney's  Kiiste  "  (CM.,  loc.  cit.). 

125.  F.  OBLONGiFOLius,  H.f.W.,  Lond.  J.  Bot.  '44. 

Vic,  nr.  Tarwin  R.  :  F.v.M.— Tas.,  Cataract  Hill:  Archer. 
Latrobe  and  Mt.  Faulkner:  Weymouth.  On  rocks,  N.  W.  Bay: 
Oldfield.— Q.,  Moreton  IsL:  H.  Tryon. 

126.  F.  obtuso-acuminatus,  CM.,  Gen.  M.,  p.  64. 
(^.,  Brisbane:  C  J.  Wild  (teste  Bailey,  Suppl.  '90). 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  27 

FissiDENS  {conthmed).  [Fissidentacece. 

127.     r.  PALLiDUs,  H.f.W.,  Fl.  N.Z. 
Tas.,  common:  Gimn  et  al. — N.S.W.,  frequent  from  Mt.  Kosciusko 
(Maiden,  '98)  to  Richmond  R.  (Watts,  96  ff.).   Bulli  Pass:  Watts, 
'95.   BlueMts.:  Betche, '95;  Forsyth, '99.  Turramurra:  Forsyth, 
'99.  Nr.  Sydney,  common. 

(a)  f.  viRiDis,  Broth. 
N.8.W.,  Richmond  R.,   Alstonville,  in  gully:  Watts,  April  '96 

(c.fr.). 

F.  2)((>rvus.     See  No.  155. 

F.  pauperrimics,  CM.     Par.  Index  gives   Australia. 
CM.,  Gen.  M.,  p.  59,  gives  S.  Africa  only. 

128.  F.  POROSUS,  CM.,  Gen.  M.,  p.  74,   Conomitrium. 
N.S.W.:  teste  CM.,  loc.  cit. 

129.  F.  puLviNATULus,  CM.,  Gen.  M.,  p.  64. 
Vic:  teste  CM.,  loc.  cit. 

130.  F.  PUNCTATUS,  Hpe.,  Jaeg.  Ad.;  F.v.M.,  Frag.  Phyt.  xi., 

Suppl. 
N.S.W. :  herb.  Melb.     (A  somewhat  doubtful  species.) 

131.  F.  PUNGENS,  Hpe.  et  CM.,  Linn.  '53;  F.v.M.,  Aust.  M., 

pi.  17;  F.  incurvus  var.,  Wils.  inFl.Tas.;  F.  bartramio- 
carpus,  CM.  ;  and  F.  incurvo-bryoides,  CM.  (teste 
CM.,  Gen.  M.,  p.  59) 
Tas.,  frequent. — S.A.,  Barossa  Range  and  Plenty  Creek:  F.v.M. 
— Vic,  stumps  of  trees,  Kew:  Reader,  July  '84. — N.S.W.,  Mos- 
man's  Bay,  Coogee  Bay,  Ball's  Head  Bay :  Whitelegge,  '84. 
Richmond  R.  and  Brunswick  R.:  Watts,  '96  if.,  frequent.  Jamieson 
Valley:  Forsyth,  '99.  Shellharbour :  E.  Cheel,  '99.  [As  F.  incurvo- 
bryoides,  frequent  in  herb.  Watts  (Richmond  R.),  also  in  herb. 
Syd.,  nr.  Syd.,  Forsyth,  and  Mt.  Warning,  Goldsmid].— Q. :  F.  M. 
Bailej^  (As  F.  incurvo-bryoides). 

F.  pygmceus,  Taylor.     See  No.  142. 


28  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

F  I  s  s  I  D  E  N  s  {continued),  [Fissidentacece. 

132.  F.  RiGiDULUS,  H.f.W.,  Fl.  N.Z. 

Vic,  Australian  Alps:  F.v.M.  and  Stirling.  Hector Travers,(loc.'?). 
— Tas.,  common.  St.  Crispin's  (We3'mouth),  Circular  Head,  etc. — 
N.S.W.,  Richmond  R.,  Wilson's  Creek:  Watts,  Dec.  '98.  Kiama  : 
Forsyth,  Dec.  '99. 

K  sarcophyllus,  CM.     See  No.  151. 

F.  scalaris.     See  iSTo.  94. 

133.  F.  SEMiLiMBATUS,  Hpe.  et  CM.,  Linn.  "53. 

Vic,  Yarra  and  Gippsland:  F.v.M.  (Aust.  M.,  pi.  18).  Austral. 
Alps:  Stirling. — Tas.:  Archer.  Nr.  Exeter,  W.  Tamar,  on  damp 
sandy  bank:  Weymouth,  '92. — N.S.W.,  nr.  Sydney,  Rookwood : 
Whitelegge,  '85  (c.fr.).  Lane  Cove:  Forsyth,  '98. 

134.  F.  SEMiRUFUS,  CM.,  Gen.  M.,  p.  76,  Conomiirmm. 
N.S.W.:  teste  CM.,  loc.  cit. 

135.  F.  SERRATO-MARGiNATUS,  CM.,  Gen.  M.,  p.  68. 
S.A.:  teste  CM.,  loc.  cit. 

136.  F.  SPLACHNOIDES,  Broth.,  A.M.  '93,  No.  27. 

Q.,  Indooroopilly,  on  ground:  Field  Naturalists  (Bail.  Bot.  Bull. 
Dec.  '93). 

137.  F.  STRiCTUS,  H.f.W.,  Fl.  Tas. 

Tas.,  Derwent  R.,  on  charred  log,  under  water:  Oldfield.  York 
Town  rivulet:  Gunn. 

138.  F.  SUBLIMBATUS,  Broth.  in  sched. 

Vic,  Lothiel,  Wimmera,  on  decaying  logs  in  a  pool:  Reader, 
Feb.  1900  (com.  as  F.  subalinibatus). 

139.  F.  suBAPPLANATUS,  CM.,  Gen.  M.,  p.  67;  F.v.M.,  Frag. 

Phyt.  xi.,  Suppl. 
N.S.W.:  teste  CM.,  loc.  cit. 

140.  F.  SuLLivANi,  CM.,  Gen.  M.,  p.  56. 

Vic,  Grampians,  Mt.  Ararat:  Sullivan.  Austral.  Alps,  up  to 
8000':  Stirling. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  29 

F I  s  s  I  D  E  X  s  {continued).  [Fissidentacece. 

141.  R  Sydneyensis,  Geh.  in  sched. 

N.S.W.,  Miiito,  on  banks  of  creek:  Whitelegge,  Feb.  '85.  Rich- 
mond R.,  on  rocks  in  creek,  Tintenbar  (fine  specimens):  Watts, 
May  '99  (det.  Broth.). 

142.  F.  Taylori,  cm.,  Syn.  1,  65;  F.  pyymceus,  Tayl. 
W.A.,  Swan   R. :  Drummond.— Tas.,  Cheshunt:   Archer.     Clay 
banks,   Mt.    Wellington:    Bastow   et   al. — Vic,   nr.   Melbourne, 
Studley  Park :  Reader,  July  '83. 

143.  F.  TENELLUS,  H.f.W^,  Fl.  N.Z. 

Tas.,  iSandstone  Hill,  Cheshunt:  Archer.  General:  Gunn,  Oldfield, 
Bastow,  Weymouth  et  al. — Vic,  Sealer's  Cove:  F.v.M.  Austral. 
Alps:  Stirling. — N.S.W.,  nr.  Sydney,  Cook's  R. :  Watts,  Forsyth. 
Richmond  and  Brunswick  Rs.,  at  many  places:  Watts,  '96  ff. — 
Q.,  Ashgrove,  Enoggera  Creek:   C.  J.  Wild. 

144.  F.  TORTUOSUS,  Hpe.  et  Geh.  Rev.  Bry.  '81. 
Tas.,  (loc.  ?). 

145.  F.  TRACHYPHYLLUS,  CM.,  Gen.  M.,  p.  76,  C o^iomitrium. 
N.S.W.,  Mosman's  Bay  '84,  Lilyvale  '91:  Whitelegge. 

146.  F.  UNDATO-DECURRENS,  CM.,  Gen.  M.,  p.  59. 
Q. :  herb.  Bailey. 

147.  F.  VicTORiALis,  Mitt.,  Cat.  '82-3. 

S.A.,  Victoria  R.:  F.v.M.  ("one  of  the  only  three  mosses  seen  by 
me  in  N.W.  Aust.  in  '55  and  56  ":  F.v.M.) 

148.  F.  viTTATUS,  H.f.W.,  Fl.  Tas. 

W.A.,  Swan  R. :  Drummond. — Tas.,  Circular  Head,  on  ground: 
Gunn.  Mt.  Faulkner,  etc. :  Weymouth.  Guy  Fawkes  Rivulet,  on 
water- washed  stones:  Watts,  '95. — Vic,  Gippsland:  F.v.M. 

149.  F.  Walteri,  cm.,  Gen.  M.,  p.  66. 
Vic,  Gippsland:  teste  CM.,  loc.  cit. 


30  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

FissiDENS  {contimied).  [Fissidentacece. 

150.  F.  Wattsii,  Broth,  in  sched. 

N.S.W.,  Sydney,  Woollahra,  damp  sandstone  rocks:  AVatts,  Mch. 
'96  (c.fr.). 

151.  F.  WeYxAIOUThi,  Par.  Index,  Suppl.  i.;  F.  sarcophylluSj 

CM.,  Gen.  M.,  p.  56  [cf.  Weymouth,  M.  Tas.   '94-5, 
p.  '21 J  F.  sarcoj^hyllus,  Burch.,  Broth,  (in  error)]. 

Tas.,  Mt.  Wellington,  on  face  of  rock  on  track,  below  the  Springs  : 

Weymouth,  '91. 

152.  F.  Whiteleggeanus,  CM.,  Gen.  M.  p.  67. 
N.S.W.,  nr.  Sydney,  Ball's  Head  Bay  (ster.),  N.  Shore,  Aug.  '84 
(c.fr.),  Lilyvale,  Sept.  '91 :  Whitelegge. — Tas.,  nr.  Hobart,  on  log, 
Huon  Rd.,  and  Mt.  Bischof,  on  ston}'-  earth-bank,  Happy  Valley, 
mixed  with  Mittenia  plumula:  Weymouth,  '92. — Q.,  Nerang:  H. 
Schneider. 

153.  F.  WiLDii,  Broth.,  A.M.  '90,  No.  5. 
■Q.j  Pimpama,  on  ground :  C  J.  Wild,  Aug.  '87. 

154.  F.  WooLLSiANUs,  CM.,  Gen.  M.  p.  66. 

N.S.  W. :  Woolls,  (loc.  ?).  Nr.  Sydney,  Northwood,  Lane  Cove  E., 
and  Mosman's  Bay:  Whitelegge,  June  '84.  Nepean  R.,  C'eeklet 
at  foot  of  Blue  Mts. :  Watts,  '99.  Katoomba  and  Shoalhaven  R. : 
Forsyth,  '99. 

[Note. — F.  deiicatidus,  Angstr.,  and  F.  pacificus,  Angstr.,  are 
both  given  for  N.S.W.  in  Mitt.  Cat.,  Jaeg.  Ad.,  and  Par.  Index, 
but  they  belong  to  the  Sandwich  Isls.     See  Rev.  Br}'.  '77,  p.  24.] 

OcTODiCERAS,   Brid.  (Sect,  of  Conomitrlum,  CM.  and  Par.) 
0.  Dillenii  (La  Pyl.),  Mont.     See  No.  155. 

155.  O.  MuELLERi,  Hpe.,  Linn.  '56  and  '60,   Conomitrium; 

Fissidens,  Mitt.  Cat.  and  Broth.  Bryales;  Octodiceras, 
Jaeg.  Ad.  ii.  In  Kew  J.  Bot.,  Mitt,  regarded  this  as 
0.  Dillenii,  a  S.  Amer.  sp.  Jaeg.,  Par.,  CM.,  and 
Broth,  separate  the  two  species. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  ."31 

OcTODiCERAS  {continued).  [Fissidentace(e. 

S.  A.,  Murray  R. :  F.v.M. — N.S.W.,  Richmond  R.,  on  rocks  under 
running  water,  Wilson's  Creek,  '97  ff.  (ster.),  Pearce's  Creek, 
Emigrant  Creek,  etc. :  Watts. 

Tribe  v.  Archidiaceae. 
A  R  c  H  1  D  I  u  M,  Brid. 

156.  A.  BRiSBANicuM,  Broth.,  A.M.  '93,  No.  24. 

Q.,  nr.  Brisbane,  Ipswich  Rd.,  on  swampy  land :  H.  Tryon,  Aug.  '90 
(c.fr.). — N.S.W.,  Richmond  R.,  on  ground,  Ballina,  Uralba,  and 
Pimlicolsl. :  Watts,  April  '99(c.fr.juv.),  July  1900,  Aug.  '01  (c.fr.). 

157.  A.  sTOLOxNACEUM,  CM.,  Gen.  M.   pp.  10-11. 
N.S.W.,    Sydney,    Moore    Park,    Greenwich,    etc.  :    Whitelegge, 
Sept.  '84.  Concord:  Watts,  July  '99  (ster.). 

Tribe  vi.  Ditrichacese  {Leptotrkhaceic,  CM.). 
B  R  u  c  H  I  A,   Schwgr.  (including  Sporledera,  Hpe.) 

158.  B.  AM(EJ^A,  CM.,  Flora  'd>^. 

N.S.W.,  nr.  Fitzroy  Falls,  on  rotten  logs  in  swamp:  Whitelegge, 
Nov.  '84  (c.fr.). 

B.  exigua  (H.f.W.),  CM.     See  No.  180. 

159.  B.  MiNUTA,  Mitt.,  Fl.  Tas.  and  Jour.  L.  Soc.  '60. 
Tas.,  Cheshunt,  on  sides  of  ditches:  Archer. 

160.  B.  Whiteleggei,  CM.,  Flora  '88,  B.  (Sporledera). 
N.S.W.,  Sydney,  Moore  Park:  Whitelegge,  July  '84  (c.fr.).— Q., 
Beenleigh:  C  J.  Wild. 

(a)  var.  minor,  CM. 
N.S.W.,  Sydney,  nr.  Randwick  Racecourse:  Whitelegge,  Aug.  '84. 

Trematodon,  Rich. 

Tr.  abrwptus.  Specimens  so  named  from  the  Bellingen 
and  Macleay  Rivers  (Aug.  Rudder)  are  in  Melb. 
herb.  (Bastow  in  MS.  list  thinks  this  may  be  Tr. 
adceqicans,  Geh.) 


32  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

T  R  E  M  A  T  0  D  0  N  (continued).  [Ditrichacece. 

161.  Tr.  ADiEQUANS,  Gch.  in  sched. 

N.S.W.,  Ball's  Head  Bay:  Whitelegge,  Oct.  '85.  Richmond  R., 
frequent:  Watts,  '96  ff. 

162.  Tr.  Baileyi,  Broth.,  A.M.  '90,  No.  2. 
Q.,  Bellenden  Ker,  Mulgrave  R. :  Bailey,  '89. 

163.  Tr.  brachyphyllus,  CM.,  Symb.,  No.  159. 

Q.,  ''tropical  Q.:  F.  M.  Bailey  in  herb.  Broth."  (teste  CM., 
Gen.  M.).   Mt.  Perry:  J.  Keys  (Bail.  Bot.  Bull.  Dec.  '91). 

16-1.     Tr.  flexipes,  Mitt.,  Jour.  L.  Soc.  '60  and  Fl.  Tas. 
Tas.,  W.  Mts.,  on  ground.  Plain  behind  Cumming's  Head:  Archer. 
[Broth.  Bryales  gives  N.Z.  only.] 

165.  Tr.  longescexs,  CM.,  Symb.  No.  160. 

N.S.W.,  nr.  Sydney:  Kayser  coll.,  '72,  Richmond  R.,  plentiful, 
especially  on  sides  of  ditches  and  banks:  Stackhouse,  '81,  Watts, 
96  ff.  Brunswick  R.,  Mullumbimby:  Bauerlen,  '95,  Watts,  '97.— 
Q.,  Beenleigh:  C  J.  Wild,  about  '88.   Brisbane:  Bailey. 

166.  Tr.  suberectus.  Mitt.,  Hdbk.  N.Z.  Fl. 

Q.,  Mt.  Perry:  Keys  (?).  See  Bail,  Syn.  Suppl.  i.  Mt.Gravatt  : 
herb.  Bailey.  [Broth.  Bryales  gives  N.Z.  only.] 

167.  Tr.  Whiteleggei,  Broth,  in  sched. 

N.S.W.,  cla}^  banks  opp.  Railway  Station,  Clifton:  Whitelegge, 
Aug.  '91. 

Pleuridium,  Schpr.  [Astomum,  CM.) 

168.  P.  BRACK YCAULON,  CM.,  Flora  '88,  Astomum. 
N.vS.W.,  Sydney,  Paddington:  Whitelegge,  Aug.  '84  (c.fr.).  Emu 
Plains,  Burwood  Park,  and  Concord  Park:  Watts,  Oct.  '99  (c.fr.). 
Warrumbungle  Ranges:  Forsyth,  Oct.  '99  (c.fr.). 

169.  P.  Breutelianum,  Hpe. 
Australia:  teste  Broth.  Bryales. 

P.  curvidum,  Taylor.     See  No.  174. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  33 

Pleuridium  (continued).  [Diirichacece. 

170.  P.  EXSERENS,  CM.,  Gen.  M.,  p.  12,  Astomum. 
Tas.:  teste  CM.,  loc.  cit.   (An  P.  sitbexset'ens  ?). 

171.  P.  GRAciLENTUM,  Mitt.,  Jour.  L.  Soc.  '60  and  Fl.  Tas. 
Tas.,  west  side  of  Tower  Hill:  Archer.  Bellerive:  A.  J.  Taylor. 
— W.A.,  King  George's  Sound:  Menzies. 

172.  P.  Krauseanum  (Hpe.  in  litt. ;  CM.,  Syn.  i.,  16, 
Astomum),  Salm.,  Rev.  Bry.  1900,  pp.  85-6.  Bruchia 
exigua  (fid.  CM.). 

S.  A.,  nr.  Kiwimanila,  Port  Adelaide:  Dr.  Behr.  —  Vic,  Yarra  R. : 
F.v.M.  '54.  Austral.  Alps:  Stirling.  Gippsland,  etc. :  herb.  Melb. 
— (I)N.S.W.,  Paddington  and  Randwick:  Whitelegge,  Aug.  '84. 
Blue  Mts. :  herb.  Melb. 

173.  P.  LAMPROPYXis,  CM.,  Symb.,  No.  85,  Astomum. 
Tas.  :  A.  Taylor,  herb.  Broth.  '90. 

174.  P.    NERVOSUM    (Hook.,  M.   Exot.,   Phascum)  H.f.W., 

SuUiv.  ;    Eccremidium    pidchellum    (fid.    Mitt.) ;     P. 
curvulum,  Tayl.  (fid.  Mitt.) 

W.A.,  Swan  R.  :  Jas.  Drummond. — Vic,  Gippsland  :  F.v.M. 
Austral.  Alps:  Stirling  [P.  curvulum). — N.S.W.,Parramatta  Park: 
Whitelegge,  Aug.  '85.  Burwood  Park,  Sydney :  Watts,  '99. 
Warrumbungle  Ranges:  Forsyth,  Oct.  '99. 

(a)  var.  iMINUS  :  Parramatta :  Woolls,  herb.  Melb. 

175.  P.  SuLLiVANi,  CM.,  Symb.,  No.  86,  Astomum. 
Vic,  Moysten:  Sullivan,  '82. 

176.  P.  SUBEXSERENS,  CM.,  Symb.,  No.  87,  Astomum. 
Tas.,  Pedder's  Field,  Kansaroa:  Weymouth,  Sept.  '89. 

177.  P.  TENELLUM,  Mitt.,  Jour.  L.  Soc.  '60  and  Fl.  Tas. 
Tas.,  Cheshunt:  Archer. 

P.  tenuissimum,  Tayl.     See  No.  179. 


34  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Pleuridium  {continued).  [Ditrichacece. 

178.  P.  viRiDE,  CM.,  Gen.  M.,  pp.  11-12,  Astomum. 

N.S.  W.,  Mosman's  Bay :  Whitelegge,  Aug.  '84  (c.fr. ).   Emu  Plains  : 
Watts,  Oct.  '99  (c.fr.). 

E  C  C  R  E  M  I  D  I  U  M,    H.f.  W. 

179.  E.  ARCUATUM,  H.f.W.,  Lond.  J.  Bot.  '46  :  Bryum 
tenuissimum,  Tayl.;  Pleuridium^  Mitt.  Cat. 

W.A.,  Swan  R. :  Drummond. 

180.  E.  EXiGUUM  (H.f.W.,  Icon.  PL,  Phascum),  8alm.,Kev. 
Bry.  1900,  pp.  85-6;  Bruchia,  CM.,  Bot.  Zeit.  '47; 
Astoniitm  Krauseanum,  Hpe.,  in  CM.,  8yn,  '49;  Pleu- 
ridium, Jaeg.  and  Par.  Wilson,  in  Kew  herb,  and 
Lond.  Jour.  Bot.  '46,  suggested  that  this  was  an 
Eccremidium;   E.  S.  Salmon  (I.e.)  confirms. 

W.A.,SwanR.:  Drummond. — ('?)N.S.W.,nr.  Sydney:  Whitelegge. 
[The  N.S.W.  record  prob.  refers  to  P.  Krauseanuni^  ^l-^'-] 

181.  E.  PULCHELLUM,  H.f.W.,  Lond.  J.   Bot.   '46;  Pleurid. 

curvulum,  Tayl.  (fid.  Mitt.  Cat.) 
W.  A.,  Swan  R. :  Drummond. — ('?)N.S.  W.,  on  earth,  nr.  Rookwood, 
Sydney:   Whitelegge,  Aug.   '85  (det.  Geh.)     [This  last  loc    may 
refer  to  Pleurid.  nervosum,  which  Mitt.  (Cat.)  identified  with  E. 
pulchellum.  ] 

182.  E.(Pseudo-Pleuridium)  Whiteleggei,  Broth.  Bryales, 

1901,  pp.  297-8,  fig.  175,  E.  S.  Salm.  del.  (subgen.  et 

sp.nov.) 
N.S.W.,  Maroubra  Bay,nr.  Sydne}^:  Whitelegge,  July  '98  (c.fr.juv.) 
and  Sept.  '98  (c.fr.matur.). 

D  I  T  R  I  c  H  u  M,  Timm.  {Leptotrichitm,  Hpe.) 

183.  D.  AFFINE,  CM.,  Bot.  Zeit. '47,  Leptotrlchum :  Tricho- 

stomum  setosum,  H.f.W.  (fid.  Jaeg.  et  al.) 
Tas.,  Sandy  Bay  Rivulet,  etc. :  Weymouth  (sub  2).  se^oso). — N.S.W,, 
common  from  Mt.  Kosciusko  to  the  Tweed  R.,  coastal  districts. 


BY  REV.   W.   WALTER  WATTS  AND  THOM  \S  WHITELEGGE.  35 

D  I T  R  I  c  H  u  M  {continued).  [Biti'ichacece. 

WaiTumbungle  Ranges:  Forsj^th, '99.  BlueMts. :  T.  Steel  (herb. 
Tech.  Mus.  Syd.)  and  Watts. — Q.  :  No  record  available,  but  no 
doubt  frequent. — Vic,  Gippsland  :  herb.  Melb. 

(a)  f.  alpina,  Broth, 
N.S.W.,  Mt.  Kosciusko:  Forsyth,  Jan.  '99. 

[CM.  in  Gen.  M.  has  D.  affine  for  Chili,  and  D.  setosum  for 
N.Z.     The  Australian  plant  may  be  one  of  his  sp.no v.] 

184.  D.  AUSTRALE,  Mitt.,  Jour.   L.  Soc.  '60  and  Fl.  Tas.; 

Lophiodon  strictus,  H.f.W. 

Tas.,  Cheshunt:  Archer.  Mt.  Wellington  and  Mt.  Bischof,  Happy 
Valley:  Weymouth.  Mt.  Wellington :  Watts  (det.  Broth.).  Huon 
Rd. :  Bastow. 

[CM.  in  Gen.  M.  gives  this  sp.  for  S.  Amer.  only.     Possibly 
the  Tasm.  plant  is  his  D.  strictiusculuin.~\ 

185.  D.  Baileyi,  cm. 

Q.:  herb.  Bailey  and  herb.  Simmonds.   Ashgrove:  Wild,  '87. 

186.  D.  BRACHYCARPUM,  CM.,  Symb.,  No.  166;  L.  cylindri- 

carpum,  Hpe. 
N.S.W.,  Blue  Mts.:  F.v.M.  '72. 

D.  hrachycarjjum,  Hpe.     See  No.  220. 
D.  capillaceum.  Mitt.     See  No.  201. 
D.  cylindricarpum^  Hpe.     See  No.  186. 

187.  D.  CYLiNDRiCARPUM,CM.,Bot.Zeit.'51;  Trichostomuvi, 

Fl.  Tas. 
Tas.,  nr.  Launceston,  on  rocks,  Cataract:  Mossman.  —  Vic,  Mt. 
WilHam:  Sullivan,  Nov.  '87.   E.  Gippsland:  herb.  Melb.  '80. 

188.  D.  ELONGATUM  (H.f.W.,  Fl.  Tas.,  Trichostomum),  Mitt. 
Tas.,  moist,  clayey  soil,  Mt,  Wellington,  Grass-tree,  etc.:  Fraser, 
J.  D.  Hooker  et  al.  Cheshunt:  Archer.  Mt.  Wellington:  Wey- 
mouth, Watts. — Vic:  teste  Melb.  Census. — E.Australia:  teste 
Par.  Index. 


36  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

D  I  T  R  I  c  H  u  M  {continued).  [Ditrichacece. 

D.  ferrugineum,  Mitt.     See  No.  220. 
D.  flaccidulum,  CM.    Given  in  Melb.  Census  for  Yic. 
(leg.  B.  Seemann,  '60).  Species  doubtful. 

189.  D.  FLAViPES,  CM.  in  Weymouth's  M.  Tas.  '95. 

Tas.,  on  earth  banks,  Cradoc  Rd.,  Huon,  Balfe's  Hill,  and 
Forestier's  Peninsula:  Weymouth,  '89. 

D.flexijolium,  Hook.     See  No.  190. 

D.  latifolium,  Schwgr.,  given  in  Melb.  Census  for  Tas. 
(Mt.  Wellington:  Bastow,'86.)  Confirmation  needed. 

190.  D.  laxifolium,  H.f.W.,  Fl.  N.Z.,   Trichostomum;  D. 
Jlexifolium,  Hook.,  M.  Exot.,  Dicranum. 

Vic:  F.v.M.— Tas.,  on  hills  about  Hobart:  J.  D.  Hooker,  Oldfield 
et  al.  Mt.  Disappointment:  Norman  Taylor. 

191.  D.  MuELLERi,  Hpe.,  Linn.  '56,  Lejnotrichum:  Lophi- 

odon,  Jaeg.  et  Par.   [Not  in  Broth.  Bryales.] 
•'From    Vic.    to    Queensland"    (teste    CM.,    Gen.    M.).— Vic, 
Grampians,  etc.:  F.v.M.   Alps:  Stirling.   Murrumbeena:  Reader, 
Nov.  '81.   E.Gippsland:  Bauerlen,  '89. — Q.:  herb.  J.H.  Simmonds. 
Ashgrove:  Wild,  '87.— N.S.W.,  New  England:  herb.  Melb. 

192.  D.  Oldfieldii,  Mitt.,  Jour.  L.  Soc.  '60,  Leptotrichum. 
Tas.,  on  ground,  (loc.  ?):  Oldfield.  St.  George's  R.,  and  Hobart 
Rivulet:  Weymouth. 

D.  phceiun,  given  in  Melb.  Census  for  Vic  and  N.S.W. 
(Blue  Mts.)   Confirmation  needed. 

193.  D.  scABRiFOLiUM,  Mitt.  Cat.  '82-3. 
Tas.:  Archer,  Oldfield,  Bastow. 

194.  D.  SEMiLUNARE,  CM.,  Symb.,  No.  167  (Sect,  of  Lepto- 
trichum). 

Vic,  Upper  Yarra,  Fernshaw  and  Gippsland  (Moe  R.) :  Luehmann, 
Jan.  '81.     N.  Gippsland,  Igers  R.  :  Tysdale  {Sesdall  in  Symb., 
Tisdall  in  Gen.  M.),  '81.  Hume  R.:  Miss  Campbell,  '81. 
D.  setosum,  H.f.W.     See  No.  183. 


BY  REV,  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  37 

D  I  T  R  I  c  H  u  M  {continued).  [Ditrichacece. 

195.  D.  STRiCTiuscuLUM,  CM.,  Symb.,  No.  164  (Sect,  of 
Leptotrichum). 

Tas.,  (loc.  '?):  Oldfield,  herb.  Broth.  '90. 

196.  D.  SUBBRACHYCARPUM,  CM.,  Symb.,  No.  165  (Sect,  of 

Leptotrichicm). 
Vic.  to  N.S.W.,  Grampians,  Mt.  Kosciusko:  Sullivan,  '84. 
Z>.  subcapillaceum,  CM.     See  No.  201. 

197.  D.  viRiDE,CM.,Symb.,No.l68  {^Qct.oi  Leptotrichum). 
Q.,  Bellenden  Ker:  Karsten,  '81. — N.S.W.,  Richmond  R.:  Stack- 
house  and  De  Camara,  '81. 

Ceratodon,   3rid. 

198.  C  coNVOLUTUs,  Reich.,  Novara  Exped.;  C.  j^u^yuraus, 

Fl.  N.Z. 
S.A..  Adelaide:  teste  F.v.M.  in  list  to  the  late  Prof.  Tate.— Tab.: 
teste  Broth.  Bryales  ("  Neuholland,  Tasm.,  N.Z.").  CM.,  Gen.  M., 
gives  N.Z.  only. 

C.  crassinervis,  Ltz. — Tas.:  teste  F.v.M.,  Frag.  Phyt. 

xi.  Suppl.     This  needs  confirmation.    Possibly  it  is 

the  var.  nervo  crassiore  of  C.  purpui-eus  in  FL  Tas., 

I  coll.   by  Oldfield  in  bogs,  N.W.  Bay.     Broth,  in 

Bryales  gives  Chili  only. 

199.  C  purpureus  (L.),  Brid. 

CosMOPOL. — Throughout  Aust.  and  Tas.  (teste  Broth.)  CM. 
regarded  our  plant  as  C.  pur  p.  var.  australis,  CM. 

200.  C  stenocarpus  (Mont.),  Br.  Eur.;  CM.,  Syn. ;  C. 
2)urj)U7'eus  var.  palustris,  Mont.  In  Gen.  M.,  CM. 
no  longer  recognised  this  sp.  Broth,  in  Bryales,  how- 
ever, retains. 

Tas.,  in  bush,  frequent:  Oldfield  et  al.  Latrobe,  etc.:  Weymouth. 
^N.S.W.,  Mt.  Dromedary:  Reader,  '80.  Richmond  R.,  Pearce's 
Creek,  on  log:  Watts,  '97. — Vic.  and  Q.:  teste  Melb.  Census. 


38  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

\^Ditrichacece. 
D  I  s  T  I  c  H  I  u  M  (Svv.),  Br.  Eur. 

201.  D.  CAPILLACEUM  (Sw.),  Br.  Eur.;  Le2:>totriGhum{Aschis- 
todon)  suhcapillaceum,  CM.  in  Gen.  M.,  p.  314; 
Sioartzia  montana,  Lindb. 

Tas.,  Mt.  Wellington,  Silver  Falls,  at  root  of  trees:  Bastow. 

[CM.  does  not  recognise  B.  capillaceum  in   Aust.  and  Tas., 
hence  his  D.  subcapill.] 

DiCRANELLA,  Schimp.   {Aongstroe77iia,  CM.) 

202.  D.AFOPiiYSATA,C.M.,Hjmb.,l>^o.lHl, {A.  apophy8atida)y 
Gen.  M.,  p.  325;  Mierodus,  Par.  Index,  Suppl.  i. 

N.8.W.,  Hurstville,  nr.  Sydney,  sides  of  creeks :  Whitelegge, 
Dec.  '84  (c.fr.).  Richmond  R.,  E.  Ballina:  Watts,  Dec.  '98  (ster.). 

203.  D.  Baileyana,  CM.,  Symb.,  No.  180,  Aongstr.  {Weisi- 

ella);  Microdus,  Par.   Index,  Suppl.  i.  D.  (Microdus)y 
Broth.  Bryales. 
Q.,  Brisbane  R.:  Bailey,  '80.   Burpengary:   Wild,  '87. 

204.  D.  BREVISETACEA,  Gch.  in  sched. 

N.S.W.,  Hurstville,  nr.  Sydney,  on  clay  banks  in  a  gully:  White- 
legge, Oct.  '84  (c.fr.). 

205.  D.  DiETRiCHiiE,  CM.,  Linn.  '67-8,  Aongstr.  (Divari- 
catella). 

q.,  Brisbane  R.:  Mrs.  Dietrich,  '64.— N.S.W.,  Ball's  Head  Bay: 
Whitelegge,  '84.  Richmond,  Brunswick  and  Tweed  Rs.,  very 
plentiful:  Stackhouse,  Watts,  Camara  et  al.  Nr.  Casino:  D.  A. 
Porter.  National  Park,  nr.  Sydney:  Forsyth. 

20G.      D.    FALLENS  (H.f.W.,  Icon.  PI.  Rar.,  Weisia),  Broth. 
Bryales;  Seligeria,  CM.;  Microdus,  Par.  Index. 
W.A.,  Swan  R.:  Drummond. 

207.     D.  PAUCiFOLiA,  CM.,  Symb.,  No.  169  (Sect,  of  Aongstr.). 
Vic,  North  East:  comm.  to  D.  Sullivan,  '83.   Omeo:  StirUng,  '83. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  39 

DiCRANELLA  {continued).  [^Ditrichacece. 

208.  D.  PELLUCiDA,  Broth,  in  sched. 

N.S.W.,  Richmond  R.,  Teven,  Sept.  '97,  Brunswick  R.,  Myocum, 
Sept.  '97 :  Watts. 

209.  D.  PERDivARiCATA,    CM.,  Symb.,  No.   173,   Aongstr. 

(Bivaricatella) 

Tas.,  Mt.  Wellington,  St.  Crispin's  Well,  about  2000':  Weymouth, 
Nov.  '90. 

210.  D.  PYCNOGLOSSA,  Broth.,  A.M.  '90,  No.  1,  Anisothecium; 
Dicraiiella,  Par.  Index  and  Broth.  Bryales. 

Q.,  Mulgrave  R.,  on  clayey  soil,  Ker  Range,  5200':  Bailey,  '89. — 
N.S.W.,  Richmond  R.  (Tintenbar:  Bauerlen,  Aug.  '95,  herb. 
Tech.  Mus.  8yd.  Alstonville  Cutting,  etc.:  Watts,  '96  ff.) 

211.  D.  RUFO-AUREA,  Hpc.,  Linn.  '60,  Aonystroemia. 

Vic,  on  porphyritic  rocks,  nr.  summit  of  Mt.  Cobboras,  5000', 
and  on  the  limestone  at  lower  levels:  F.v.M.,  Stirling  et  al. 

212.  D.  ScHREBERi  (Sw.),  Schimp.;  Anisothecium  crispum 
(Schreb.),  Lindb.;  D.  {Anisotheciurn),  Broth.  Bryales. 

Tab.,  on  rocks.  Newtown  Falls,  '89,  and  St.  Crispin's,  Mt. 
Wellington,  '90:  Weymouth. 

(a)  f.  procera. 

Tas.,  Mt.  Wellington:  Weymouth,  Mch.  '91. 

213.  D.   Stackhousiaxa,  CM.,  Symb.,  No.   179,  Aongstr. 

(Weisiella)  ;    Microdus,    Par.    Index,    Suppl.    i. ;    D. 
(Mic7'odus),  Broth.  Bryales. 

N.S.W.,  Richmond  R.:  Stackhouse,  '81. 

214.  D.  TENAX,  CM.,  Symb.,  No.  178,  Aongstr.  (Weisiella); 

Microdus,  Par.  Index,  Suppl.  i.;  D.  {Microdus),  Broth. 
Bryales. 

Q.:  F.  M.  Bailey, '84.   Breakfast  Creek  :  Tryon. 


40  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

DiCRANELLA  (continued).  [Ditrichacece. 

(a)  var.  longipes,  CM.,  ibid. 
Q.,    between    Cooktown    and    Douglas,    assoc.    with    Philonotis 
graminea:  Dr.  Lucas,  '84. 

215.  D.  TRiCHODONTOiDEA,  CM.,  Rev.  Bry.  '76  (name).  Not 
in  CM.,  Gen.  M.,  apparently. 

N.S.W,,  nr.  Sydney:  Kayser  coll.  Lawson,  Blue  Mts.,  and  Gore 
Cove,  Sydney:  Whitelegge,  '84.  Brunswick  R.,  Mullumbimb}^: 
Bauerlen,  Sept.  '95. 

216.  D.  TRiCRURis,  CM.,  Linn.  '68,  Aongstr.  [Dobelon); 
Gen.  M.,  Aongstr.  (Weisiella);  Dichodontium,  Jaeg., 
Par. 

Q.,  Brisbane  R.:  Mrs.  Dietrich. — N.S.W.,  Lilyvale:  Whitelegge, 
'91.  Richmond  R.,  and  Cook's  R.,  nr.  Sydney:  Watts.  Buningbar 
and  Turramurra:  Forsyth. 

217.  D.  Wattsii,  Broth.,  A.M.  '98,  No.  95. 

N.S.W.,  Richmond  R.,  road  from  Brooklet  to  Fernleigh,  damp 
places:  Watts,  Nov.  '96  (c.fr.).  Later  at  Pearce's  Creek  and 
Alstonville,  and  prob.  Wilson's  Creek:  idem. 

Tribe  vii.  Dicranaceae. 
B  L  I  N  D  I  A,  Br.  Eur. 

Bl.  acuta,  Br.  et  Schimp.     See  No.  219. 

218.  Bl.  arcuata,  Mitt.,  M.  Aust.  Amer.,  Bl.  Magellanica, 

H.f.W.  (fid.  CM.,  Gen.  M.,  p.  245) 
Tas.:  Archer.  St.  Crispin's  Well,  Mt.  Wellington,  on  masonry: 
Weymouth  (c.fr.).  Watts.   Rocks,  Millhouse's  Falls,  Huon  Rd.: 
Weymouth. 

219.  Bl.  curviseta.  Mitt.,  M.  Aust.  Amer.,  Bl.  acuta  var. 
curviseta,  Mitt.,  Jour.  L.  Soc.  '60;  Bl.  acuta  (H.f.W., 
Fl.  Ant.). 

Tas  ,  on  rocks,  Johnny's  Creek :  Oldfield.  On  stones,  rivulet 
behind  Cumming's  Head,  W.  Mts.:  Archer. — Vic:  teste  Melb. 
Census  (as  Bl.  acuta). 


BY  REV.   W.  AVALTER  WATTS  AND  THOMAS  WHITELEGGE.  41 

B  L  I  N  D  I  A  (continued).  [^Dicranacece. 

220.  Bl.  ferruginea,  (Mitt.,  Jour.  L.  Soc.  '60,  Dicraniim 
{Leptot7'ir.hum),)  Broth.  Bryales,  '01  ;  A7i{sothecium, 
Mitt.  Cat.  ;  Dichodontium,  Jaeg.  Acl.,  Par.  Index  ; 
Ditrichiun  brachycar^jum,  Hpe.,  Linn.  '71-2  (Par. 
Index,  Suppl.  i.). 

Tas.,  on  ground,  plain  behind  Cumming's  Head,  W.  Mts.:  Archer. 
Boggy  soil  top  of  Mt.  Wellington:  Weymouth,  Nov.  '90.— N.S.W., 
Blue  Mts.:  Woolls.— Vic:  teste  Melb.  Census  and  CM.,  Gen.  M., 
p.  313. 

£1.  Magellanica,  H.f.W.     See  No.  218. 

221.  Bl.  robusta,  Hpe.,  Linn.  '60;  F.v.M.,  Aust.  M.,  pi.  7. 

Vic— N.S.W.,  Munyang  Mts.  in  bogs,  6000-7000',  also  in  the 
Grampians:  F.v.M.  "From  shaded  sidelings  of  mica  schist  nr. 
Omeo,  2000',  to  the  summits  of  Mt.  Kosciusko  on  granitic  rocks, 
7000'":  Stirling, '80-88.  Blue  Mts.:  herb.  Melb.  (c.fr.).  Also  coll. 
by  Forsyth  on  Mt.  Kosciusko,  Jan.  '99  (herb.  Syd.).— Tas.,  top 
of  Mt.  Wellington:  Weymouth,  '90.  Wellington  Falls:  Bastow, 
'87. 

222.  Bl.  tenuifolia,  H.f.W.,  Lond.  Jour.  Bot.  '46  (1), 
Dicranum. 

Tas.,  rock,  behind  Cumming's  Head.  W.  Mts.:  Archer.  Swampy 
ground,  top  of  Mt.  Wellington:  Weymouth. — Vic:  teste  Par. 
Index,  Suppl.  i. 

[In  herb.  Melb.  identified  with  £,  robusta.] 

223.  Bl.  Wellingtonii,  CM.,  Gen.  M.,  p.  245. 
Tas.,  Mt.  Wellington,  assoc.  with  Bl.  robusta. 

Dicranoweisia,  Lindb. 

D.  cirrhata  (Hedw.),  Lindb.  A  N.  Hemisphere  sp. 
recorded  for  Tas.  in  Par.  Index — apparently  in 
error. 


42  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

D  I  c  R  A  N  o  w  E  I  s  I  A  {continued).  [Dicranacew. 

224.  D.  MicROCARPA  (H.f.W.,  Fl.  Tas.,  Weisia),  Par.;  Holo- 
niitriuni  cirrhatiwi,  Mitt.;  Dic7'anum,  Mitt.  Cat. 

Tas.,  Cheshunt:  Archer.  Glen  Leith  and  Mt.  Nelson:  J.  D. 
Hooker.  Kangaroo  Pt.  (teste  Bastow,  M.  Tas.). — Vic:  F.v.M. 
(teste  Mitt.,  Jour.  L.  Soc.  '60),  (loc.  1) 

H  0  L  o  M  I  T  R  I  u  M,  Brid. 

225.  H.  CORTICOLUM,  Broth.,  in  CM.,  Gen.  M. 
(^.,  nr.  Brisbane  (Bail.,  Bot.  Bull.  '91). 

226.  H.  DiETRicHLE,  CM.,  Linn.  71-3. 

Q,.,  Brisbane  R.:  Mrs.  Dietrich,  '64.  Severn  R.:  Hartmann  (see 
Rev.  Bry.  '77,  p.  43,  for  Hartmann's  coll.  nr.  Toowoomba). 
Russell  River:  Sayer,  '86. 

227.  H.  HoDGKiNSONLE,  CM.,  Symb.,  No.  78. 
N.S.W.,  Richmond  R.:  Miss  Hodgkinson  (herb.  Melb.  '79). 

(a)  var.  virescens,  CM.,  ibid.;  H.    Whiteleggei,  CM., 

olim. 
N.S.W.,   Sydney,   on   rocks,   Greenwich:   Whitelegge,  June  '84. 
Also  Ball's  Head  Bay,  Gore  Cove,  Manly,  and  St.  George's  R.: 
idem. 

228.  H.  MuELLERi,  Hpe.,  Linn.  '70. 

Q.,  Rockingham  Bay:  Dallachy, '68.  Bellenden  Ker:  S.Johnson, 
'91.— N.S.W.,  Blue  Mts.,  Mermaid's  Glen:  Whitelegge,  Oct.  '85. 

229.  H.  Nov^  Valesle,  CM.,  Rev.  Bry.  76.  (Not  in  CM., 

Gen.  M.). 
N.S.W.:  Kayser  coll.,  "  nr.  Sydney."  (A  doubtful  sp.) 

230.  H.   perich.etiale  (Hook.,  M.   Exot.,  Trichostomuni), 

Brid.;  JSpnblepharis,  Wils.,  Fl.  N.Z. 
Tas. — Vic— N.S.W.  — Q.,  fairly  frequent,  mostly  on  trees,  but  in 
E.  Ballina,  Richmond  R.,  on  sandy  soil  (Watts,  '96).  Tasman's 
Peninsula:   Weymouth. 


¥ 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  4S 

H  0  L  0  M  I  T  R  I  u  M  (continued).  [Dicranacece. 

H.  vaginatum,  Brid.      Given  in  Melb.  Census  for  Q. 

(Condamine  Scrub:  Hartmann,  '75);  also  in  Hpe.'s 

List  of  '80;  record  very  doubtful. 
H.  Whiteleggei,  CM.     See  No.  227  var. 

C  A  M  p  Y  L  0  p  u  s,  Brid.   (Sect,  of  Dicranum,  CM.  et  al.) 

231.  C  APPRESSiFOLius,  Mitt.,  Hdbk.  N.Z.  FL;  C.  clavatu&, 

H.f.W. 
Tas.  and  Q.:  teste  Melb.  Census. — N.S.W.  (loc.  ?):  Cunningham. 
Burns'  Bay,  on  stones  in  creek:  Whitelegge,  'Aug.  '84  (c.fr.) 

232.  C  AusTRALiENSis,  Duby,  '69. 

Vic,  (loc.  '?) :  "  Garcon  Maria  et  F.  Mueller"  (teste  Jaeg.,  Ad. 
Suppl.) 

233.  C.  BicoLOR,  Hsch.,  M.  Sieb.,  Dicranum. 

Tas. — W.A. — Vic. — N.S.W.,  many  Iocs.  Frequent  about  Port 
Jackson,  \vhere,  apparently,  Sieber  first  found  it. 

234.  C  BRUNNEUS,  CM.,  Symb.,  No.  51. 

Tas.,  Mt.  Wellington:  Beccari,  Feb.  .'78.  Also  a  form:  Mrs. 
Kayser,  in  herb.  Melb.  '83. 

235.  C  CADUCiPiLUs,  Besch. 

"  E.  Australia":  teste  Broth.  Bryales. — N.S.W.,  nr.  Sydney: 
herb.  Cardot. 

236.  C  CAPiLLATUS,  H.f.W.,  Fl.  Tas. 

Tas.,  bogs.  Brown's  R. :  Old  field.  South  Port:  Stuart.  Happy 
Valley,  Mt.  Bischof :   Weymouth  (f.  fol.  integerrimis). 

237.  C.  CLAVATUS,  R.Br.;  C.Jlexuosus,  Brid.  (fid.  Fl.  Ant.) 

Tas.:  R.  Brow^n,  Sieber,  Oldtield,  Bastow,  Weymouth  (Watch- 
horn's  Hill,  etc.)  et  al. — N.S.W.,  nr.  Sydney:  Jupp,  Watts  et  al. 
Menangle:  Harper.  Monga  and  Grosse  Valley:  Bauerlen.  Rich- 
mond R. :  Watts.  Nepean  R. :  A.  Sharpe. — S.A.,  Adelaide: 
Schauinsland,  '96-7. 


44  CENSUS    MUSCORUM  AUSTRALIENSIUM, 

Campylopus  {continued).  [Dici'anacece. 

238.  C.  DEPiLis,  CM.,  Gen.  M.,  p.  276. 
Vic,  Mt.  William:  Sullivan  (?). 

C.  Dietrichice,  CM.,  Jour.  Godeffroy.  Recorded  for  Q. 
by  F.v.M.  in  Frag.  Phyt.  xi.  Suppl.,  and  Bail.  Syn. 
'83.  Evidently  an  error.  Jaeg.  Ad.  gives  "insul. 
austral.  Tongatabu:  A.  Dietrich,  '72."  Par.  Index 
gives  Tonga  only. 

239.  C  ERYTHROPOMA,  Duby  '69,  Mem.  Soc.  Phys.  Geneva. 
Vic:  "Garcon  Maria  et  F.  Mueller"  (Jaeg.,  Ad.  Suppl.). 

C.Jlexuosus  (Hedw.),  Brid.  Given  for  Vic  (Sealer's 
Cove:  F.v.M.)  in  Frag.  Phyt.  xi.  Suppl.  Broth. 
Bryales  gives  N.  Hemisphere  only.  Prob.  C. 
clavatus,  q.v. 

C.Jlexuosus,  CM.     See  No.  254. 

24:0.     C  HoLOMiTRiUM,  CM.,  Syn.;  C.  cajnllaceus,  H.f.W., 
Lond.  J.  Bot.  '44.   (Not  in  CM.,  Gen.  M.) 
Vic:  teste  F.v.M.,  Frag.  Phyt.  xi.  Suppl. 

241.  C  HOMALOBOLAX,  CM.,  Symb.,  No.  45. 
W.A.,  King  George's  Sound:  Webb  '82,  herb  Melb. 

242.  C  iNSiTiTius,  H.f.W.,  Fl.  Tas.  (?  C.  insidiosus,  Mitt. 
Cat.)  "This  appears  to  me  to  be  identical  with 
Brown's  C.  clavatus;' Mitt., ^\.  Tas.  '60.  Broth.  Bryales 
retains  the  sp. 

Tas.,  bogs   in   gullies  about  Brown's  R. :  Oldfield.    South  Port: 
Stuart. 

243.  C  INTROFLEXUS  (Hedw.,  Dicranum),  Mitt. 
Common  throughout  Australia  and  Tasmania.   Ascends  to  6000', 
Austral.  Alps:  teste  Stirling. 

244.  C  KiRKii,  Mitt.;  Dicnemos  Moorei,  Broth.,  Geh.  in 

Rev.  Bry.  '97,  p.  67  (teste  Broth.,  A.M.  '98,  p.  74). 
Tas.:  teste  Broth,  loc.  cit. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  45 

C  A  M  p  Y  L  o  p  u  s  (continued).  [Dicranaceoi. 

245.     C.    LEPTOCEPHALUS,   CM.,   Linii.    '55,  Dicranum;   C. 
introfleoms  (fid.  Mitt.  Cat.). 
W.A.  and  Vic:  teste  F.v.M.,  Frag.  Phyt.  xi.  Suppl.  — S.A.,  Mt. 
Gambier:  teste  F.v.M.  in  list  to  the  late  Prof.  Tate. 

C.  Martince  (Vic.)  and  C.  microgaster  (S.A.)  are  re- 
corded in  Melb.  Census.  Particulars  needed.  C. 
Martensi  is  given  bj-Mr.  Stirling  for  Austral.  Alps, 
ascending  to  4000'.     Is  this  C.  Martince  ? 

24G.     C.  NiGRO-FLAVus,  CM.,  Symb.,  No.  46. 
W.A.,  Mt.  Lindsay:   Webb  '82,  herb.  Melb. 

247.  C  Nov^  Valesle,  Broth.,  A.M.  '98,  No.  97. 
N.S.W.,  Bulli  Pass:  Watts,  Nov.  '95  (ster.). 

248.  C  NUDUS,  Hpe.,  Linn.  '60. 
Vic,  Tarwin  R.:  F.v.M. 

249.  C  PERAURicuLATUs,  Brotli.,  A.M.  '99,  No.  142. 
N.S.W.,  Richmond  R.,  Granuaille  Rd.:  Watts,  Aug.  '98. 

250.  C  PUDicus  (Hsch.,  M.  Sieb.,  Dicranum)^  Jaeg. 
N.S.W.,   nr.    Sydney:    Sieber,    Kayser.     Richmond   R.:    Watts, 
June'96.— Vic,  Bunyip  Creek:  F.v.M.— Tas.,  Peppermint  Bay '89, 
Mt.  Wellington  (Guy  Fawkes  Rivulet),  and  Mt.  Bischof :  Wey- 
mouth.— Q.,  Brisbane  River  Scrubs:  herb.  Bailey. 

251.  C  SENEX,  CM.,  Symb.,  No.  50. 

Vic,  Gippsland,  nr.  Moe  R.:  Luehmann  '81,  herb.  Melb.  Hume 
R.:  Miss  Campbell  '81,  herb.  Melb.— N.S.W.:  teste  CM.,  loc.  cit. 

252.  C  SUBBICOLOR,  Broth.,  Geh. 

Vic,  Mt.  Arapiles:  Reader,  Aug.  '96.  Mt.  William:  teste  CM., 
Gen.  M. 

C.  subcapillatus.     Given  in  Melb.  Census  for  N.S.W. 
^  and  Q.     Particulars  needed. 

C.  subto7'quatus,  CM.     See  No.  257,  var. 
[In  herb.  Bailey,  Q.,  is  a  C.  suhtorquatus  var.  flavifoliiis,  but 
this  var.  is  not  found  in  CM.,  Symb.,  under  C.  WooUsii.] 


46  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Campylopus  {continued).  \^Dicranacece. 

253.  C.  TASMANicus  (Scliinip.),  CM.,  Symb.,  No.  49  (received 
by  Schimp.  from  Tas.  "  sub  C.  introjfexus,^'  herb. 
Hooker). 

Common  in 'Tas.,  Vic,  KS.W.,  and  W.A.:  teste  CM.,  loc.  cit. 
**E.  Australia,  Tas.  and  N.Z.":  teste  Broth.  Bryales.  On  earth, 
Fitzroy  Falls :  Whitelegge,  Nov.  '84. 

C.  torfaceus,  Mitt.     See  No.  254. 

254.  C.  TORQUATUS,  Mitt.,  Jour.  L.  Soc.  '60,  and  Fl.  Tas.; 

C.flexuosus,  CM.,  Bot.  Zeit.  '51;  C.  torfaceus,  Mitt., 

Kew  Miscell.  '56. 
Tas.,  on  decaj^ed  logs  nr.  Cheshunt:  Archer.  Mt.  Wellington, 
etc.:  Weymouth,  Watts  et  al. — Vic:  F.v.M.  Austral.  Alps,  up 
to  5000':  Stirling. — Q.,  Bellenden  Ker:  Karsten.— N.S.W.,  Rich- 
mond R.:  Camara.  [The  Richmond  R.  and  Q.  records  are  doubtful; 
all  specimens  in  herb.  Watts  named  at  first  C  torquatus  by  Broth, 
were  corrected  to  C.  WooUsii.   Broth,  in  Bryales  gives  Tas.  onl3\] 

255.  C  viRiDiCATUs,  CM.,  Symb.,  No.  52. 

N.S.W.,  Sydney,  North  Shore,  North  wood,  and  Waterloo:  White- 
legge, June  and  July  '84. — Q.,  (loc.  ?):  Bailey  (a  form). 

256.  C  Whiteleggei,  CM.  in  herb.,  Gen.  M.,  p.  267. 
N.S.W.,  Head  of  Blue's  Bay,  '84,  Ball's   Head  Bay,   Feb.  '^d: 
Whitelegge. — Q.,  Stanthorpe,  on  rocks. 

257.  C  WooLLSii,  CM.,  Symb.,  No.  44. 

N.S.W.,  Sydney:  Woolls,  herb.  Melb.'Sl.  Little  R.:  R.  Collie, '90. 
Brunswick  R.  :  Biiuerlen,  '95.  Richmond  R.  :  Watts,  '96  ff., 
common  on  logs  and  ground.  Mt.  Warning,  Turramurra,  Ourimbah, 
National  Park,  etc.:  Forsyth.  Wyong:  Hamilton.  Shellharbour: 
Cheel. — Q.,  Nerang  Creek:  Schneider.   Brisbane:  Bailey. 

(a)  var.  cylindrica  (CM.  ibid.,  "  theca  perfecte  cylin- 
drica"),  Par.  Index,  Suppl.;  C.  subtorquatus,  CM.  in 
sched. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS   WHITELEGGE.  4/ 

Campylopus  {continued).  [Dicranaceai, 

N.S.W.— Sydney:  Kayser  coll.  75.  Woolls,  '81.  Manly,  Botany, 
and  Clifton:  Whitelegge,  '84.— Q.,  Brisbane:  Bailey,  '89.— Vic, 
Mt.  William:  G.  G.  Miller,  '83.  {C.  snbforquatus  ya.i\  Jiavi/olius, 
q.v.,  was  coll.  by  C.  J.  Wild  at  Pimpama  and  Burpengary) 

[Note. — C.  densifolius,  Angstr.,  and  C.  geniculatus,  Angstr., 
have  been  recorded  for  N.8.W.  in  error.  They  belong  to  the 
Sandwich  I.sls.     See  Rev.  Bry.  '77,  p.  24.] 

D  I  c  R  a  N  0  D  0  N  T  I  u  M,  Br.  Eur. 

258.  D.  TAPES,  CM.,  Symb.,  No.  42,  Sect,  of  Dicranum. 
Tas.,  Deal  Island:  Judge  Dobson,  Mch.  '72.   Herb.  Melb.  '84. 

M  E  s  o  T  u  s,  Mitt. 

259.  M.  ACUTUS,  Mitt.  Cat. 

*' Australia,  inter  SphcHrophoron,  from  Borrer's  collection"':  teste 
Mitt.,  loc.  cit.— Tas.  {X)\  teste  Par.  Index. 

Leucoloma,  Brid. 

[Some  difficulty  is  experienced  in  separating  this  genus  from 
Dicrajium.  Dr.  C.  Mueller  refused  to  see  in  the  hyaline  leaf- 
margin  a  ground  of  generic  distinction.  In  his  '  Gen.  M.,' 
Leucoloma,  Dicnemonella,  and  Oncophorohma  are  sections  of 
Dicranum,  but  in  '  Symbolae  '  the  section  Leucolotna  includes  the 
other  two  as  subsections.  Dr.  Brotherus,  in  '  Bryales,'  makes 
Leucoloma  a  large  genus  covering  so  much  additional  ground 
that  all  our  Australian  Dicrana  become  Leucolomata.  For  the 
present  we  have  used  Leucoloma  in  Mueller's  sense,  including 
Dicnemonella  and  Oncophoroloma,  and  have  retained  Dicranum 
for  the  remaining  species.] 

260.  L.  austro-scoparium,  CM., in  Broth.,  A.M.  '95,  No.  54, 

DicranuTii. 
Q.,  summit  of  Bellenden  Ker,  4-5000':  Bailey.   "  Bloomfield  R.": 
teste  CM.,  Gen.  M. 


48  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Leu  COLO  MA  {continued).  [Dicranacece. 

261.  L.  BAUER.E,  CM.,  Symb.,  No.  68,  Sect,  of  Dicranum. 
Q.,  Bloomfield  R.:  Miss  Bauer,  herb.  Melb.  '84. 

262.  L.  BRACHYPELMATUM,  CM.  in  sched.,  Sclerodontium. 
N.S.W.,  Gore  Cove,  nr.  Sydney:   VVhitelegge.  (Not  in  Gen.  M.) 

263.  L.  CALYMPERACEUM  (CM.,  Sjmib.,  No.  62,  Dicranum), 

Broth,  Bryales;  D.  calymj^eridum,,  Bail.,Syn.  Suppl.  '84. 

Q.,  tropical:  Bailey  in  herb.  Kiaer,  '84. 

264.  L.  CLAViNERVE,  CM.,  in  Broth.,  A.M.  '93,  No.  26. 

Q.,  Petrie's  Quarries,  Brisbane,  on  rocks:  H.  Tryon,  Aug.  '90. 
''Tropical  Q.":  teste  Bail.,  Sjm.  Suppl.  '88. 

265.  L.  DiCARPUM  (Hsch.,  M.  Sieb.,  Dicranum),  Broth. 
Bryales;  D.  leucolomoides,  CM.,  Bot.  Zeit.  '51  (fid. 
Mitt.). 

N.S.W.,  nr.  Sydney:  Sieber.  Maitland:  Vickary.  New  Jerusalem, 
William's  R.:  J.  Lamont,  '90.  Monga:  Bauerlen.  Menangle:  herb. 
Tech.  Mus.  Syd.  Richmond  and  Brunswick  Rs.,  frequent:  Watts. 
— Q.  :  teste  Bail.,  Syn.  '83.— Vic,  Gippsland:  F.v.M.  Austral. 
Alps,  2000'-3500':  Stirling.  (Tas.  records  in  Fl.  Tas.,  etc.,  are 
apparently  D.  polysetum,  especially  the  var.  spinosum,  H.f.W.) 

266.  L.  Fraseri,  Mitt.,  Dicnemonella  and  Sclerodontium. 
N.S.W.:  Fraser.   Parramatta:  Woolls  (Mitt.  Cat.). 

267.  L.  Harrisii,  Geh.  in  sched.;  Poecilophyllum  in  Melb. 

Census. 
N.S.W.,  Cambewarra:  C  Harris,  Sept.  '85. 

268      L.  IMBRICATUM,  Brotli., Geh. in  Broth.,  A.M.  '98,  No.  91. 
N.S.W.,  Mt.  Dromedary:  Reader. 

269.     L.  iNCANUM,  Mitt.,  Hdbk.  N.Z.  Fl. 
Australia:  teste  Broth.  Br3'ales. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  AVHITELEGGE.  49 

Leucoloma  (continued).  \_Dicranacece. 

270.  L.  LEicHHARDTi,Hpe.,Linn. '70;  Poecilophyllum,  Mitt. 

Cat. 

N.S.W.:  Leichhardt.  Armidale:  Perrot.  IllawarraR.:  S.Johnson, 
herb.  Melb. — Vic:  F.v.M.— Q.,  Brisbane  R.:  Baile3^  Cunning- 
ham's Gap:  Hartmann. 

271.  L.  LiMBATUM,  Broth.  (CM.,  Gen.  M.,  Dicranum). 

S.A.:  teste  CM.,  op.  cit.,  p.  285.— Vic,  Blacks'  Spur:  Miss 
Campbell. 

272.  L.  SERRATUM,  Broth.,  A.M.  '90,  No  3. 

Q.,  Mt.  Mistake,  on  trees :  Bailey,  June  '85. — N.S.W.,  Cambewarra : 
C  Harris,  '85.  Lilyvale:  Whitelegge,  Sept.  '91.  Richmond  R.,  on 
trees  in  scrubs  at  many  places:  Watts,  '96  ff.  On  sandy  soil  on 
rocks,  Roseville,  nr.  Sydney,  '96,  and  Blue  Mts.  '02:  Watts. 

273.  L.  SiEBERiANUM,  Hsch.,  M.  Sieb.;  Leucodon  pallidus, 

Hook.;  Sclerodo7itium,  pallidurn,  Schwgr. 

Common  on  the  E,  Coast  of  Australia,  in  N.S.W.  and  Q.  Also 
found  in  Vic,  S.A.,  and  Tas.     Sieber's  locality  was  nr.  Sydney. 

(a)  var.  rigida.  Broth. 
Q.,  Mt.  Archer:  C  T.  Musson. 

274.  L. STRiCTiPiLUM, CM., Symb.,  No.  71,  Sect. of  Dicranum. 
N.S.W.,  nr.  Sydney:  Kayser  coll.  (ster.) 

275.  L.  suBiNTEGRUM,  Broth.,  A.M.  '90,  No.  4. 

Q.,  Bellenden  Ker,  Palm  Camp,  4000',  on  tree:  Bailey,  '89  (pi. 
fern.  ster.). 

276.  L.  SuLLivANi,  CM.,  Symb.,  No.  69,  Sect,  of  Dicranum. 

Vic,  Mt.  William,  Grampians:  Sullivan,  herb.  Melb.  '82.  Austral. 
Alps,  2000':  Stirling.  Healesville,  nr.  Melbourne:  Schauinsland, 
'96-7. 

4 


50  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

Leucoloma  (continued).  [Dicranacece. 

277.  L.  WhitelegCxEI,  CM.,  Symb.,  No.  70,  Sect,  of 
Dicranum ;  Broth.,  A.M.  '95,  No.  55  (Dicranum), 
Bryales  (Leucoloma). 

N.S.W.,  Fitzroy  Falls:  Whitelegge,  Nov.  '84. 

D  I  c  R  A  N  u  M,  Hedw.,  Schimp. 

278.  D.  ANGUSTiFOLiuM,  H.f.W.,  Fl.  Tas. 
Tas.,  in  bogs,  Mt.  Wellington:  Gunn,  Oldfield  (ster.). 

279.  D.  angustinerve,  Mitt.,  Jour.  L.  Soe.  '60,  and  Fl.  Tas. 
Tas.,  on  dead  wood,  Cheshunt:  Archer.  Also  collected  by  Gunn, 
Nowell  et  al.— Yic:  F.v.M.— N.S.W.,  Blue  Mts.:  herb.  Melb. 

280.  D.  ARGUTUM,  Hpe.,  Linn.  '68-70. 

N.S.W.:  Hooker's  Coll.  (Mitt.,  Cat.).     Shoalhaven  R.:  herb.  Melb. 

281.  D.  AUSTRixuM,   Mitt.  Cat.;    D.  sphagni  var.,  Fl.  Tas. 

(Sp.  not  in  Par.  Index) 
W.A.,  King  George's  Sound:  Cunningham. — Q.:  teste  Bail.  Syn. 
'83.   "Interior  of  subtropical  Australia ":  Sir  T.  Mitchell. —Tas.: 
teste  Mitt.  Cat.,  F.v.M.,  and  Bastow,  M.  Tas. 

282.  D.  austro-congestum,  CM.,  Symb.,  No.  61. 
N.S. VV.,  on  rocks,  Fitzroy  Falls:  Whitelegge,  '84  (ster.). 

283.  D.  Baileyanum,  CM.,  Symb.,  No.  60. 
Q.:  Bailey  in  herb.  Kiaer,  '83. 

284.  D.  Billardierii,  Schwgr.;  D.  truncorum^  Schimp. 
Australia,  (loc.  *?):  La  Billardiere. — Vic:  F.v.M.  Blacks'  Spur: 
Miss  Campbell,  '84.  Austral.  Alps,  2-3000':  Stirling,  "80  ff.— Tas., 
common  on  roots  of  trees,  on  logs,  etc.,  Mt.  Wellington,  etc. : 
Archer,  Gunn,  Stuart,  Oldfield,  Hooker,  Bastow,  Weymouth 
(St.  Crispin's,  Mt.  Nelson,  etc.)  et  al. — N.S.W.,  Port  Jackson: 
Gaudichaud.   Monga,  on  rocks :  Bauerlen  (herb.  Tech.  Mus.  Syd.). 

D.  hrachyjjelma,  CM.     See  No.  292. 


BY  REV.   W.   WALTER  WATTS   AXD  THOMAS  WHITELEGGE.  51 

D  I  c  R  A  N  u  M  {continued).  [^Dicranacece. 

285.  D.  BuRCHARDi,  Par.  Index,  Suppl.  i.;  D.  rige^is,  CM., 
Symb.,  No.  57;  D.  rigens,  Burch.,  Broth,  (in  error), 
Weymouth,  M.  Tas.;  L.  rigens  (CM.),  Broth.  Bryales. 

Tas.,  Mt.  Wellington,  "Ploughed  Fields,"  on  rocks,  4000',  '91, 
and  on  log,  Lauriston  Gully,  '89:  Weymouth  (herb.  Burch.  et 
Broth.). 

D.  chlorocaulon,  CM.  Name  in  F.v.M.,  Frag.  Phyt.  xi., 
Suppl.  for  N.S.W.,  but  not  in  CM.,  Gen.  M. 

286.  D.  CHLOROCLADUM,  CM.,  Rev.  Bry.  '76,  and  Symb., 

No.  74. 
N.S.W.,  nr.  Sydney:  Kayser  coll.  Hunter  R.:  herb.  Melb. 

287.  D.  CONFINE,  Hpe.  et  CM.,  Linn.  '56. 

Vic,  Sealer's  Cove:  F.v.M.  Oakleigh:  Reader,  Aug.  '86.— Tas.: 
teste  CM.,  Gen.  M.  (Melb.  Census  gives  Vic.  only.) 

288.  D.  DiAPHANONEURON,  Hpe.,  Linn.  '70. 

Vic,  Stirling's  Range:  F.v.M.— W. A.:  teste  F.v.M.,  Frag.  Phyt. 
xi.,  Suppl. 

289.  D.  EUCAMPTODONTOiDES,  Broth.,  Geh.  in  Broth.,  A.M. 

'95,  No.  58. 
Tas.,  Jones'  track,  Macquarie  Harbour,  on  sandy  button-grass 
hills:  T.  B.  Moore,  July '93. 

D.  fasciatum,  Hed.  A  N.Z.  sp.  recorded  for  Tas.  in 
Par.  Index  and  for  N.S.W.  in  Melb.  Census  (with 
a  1).  The  Tas.  record  also  is  doubtful. 

D.Jlav>folium,CM.  |  Names  in  F.v.M.,  Frag.  Phyt.  for 
D.  Geheebii,  CM.      i      N.S.W.  Not  in  CM.,  Gen.  M. 

290.  D.  INTEGERRIMUM,  Broth.,  Geh.  in  Broth.,  A.M.  '95, 

No.  57. 
Tas.,  Jones'  track.   Sprent  R.,  W.   Coast,  on  sandy  button-grass 
plains:  T.  B.  Moore. 

D.  Kaipare7ise.     See  No.  292. 


52  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

D  I  c  R  A  N  u  M  {continued).  [Dicranacece. 

291.  D.  Kroneanum,  CM.,  Symb.,  No.  65. 

Vic,  on  eucalypts  and  tree-ferns,  Fernshaw  (Venus  Expedition): 
H.  Krone,  April  75.  — Tas.,  Mt.  Wellington:  Dr.  Beccari,  Feb. 
78.  Fern  Gully,  Mt.  Wellington:  Weymouth,  Jan.  '88.— N.S.W.: 
teste  F.V.M.,  Frag.  Phyt.  xi.,  Suppl. 

292.  D.   Menziesii,  Ta}^.,  Phytol.;  D.  Kaiparense  (iormerly 

D.  bmchypehna),  CM.,  fid.   Mitt.,  Jour.  L.   Soc.  '60 

(cf.  CM.,  Gen.  M.). 
Tas.,  frequent:  Archer,  Gunn,  Bastow,  Weymouth  (Bo wen  Creek, 
Henty  R.,  etc.)  et  al. — Vic,  Grampians,  Victoria  Range,  Tarwin 
R.,  and  Sealer's  Cove:  F.v.M. — N.S.W.,  S3^dney,  Gore  Cove,  on 
rocks  and  trees:  Whitelegge  (c.fr.).  Richmond  R.,  Wilson's 
Creek:  Watts  (ster.).  BlueMts.:  Schauinsland, '967.  Katoomba: 
Forsyth,  Dec.  '99  (c.fr.). 

(a)  var.  rigidum,  Mitt.,  Jour.  L.  Soc.  '60. 
Tas.:  Archer  and  Gunn  apparently  coll. 

D.  viicrocarpwm.     See  No.  224. 

293.  D.  MONOCARPUM,  CM.  in  sched. 

N.S.W.,  Blue  Mts.,  Lawson,  on  trees  in  a  gully:  Whitelegge, 
June  '85  (c.fr.). 

294.  D.  Nelsoni,  CM.,  Symb.,  No.  58. 
Tas.,  Facy's  gully,  Mt.  Nelson:  Weymouth. 

295.  D.  (Edithecium,  CM.,  Symb.,  No.  63. 
N.S.W.,  Fitzroy  Falls:  Whitelegge,  Nov.  '84  (c.fr.). 

296.  D.  POLYCHiETUM,  Hpe.,  Linn.  '60. 
Vic,  Yarra  R.  and  Grampians:  F.v.M. 

297.  D.  POLYSETUM,  Hpe.,  Linn.  '60. 

Vic,  Dandenong  Ranges,  on  rocks,  etc.:  F.v.M. — Tas.,  Buckland  : 
W.  Turvey,  '87.  Guy  Fawkes  Rivulet,  Mt.  Wellington,  Russell's 
Falls,  etc.:  Weymouth,  Watts  et  al.  (All  Tas.  records  of  L. 
dicarpum  apparently  belong  here) — N.S.W.:  teste  Melb.  Census. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  53 

D  I  c  R  A  X  u  M  [continued).  [^Dicranacece. 

298.  D.  PUNCTULATUM,  Hpe.,  Linn.  '60. 

Vic,  Austral.  Alps:  F.v.M.,  Stirling  et  al.  "On  metamorphic 
schists  nr.  Omeo  between  2000'  and  3500'"  (Stirling)  —KS.W.: 
teste  F.V.M.,  Frag.  Phyt.  xi.,  Suppl. 

299.  D.  PUNGENS,  H.f.W.,  Lond.  J.  Bot.  '4-4,  and  Fl.  Ant.; 

D.  Novce  HoUandice,  Hsch.,  partim. 
Tas.,    Cheshunt:    Archer.    Summit  of  Mt.    Wellington:    Gunn, 
Oldfield  et  al. — Vic:  teste  Melb.  Census.   [Broth.  Bryales,  gives 
this  sp.  for  Fuegia  and  the  Antarctic  isls.  only.  The  Vic.  record, 
at  any  rate,  is  jDrob.  D.  suhjningens.^ 

300.  D.  PUNGENTELLA,  CM.,  Symb.,  No.  59. 
Tas.,  Mt.  Wellington:  herb.  Melb. 

301.  D.  ROBUSTUM,  H.f.W.,  Lond.  J.  Bot.  '44,  and  Fl.  Tas. 
Tas.,  Cheshunt:  Archer. — "Australia":  teste  Jaeg.  Ad.  [Broth., 
Bryales,  gives  Chili,  Fuegia,  and  N.Z.  only.] 

var.  pungens.     See  No.  299. 

302.  D.  SETOSUM,  H.f.W.,  Lond.  J.  Bot.  '44,  and  Fl.  Ant. 
Tas.,  Mt.  Wellington,  not  uncommon:  J.  D.  Hooker,  Oldfield, 
Bastow,  Weymouth  (Fork  Creek  and  "  Ploughed  Fields  "),  Watts 
(track  to  St.  Crispin's,  det.  Broth.),  et  al.   [Broth.,  Bryales,  gives 
Fuegia  and  Antarctic  isls.  only.] 

(a)  var.  attenuatum.    Given  in  Fl.  Ant.    Par.  Index 
says  "c.  typ./'butoccurrencein  Tas.  needs  confirmation. 

303.  D.  SORDIDE-VIRIDE,  C.  M.,  Rev.  Bry.  '76;  "  voisin  du  D. 

Menziesii  Tayl^  (Geh.,  loc.  cit ). 
N.S.W.,  nr.  Sydney:  Kayser  coll.  Blue  Mts.,  Blackheath  Glen: 
Whitelegge,  Sept.  '85  (c.fr.). 

B,  Starkii,  W.M..  Bastow,  M.  Tas.,  gives  "Brown's 
R.,  Tas.";  but  the  record  is  more  than  doubtful. 
Barren  and  dwarf  specimens  only  were  referred  to 
in  Fl.  Tas.,  and  named  with  a  1 


54  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

D  I  c  R  A  N  u  M  {continued).  [Dicranacece. 

D.  sphagni,  Wahl.  Given  in  Fl.  Tas.  for  barren  speci- 
mens collected  by  Gunn.     See  No.  281. 

304.  D.  suBERECTUM,  Hpe.,  Linn.  '60. 

Vic,  Tarwin  R.,  "  in  valle  ":  F.v.M.— N.S.W.:  teste  CM., Gen.  M. 
Nellie's  Glen,  on  moist  sandstone:  T.  Steel,  Dec.  '98  (ster.). — Q., 
Palm  Camp,  Bellenden  Ker,  4000':  herb.  Bailey. 

(a)  var.  fol.  angustioribus. 
Vic,  Tarwin  R.:  F.v.M.,  herb.  Melb. 

305.  D.  SUBPUNGENS,  Hpe.,Linn.  '60;  D.pungens  (fid.  Mitt.). 
Vic,  Grampians,  "William  and  Victoria  Ranges:  F.v.M. — Tas.: 
teste  Broth.  Brj^ales. — N.S.W.,  Tingiringi:  Bauerlen. 

306.  D.  suBSETOSUM,  CM.,  Symb.,  No.  54. 

Tas.,  Mt.  Wellington:  Weymouth,  Jan.  '88. — Australia  (a  form 
sent  to  CM.  in  '67  hjJ.  E.  Zetterstedt,  but  without  specified  loc). 
D.  sicbviride,  CM.,  Linn.  '72.  Given  in  Mitt.  Cat.  for 
N.S.W.  ("nr.  Sydney:  Kayser").  Not  in  Geheeb's 
list  of  the  Kaj^ser  mosses  in  Rev.  Bry.  '76,  nor  in 
Jaeg.  Ad.  Possibly  an  error  for  D.  sordide-mride. 
We  have  not  access  to  Linn,  of  '72. 

307.  D.  TRicHOPHYLLUM,  Hpe.,  Linn.  37,  p.  515. 
Australia:  teste  Broth.  Bryales.     In  Jaeg.  Ad.  "  Australia,  Chat- 
ham Isl." 

308.  D.  Walteri,  Hpe. 
Vic:  C  Walter. 

309.  D.  Weymouthi,  CM.,  Symb.,  No  56. 
Tas.,  Southdale:  Weymouth,  Oct.  '89. 

D  I  c  N  e  M  0  s,  Schwgr. 

310.  D.  CALYCiNUS  (Hook.,  M.  Exot.,  Leucodon),  Schwgr. 
"Australia":  herb.  Dickson  (Mitt.  Cat.).— N.S.W.:  teste  Melb. 
Census. — Vic,  AustraL-  Alps,  up  to  3000':  Stirling.  [CM.  and 
Broth,  give  N.Z.  onl3\] 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  55 

D  I  c  N  E  M  0  s  (continued).  \^Dicranacece. 

D.  e7ie7'vis,  Geh.     See  No.  312. 
D.  Moorei.     See  No.  244. 

D.  Muellerianus.     See  No.  312. 

311.  D.  RUGOSUS  (Hook.,  M.  Exot.,  Leucodon),  Scliwgr. 
"Australia":  herb.  Dickson  (Mitt.  Cat.).— N.S.W.:  teste  Melb. 
Census. 

[Occurrence  in  Australia  doubtful.  Dickson  sent  the  moss  to 
Turner  marked  "  Australia,"  but  no  other  botanist  has  ever  coll. 
it  here.    Possibly  from  Tahiti.    See  Salmon,  Jour.  Bot.  Jan.  1902.] 

EucAMPTODON,  Mont. 

E.  Hampeamis,  CM.     See  No.  312. 

312.  E.  MuELLERi,  Hpe.  et  CM.,  Linn.  '70;  Dicneiuon 
enerve,  Geh.  in  Rev.  Bry.  76  (fid.  CM.,  Gen.  M.); 
E.  Hampeanus^  CM.,  loc,  cit.,  evidently.  Dicnemos 
Muellerianus. 

Yic,  Austral.  Alps:  F.v.M.  et  al.— N.S.W.,  New  England,  '69 
(collector  X).  Tenterfield:  Bauerlen,  Aug.  '01.  S.  Coast,  Fitzroy 
Falls,  Cambewarra:  Whitelegge,  Nov.  '84.  Blue  Mts.,  common 
in  glens :  Whitelegge  '84,  Forsyth  '99,  Watts  '02.  N.  Coast, 
Richmond  R.,  frequent  (Hunter's  Scrub,  Alstonville,  etc.):  Watts 
'96ff.   Mt.  Warning  :  Forsyth  '98. 

313.  E.  spiNicusPES,  Broth,  in  sched. 

N.S.W,,  East  Ballina,  swampy  ground  on  heath:  Watts,  Nov. 
'97fr.  (ster.). 

Tribe  viii.  Leucobryacese. 

Leucobryum,  Hpe. 

L.  australe.     See  No.  330. 

314.  L.  Baileyanum,  CM.,  Gen.  M.,  p.  80. 

Q.:  herb.  Bailey.— N.S.W. ,  Ball's  Head  Bay,  Mosman's  Bay,  and 
Head  of  Double  Bay,  nr.  Sydney  :  Whitelegge,  '84. 


56  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Leucobryum  {continued).  [Leucohiyacecn. 

315.  L.  BALLiNENSE,  Broth.  in  sched. 

N.S.W.,  Richmond  R.,  East  Ballina  on  trees  :  Watts,  '99ff. 
L.  hrachy^yhyllum,  Hsch.     See  No.  318. 

316.  L.  BRACK YPHYLLUM,  Hpe.,  Linn.  '39. 

N.S.W.,  common  from  the  S.  Coast  to  the  Tweed. — Q.,  Moreton 
Bay:  Bailey.  BellendenKer:  Karsten. — Vic:  teste  Melb.  Census. 

(a)  var.  minus.  Name  in  Geheeb's  paper  on  the  Kayser 
coll.,  Rev.  Bry.  76. 

N.S.W.,  "nr.  Sydney":  Kayser  coll. 

317.  L.  Camar^,  cm.,  Gen.  M.,  p.  80. 
"Australia":  teste  CM.,  loc.  cit.;  loc.  not  recorded. 

318.  L.   candidum,  Brid.     [In  Par.  Index,  "Z.  candidum 

(Brid.),  Jaeg.,"  covers  L.  brachyjyhyllum,  Hpe.,  as  well 
as  L.  hrachyphyllum,  Hsch.,  and  "  var.  major "  is 
prob.  L.  candidum  proper.  Broth,  returns  both  species.] 

Tas.,  Cheshunt :  Archer.  Huon  R.,  Brown's  R.,  etc.:  Oldfield,  J. 
D.  Hooker,  Bastow  et  al.  Guy  Fawkes  Rivulet:  Watts. — Vic: 
A.  H.  Lucas  and  Miss  Campbell.  Austral.  Alps  :  Stirling.  Heales- 
ville,  on  tree  stem :  Schauinsland,  '96-7.  Dandenong  Ranges : 
Luehmann. — N.S.W.,  Sugar  Loaf  Mt.,  Braidwood  district: 
Bauerlen,  '90.  Lilyvale :  Whitelegge,  Sept.  '91.  Blue  Mts.: 
Schauinsland.  Jamieson  Valley  :  Forsyth,  '99.  Richmond  R.,  very 
scarce  :  Watts  (nr.  Ballina).  Parramatta  :  F.v.M.  (earliest  N.S.W. 
record) . 

319.  L.  conocladulum,  CM.,  Gen.  M.,  p.  80. 
Q.,  Gympie :  C  T.  Musson,  herb.  Bailey. 

L.  cy cadis.     See  No.  331. 

320.  L.  FLAVESCENS,  CM.,  Gen.  M.,  p.  80. 
**  Australia":  loc.  not  recorded. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  57 

Leucobryum  {continued).  \Leucohryacece. 

321.  L.  MiNUTUM,  CM.,  Gen.  M.,  p.  80. 
"Australia":  teste  CM.,  op.  cit.;  loc.  not  recorded. 

L.  sanctum  (Nees,  Dicramun),  Hpe.,  Linn.  xiii.  Given 
in  Hpe.  list  '80  for  Australia,  and  in  Melb.  Census 
for  Q.  with  a  ?  ["  Von  Nepal  bis  Malesise":  Broth. 
Bryales.  Occurrence  in  Australia  needs  confirma- 
tion.] 

L.  Schimpf.rl.  In  Melb.  Census  with  a  1  for  Vic.  and 
Q.  (Johnston  R.)  Arthrocormiis  /Schimjyeri,!).  et 
M.,  belongs  to  the  Ind.  Archipel.  TheQ.  specimen 
in  herb.  Melb.  is  prob.  Octohl.  albidum. 

322.  L.  SPEIROSTICHUM,  CM.,  Rev.  Bry.  '76,  and  Gen.  M., 

p.  80;  L.  Teysmannianum  forma  (fid.  Hampe). 

N.S.W.,  nr.  Sydney  :  Kayser  coll.  Blue  Mts.:  Whitelegge,  Watts, 
Mrs.  King  et  al.,  common  in  gullies.  Richmond  R.  and  Brunswick 
R.,  frequent:  Watts,  '96fF.  (c.fr.).  Clarence  R.,  Grafton:  herb. 
Melb.  Tweed  R.  '98,  National  Park,  Sydney  and  Kiama,  '99  : 
Forsyth.  Fruit  rare. 

323.  L.  SPINIDORSUM,  CM.,  Symb.,  No.  1. 

Tas.,  Grove  Creek,  Laus  Bay,  Mch.  '91  (c.fr.),  Port  Cygnet,  Sept. 
'89:  Weymouth. 

324.  L.  STRiCTiFOLiUM,  Broth.,  A.M.  '98,  No.  90. 

Q.,  Eumundi:  Bailey.— N.S.W.,  Blue  Mts.,  Lawson :  Betche, 
Aug.  '95;  Katoomba  Falls:  Watts,  '02.  Richmond  R.,  in  many 
places:  Watts,  '96ff.  (c.fr.  rare).  Cowan  Creek:  Hamilton,  '99. 
Morisset :  J.  L.  Boorman,  Oct.  '99  (herb.  Syd.). 

325.  L.  SUBCHLOROPHYLLOSUM,  Hpc.,  Linn.  '76. 
N.S.W.,  N.  Coast,  Mt.  Warning:  Guilfoyle,  '71. 

326.  L.  TURGiDULUxM,  CM.,   "M.  Schauinslandiani,"  CM. 
et  Broth.,  1900. 

N.S.W.,  Blue  Mts.,  Katoomba:  Schauinsland,  96-7. 


58  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Leucobryum  {continued).  [^Leucohryacece. 

327.  L.  vEsicuLOSUM,  CM.,  Gen.  M.,  p.  83. 

Q.,  Whelanian  Pools,  Palm  Camp  and  Harvey's  Camp,  Bellenden 
Ker :  herb.  Bailey.  Trinity  Bay  :  Mrs.  Gribble,  '94. 

L.  viride,  Mitt.  Given  in  Melb.  Census  for  (Vic.  and) 
Q.  (Daintree  R,),  but  confirmation  needed.  It 
appears  to  be  a  well-defined  species. 

328.  L.  Wattsii,  Broth,  in  sched. 

N.S.W.,  Richmond  R.  and  Brunswick  R.,  at  several  places  on 
ground  and  rocks  :  Watts  (ster.).  First  found  at  Killen's  Falls, 
Emigrant  Creek  :  Watts,  June  '96.  [A  specimen  coll.  by  Dallachy 
at  Rockingham  Bay,  Q.,  and  named  L.  sanctum  var.  in  Melb.  herb, 
belongs  here.] 

Leucophanes,  Brid. 

329.  L.  (Tropinotus)  apiculatum,  Hpe.     Name  in  CM., 
Gen.  M.,  p.  86. 

(^.,  "Coasts  of  tropical  Australia":  teste  CM.,  loc.  cit. 

330.  L.  (Tropinotus)  australe.  Broth,,  A.M.  '90,  Ko.  8. 
Q.,  Freshwater  Creek,  Trinity  Bay,  on  tree  :  Bailey  (pi.  fem.  ster). 
"Ker  Range":  teste  CM.,  Gen.  M. 

L.  octoblepharoides,  Brid.,  is  recorded  in  Melb.  Census 
for  Johnston  River  (Berthaud)  and  Daintree  R. 
(Pentzske).     Confirmation  needed. 

Octoblepharum,  Hedw. 

331.  O.  albidum  (L.,  Bryum)^  Hedw. 

Q.,  Fern  root  under  large  stone,  Coldstream  Mt.,  Miriam  Yale, 
Gladstone  and  Port  Curtis  :  C  T.  Musson,  Aug.  '87  (c.fr.).  On 
epiphytic  orchids,  trop.  Q. :  F.  M.  Bailey.  Mt.  Perry  :  J.  Keys. 
Rockingham  Bay,  '64.  Endeavour  R. :  W.  Persich,  '87.  Goode 
Isl.:  Powell. — S. A.,  Port  Darwin  :  Prof. Tate,  (l!!^ ame Leucobryum 
cy cadis  suggested  in  error  by  F.v.M.) 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  59 

Tribe  ix.  Calymperacese. 

C  A  L  Y  M  P  E  R  E  S,  Sw. 

C.  armatum,  Broth.     See  No.  338. 

332.  C.  Kennedyanum,  Hpe.,  Linn.  '76. 

Q.,  Rockingham  Bay:  E.  B.  Kennedy.  Goode  Isl. :  Powell,  '83. 
Helidon  :  Wild,  '88. 

333.  C.  LATiFOLiUxM,  Hpe.,  PI.  Preiss.  '46. 

W.  A.,  on  trunk  of  Macrozamia  Preissii  with  Fahronia  HamjJeana 
and  Calymperes  Moliiccense  (I)  nr.  Perth:  Preiss,  '39-40  (ster.). — 
N.S.W.:  teste  Mitt.  Cat.— Q.:  teste  Melb.  Census. 

C.  Mok'.ccense,  Schwgr.  Given  in  Hpe.,  PI.  Preiss.  for 
W.A.,  but  not  in  Jaeg.  Ad.,  Mitt.  Cat.,  etc. 

334.  C.  Motleyi,  Mitt.,  Dozy  et  Mlkb.,  Bryol.  Jav. 
Q.,  Cape  York:  W.  Micholitz,  Aug.  '97  (c.fr.). 

335.  C.  nigrescens,  Broth., Geh.  in  Broth.,  A.M. '99,  Ko.l48. 
Q.,  Bellenden  Ker  :  Mrs.  Gribble,  herb.  Melb. 

336.  C.  PANDUR.EFOLiuM,  Broth.,  A.M.  '98,  No.  100. 
Q.,  Cape  York  :  W.  Micholitz. 

C.  repens,  Harv.     See  No.  342. 

Syrrhopodon,  Schwgr. 

337.  S.   CLAVATUS,   Schw.     S.  fasiciilatus,  Hook,  et  Grev. 

(fid.   Mitt.   Cat.,  Thyridium).    Par.  Index  separates 
the  two. 
N.S.W.:  Sieber.— Q.,  Johnstone  R. :  Berthaud,  Dec.  '82. 
S.  fasciculatus.     See  No,  337. 

338.  S.   FiMBRiATULUS,   CM.,   Linn.   '75,  p.  557 ;    Sauerb., 
Jaeg.  A.d.  ii.;  S.fimhriatus,  CM.  at  first  in  Linn.  '71-3. 

Q.,  Brisbane  R. :  Mrs.  Dietrich  '64,  with  Isopterygium  austro- 
pusillum.  Pimpama,  Burpengary,  and  Ashgrove  :  C  J.  Wild. — 
N.S.W.,  Richmond  R.,   Hunter's  Scrub,  Tuckombil,  and  Three 


60  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Syrrhopodon  (contimted).  [Cali/mperacece. 

Mile  Scrub,  on  tree  and  on  Platy cerium,  '96ff.  :  Watts.  Also  on 
sandy  ledges  of  cliffs,  ^Yatson's  Bay,  nr.  Sydney  {Calymperes 
armatum,  Broth,  in  sched.  at  first) :  Watts. 

339.  S.  MAMMiLLOSus,  CM.,  Linn.  '75. 

Q.,  Nerang  Creek :  herb.  Bailey.  Pimpama :  Wild,  July  '87. 
[Broth.,  Bryales,  gives  New  Caledonia  only.] 

340.  S.  Nov^  Valesi^.  CM.,  Rev.  Bry.  '76;  Symb.  '98, 

No.  182. 
N.S.W.,  nr.    Sydney:    Kayser  coll.    Herriott,   herb.    ]\]elb.    '81. 
Richmond   R. :    Fawcett  '81,  Tintenbar   '95,   Bauerlen,   on   tree 
trunks,  Watts  '96ff.   (mostly  on  Platy  cerium).  Fruit  mature  in 
Oct.  and  Nov. 

(a)  f.  microcarpa. 
Q.,  Trinity  Bay:  Sayer.  '86. 

S.  {Thyridiur}i)  huinile.  Given  in  Melb.  Census  for 
Q.  (Johnson  R. :  Berthaud,  Dec.  '82).  Confirmation 
required. 

341.  S.  PLATYCERII,  Mitt.,  Fl.  Vit. 

N.S.W.,  on  Platyceriuin  alcicorne,  Port  Hacking:  Whitelegge, 
April  '85  (c.fr.).   Broth.,  Bryales,  gives  Lord  Howe  Lsland  only.] 

342.  S.  REPENS,  Harv.,  Lond.  J.  Bot.  '40;  Calymperes,  J a,eg, 

Ad.  i.;  Thyridium,  Jaeg.  Ad.  ii. 
Australia:  teste  Jaeg.   Ad. — Q. :   teste  F.v.  M.,  Frag.  Phyt.  xi. 
Suppl.,  and  Bail.  Syn.  Suppl.  '83.   [Broth.,  Brj^ales,  gives  Penang 
and  Ceylon  only.  Par.  Index,  "Asia."] 

343.  S.  SUBFASCICULATUS,  Hpe.,  Linn  '76,Codonoblepharum. 
Q. ;  teste  F.v.M.,  Frag.  Phyt.  xi.  Suppl.,  and  Bail.  Syn.  Suppl.  '83. 
[Jaeg.  Ad.  Suppl.  gives  "Australia,  insul.  Tuokuro  ?"  herb.  Melb. 
Broth.,  Bryales,  gives  "Tuo-kuro"  only.] 

344.  S.  UNDULATULUS,   Broth.,  Geh.  in  Broth.,  A.M.   '99, 

No.  147,  and  Bryales,  p.  369. 
Q.,  Bellenden  Ker :  Mrs.  Gribble,  herb.  Melb.  (ster.) 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  61 

S\RRHOPODON  {continued).  [Calymperacece. 

345.  S.  Wattsii,  Broth.,  A.M.  '99,  No.  143. 

N.S.W.,  Richmond  R.,  Wardell  Rd.,  on  saplings  and  scrub  vines  : 
Watts,  June  '98  and  Aug.  '01  (ster.).  Also  on  heath,  E.  Ballina 
'01  (ster.),  and  prob.  on  tree,  Fernleigh. 

Tribe  x.  Pottiacese. 

Admittedl}'-  a  very  difficult  family.  Jaeger  (Adumbr.)  separated 
the  Weisiacece  and  placed  them  first  of  the  Acrocarps.  Dr. 
Brotherus'  publications  hitherto  have  followed  Jaeger  in  this  and 
in  several  other  respects.  Dr.  C.  Mueller,  Mitten,  and  now  Dr. 
Brotherus  ('Bryales')  include  Weisia  and  allied  genera  under 
Pottiacece.  CM.,  in  Gen.  M.,  Jaeg.,  Paris  (Index)  et  al.  make 
Barhula  a  comprehensive  genus,  including  Tortula,  Tortella,  etc. 
Broth.  ('Bryales,  Part  i.')  forecasts  a  very  complete  analysis  of 
the  Pottiacece,  wndev  the  well-known  divisions  of  (i.)  Trichostomece 
(leaves  mostly  narrow,  elongate,  and  never  broader  in  the  upper 
half);  (ii.)  PottiecB  (leaves  mostly  broad,  oval  to  spathulate);  to 
which  he  adds  (iii.)  Encalyptece,  usually  regarded  as  a  separate 
family.  Unfortunately,  his  complete  account  of  the  Pottiacece  is 
not  to  hand.  To  his  outline  of  the  generic  distinctions  of  the 
subfamily,  TrichostoTnece,  we  acknowledge  our  great  indebtedness. 
Our  want  of  descriptions  of  some  of  the  species,  however,  and  the 
absence  of  material  for  examination  in  many  cases,  make  accuracy 
in  every  detail  scarcely  possible. 

i.  Trichostomeze. 
A  s  T  o  M  u  M,  Hpe. 

346.  A.BRACHYPELMA,  CM., Flora '88, P/iascitm(>S'ys^e6^m?w). 
S.A.,  Mt.  Lofty:  Tepper  '84,  herb.  Melb. 

347.  x\.  BRiSBANicuM  (CM.,  Linn.  '71-2,  Acaidon),  Broth. 

Br3^ales. 

Q.,  Brisbane  R. :  Mrs.  Dietrich,  '64. 


62  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

A  s  T  0  M  u  M  {continued).  [Pottiacece 

348.  A.  CYLiNDRicuM,  Tayl.,  Lond.  J.  Bot.  '46,  Phascum; 
Phascicm  {Astomum),  Mitt.,  Jour.  L.  Soc.  '60;  Ph. 
tetragonum  var.  cylindricum,  CM.,  Syn.  (cf.  Gen.  M., 
p.  23);  Tetrapternin,  J ?Le,g.;  Tetrapt.  australe,  Hpe.  (fid. 
Jaeg.) 

W.A.,  Swan  R.  :  Drummond.^N.S.W.,  Tarn  worth :  Musson. 
Narromine  :  Maiden,  '98.  Warrumbungle  Ranges :  Forsyth,  '99. 
Emu  Plains  :  Watts  Blue  Mts.  :  herb.  Melb.— Vic,  Yarra  R.  : 
F.v.M.  and  Reader.  Gippsland  :  F.v.M.  '55.— Tas.,  Cheshunt : 
Archer.  Kangaroo  Point,  Bay  View  Paddock,  etc.  :  Bastow, 
Weymouth,  A.  J.  Taylor,  McLeod. — Q.,  on  Southern  border  :  J. 
F.  Shirley. 

(a)  f.  laxior.   Yarra  R.,  Vic.  :  F.v.M. 

349.  A.  Nov.E  Valesli:,  Broth,  in  sched. 
N.S.W.,  in  park,  Emu  Plains:  Watts,  Oct,  '99. 

350.  A.  PERPUSiLLUM  (CM.,  Linn.  '71-2,  Phascum),  Hyrneno- 

stomum,  Par.  Index. 
Q.,  Brisbane  R.  :  Mrs.  Dietrich,  '64.  North  Pine  and  Humpybong  : 
Wild,  '88. 

351.  A.  SuLLiVANi,  CM.,  Flora  '88,  Phascum  (Sysfegium). 
N.S.W.,  Hunter  R.  :  Miss  Carter,  '82.— Vic,  nr.  Studley  Park, 
Melbourne :  Reader,  Sept.  '84.  First  found  by  D.  Sullivan  at 
Moysten  in  '82. 

352.  A.  Weymouthi,  CM.,  Symb.,  No.  80,  Phascum. 
Tas.,  Pedder's  Field :  Weymouth,  Sept.  '89. 

H  Y  M  E  N  0  S  T  O  M  U  M,  R.  Br. 

353.  H.  AMCENUM,  Geh.  in  sched. 

N.S.W.,  Ball's  Head  Bay,  on  logs  :  Whitelegge,  Aug.  '84  (c.fr.). 

354.  H.  ANGUSTATUM,  Broth.  in  sched. 

N.S. W.,  Three  Mile  Scrub,  nr.  Byron  Bay:  Watts,  Sept.  '97  (c.fr.). 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  63 

Hymenostomum  {continued).  [Poitiacece. 

355.  H.  OLiYACEUM,  CM.  in  sched. 

N.S.W.,  Clifton:  WhiteJegge,  Aug.  '91  (c.fr.).  Brunswick  R., 
Aug.  '98  (c.fr.)  and  Burwood  Park,  Sydney  :  Watts.  Nepean  R. 
and  National  Park,  Sept.  and  Dec.  '98  (c.fr.),  Warrumbungle 
Ranges,  Kiama,  Barber's  Creek, and  Valle}^  of  Waters  (Blue  Mts.): 
Forsyth.  Tweed  R.  :  Goldsmidt. 

356.  H.  PERTORTiLE,  CM.  in  sched.  :  Par.  Index,  Suppl.  i. 
Q.  :  teste  Par.,  loc.  cit. 

357.  H.  SUBOLIVACEUM,  CM.  in  sched. 

N.S.W.,  Burwood  Park,  Sydney:  Watts,  Mch.  '99.  First  collec- 
tor (?). 

358.  H.  SuLLivANi,  CM.  (teste  F.  M.  Reader). 

Vic,  on  soil,  nr.  Kew,  Melbourne  :  Reader,  July  '84.  Mt.  William 
and  Mt.  Ararat :  Sullivan,  '82-3.  Port  Phillip  :  C  French. 

359.  H.  TORTILE,  Br.  Eur. 

Tas.,  Ocean  Grove  Rd.  :  Bastow,  '88. — N.S.W.,  Parramatta : 
F.V.M.,  herb.  Melb. 

W  E  I  s  I  a,  Hedw. 

W.  cirrhata.     See  No.  224. 

W.  coarctata  in  Melb.  herb.  (Port  Phillip  :  C  French) 
needs  confirmation. 

W.  controversa.     See  No.  369. 

360.  W.  FLAviPES,  H.f.W.,  Fl.  N.Z. 
Tas.,  Vic,  N.S.W.,  Q.,  frequent. 

(a)  var.  longifolia,  Broth. 
N.S.W.,  Wilson's  Creek,  Richmond  R.  :  Watts,  Sept.  '97. 

361.  W.  GRAciLiSETA,  Broth.  in  Bail.  Bot.  Bull.,  Dec.  '91. 
Q.,  Mt.  Perry :  J.  Keys. 

W.  microcarpa,  H.f.W.     See  No.  224. 


64  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

W  E  I  s  I  A  (continued).  [Pottiacece. 

362.  W.   NUDA,  Mitt.  Cat.;  Pottia,  F.v.M.,  Frag.  Phyt.  xi., 

►Suppl. 
Q.,  Moreton  Bay:  F.v.M.  (c.fr.).   Hamilton:  Wild,  Aug.  '87. 

363.  W.  NUDiFLORA,  CM.  et  Hpe.,  Linn.  '53. 

S.A.,  Bugle  Range:  F.v.M. — Vic,  Yarra,  Dandenong  Ranges, 
Mt.  Abrupt,  etc.:  F.v.M.  Austral.  Alps,  "similar  Iocs,  with 
Phascwn  disriunijens,  viz.,  Livingstone  Creek,  Omeo  "  :  Stirling, 
'80-88.   Mt.  William  :  Sullivan. 

(a)  var.  major,  Hpe.,  Linn.  '56  (?). 
Yic,  with  type. 

364.  W.  PERLiNEARis,  CM.  in  Bail.  Bot.  Bull.  Dec.  '9L 
Q.,  Mt.  Perry:  J.  Keys. 

365.  W.  PiMPAM^,  CM.  in  Bail.  Syn.  Suppl.  3. 

Q.,  Pimpama  Creek  and  Enoggera  Waterworks :  C  J.  Wild, 
Aug.  '87. 

366.  W.  RiPARiA,  Hpe.,  Linn.  '60. 

S.A.,  on  banks  of  Gawler  R.,  Lofty  Range  :  F.v.M.  '50. 

367.  W.  TORTELLOiDES,  Broth.  in  sched. 

N.S.W.,  on  rocky  bank,  Uralba  Rd.,  Richmond  R.  :  Watts,  Oct. 
'98  (c.fr.). 

368.  W.  TRUNCATA,  CM.  in  sched.;    W.  controversa,  Hedw. 
(fid.  Bastow  in  MS.  list). 

N.S.W^,  on  rocks  in  a  gully,  Hurstville  :  Whitelegge,  Oct.  '84 
(cfr.). 

369.  W.  viRiDULA  (L.,  Bryum),  Brid.;  W.  controversa,'H.ed\v . 

Tas.,  Port  Sorell,  Long  Hill,  and  Cheshunt :  Archer.  Frequent 
throughout  the  Isl.  :  Oldfield,  Bastow,  We3^mouth  et  al. — N.S.W., 
Bulli  Pass,  Sydney,  and  Richmond  R. :  Watts. — Q.,  Mt.  Perry: 
J.  Keys. 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  65 

W  E  I  s  I  A  {continued).  [Pottiacece. 

370.  W.   Whiteleggeana,   CM. ;    W.   inflexa,   Hpe.    (fid. 

Bastow  in  MS.  list.) 

N.S.W.,  walls  of  old  dam,  Ball's  Head  Bay  and  Middle  Harbour : 
Whitelegge,  July- Aug.  '84  (c.fr.). 

G  Y  M  N  0  s  T  0  M  u  M,  Hedw. 

G.  hicolor,  Br.  et  Schpr.;  Barhula^  Lindb.,  Paris.  A 
barren  specimen  collected  by  Oldfield  was  so  named 
with  hesitancy  in  Fl.  Tas. 

371.  G.  calcareum,  Nees  et  Hsch. 

Tas.,  on  rocks,  N.W.  Bay  :  Oldfield.  Hobart  Waterworks  :  Wey- 
mouth.— N.S.W.,  on  a  native  kitchen  midden,  George's  B.,  Como, 
and  on  walls  below  Falls,  Mosman's  Bay  :  Whitelegge,  '85. — Q., 
Gladfield  :  C.  J.  Gwyther. 

(a)  var.  australe.  Broth.,  Geh.  in  Bev.  Bry.  '97. 
Vic,  Moysten  :  Sullivan,  '87.— N.S.W.,  Botany  Bay  :  Whitelegge. 
G.  inflexum.     See  No.  292. 

372.  G.  SCABRIDORSUM,  Broth.  in  sched. 

N.S.W.,  Richmond  R.,  on  damp  walls  and  roof  of  cave  in  sea 
cliff,  Skinner's  Head  :  Watts,  April  1900  (ster.),  Feb.  '02  (ster.). 

T  R  I  c  H  0  s  t  o  M  u  M,  Hedw. 

Tr.  calcicolum,  Hpe.     See  No.  401. 

373.  Tr.  cirrhatum,  Hpe.,  Icon. 

W.A.,  Swan  R.,  mixed  with  Barhula  calycina :  Preiss,  39. — 
[N.S.W. :  teste  Melb.  Census,  but  referring  to  a  Lord  Howe  IsL 
record.] 

Tr.  elongatum.     See  No.  188.  y*^G^!  ^CA^ 

Tr.  laxifolium.     See  No.  190.  ^^^"^O^^N^^^^^ 

Tr.fuscescens,     See  No.  398.  fa^  -^    -^e^^  ^^;^  <P' 

LIBRARY^) 
-^©•%.         ^/ 

•   W5^ 


66  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

T  R  I  c  H  0  s  T  0  M  u  M  {continued).  [Pottiacea^. 

374.  Tr.  leptotheca,  CM.,  Linn.  '67-<S. 

Vic,  Gippsland,  mixed  with  lliuidmm  snherectum :  F.v.M. 
Austral.  Alps :  Stirling. 

Tr.  pachyneuriis.,  pachyneuroides,  and  rcjlexid ens .     See 
Nos.  436-438. 

375.  Tr.  rubiginosum,  CM.,  Linn.  '67-8;  Tortula,  Mitt. 
Vic,  Yarra  R.:   F.v.M. 

[Note.  —  Melb.  Census  gives  Tr.  horhonicum  (i  brishanicum)  for 
Q.     Information  needed.] 

T  0  R  T  E  L  L  A  (CM.),  Limpr. 

376.  T.   ANGUSTiCAULis,  CM.  in  sched.,  Barbula.  (Not  in 

Gen.  M.). 

N.S.W.,  nr.  old  rifle  range,  Moore  Park,  Sydney:  Whitelegge, 
Sept.  '84  (c.fr.). 

377.  T.  ARISTATULA,  Brotli.  in  sched. 

N.S.W.,  in  bank  of  Alstonville  Cutting,  5Jm.  from  Ballina  : 
Watts,  April  '98ff.  (ster.). 

378.  T.  Knightii,  Mitt.,  Jour.  L.  Soc.  '60,  Tortida. 

Tas.,  not  uncommon  on  Mt.  Wellington  :  Bastow,  We3'mouth, 
Watts  et  al. — N.S.W.,  Longueville,  Lane  Cove  R.,  Sydney  : 
Whitelegge,  Sept.  '85  (c.fr.). 

379.  T.  NANO-TORTUOSA,  CM.,  Symb.,  No.  209. 
N.S.W.,  Ball's  Head  Bay  :   Whitelegge,  Sept.  '84  (c.fr.). 

380.  T.  Nov^  Valesi^,  Broth,  in  sched. 

N.S.W.,  Greenwich,  nr.  Sydney,  Jan.  '98  (c.fr.),  and  Wilson's 
Creek,  Richmond  R.,  Aug.  '98 :  Watts. 

381.  T.  PERHUMiLTS,  Broth.  in  sched. 

Q.,  Gladfield  :  Gwyther  (Bail.  Bot.  Bull.  A  pi.  '96). 


BY  REV.  W.   AVALTER  WATTS  AND  THOMAS  WHITELEGGE.  67 

[Pottiacece. 
T  R  I  Q  u  E  T  R  E  L  L  A,  CM.  (gen.iiov.  '97). 

382.  Tr.  filiformis,  CM.,  MSS.;  Par.  Index. 
S.A.,  Adelaide:  teste  Par.,  op.  cit. 

383.  Tr.  fragilis,  CM. 

Yic,  on  soil  in  woods,  Dimboolashire  :  Reader,  July  '96. 

384.  Tr.     papillata     (H.f.W.,    Fl.  N.Z.,    Lejnodontmm)  ; 

Didymodon,  Fl.  Tas.;  Leskea  rubricaidis,  Tayl. 
Tas.,  frequent :  Hooker,  Oldfield,  Archer,  Weymouth,  Bastow  et 
al. — Vic  ,  Adamson  (Melbourne),  F.v.M.  (Wangaratta),  C  French 
(Brighton),  Spence  (Ballarat). — S.A.,  Yorke  Peninsula:  Tepper. 
N.S.W.,  Tam worth  :  Musson.   Emu  Plains  and  Goulburn  :  Watts. 

385.  Tr.  Preissiana  (Hpe.,  Linn.  '60,  Zygodon),  C  M.;  Mitt. 

identifies  with  the  preceding. 
W.A.,  Mt.  Kaiserstuhl :  Preiss. 

386.  Tr.  Richardsle,  CM. 

NSW.  and  S.A.  :  teste  Par.  Index,  Suppl  i. 

387.  Tr.  scabra,  CM. 
Yic.  :  teste  Par.  Index,  Suppl.  i. 

388.  Tr.  triquetra,  CM.,  MSS.;  Par.  Index. 
"  Australia  "  :  teste  Par.,  op.  cit. 

Leptodontium,  Hpe. 

389.  L.  australe,  CM.,  Gen.  M.,  p.  409. 
Q.  :  teste  CM.,  loc.  cit. 

L.  papillatum.     See  No.  384. 

T  R  I  D  O  N  T  I  U  M,   Hook.fil. 

390.  T.  TASMANicuM,  Hook.f.,  Fl.  N.Z.;  Dicranum,  Hook.f., 
Icon.  PI.;  Dichodontium,  Schimp.;  Didymodon,  Mitt., 
Jour.  L.  Soc.  '60. 

Tas.,  on  water-washed  rocks,  Ben  Nevis  :  Gunn.  Oldfield.  Fossili- 
ferous  limestone.  Meander  R.  :  Archer.  Guy  Fawkes  Rivulet, 
Mt.  Wellington  :  Weymouth,  '97.  Newtown  Falls,  etc.  :  Bastow, 
'85-7. 


68  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

[Pottiacece. 
H  Y  o  p  H  I  L  A,  Brid. 

391.  H.  ANGUSTiFOLiA,  CM.,  Gen.  M.,  p.  395. 
Q.,  Toowoomba:  C.  J.  Wild,  Dec.  '88. 

392.  H.   INFLEXA  (Tayl.,  Gymnostomum);  Pottia,  Hpe.  et 
CM.,  Linn.  '53;    Weisia  al. 

W.A.,  Swan  R. :  Drummond.— S.A.,  Gawler  R.  :  F.v.M. '46.— 
Tas.  :  herb.  Melb.  and  CM.,  Gen.  M.— Yic,  Gippsland  :  F.v.M. 
'55. — N.S.W,  Parramatta:  Woolls. 

H.  Muelleri  occurs  in  Melb.  Census  for  Rockhampton, 
Q.     Confirmation  needed. 

393.  H.  puGiONiFOLiA,  CM.,  Gen.  M.,  p.  395. 

Q.,  Dawson  and  Burnett  Rs. :  Musson.  Brisbane  R. :  H.  Tryon. 
Ashgrove :  Wild,  '87. 

H.  squarrosa,  Broth.     Name  in  herb.  Bail,  for  Q. 

394.  H.  TENERA,  Broth. 

Q.,  Brisbane  R.,  nr.  St.  Lucia  Estate,  and  Petrie's  Quarries  :  H. 
Tryon  (Bail.  Bot.  Bull.,  Mch.  '91). 

D  I  D  Y  M  0  D  0  N,  Hedw. 

D.  papillatus.     See  No.  384. 

395.  D.  RUBELLUS  (Hoffm.,  Bryiim),  Br.Eur;  Barhula,  Mitt.; 

Trichostomum,  CM. 
Tas.,  on  dead  log,  Lauriston  Gully,  Kangaroo  Pt.  '91,  on  rocks, 
Hobart  Rivulet  '92,  Newtown  Rivulet  '93  :  Weymouth. 

B  A  R  B  u  L  A,  Hedw. 

396.  B.  ACROPHYLLA,  CM.,  Symb.,  No.  199. 

Vic,  Moysten.  Mt.  William:  Sullivan, '83  (N.S.W.:  teste  Par., 
prob.  in  error). 

397.  B.  AMa<:NA,  CM.,  Symb.,  No.  201. 

Vic,  Mt.  William,  with  B.  acrophylla  :  Sullivan,  '83. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  69 

B  A  R  B  u  L  A  {continued).  [Pottiacece. 

39S.  B.  AusTRALASi^(Hook.  et  Grev.,  Tricliostoimtm), Brid.; 
Tortula  Australasica,  Mitt.  Cat.;  T .  fuscescens,H.i.^ .', 
B.  rufiseta,  Tayl. :  CM.,  Syn.  i.,  618. 

W.A  ,  King  George's  Sound:  Menzies.  Swan  R. :  Drummond. 
Porongorup:  F.v.M.  '67. — Tas.,  west  side  of  Cataract,  Launces- 
ton :  Archer.  Nr.  Hobart :  Lyall,  Bastow  et  al. — S.  A.,  Rivoly 
Bay,  Glenelg  R.,  Gawlertown,  Barossa  Range,  Mt.  Gambier  : 
F.V.M.— Vic,  Gippsland  :  F.v.M. 

%  (a)  var.  fol.  laxioribus,  mentioned  by  Wils.  in  Fl.  Tas. 
as  having  been  coll.  by  Oldfield  in  Tas. 

399.  B.  AUSTRo-uNGUicuLATA,  CM.,  Gen.  M.,  p.  436. 
N.S.W.,  on  earth,  head  of  Double  Bay:  Whitelegge,  July  '84  (c.fr.). 

400.  B.  AUSTRO-UNGUICULATULA,  Watts;  B.  austro-ungnicu- 
lafa,  CM.,  Gen.  M.,  p.  440  (not  of  p.  436). 

Vic,  Alptrington,  June  '84.  Melbourne,  '92  :  Reader. 
B.  brachyphylla,  Hpe.     See  No.  408. 

401.  B.    CALCicoLA    (Hpe.    Icon.,    2'richostomum),    Broth. 

Bryales;  Tortula,  Mitt.  Cat. 
W.A.,  on  calcareous  rocks,  nr.  Fremantle  :  Preiss,  Aug.   '39. — 
Vic  :  teste  Par.  Index,  prob.  in  error. 

402.  B.  CALYCiNA,  Schwgr. 

Throughout  Australia  and  Tas.  Apparently  first  found  in  Aus- 
tralia by  La  Billardiere. — W.A.,  Swan  R. :  Preiss,  '39.  Upper 
SwanR. :  Miss  Se well.  St.  George's  Sound  :  Cunningham. — S.A., 
Barossa  Range,  etc.:  F.v.M. — Vic,  Austral.  Alps:  Stirling, 
F.v.M.  et  al.  Dimboola:  Reader. — N.S.W.,  Tumberumba :  W. 
Bull.  Richmond  R. :  Watts.  National  Park  and  Shoalhaven  R. : 
Forsyth.— Q.:  Miss  Gore  et  al. — Tas.,  common.  King  Isl, :  A. 
Neate. 

(a)  var.  brevicaulis  (Hook.),  Hpe. 

Vic,  Sealer's  Cove:  F.v.M. 


70  CENSUS    MUSCORUM    AUSTKALIENSIUM, 

B  A  K  B  u  L  A    {continued).  [Pottiacect. 

1  (b)  var.  ATTENUATA  ("var.  fol.  longioribus  acutioribus 
attenuatis,"  H.f.W.,  Fl.  Tas.). 

Tas.  :  Gunn. 

403.  B.  CHLOROTRICHA  (Broth.,  Geh.  in  Broth.   A.M.  '93, 
No.  29,  Tortula),  Par.,  Broth.  Bryales. 

N.S.  W.,  Tilba  :  Reader.  Nepeaii  R.,  Emu  Plains  :  Watts,  Oct.  '99 
(ster.).-  Vic,  banks  of  Yarra  R.:  Watts,  '02. 

404.  B.  CHRYSOCH.ETE,  CM.,  Symb.,  No.  197. 
Vic,  on  sandy  soil,  nr.  Dimboola  :  Reader,  July  '90. 

405.  B.  CHRYSOPUS,  CM.,  Symb.,  No.  200. 

Tas.,   Blackman's    Bay,    road    from    Durally   to    Bream    Creek : 
W^eymouth,  Oct.  '89. 

B.  Commersonii,  Mitt.  Given  in  Frag.  Phyt.  xi.,  Suppl. 
for  N.S.W.  and  Q.     Particulars  needed. 

B.  Crmvfordi,  Par.     See  No.  450. 

B.  crispifolia  (Mitt.,  2'ortula),  Jaeg.  Ad.  A  N.Z.  sp. 
given  in  Melb.  Census  for  Tas.  Mt.  Dromedary  : 
Bastow,  '86.     Confirmation  needed. 

406.  B.  CYLiNDRANGiA,  CM.,  Symb.,  No.  198. 

Vic,  Ballarat :  Way,  herb.  Melb.  '81.  Daylesford  :  Wallace,  herb. 
Melb.  '77. 

407.  B.  GEMiNATA,  CM.,  Gen.  M.,  p.  454. 
Vic,  Mt.  William  :  Sullivan,  '83. 

408.  B.   Hampeana,  Par.    Index ;  B.   hracJiyphylla,   Hpe., 
Linn.  '60. 

Vic,  Gippsland:  F.v.M.  '55.— Tas.:  teste  F.v.M.,Frag.  Phyt.  xi., 
Suppl.  {B.  brachy2)fiylla). 

B.  inclinata,  Schwgr. ;  Tortella,  Limpr.  Given  for 
"  Australia  "  in  Par.  Index,  Suppl.  Confirmation 
needed. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  71 

B  A  R  15  u  L  A  {continued).  [Pottiacece. 

409.  B.  LuEHMANNi,  Broth.,  Geh.  in  Broth.  A. M.  '95,  No.  68. 
Vic,  Loutit  Bay:  Luehmanri,  '91. 

B.  micro2:)hylla.     Given  by  Stirling  for  Austral.  Alps. 
Information  needed. 

410.  B.  NANOCAULis,  CM.,  Synib.,  No.  195. 
Tas.,  Creek,  Eagle  Hawk:  Weymouth,  Oct.  '89. 

411.  B.  Parramattana  (Mitt.  Cat.,  Tortula). 
N.S  W.,  Parramatta  :  Woolls. 

412.  B.  PSEUDO-piLiFERA,  Hpe.  et  CM.,  Linn.  '53;  Tortula 
pungens  and  luteola  (fid.  Mitt.  Cat.). 

Vic,  Mt.  Ararat:  Sullivan.  Austral.  Alps:  Stirling.  Prob.  first 
found  by  F.v.M.  in  '55.— Tas.,  Wattle  Hill,  Sorell :  Weymouth, 
Jan.  '91.— N.S. W.,  Blue  Mts.  :  herb.  Melb.  Barber's  Creek: 
Forsyth,  Sept.  '99. 

B.  rubella,  Hoffm.     See  No.  395. 

413.  B.  SPEIROSTEGA,  CM.,  Symb.,  No.  196. 
N.S.W.,  Lawson,  Blue  Mts.:  Whitelegge,  June  '84. 

414.  B.  suBCALYCiNA,  CM.,  Linn.  '67-8. 

Q.,  Brisbane  R. :  Mrs.  Dietrich,  Wild  et  al. — Vic,  Dimboola : 
Reader, Sept.  '94. — N.S.  W.,  Richmond  and  Clarence  Rs.,  common : 
Watts, '96ff.  Nr.  Sydney:  Forsyth  (Lane  Cove),Cheel(Rookwood), 
Watts  (Waverley).  Emu  Plains:  Watts.  Tentertield:  Bauerlen, 
Aug.  '01  (herb.  Tech.  Mus.  Syd.). 

415.  B.  suBSPiRALis,  Hpe.,  in  F.v.M.,  Aust.  M.  pi.  2. 
Vic,  Gippsland  :  F.v.M. 

416.  B.  SUBTORQUATA,  CM.  et  Hpe.,  Linn.  '53. 

S.A.,  Mt.  Gambier:  F.v.M.  (Austr.  M.  pi.  3).— Tas.,  on  sandy 
bank.  Pirate's  Bay,  E.  Coast:  Weymouth,  '89.— Vic  :  teste  Melb. 
Census. 


72  CENSUS    MUSCORUM  AUSTRALIENSIUM, 

B  A  R  B  u  L  A  {continued).  [PoUiacece. 

417.  B.  TORQUATA,  Tayl.,  Lond.  J.  Bot.  '46. 

W.A.,  Swan  R. :  Drummond. — Tas.,  Cheshunt:  Archer. — S.A., 
Mt.  Gambier,  Glenelg  R.,  etc.:  F.v.M.— N.S.W.,  Parramatta : 
Woolls.  Tomingley,  Daffodil  Creek:  Maiden,  Sept.  '98.— Vic, 
Mallee  Country,  Wimmera  '96,  nr.  Doncaster  '83 :  Reader. 
Gippsland :  F.v.M.  and  Lucas.  Mt.  William  :  Sullivan  (f.  seta 
breviore  et  crassiore). 

418.  B.   WiLDii,  Broth.,  A.M.   '90,   No.   9;   Trichostomum 

{A7iacali/pta),  CM.,  Gen.  M. 
Q.,  Highfields,  1500',  on  calcar.  ground:  C.  J.  Wild,  Dec.  '88. 

ii.  PoTTiE.i:. 
A  c  A  u  L  0  N,  CM.  {S2}hceran(/iuni,  Schimp.). 

419.  A.  APicuLATUM  (H.f.W.,  Fl.  N.Z.,  Phascum),  Jaeg.  M. 

Cleist.;  Sphccrangium,  Jaeg.  Ad. 
Tas.,  on  hard  track,  nr.  Brighton:  Weymouth,  No.  1130. 

(a)  var.  turgidum,  Mitt.,  Jour.  L.  Soc.  '60,  and  Fl.  Tas.; 
A.  turgidum,  Mitt.  Cat. 
Tas.,  Cheshunt:  Archer.   Brooke's  Field:  Bastow,  July  ^^Q. 

420.  A.  AUSTRO-MUTicuM,  Geh,  in  sched. 

N.S.W.,  on  clay,  Parramatta  Park:   Whitelegge,  Aug.  '85  (c.fr.). 
A.  brisbanicu7n,  CM.     See  No.  347. 

421.  A.  iNTEGRiFOLiuM,  CM.,  Bot.  Zeit.  '55. 
Vic,  on  bare  ground,  Yarra  R.,  nr.  Melbourne:  F.v.M. 

422.  A.  SuLLivANi,  CM.,  Flora  '88. 

Vic,  Mt.  Ararat:  Sullivan,  '82.— N.S.W.,  Centennial  Park, 
Sydney:  Aug.  '99  (c.fr.)  and  Nowra,  Badgery's  Crossing,  Sept. 
'99  (c.fr.) :  Forsyth.  Emu  Plains,  on  dry  bank  :  Watts,  Oct.  '99 
(c.fr.). 

A.  turgidum.     See  No.  419,  var. 


BY  REV.   W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  73 

[^Pottiacece. 
P  H  A  s  c  u  M,  L.  {EuphasGum,  CM.). 

Ph.  disrumpens.     See  No.  428. 

423.  Ph.  Drummondii,  Wils.,  Lond.  J.  Bot.  '48. 
W.A.,  Swan  R. :  Drummond. — Vic,  Melbourne:  Adamson. 

424.  Ph.  loreuMj  Wils.,  in  herb.  Brit.  Mus. 
W.A.,  Swan  R.:  teste  Par.  Index,  Suppl.  i. 

425.  Ph.  molle,  CM. 

Vic,  sandy  desert,  Dimboolashire  :  Reader,  Aug.  '97. 

Ph,  tetragonum  var.  cylindricum.  See  No.  348.  Melb. 
Census  gives  Ph.  tetragonum  itself  forTas.,  in  error. 

426.  Ph.  tetrapteroides,  CM.,  Symb.,  No.  82. 
Vic,  on  damp  soil,  Dimboola  :  Reader,  Oct.  '93. 

[Note. — For  Ph.  hrachypelma,  cglindricum,  perpusillum,  SulH- 
vani,  and  Weymouthi,  see  Nos.  346,  348,  350,  351,  352.] 

Tetrapterum,  Hpe.  In  Gen.  M.,  CM.  thinks  that 
Hampe's  T.  australe  (usually  made  a  syn.  of  2\ 
cylindricum,  see  No.  348)  is  probably  Phas.  tetra^)- 
teroides.  Taylor's  Ph.  cylindricum  he  places  under 
his  Section  Systegium  (see  Astomum). 

Pleurophascum,  Lindb.  (The  proper  position  of  this 
remarkable  moss,  usually  placed  here,  is  prob.  among  the 
Pleurocarps.) 

427.  P.  GRANDiGLOBUM,  Lindb.,  Lond.  Jour.  Bot.  '75,  and 

Rev.  Bry.  '76. 
Tas.,  Alps,  Picton  R. :  R.  Johnston  and  Rev.  J.  Bufton.     Mac- 
quarie  Harbour,  W.  Coast:  T.  B.   Moore,  Sept.-Oct.  '96  (herb. 
Weymouth). 

ScHIZOPHASCUM,   CM. 

428.  S.  disrumpens,  CM.,  Flora  '88  (Sect,  of  Phascum). 
Vic,  Moysten  :  Sullivan,  '82.  Austral.  Alps,  Livingston  Creek, 
Omeo  :  Stirling  (ascends  to  3000'). 


74  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

[Pottiacece. 
P  0  T  T  I  A,  Ehrh. 

429.  P.  BRACHYODUS,  Hpe.,  Linn.  '60,  Anacalypla;  Desmato- 

don  adu^tiis,  Mitt. 
Vic,  Arthur's  Seat  and  Yarra  R.  :  F.v.M.   '54.   Austral.  Alps  : 
Stirling,  '80-90.— N.S.W.,  Blue  Mts.  :  herb.  Melb. 

430.  P.  BRACHYPHYLLA,  Hpe.,  in  herb.  Melb.;  CM.,  Gen.  M., 

p.  389. 
Vic,  "on  Quercus  CUffortiana^^ :  teste  CM.,  loc.  cit.     In  sandy 
soil,  Dimboolashire  :  Reader,  Sept.  '98  and  July  1900. 

431.  P.   BREViCAULis  (Tayl.,  Lond.  J.  Bot.  '46,  Gymnosto- 
mum),  CM.,  Syn. 

W.A.,  Swan  R. :  Drummond. — Vic,  Queenscliff:  F.v.M.  '67. 

432.  P.  c^spiTULOSA,  Hpe.  et  CM.,  Linn.  '53,  Anacalypta: 

Pottia,  CM.,  Gen.  M. 
S.A.,  Mt.  Lofty  :  F.v.M.— Vic,  Yarra  E. :  F.v  M.   On  clayey  soil, 
Salt  Lake,  Lothiel,  Wimmera  :  Reader,  96. 

433.  P.  Readeri,  cm.,  Symb.,  No.  212. 
Vic,  Alptrington  :  Reader,  June  '84  (c.fr.). 

434.  P.  RECURVIMUCRONATA,  CM.,  Gen.  M.,  p.  388. 
Vic:  teste  CM.,  loc.  cit.  (loc.  not  specified). 

P.  Stackhousii.     Given  in  Melb.   Census  for  N.S.W. 
in  error.     The  specimen  is  not  a  Pottia  (W.W.W.). 

435.  P.  TASMANiCA,  Broth.,  A.M.  '95,  No.  69. 
Tas.,  Jordan  R.,  nr.  Brighton:  Weymouth. 

D  E  s  M  A  T  o  D  o  N,  Brid. 

D.  adustus.     See  No.  429. 
D.  nervosus.     See  No.  442. 

436.  D.  PACHYNEUROiDES,  CM.,  Gen.  M.,  p.  414  (Sect,  of 

T7'ichostomum). 
N.S.  VV.,  on  walls  of  Domain,  S3'^dney :  Whitelegge,  Aug.  '85  (c.fr,). 
Cook  Park,  Sydney,  July  '84  :  idem. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  75 

D  E  s  M  A  T  o  D  o  N  {coiifimied).  \Pottiacea\ 

437.  D.  PACHYNEURUS,  CM.,  Gen.  M.,  p.  414  (Sect,  of 
Trichostomiim). 

N.S.W. :  teste  CM.,  loc.  cit. 

438.  D.  REFLEXTDENS  (Hpe.,  Linn.  '60,  Trichostomum) ,  Jaeg. 
Ad. 

Vic,  YaiTca  R.  and  Avon  R. :  F.v.M.  '54. 

439.  D.   Weymouthi,    CM.,    Gen.   M.    p.    414    (Sect,   of 

Trichostomum). 
Tas.  :  teste  CM.,  loc.  cit. 

T  0  R  T  u  L  A,  Hedw.  (emend.  Lindb.). 
T.  adusta.     See  No.  429. 

440.  T.  ANDROGYNA,  CM.,  Sjmb.,  No.  191,  Barhula. 
Vic,  on  sandy  soil,  Dimboola :  Reader,  July  '90  (c.fr.)  and  '94. 

T.  antarctica^  Hpe.  Mitt.  Cat.  gives  as  syn.  of  T. 
pri7iceps.  CM.,  Gen.  M.,  gives  as  separate  sp.  for 
Falkland  Isls.  only. 

441.  T.  ARiSTATULA,  CM.,  in  Jaeg.  Ad.  &  Gen.  M.,  Barhida; 
Encalijjda  aristata,  Hpe.,  Linn.  '71-3;  Pyramitrium, 
Mitt.  Cat.  [cristatuin  in  error). 

N.S.W.,  Blue  Mts,:   Mrs.   Calvert.— Vic :   teste  Melb.  Census, 
prob.  in  error. 

442.  T.    atrovirens    (Sm.,  Eng.   Bot.,    Grimmia),   Lindb.; 

Desmatodo7i  nervosus,  Br.  Eur.  ;    Trichostomum  con- 

volutu7n,  Brid.  (fid.  Jaeg  ). 
Tas.,  frequent:  Hooker,  Archer,  Bastow  et  al. — Vic,  Avon  R. 
and  Yarra  R.:  F.v.M.  Nr.  Melb.:  Adamson,  also  Reader,  '83. 
Austral.  Alps :  Stirling.  Salt  Lake,  Lothiel,  July  '98,  Dimboola, 
1900:  Reader.— Q.,  Gladfield  :  Gwyther.— N.S.W.,  Shoalliaven 
R. :  Forsyth,  Sept.  '99.  Bank  of  Nepean  R.,  Emu  Plains:  Watts, 
Oct.  '99. 

(a)  f.  nervo  excedente. 

Vic,  in  wet  sandy  places,  Dimboolashire  :  Reader,  Aug.  1900. 


76  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

T  o  E  T  u  L  A  (continued).  [Pottiacece. 

443.  T.  AUSTRO-MURALis,  CM.,  Symb.,  No.  207,  Barhula. 
N.S.W.,  on  walls  of  Australian  Museum,  Sydney,  and  at  Ball's 
Head  Bay  and  Rand  wick  Rd.:  Whitelegge, '84. — Q. :  teste  Melb. 
Census. — Vic,   on   stone   walls,    Melbourne:    Reader,    Feb.    '83. 
Pyrenees:  Sullivan,  '87. 

444.  T.  AUSTRO-RURALis,  CM.,  Symb.,  No.  189,  Barhula. 
Q.,  York  Peninsula:  Tepper,  '79. 

445.  T.  Baileyi,  Broth.  A.M.  '90,  No.  10.' 
S.A.,  Adelaide:  Bailey  '88,  on  bark  of  tree. 

446.  T.  BRACHYTRICHA,  CM.,  Symb.,  No.  194,  Barhula. 
Tas.,  Creek,  Eagle  Hawk:  Weymouth,  Oct.  '89. 

447.  T.  BREviPiLA,  Hpe.  et  CM.,  Linn.  '53,  Barhula. 
Vic,  Sealer's  Cove:  F.v.M.— S.A.,  Mt.  Gambier:  F.v.M. 

448.  T.  BREViSETACEA,  Hpe.  et  CM.,  Linn.  '53  (Barhula 

hreviseta);  F.v.M.,  Austr.  M.  pi.  4.     Name  corrected 

by  Jaeg.  and  CM. 
S.A.,     Mt.     Gambier:     F.v.M. — Vic,    Austral.    Alps:     Stirling, 
'80-88.  Sealer's  Cove:  F.v.M.  '54. 

T.  chlorotricha.     See  No.  403. 

449.  T.  CRASSINERVIA,  Tayl.,  Linn. ' 53,  Desmatodon;  Barhula 

Sidlivaniana,  CM.  in  sched. 
W.A  ,  Swan  R. :  Drummond. — S.A  ,  Gawler  town:  F.v.M.- — Vic, 
Yarra  R.  and  Avon  R.:  F.v.M.   Austral.  Alps:  Stirling,  "80-88. 
Coburg:  Reader,  '88. 

(a)  var.  longiseta,  CM. 
Vic,  on  stones  nr.  Alptrington:  Reader,  June  '84. 

450.  T.  Crawfordi  (Par.  Index,  Suppl.  i.,  Barhula),  Watts; 

Barhula  asperifolia,  CM.,  Symb.,  No.  188. 

N.S.W.,  Macleay  R.:  A.  R.  Crawford,  '84. 


\ 


BY  REV.   W.   WALTER  WATTS  AND  THOMAS  WHITELEGGE.  77 

T  0  R  T  u  L  A  {conthiued).  [Fottiacece. 

451.  T.  CRISPATA,  Hpe.,  Linn.  '76,  Barhula;  Streptojoogon, 
Jaeg.;  *S'.  mnioides,  Schwgr.,  fid.  Wilson. 

Vic,  Mt.  Macedon,  on  branches  mixed  with  Orthotrichwm  Interim  : 
herb.  Melb.  (Lucas  ?). — Tas.,  on  willow,  Johnny's  Creek,  nr. 
Hamilton  Rd.  '93,  Hobart  Rivulet :  Weymouth.  Cheshunt,  Falls 
Run :  Archer.  Mt.  Wellington :  Mossman  (as  S.  mnioides). 
Cascades  :  Bastow. 

452.  T.  CUSPIDATA,  H.f.W.,  Fl.  Tas.;  7\  princejis,  e.p.  (fid. 

Mitt.). 
Tas.,  moist  earth,  woods,  Mt.  Wellington  :  Oldfield.  Cheshunt : 
Archer.  Kangaroo  Bottom  :  J.  D.  Hooker.  King  Isl. :  A.  Neate. 

453.  T.  ELAPHROTRICHA,  CM.,  Symb.,  No.  205,  Barhida. 
Tas,  wall  of  R.C.  Cemetery,  Brisbane  St.,  Hobart:  Weymouth, 
Sept.-Oct.  '90  (c.fr.). 

(a)  var.  incana,  CM.,  ibid. 
Tas.,  top  of  stone  wall,  Reservoir,  Hobart :  Weymouth,  Oct.  '90. 

454.  T.  fleximarginata,  Hpe.  et  CM.,  Linn.  '53;  F.v.M., 

Austr.  M.  pi.  6:  Barhida. 
Vic,  Gippsland  :  F.v.M. 

T.  hystrichosa.  Given  in  Melb.  Census  for  S.  A.  Yorke 

Peninsula  :  Tepper.     Confirmation  needed. 
T.  involucrata.     See  No.  478,  var. 
T.  Icevipila.     See  No.  465. 
T.  lamellosa.     See  No.  476. 

455.  T.  Latrobeana,    CM.,   Bot.   Zeit.    '64,  Barhida;    B. 

Preissiana,  Hpe.  (not  of  CM.). 
Vic,  Latrobe  and  Axe  Rs. :  F.v.M. -^SS.   Austral.  Alps:  StirHng. 
— Tas.:  Archer. 

T.  luteola.     See  No.  468. 

456.  T.  Melbourniana,  CM.,  Symb.,  No.  206,  Barhula. 
Vic,  on  stone  walls,  Fitzroy,  Melb.:  Reader,  Feb.  '83  (c.fr.). 

T.  mnioides.     See  No.  451. 


78  CENSUS    MUSCORUM    AUSTRALIEXSIUM, 

T  o  R  T  u  L  A  {continued).  [Pottiacece. 

457.  T.  MOLLissiMA,  Broth,  in  sched. 

N.S.W.,  on  ground,  "Yodalla,"  Emu  Plains:  Watts,  Oct.  '99. 
T.  Muelleri.     See  No.  46G. 

458.  T.  MURALis  (L),  Hedw.;  Barhula,  Timm. 

Tas.,  stone  wall,  Garden  Crescent  Reservoir,  '90 :  We^^mouth. 
Newtown  Greek  :  AVatts,  '95. — N.S.W.,  common  about  Sydney 
on  masonry:  Whitelegge,  Watts.  (Good  fruit  Sept.-Dec).  [CM. 
not  recognising  this  Europ.  sp.  in  Austr.,  it  is  prob.  his  B. 
austro-muralisA 

459.  T.  murina,  CM.,  Symb.,  No.  192,  Barhula. 
N.S.W..  Lower  Hunter  R. :   Miss  Carter,  '82. 

460.  T.  xano-subulata,  CM.  in  sched.,  Barhula. 

N.S.W.,  on  wall  of  culvert,  Randwick  Rd.,  nr.  Syd. :  Whitelegge, 
Sept.-Oct.  '84  (c.fr.). 

461.  T.  cedineura,  CM.,  Barhula. 

Vic,  on  sandy  soil,  Coker  Dam,  Wimmera  :  Reader,  July  '96. 

462.  T.  pandur.efolia,  Hpe.  et  CM.,  Linn.  '53,  Barhula; 

F.v.M.,  Austr.  M.  pi.  5. 

Vic,  Gippsland  :  F.v.M.— N.S.W.,  Walcha  :  E.  Betche,  Dec. '98. 
Stewart's  Brook :  Maiden,  Aug.  '99.  Warrumbungle  Ranges : 
Forsyth,  Oct.  '99.— Tas.:  teste  CM.,  in  Gen.  M. 

463.  T.  PAPiLLOSA,  Wils.,  Lond.  J.  Bot.  '45. 

A  wide-spread  sp.  first  found  in  England. — Vic,  Sealer's  Cove  : 
F.v.M.  Austral.  Alps:  Stirling, — Tas.,  on  trees  and  in  swampy 
places  :  Gunn.  Mt.  Rumney  :  Bastow,  '85. — N.S.W.,  Parramatta  : 
F.v.M. 

464.  T.  PERBREViFOLiA,  CM.  (Si/ntrickia). 

Vic.j  on  sandy  soil  in  woods,  Dimboolashire  :  Reader,  July  '96. 


BY  REV.  W.   WALTER  WATTS  AND  THOMAS   WHITELEGGE.  79 

T  0  R  T  u  L  A  (continued).  [Fottiace(fi. 

465.  T.  Preissiana,  CM.,  Syn.,  ^a?-6?6^rt;   BJcevipila^Hpe.; 

B.  ruralis  (ace.  to  some  detenus.). 
W.A.,  on  deca3dng  branches,  isl.  of  Rottnest,  Swan  R. :  Preiss, 
Aug.  '39. — S.A. :  teste  Melb.  Census  (Bugle  Ranges  :  F.v.M.)  and 
Par.  Index  (sub  B.  ruralis). — Vic,  Latrobe  R. :  F.v.M.  '55. 

466.  T.  PRiNCEPS,  De  Not.;  B.  Muelleri,  Br.  et  Schimp. 
CosMOPOL. — Tas.,  Port  Sorell :  Archer.  Nr.  Hobart  :  J.  D. 
Hooker,  Mt.  Rumney,  etc.,  frequent. — Vic. :  F.v.M  et  al.  Studley 
Park  :  Reader,  '83.  Mt.  Ararat :  Sullivan,  '87.— KS.W.,  Jenolan 
Caves  :  Maiden,  Aug.  '98,  Blakely,  '99.  Jerarie  Creek  :  Forsyth, 
'99.  Tenterfield:  Bauerlen,  Aug.  '01.  Blue  Mts.  :  herb.  Melb. 
(an  early  specimen\  —  S.A.,  Clarendon:  Tepper. 

467.  T.  propinqua,  CM.,  Symb.,  No.  190,  Barhula. 

Vic,  Moysten  :  Sullivan,  Oct.  '83.  Austral.  Alps  :  Stirling.  Mt. 
Ararat :   Sullivan.   Port  Phillip  :  C  French. 

468.  T.  PUNGENS,  H.f.W.,  Fl.  Tas.;  T.  luteola,  Mitt. 

Tas.,  Cheshunt :  Archer.  Common  on  rocks.  Grass  Tree  Hill,  etc. : 
J.  D.  Hooker,  Gunn,  Oldfield  et  al. — Vic,  Darebin  Creek  and 
Gippsland  :  F.v.M.   (An  B.  pseudo-pilifera  ?). 

469.  T.  Readeri,  CM.,  Symb.,  No.  193,  Barhula. 
Vic,  on  ground,  nr.  Melton  :  Reader,  Aug.  '86  (c.fr.). 

470.  T.  recurvata.  Hook.,  M.  Exot.,  Barhula. 

Tas.,  on  wet  ground,  Mt.  Dromedary:  Oldfield  (teste  Fl.  Tas.). 
[CM.,  in  Gen.  M.,  gives  Desmatodon  recni'vatus,  Hook.,  for  the 
Cape  of  Good  Hope  only.] 

471.  T.  robusta.  Hook,  et  Grev.,  Barhula-. 

Vic,  Cobberas  Mts.:  F.v.M.  (CM.,  in  Gen.  M.,  gives  Patagonia 
and  the  Antarctic  Isls.  only.) 

T.  rotundata,  Geh.,   iued.    (a  doubtful   sp.,  prob.    1\ 
pandurwjolia). 
N.S.W.,  on  earth.  Port  Hacking :  Whitelegge,  April  '85. 


80  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

T  0  R  T  u  L  A  (contimied).  [Pottiacece. 

472.  T.  RUBELLA,  H.f.W.,  Fl.  Tas.;  B.  2:>rinceps,  e.p.  (fid. 

Mitt,  in  Jour.  L.  Soc.  '60). 
Tas.,  Glen  Leith :  J.  D.  Hooker,  Gunn  et  al.   [CM.,  in  Gen.  M., 
gives  N.Z.  only.  The  Tas.  Iocs,  may  belong  to  T.  princeps  or  T. 

papiUosa.] 

473.  T.  RUBRA,  Mitt.,  Hdbk.  N.Z.  Fl.;  Desmatodon,  F.v.M., 

Frag.  Phyt.  xi.,  Suppl. 
Vic,  Austral.   Alps:  F.v.M.,  Stirling  et  al.    "Common  on   the 
Mitta  Mitta,  Mitchell  and  Tambo  River  sources,  between  2000' 
and  6000'"  (Stirling).— N.S.W.,  Mt.  Kosciusko:  Findlay,  herb. 
Melb. 

T.  ruralis.     See  No.  465. 

T.   scabrifolium.     Given    in    Melb.   Census    for    Yic. 

Information  needed. 
T.  schizopogoniacea.     See  No.  475. 

474.  T.  SERRULATA  (Hook.  et  Grev.,  Brewst.  Jour.,  Barhula) 

Mitt.;  T.  rubra  (fid.  Mitt.). 
"  East  Australia  "  :  teste  Par.  Index,  Suppl.  i.  [An  Antarctic  and 
N.Z.  sp. — CM.,  in  Gen.  M.,  gives  Fuegia  only.] 

475.  T.  STREPTOPOGONiACEA,  CM.,  Symb.,  No.  196,  Barhida; 

B.  schizopogoniacea,  CM.  in  sched. 
N.S. W.,  Moss  Vale:  Whitelegge,  Nov.  '84  (c.fr.).     Hinton:  Watts. 

476.  T.  SuLLiVANiANA,  CM.,  Symb.,  No.  208,  Barhula;  B. 
lamellosa,  CM.  in  sched.  (teste  CM,  Gen.  M.) 

Vic,  Moysten  :  Sullivan,  '83,  Austral.  Alps  :  Stirling.  Wimmera 
(teste  F.  M.  Reader). — N.S.  W.,  Warriimbungle  Ranges  :  Forsj^th, 
Oct.  '99  (c.fr.). 

T.  SidHvaniana,  CM.  in  sched.  (Barbiola).  See  No.  449. 

477.  T.  TASMANICA,  Hpe.,  Linn.  '52,  Barhula;  ?  B.  pungens, 

H.f.W. 
Tas.  :  Stuart. — "  E.    Australia  "  :   teste   Par.   Index   (a  doubtful 
record.   Melb.  Census  gives  Tas.  only).  A  specimen  from  Gippsland 
(F.v.M.)  is  probably  T.  j)seiido-pilifera. 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE.  Si 

T  o  R  T  u  L  A  {continued).  [Pottiacece. 

478.  T.  VESICULOSA,  CM.,  Symb.,  No.  185,  Barbula. 
Vic,  Mt.  Ararat,  nr.  Moysten  :  Sullivan,  Oct.  83.    Austral.  Alps: 

Stirling. 

(a)  var.   involucrata,  CM,  loc.  cit.;  B.  involucrata, 
CM.,  in  herb.  Melb. 
Vic,  Grampians  :  herb.  Melb.  '81.  Mt.  Ararat:  Sullivan,  Nov.  '87. 

479.  T.  AViLHELMii,   CM.,   Hedw.    '97,  p.    104,    Barbula; 

Streptopogon,  Par.  Index. 
Vic  and  Tas.  :  teste  Par.,  op.  cit. 

iii.  Encalypte.e. 
Encalypta,  Schreb.  (including  Pyramilrium). 

E.  aristata,  Hpe.;  Pyramitrluin^  Mitt.  Cat.   See  No.  441. 

E.  australis,  Mitt.     See  No.  481.    [CM.,  in  Gen.  M., 

gives  E.  australis,  Mitt.,  for  the  isl.  of  Chiloe  alone]. 

480.  E.  Nov^-Valesi^,  Hpe.,  Linn.  '71-3;  Pyramitrium, 

Mitt.  Cat.;  P.  ISfovce-Camhrice,  Hpe.  in  List,  '80. 
N.S.  W.,  Blue  Mts. :  Mrs.  Calvert.   "  Other  localities  "  :  teste  CM., 
in  Gen.  M.— Vic,  Gippsland :  F.v.M.  '55. 

481.  E.  TASMANICA,  Hpe.  et  CM.,  Linn.  '53;  E.  aust7'alis, 

Mitt.,  et  vulgaris  var.  Fl.  Tas.  and  Hpe.;  E.  ciliaia, 
Hedw.,  Mitt,  in  Kew  J.  Bot.  viii.  (fid.  Mitt.  Cat.). 
Tas.,  Launceston,  nr.  Cataract,  and  nr.  Cheshunt :  Archer.  Mt. 
Rumney,  and  Johnny's  Creek,  Derwent  Valley:  Weymouth.  Also 
coll.  by  Stuart  et  al.  Summit  of  Mt.  Rumney :  Bastow. — Vic, 
N.S.W.,  frequent  in  alpine  and  subalp.  regions,  esp.  on  Mt.  Ararat 
(teste  CM.) :  Sullivan,  Stirling  et  al.  ("  the  most  widely  distri- 
buted, 2000'-7000"' :  teste  Stirling  in  litt.).  Jenolan  Caves: 
Maiden,  Aug.  '98  (c.fr.),  Blakely,  Oct.  '99.  Warrumbungle 
Ranges  :  Forsyth,  Oct.  '99.  Coll.  at  Buchan  R.,  etc.,  by  F.v.M. 
in  '54-'55. 

E.  vulgai'is  var.     See  No.  481, 
6 


82  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

Tribe  xi.  G-rimmiacese. 
G  R I  M  M  I  A,  Ehrli. 

482.  G.  ANTiPODA,  CM.,  Symb.,  No.  285. 
Tas.,  Facy's  Gully :  Weymouth,  No\'.  '90. 

483.  G.  APOCARPA  (L.,  Bryum),  Hedw. 

Tas.  and  Vic,  frequent.  Austral.  Alps :  F.v.M.,  Stirling  et  al. 
"  On  gnoissic  rocks  nr.  Omeo,  2000' "  (Stirling).  On  bark  of 
Eucalypt,  Yarra  R.,  nr.  Melb. :  Reader,  '83.— N.S.W.,  Parramatta 
(or  N.  Shore,  S3^dney) :  Whitelegge  (fid.  Geheeb). 

(a)  f.  fol.  breviter  piliferis. 

N.S.W.,  Merritt's  Camp,  Mt.  Kosciusko,  Jan.,  '99,  and  Warrum- 
bungle  Ranges,  Oct.,  '99:  Forsyth. 

(b)  f.  fol.  brevissime  piliferis. 

Tas.,  Johnny's  Creek  :  Weymouth. — N.S.W.,  Creeklet,  Nepean 
R.,  Blue  Mts. :  Watts,  Oct.  '99  (or  possibly  the  f.  submutica). 

(c)  f.  submutica. 
Tas.,  Sorell :  Weymouth.— N.S.W J     See  under  f.  "b." 

484.  G.  ATRiCHA,  CM.,  Symb.,  No.  280. 
Vic,  on  stones,  nr.  Melton :  Reader,  Aug.  '86. 

G.  australasica,  Mitt.     Given  in  Bastow's  MS.  list  of 

Austr.  Mosses  for  N.S.W.     Information  needed. 
G.  austro-apocarpa,  CM.  in  sched.     This  seems  to  be 
G.  truncato-apocarp)a.     Par.  Index  Suppl.  gives  G. 
austro-apocarpa  for  N.Z. 
N.S.W.,  on  dry  rocks  in  creek,  nr.  Parramatta  Park:  Whitelegge, 
Oct.-Dec.  '84  (c.fr.). 

485.  G.  AUSTRO-FUNALis,  CM.,  Symb.,  No.  286. 
Vic,  Mt.  William :  Sullivan,  '83,   Austral.  Alps :  Stirling. 

486.  G.  BASALTiCA,  Mitt.,  Hdbk.  N.Z.  Fl. 

Tas.,  Mt.  Rumney,  on  rocks  :  Bastow. — Vic. :  teste  Melb.  Census. 
(Weymouth,  in  herb.,  gives  as  syn.  of  G.  cygnicollis). 


tJY  REV.   W.  WALTKK  WATTS  ANt)  THoMaS  WHITELEGGE.  So 

Grim  m  i  a  {continued).  [Grimmiacett. 

487.     G.  callosa,  CM.  et  Hpe.,  Linn.  '53. 
8.  A.,  Barossa  Range  and  Brown  Hill  Creek  :  F.v.M. 

ii^^.     G.  Camphelli^,  cm.,  Symb.,  No.  278. 
Vic,  Hume  R.:  Miss  Campbell,  herb.  Melb.  '81. 

489.  G.  compactula,  CM.,  Symb.,  No.  289. 
N.S.W.,  Mt.  Kosciusko:  Sullivan,  herb.  Melb.  '84. 

490.  G.  CRISPATULA,  Hpe.  et  CM.,  Linn.  '53;  G.  crisjyata  in 

Mitt.  Cat. 
W.A.:  teste  Melb.  Census.— S.  A.,  Flinder's  Range,  Fifth  Creek, 
Mt.  Lofty:  F.v.M.  (Tate,  List). 

49 L     G.  CYATHOCARPA,  Hpe.,  Linn.  '74,  Gue^nhelia. 
Vic,  Gippsland:  F.v.M.   Austral.  Alps  :  Stirling.— N.S.W.,  Blue 
Mts.:  Mrs.  Calvert. 

492.  G.  CYGNicoLLis,  Tayl.,  Lond.  J.  Bot.  '46;  G.  pulviriata 

var.  obtusata,  Brid.  (fid.  Mitt.  Cat.,  and  Par.  Index); 

var.  Africana  (Hedw.),  H.f. W.  ("  formerly  called  G. 

basaltica":  Weymouth  in  herb.). 
Tas.,  plentiful. — Australia,  from   W.  A.  to  Q.,  frequent,  esp.  in 
mountainous  districts.     Found  by  Preiss  in  W. A.   (Mt.  Brown, 
York),  Sept.  '39. 

493.  G.  CYLiNDROPYxis,  CM.,  Symb.,  No.  287. 

Vic,  Mt.  William:   Sullivan,  '83.   Austral.    Alps,   up  to  3000': 
Stirling. 

494.  G.  ENCALYPTOIDES,  CM.  (teste  F.  M.  Reader) 
Vic. :  D.  Sullivan. 

495.  G.  FLEXIFOLIA,  Hpe.,  Linn.  '(30. 

Vic,  Austral  Alps:  F.v.M.  (Snowy  R.),  Stirling. 

G.fu7ialis  (Schwgr.),is  recorded  f ro  m  Tas.  (Gunn  lect.), 
but  the  specimens  were  poor,  and  the  record  is 
doubtful.     See  Fl.  Tas. 


8^  CENSUS    MUSCORUM    AUStRALIENSIUM, 

G  K  I  M  M  I  A  {continued).  [Grimmiacece. 

496.  G.  HiSTRioiPiLA,  CM.  (teste  Reader). 
Vic,  among  lichens,  Mt.  Arapiles :  Reader,  Aug.  '96. 

497.  G.  LEiocARPA,  Tayl.,  Lond.  J.   Bot.  '46;  G.  leueojjhcea 

var.  subrotunda,  Wils.,  op.  cit. 

Tas.,  not  uncommon. — Australia,  frequent,  from  W.A.  to  Q. 
(Barossa  Range,  S.A. :  F.v.M.  Austral  Alps,  up  to  3000':  Stirling. 
Girilambone,  dry  interior  of  N.S.W. :  Biiuerlen.  Warrumbungle 
Ranges:  Forsyth.  Goulburn  and  Richmond  R.:  Watts.  Q.,  Glad- 
field  :  C.  J.  Gwy ther.  Many  other  Iocs,  known). 

G.  leucophcea,  Grev.     See  No.  497. 

498      G.   MUTICA,    Hpe.,   Linn.   '60 ;   G.   apocarpa  var.   fol. 
muticis,  Hook,  f.,  Fl.  Tas.,  etc. 

Tas.,  wet  rocks,  Mt.  Faulkner  '92,  Millhouse's  Falls  '93,  Hobart 
Rivulet,  etc.:  Weymouth.— Vic,  Mitchell  R.,  Mt.  Wellington, 
and  Sealer's  Cove  :  F.v.M.  Alps  :  Sullivan  and  Stirling. — N.S.  W., 
Parramatta :  Woolls.  Richmond  R.,  Killen's  Falls,  Emigrant 
Creek  :  Watts,  '96.   Shoalhaven  R. :  Forsyth,  '99. 

499.  G.  OBTUSATA,  Hpe.  et  CM.,  Linn.  '53,  Guemhelia. 
Vic,  Gippsland  :  F.v.M.— N.S. W.,  Mt.  Kosciusko:  Forsyth, '99. 

500.  G.  PARRAMATTENSis,  CM.,  Symb.,  No.  276. 
N.S.W.,  Parramatta:  Whitelegge,  Oct.  '84  (c.fr.). 

501.  G.  PROCUMBENS,  Mitt.  Cat.  '83. 

Vic,  Snowy  R. :  F.v.M.  Austral.  Alps:  Stirling. 

G.  jjulvinata  var.  ohtusata,  and  var.  AJricana.     See 
No.  492. 

502.  G.   PYGMiEA,  CM.,  Syn.  i.,   787;  G.  irichophylla  var. 

australis,  Hpe.,  PI.  Preiss. 

"  Australia,"  on  granitic  rocks  :  Preiss.  —  S.A.,  Darebin  Creek  : 
F.v.M.— Tas.  :  Stuart. 


IJY  REV.   W.   WAiiTEK  WATTS  AND  THOMAS  WHITELKCJGK.  85 

G  K I  M  M  I  A  {continued).  [Grimmiacece. 

503.  G.  STKNOPHYLLA,  CM.,  Symb.,  No.  271. 

Tas..  rocks,  Sugar  Loaf,  Green  Fonts,  May  '92  :  Weymouth  in 
herb.  Broth.  '93  (sub  G.  trichophylla,  Grev.). 

504.  G.  Stirlingi,  CM.,  Symb.,  No.  281. 
Vic,  Omeo,  2500'-3000',  on  rocks  :  Stirling,  '83. 

505.  G.  suBCALLOSA,  CM.,  Symb.,  No.  277. 
Vic,  Mt.  Cole,  Pj^renees  :  D.  Sullivan,  '83. 

506.  G.  SuLLiVANi,  CM.,  Symb.,  No.  288. 

Vic,  common  all  over  the  Alps,  up  to  6000',  chiefly  on  the  meta- 
morphic  areas;  first  found  nr.  Omeo  by  D.  Sullivan.  Also  coll. 
by  Stirling.     CM.  gives  Sullivan's  date  as  '83;  Reader  gives  '76. 

507.  G.  TASMANicA,  CM.,  Symb.,  No.  272. 

Tas,,  loc.  not  specified  :  Stuart,  herb.  Sonder,  '40.  On  wall  of 
II. C  Cemetery,  Brisbane  St.,  Hobart :  Weymouth,  Oct.  '90. 

508.  G.  TRICHOPHYLLA,  Grev.,  Scot.  Crypt.  Flor. 

Tas.,  Cheshunt :  Archer.  Mt.  Wellington,  etc. :  Hooker,  Oldfield, 
Weymouth  (Pinnacle),  Watts  (St.  Crispin's),  Bastow  et  al., 
frequent.  — N.S.W.,  Mt.  Kosciusko:  Forsyth,  Jan.  '99.— S.A., 
Adelaide  :  Schauinsland,  '96-7. — Q. :  teste  Melb.  Census. 

509.  G.  TRUNCATO-APOCARPA,  CM.,  Symb.,  No.  282. 
N.S.W.,  Parramatta :   Whitelegge,  Oct.    '84   (c.fr.).     See  under 
No.  4S4. 

510.  G.  WooLLsiANA,  CM.,  Symb.,  No.  279. 

N.S.W.,  Sydney  :   Woolls,  '81  (c.fr.).   Cambewarra :  Whitelegge. 

R  H  A  c  0  M  I  T  R I  u  M,  Brid.  (Sect,  of  Grimmia,  CM.) 

511.  R.  AMCENUM,  Broth.,  A.M.  '99,  No.  151. 

N.S.W.,  Merritt's  Camp,  Mt.  Kosciusko:  Maiden  and  Forsyth, 
Jan.  '99. 

a  aquaticum,  CM.     See  No.  517. 

R.  austro-sudetictim.     Given  in  Melb.  Census  for  Vic 
Particulars  needed. 


86  CENSUS    MUSCORUM    AUSlRALiENSiU^}, 

R  H  A  c  o  iM  I  T  R  I  u  M  {continued).  .[Grimmiacece. 

512.  R.  CHLOROCARPUM,  Mitt.  Cat.,  Grimmia  {Dryptodon)] 
R.  crispulum  (fid.  Jaeg.,  but  not  Mitt.). 

Tas.  :  Archer. 

513.  R.  CRISPULUM,  H.f.W.,  Fl.  Ant,  N.Z.,  and  Tas.,  A'. 

(Di'yptodori). 
Tas.,   on   rocks,    rivulet,   south  of   Cumming's    Head,   W.    Mts. : 
Archer.      Also  coll.  by  Hooker,  Gunn  et  al. 

(a)  var.  fol.  brevioribus  muticis,  Fl.  Tas. 

Tas.,  on  trap  rocks,  Glen  Leith,  rare;  on  stones  nr.  Lake  Echo  : 
Hooker,  f. 

(b)  var.  fol.  longioribus  muticis,  Fl.  Tas. 

Tas.,  St.  Patrick's  R. :  Gunn.  Rocks,  N.  and  S.  Huon  :  Oldfield. 
Rocks  and  stones,  rivulet,  nr.  Cumming's  Head,  W.  Mts. :  Archer. 

514.  R.    EMERSUM,    CM.,    Bot.    Zeit.    '51,    Grimmia;    R. 
symphyodontum  (fid.  Mitt.). 

Tas.,  on  trunks  of  trees  about  Mt.  Wellington  :   Mossman. 
R.  fasiculare  var.     See  No.  523. 

515.  R.  heterostichum  (Hedw.,  Trichostomum),  Brid. 
Tas.,  Mt.  Wellington,  above  Springs,  common. 

(a)  var.  tasmanicum,  Hpe.,  Linn.  '52. 
Tas.  (loc.  1) 

[The  vars.  alopecurwm  and  gracilescens  are  also  recorded  for 
Tas.  (vide  Fl.  Tas.  and  Bastow,  M.  Tas.),  but  Jaeg.  Ad.  gives  Tas. 
for  var.  tasm,anicuni  only.] 

516.  R.  LANGUiNOSUM  (Hedw.,  Trichostomum)^  Brid. 

Vic. — N.S.W.,  Australian  Alps  :  F.v.M.  "  Either  this  or  an  allied 
sp.  nr.  the  summit  of  Mt.  Kosciusko,  7000':  Stirling. — Tas.,  top 
of  Mt.  Wellington  :  Bastow,  Weymouth,  Watts  (on  ground  close 
to  pinnacle)  et  al. 

517.  R.  PROTENSUM,  Braun;  G.  aqaatica  (Brid.),  CM.,  fid. 
Jaeg.,  Broth. 

Tas.,  Mt.  Wellington. 


BY  RRV.   W.   WALTER,  WATTS  AND  THOMAS  WHITELEGGE.  <S7 

R  H  A  c  0  M  I T  R  I  u  M  (continued).  \Grimmiacece. 

518.  R.  PRUiNOSUM,  CM.;  R.  languinosum,  H.f.W.,FI.  Ant.; 

var.  2)rui7iosuni,  Fl.  Tas. 
Tas.,  Mt.  Wellington:  Gunn.  Weymouth  et al. (on  rock,  "Ploughed 
Field  "). 

519.  R.  PSEUDO-PATENS,  CM.,  Rev.  Bry.  76,  Symb.,  No.  290, 

G.  {Dryptodo7i). 
N.S.W.,  nr.  Sydney  :  Kayser  coll. 

520.  R.  PYCNOTRiCHUM,  CM.,  Symb.,  No.  291. 

Tas.,  Guy  Fawkes  Rivulet,  Sept.  '90,  top  of  Mt.  Wellington,  Feb. 
'88  (/.  nigrita):  Weymouth,  herb.  Burchard.  Also  Oldfield  in 
herb.  Broth.  [This  is  prob.  the  sp.  returned  by  other  bryologists 
as  R.  heterostichum]. 

521.  R.  RUPESTRE,  H.f.W.,  Fl.  N.Z.;  Drypiodon,  Fl.  Ant. 
Tas.,  on  moist  rocks  on  mountains  :  teste  Bastow,  M.  Tas. — Vic. : 
teste  Melb.  Census. 

522.  R.  SuNDAicuM,  CM.,  Verhandl.  Gesell.  in  Wien  '69. 
Q. :  teste  Mitt.  Cat.,  and  Bailey,  Syn.  '83.   [Jaeg.  gives  only  Java.] 

523.  R.  SYMPHYODONTUM,  C  M.,  Syn.,  Grimmia;  R.  fasiculare 

var.,  H.f.W.,  Fl.  Ant.;  R.  symphyodoyi,  Jaeg.  Ad. 
Tas.,  on  dry  rocks  at  the  back  of  Hobart :  J.  D.  Hooker.  On  rocks, 
the  Falls  :  Archer.   Mt.  Wellington  :  Mossman. 

G  L  Y  P  H  0  M  I  T  R  I  u  M,  Brid.  {Ptychomitrium,  Par.  Index) 

524.  G.  acutifolium,  H.f.W.,  Fl.  Tas. 

S.A.:  C  Prentice. —Vic,  nr.  Melbourne:  Reader,  '84. — Tas.,  on 
rocks,  Port  Sorell :  Archer.  Glen  Leith  :  J.  D.  Hooker.  On  banks 
and  trap  rocks,  nr.  the  Derwent.  Sugar  Loaf,  Kempton  :  Wey- 
mouth. 

525.  G.  Adamsoni,  Mitt.,  Jour.  L.  Soc.  '60. 

Vic,  nr.  Melb..  common  on  trap  rocks  :  Adarason, — N.S.W,  ; 
teste  Melb.  Census, 


88  CENSUS    MUSCORUM    AUSTRALIENSIUM, 

G  L  Y  p  H  o  M  I  T  R I  u  M  (continued).  [Grimmiacece. 

526.  G.  AUSTRALE,  Hpe,,  Linn.  '55,  Brachysteleum. 
Vic,  Alps,  towards  Buchan  R. :  F.v.M. 

527.  G.  COMMUTATUM,  CM.,  Symb.,  No.  269,  Brachystelenm. 
Q.,  Brisbane  R. :  Mrs.  Dietrich.  '64.  Toovvoomba:  Hartmann,  '80 
(eir.).  Bunya  Mts.  :  Bailey,  '85.— N.S.W.,  Tarn  worth:  C.  T. 
Musson,  '90.  Lower  Hunter  R. :  Miss  Carter,  '82  (f.  cal.  operculo 
parum  majore). 

528.  G.  LATiFOLiuM,  Broth.,  A.M.  '95,  No.  70. 

Tas.,  Newtown  Rivulet,  on  water-washed  rocky  bank :  Weymouth, 
Nov.  '95. 

529.  G.  LAxiFOLiUM,  C.  M.,  Symb.,  No.  268,  5mc7i2/s^e?ew??2. 
N.S.W.,  Parramatta  :  Whitelegge,  Dec.  '84  (e.fr.).  South  Head, 
Port  Jackson  :  J.  B.  Ward,  July  '99.  Shoalhaven  R.,  Sept.  '99, 
and  Kiama,  Dec.  '99  :  Forsyth. 

530.  G.  MiCROBLASTUM,  C. M., Symb.,  No.  270,  Brachysteleum. 
N.S.W.,  nr.  Sydney:  Kayser  Coll. 

531.  G.   MiTTENii   Jaeg.,  Ad.,  Ptychomitriiim;  G.  serratum 

Mitt.,  Jour.  L.  Soc.  '60. 
Tas.,   on   rocks,   north   side   of   Cataracts,   Launceston:    Archer. 
Nelson  Range:  Weymouth  (rocks.  Proctor's  Rd.,  etc.). — N.S.W., 
Cambewarra:  Thorpe,  '84. 

532.  G.    MUELLERi,   Mitt.,   Jour.    L.    Soc.    '60;   Brachystel- 

enni  Howeanum,  Hpe. 
Q.,  Glasshouse  Mts.  and  Brisbane  R.:  F.v.M.  Mt.  Archer: 
Mus.son. — N.S.  W.,  very  common  on  basaltic  rocks  on  the  Richmond 
R.;  frequent  nr.  S3^dne3';  Nepean  R.:  Watts,  '99.  Barber's 
Creek,  Kiama,  and  Warrumbungle  Ranges:  Forsj^th,  '99. — Vic, 
nr.  Melb.:  Reader. 

G,  serratum,  Mitt.      See  No.  53 L 


BY  REV.  W.  WALTER  WATTS  AND  THOMAS  WHITELEGGE. 


89 


ADDENDA  (Oct.  29th,  1902). 

r)33.     Sphagnum  commutatum,  Wariist.,  sp.n.,  Magy.  Botan. 
Lapok.  1902,  p.  45. 

N.S.W.,  Swainp  at  north  end  of  Three  Mile  Scrub,  nr.  Byron 
Ba}'-,  and  on  edge  of  Fresh  Water  Lake,  Seven  Mile  Beach,  N. 
Coast:  Watts,  May  2nd  and  29th,  1900. 

3  bis.     (a)  Sphagnum  Brotherusii,  Warnst.,  (a)va,r.  plumo- 
SULUM,  Warnst.,  in  litt.  Aug.  1902. 
N.S.W.,   in   dripping   water,   cliff,   below    *S^;/i.    drepanodadumy 
Shaw's  Bay,  Richmond  R.:  Watts,  May  26th,  1900. 

534.     FissiDENS    ASPLENioiDES  (Sw.),   Hedw.,    M.    Frond.; 
Hypnum,  Sw.,  Prodr.  p.  140. 
N.S.W.,  Wilson's  Creek,  Richmond  R.,  Oct.  1901,  and  BlueMts., 
nr.  Emu,  Nov.  1900:  Watts. 

[CM.,  Gen.  M.  gives  Cuba  only;  Broth.  Bryales,  West  Indies 
and  S.  Amer.  A  recent  return  from  Broth,  contains  the  name 
for  specimens  coll.  as  above.] 


INDEX  OF  GENERA. 

(Names  in  italics  are  Synonyms.) 


Acaulon 

Anacalypta,  vide  Pottia. 

Andreaea    ... 

Auisotheciiim,  vide  Dicranellaand 

Blindia. 
Aongstrcemia,  vide  Dicranella. 

Arehidium  

AschModoii,  vide  Distichium. 

Astomura  ... 

Astomum,  vide  Pleuridium 

Atrichum 

Barbula     ... 

Blindia 

Braclnjateleum,  videGlyphomitrium. 

Bruchia     ... 

Buxbaumia  

Calymperes  


PAGE 

PAGE 

72 

Campy  lopus         43 

Catharinea,  vide  Atrichum. 

12 

Catharinella,  vide  Pogonatum. 

Ceratodon 37 

Codonoble2)hartim,Yk\eSyvYh.OT^odon. 

Conomitrium,  vide  Fissidens. 

31 

Dawsonia ...     14 

Desmatodon         74 

61 

Dkhodontiuin,    vide    Dicranella 

and  Blindia. 

15 

Dicnemonella,  vide  Leucoloma. 

68 

Dicnemos   .          54 

40 

Dicranella 38 

jm. 

Dieranodontium 47 

31 

Dicranoloma,    vide    Leucoloma 

14 

(note). 

59 

Dicranoweisia      41 

90 


CENSUS    MUSCORUM    AUSTRALIEXSIUM. 


PAGE 
47,50 

67,  68 


Dicranum 

Didymodon  

Diohelon,  vide  Dicranella. 

Distichium 

Ditrichum... 

Divaricatella,  vide  Dicranella. 

Dryptodon,  vide  Rhacomitrium 

Eccremidium 

Encalypta 

Eucamptodon 
Fissidens  ... 
Glyphomitrium    ... 
Giimmia  ... 

Gymnostomum    

Holomitrium 

Hymenostomum 

Hyophila   ... 

Leptodontium      

Leptotrichum,  vide  Ditrichum. 
LesA;ert, vide  Triquetrella  papillata.  67 


34 

81 
55 
89 
87 
82 
65,  74 
42 
62 
68 
67 


21 


Leucobryum 

Leucodon,  vide   Leiicoloma   and 

Dicnemos, 
Leucoloma 
Leucophanes 
Lophiodon,  vide  Ditrichum, 

Mesotus     

Microdns,  vide  Dicranella. 
Octoblepharum    ... 
Octodiceras 

Oligotrichum        

Oncoplwroloma ,  vide  Leucoloma 

(note). 
Phascum  ..  


55 


47 
58 

47 

58 
80 
16 


73 


PAGE 

Pleuridium  ...         ...         ...     32 

Pleurophascum 73 

PcccilopJiylliim,  vide  Leucoloma. 
Pogonaturn  ..  ...         ...     17 

Polytrichadelphus  ..  ...     17 

Polytrichum         19 

Pottia        74 

Psilopilum  16,  18 

Pti/chomitrium,  vide  Glyphomitrium. 
Pyramitrium,  vide  Encalypta. 
Rhacomitrium      ..  ...         ...     85 

Schizophascum 73 

Sclerodontium,  vide  Leucoloma. 
Seligeria,  vide  Dicranella. 
Spharangium,  vide  Acaulon. 
Sphagnum  . . ,         . .  8,  89 

Sporledera,  vide  Bruchia. 
Streptopogon,  vide  Tortula. 
Swartzia,  vide  Distichium. 
Symhlepharis,  vide  Holomitrium. 
Syntrichia,  vide  Tortula. 

Syrrhopodon        59 

Systegium,  vide  Astomum. 

Tetrapterum         73 

Thi/ridium,  vide  Syrrhopodon. 

Toitella 66 

Tortula 75 

Trematodon  31 

Trichostomum     34,65,86 

Tridontium  67 

Triquetrella  ...         ...         ...     G7 

Tropinotji!^,  vide  Leucophanes, 

Weisia       38,  G3,  68 

WeUiello,  vide  Dicranella. 


[Printed  off  December  8th,  1902.; 


493 


NOTES  OX  SOME  NEW  SOUTH  WALES  HEPATICS. 

By  the  Rev.  W.  Walter  Watts. 

(Continued  from  Proceedings,  1901,  j)^.  215,  633.) 

I  beg  to  place  on  record  the  following  additional  Hepatics, 
named  in  a  recent  return  from  M.  Stephani,  through  the  good 
offices  of  Dr.  E.  Levier,  to  whom  they  were  sent  in  November, 
1901. 

(1)  Species  collected  by  me  on  the  Richmond  and  Brunswick 
Rivers  : — 

Aneura  pinnatifida,  Nees;  East  Ballina,  March,  1901. 
H    Chiloscyphus  argutus,  Nees;  Ballina,  July,  1901,  Alston ville, 
"  Sept.,  1901. 

,,  limosus,  Carr.  et  Pears.;  Ballina,  July,  1901. 

Fimhriaria  co7iocephala,  Steph.;  bank  of  cutting,  Rous,  Sept., 

1901. 
FruUania  megalocarpa,  Tayl.;  East  Ballina,  March,  1901. 

,,  clavata,  Tayl.;  East  Ballina,  May,  1901. 

Jungermannia  montana,  Steph.;  Alston  ville  Cutting,  March  and 

June,    1901,   Shaw's    Bay,   Ballina, 
April,  &c.,  1901. 
Lejeunea  (Eulejeunea)  cuspjidistipula,^te'ph.;  on  "tea"  tree,  nr. 

Ballina,  May,  1901. 
,,  ,,  elobata,   Pears.  ;   on  tree,    East   Ballina, 

March,  1901. 
,,  ,,  tasmanica,  Gottsche ;    brooklet,   Alston- 

ville  and  Rous,  May,  1900,  etc. 
33 


494  NOTES  ON  SOME  NEW  SOUTH  WALES  HEPATICS. 

Lepidozia  capilligera,  Lelim.  et  Lindenberg;  Alston  ville  Cutting, 

June,  1901,  Tuckombil,  April,  1901. 
,,        glaucophylla,   Tayl. ;   Tuckombil,    Alstonville,    April, 

1901. 
,,        gracillima,  Carr.   et  Pears.;    nr.   Ballina,  March  and 

July,  1901. 
Marchantia  cephaloscypha,  Steph.;  Byron  Bay,  Oct.,  1901. 
Metzgeria  crassicostata,  Steph.;  Tuckombil,   Alstonville,   Sept., 

1901. 
,,  furcata,  (L.)  S.  O.  Lindb.;  Myocum,  Brunswick  R., 

Oct.,  1901. 
Plagiochilafascictilata,,  Ldnbg. ;  Head  of  Wilson's  Creek,  Oct., 

1901. 
Fycnolejeunea  lo7}gidens,  Steph.;  on  trees,  E.  Ballina,  May,  1901. 
Radula  Mittenii,  Steph.;  Head  of  Wilson's  Creek,  Oct.,  1901. 
Trachylejeunea  eUgantissima,  Steph.;  growing  on  Pteris  falcata, 

Tuckombil,  Alstonville,  Sept.,  1901. 

There  are  included  in  the  return  six  new  species,  five  of  which 

M.  Stephani  has  done  me  the  honour  of  dedicating  to  myself. 

These  will  be  recorded  when  described. 

(2)  The  following  were  also  returned  : — 

Fimhriaria  Drummondii,  Tayl.;  Albury,  leg.  Arthur  Sharpe, 

July,  1900. 
Frullania    pycnantha,   Tayl.;    Tam worth,    leg.  C.    T.    Musson 

(No.  658). 
Trichocolea    tomentella,  (Huds.)   Nees;     Kurrajong,  leg.  C.    T. 

Musson. 

Corrigenda. — My  last  list  of  records,  by  an  unfortunate  slip, 
gave  Frullania  Hampeana,  Nees;  it  should  read  F.  Hampeana^ 
Lindenberg.  A  slip  of  Dr.  Levier's  led  me  to  record  a  Tasmanian 
species,  Marsupidium  setulosum  ;  this  should  be  corrected  to 
Acroholhus  unguiculatus  (H.  f.  Tayl.). 


495 


NOTES    AXD    EXHIBITS. 

Mr.  W.  S.  Dun  exhibited  specimens  of  Conularia  inornata, 
Dana,  0.  Icevigata,  Dana,  and  C.  tasvianiensis,  Johnston  (possibly 
an  acute  form  of  inornata)  from  the  Lower  Marine  Beds  (Permo- 
Carboniferous)  of  Ravensfield  and  Harper's  Hill,  iST.S.W.  An 
imperfect  specimen  of  C.  inornata  21  cm.  in  length  was  shown; 
a  smaller  specimen  of  the  same  species  showed  the  inbent  and 
triangular  apex  of  the  sides  very  clearly.  The  genus  is  but 
seldom  met  with  in  the  Upper  Marine,  but  is  not  rare  in  the 
Lower  Series. 

Mr.  Hedley  exhibited  a  series  of  mollusca  lately  dredged  in 
100  fathoms  off  WoUongong  by  Mr.  Halligan  and  himself, 
including  Lotorium  nodocostatum,  hitherto  only  known  from 
Tasmania,  Chlamys  Jenesirata,  Hedley,  doubtfully  distinct  from 
a  Tertiary  fossil,  and  a  species  of  Dymia  lately  described  by  himself 
in  the  "  Thetis  "  Report. 

Mr.  Stead  showed  a  preparation  of  the  first  and  only  English 
Lobster  {Homarus  vulgaris)  to  reach  Australian  shores  alive.  It 
arrived  with  the  shipment  of  English  Plaice  recently  introduced, 
but  it  did  not  rally  from  the  effects  of  the  voyage,  and  died  a  few 
hours  after  removal  to  Port  Hacking.  As  it  was  an  ovigerous 
female  this  was  to  be  regretted. 

Mr.  R.  Greig  Smith  exhibited  a  young  specimen  of  the  Sock- 
eyed  Salmon  affected  with  Saprolegnia  ferax,  forwarded  to  him 
from  Tasmania.  Also  various  cultures,  chemical  products,  and 
microscopic  preparations  in  illustration  of  his  paper  on  Wattle 
gums. 

Mr.  Baker  exhibited  a  specimen  of  Ardisia  raceinosa,  n.sp.,  in 
illustration  of  his  paper.  He  also  offered  some  observations  on  the 
desirability  of  collecting  data  relating  to  the  effects  of  the  present 


496  NOTES    AND    EXHIBITS. 

drought  on  the  flora;  and  he  stated  that  he  was  moving  in  the 
matter  through  the  Department  of  Public  Instruction.  He  had 
received  information  that  one  of  the  plants  which  was  suffering 
most  severely  in  some  districts  was  the  Currawong  (Acacia 
doratoxylon).  a  useful  fodder  plant  in  time  of  drought. 

The  President  emphasised  the  importance  of  the  subject,  and 
intimated  that  he  too  had  already  taken  some  steps,  in  conjunction 
with  the  officers  of  the  Forestry  Branch,  to  obtain  and  record 
information. 

Mr.  Carne  also  spoke  fi-om  recent  personal  experience  of  the 
destructive  effects  of  the  drought  in  several  districts;  and  he  was 
able  to  testify  to  the  value  of  Acacia  doratoxylon  for  dairy  cattle 
as  a  stand-by  during  drought. 


WEDNESDAY,  OCTOBER  29th,   1902. 


The  Ordinary  Monthly  Meeting  of  the  Societ}^  was  held  in 
the  Linnean  Hall,  Ithaca  Road,  Elizabeth  Bay,  on  Wednesday 
evening,  October  29th,  1902. 


Mr.  J.  H.  Maiden,  F  L.S.,  Arc,  President,  in  the  Chair. 


The  Donations  and  Exchanges  for  the  month,  amounting  to  9 
Vols.,  73  Parts  or  Nos.,  15  Bulletins,  6  Reports,  22  Pamphlets, 
and  1  Miscellanea,  recei\'ed  from  43  Societies,  itc,  and  2  Authors, 
were  laid  upon  the  table. 


497 


ON    TWO    REMARKABLE    SPOROCYSTS    OCCURRINU 

IN  MYTILUS  LATUS,  ON  THE  COAST  OF 

NEW  ZEALAND. 

By  Professor  W.  A.  Haswell,  M.A.,  D.Sc,  F.R.S. 

(Plates  xix.-xx.) 

The  parasites  which  form  the  subject  of  the  present  communica- 
tion were  found  by  me  last  summer  in  specimens  of  a  species  of 
Mytilus  (M.  latus)  living  on  the  coast  of  New  Zealand.  I  am 
greatly  indebted  to  Dr.  Chas.  Chilton,  Acting  Professor  of  Biology 
at  Canterbury  College,  Christchurch,  for  subsequently  procuring 
and  sending  me  a  large  consignment  of  live  specimens  of  the 
mussel. 

The  Spokocysts  of  an  Echisostomum. 

The  Sporocysts,  in  which  the  Cercarise  are  developed,  were 
found  in  a  fairly  large  proportion  (about  10  per  cent.)  of  the 
mussels  examined,  and  always  in  large  numbers.  They  infest  the 
mantle-folds  and  the  region  immediately  in  front  of  the  posterior 
adductor  muscle,  and  are  most  abundant  in  the  gonads  and 
nephridia.  In  infected  mussels  this  region  of  the  body  is  coloured 
bright  red  owing  to  the  presence  of  hundreds  of  the  Sporocysts, 
which  contain  a  conspicuous  red  substance  to  be  referred  to 
below. 

From  the  fact  that  such  large  numbers  of  the  Sporocysts  occur 
together,  it  is  to  be  inferred  that  they  multiply  actively  in  the 
tissues  of  the  mussel.  This  multiplication  of  Sporocysts  takes 
place  not  only  by  budding,  or  rather  binary  fission,  but  also, 
though  comparatively  rarely,  by  a  process  corresponding  to  that 
by  which  in  many,  if  not  most,  Sporocysts  Redia^  are  developed. 


498  ON    TWO    REMARKABLE    SPOROCYSTS, 

The  Sporocysts  (figs.  1  and  2)  reach  a  maximum  length  of  about 
3  mm.  They  are  able  to  creep  about  by  alternate  elongations  and 
contractions.  During  these  movements  the  shape  necessarily^ 
varies  greatly,  but  a  "head,"  directed  forwards,  in  locomotion,  is 
always  recognisable,  being  constricted  off  as  a  rounded  knob.  As 
Braun  (3)  points  out,  it  is  erroneous  to  refer  to  this  as  a  sucker. 
The  muscular  layers  in  this  position  are  not  thickened  or  modified 
in  any  way.  At  its  apex  this  is  capable  of  being  deeply  involuted, 
the  result  being  the  development  of  a  pit  with  a  terminal  opening. 
Since  it  is  usually  in  this  position  that  the  fully  developed 
Cercaria  escapes,  an  actual  perforation  sometimes  is  found  to 
occur,  but  this  is  of  an  entirely  temporar}^  character. 

In  many  cases,  applied  to  the  outer  surface  of  the  Sporocyst 
were  groups  of  amoeboid  cells  emitting  pseudopodia.  Such  groups 
of  cells  are  stated  by  Biehringer  to  occur  generally  in  the  Sporo- 
cysts studied  by  him;  the  cells,  which  are  blood-corpuscles  of 
the  host,  eventually  giving  rise  to  a  layer —  "  the  paletot "—  which 
completely  invests  the  parasite.  Such  a  continuous  layer  I  have 
never  found.  The  groups  of  cells  were  never  found  in  entire  pre- 
served specimens  or  in  sections,  having  always  become  detached, 
apparently'',  in  the  process  of  preparation. 

The  Sporocyst  is  enclosed  in  a  non-cellular  cuticular  laj^er  in 
which  no  trace  of  nuclei  occurs.  This  la3'er,  which  is  0'003  mm. 
in  thickness,  exhibits  a  fine  vertical  striation,  the  significance  of 
which  will  be  referred  to  presently.  Beneath  it  are  the  layers  of 
muscle,  which  are  extremely  thin,  an  outer  layer  of  circularly 
and  an  inner  of  longitudinally  running  fibres. 

The  outer  layer  (cuticle  of  most  authors)  is  described  as  a  thin 
structureless  membrane.  Biehringer  (2),  Heckert  {teste  Braun) 
and  Loos  (9)  describe  it  as  containing,  in  the  young  condition, 
sparsely  distributed  nuclei.  In  the  Sporocyst  from  Mytihis  latus 
nuclei  were  not  found  in  the  cuticle  of  any  of  the  numerous 
specimens  which  were  examined  for  them  :  it  is  possible,  however, 
that  the  same  may  not  hold  good  of  those  Sporocysts  that  are 
developed  directly  from  the  Miracidium  embrj^o.     In  the  Sporo- 


BY    PROFESSOR    AV.    A.    HASWELL.  499 

cyst  of  Cerca7'ia  armata,  also,  nuclei  are  absent  in  the  cuticle 
according  to  Schwarze  (12). 

The  germinal  epithelium  (fig.  8)  lines  the  whole  interior  of  the 
Sporocyst.  In  the  greater  part  of  its  extent  in  a  mature  Sporo- 
cyst  it  is  a  thin  layer  of  flattened  cells,  most  of  which  are  stellate, 
the  processes  of  neighbouring  cells  anastomosing  to  form  a  net- 
work. At  the  anterior  and  posterior  ends  it  is  thickened,  the 
thickening  consisting  of  a  layer  of  vertically  elongated  cells.  At 
its  outer  end  each  cell  of  these  thickened  regions  terminates 
in  one  or  several  very  fine  processes,  which  are  clearly 
traceable  through  the  layer  of  muscle,  and  are  seen  to 
be  continuous  with  the  vertical  striations  of  the  cuticle.  At 
its  opposite  free  end  each  cell  projects  into  the  cavity  of  the 
Sporocyst.  In  the  living  specimen  globules  were  seen  to  be  given 
off  occasionally  from  the  inner  ends  of  these  cells,  becoming  free 
in  the  interior  of  the  cavity,  and  often  becoming  collected  in 
masses.  These  globules  were  swallowed  by  the  more  mature 
Cercaripe,  in  the  intestine  of  which  considerable  numbers  of  them 
were  nearly  always  to  be  detected.  It  might  be  supposed  that 
this  phenomenon  is  not  a  natural  one,  but  brought  about  by  the 
pressure  to  which  the  living  Sporoc3^st  is  naturally  subjected  in 
order  that  its  structure  may  be  examined.  But  such  an  explana- 
tion cannot  be  entertained  in  view  of  the  results  obtained  from 
the  examination  of  sections  of  specimens  fixed  by  various  reagents. 
In  such  sections  the  epithelium  has  the  appearance  represented  in 
in  fig.  8,  the  cells  presenting  a  variety  of  stages  in  the  process  of 
formation  of  the  globules;  and  many  of  the  latter  are  to  be  found 
free  in  the  cavity  of  the  Sporocyst  and  in  the  intestine  of  the 
Cercaria3.  Among  the  globules  in  the  latter  position  entire 
nucleated  cells  are  frequently  to  be  observed. 

It  thus  appears  to  be  almost  certain  that  the  germinal  epithe- 
lium has  an  important  secondary  function  in  providing  nutriment 
for  the  Cercarise  with  fully  developed  and  functional  digestive 
canal.  To  nourish  the  many  growing  embryos  which  it  contains, 
the  wall  of  the  Sporocyst  must  absorb  nutrient  matter  in  large 
quantity  from   the  tissues  of  its  host.       This   is   taken   in,    un- 


.500  ON    TWO    REMARKABLE    SPOROCYSTS, 

doubtedly,  by  the  cells  of  the  germinal  epithelium  through  the 
pores  of  the  cuticle,  and  most  of  it  must  be  given  off  in  a  state  of 
solution  into  the  fluid  in  which  the  developing  embrj^os,  in  all  but 
their  earliest  stages,  float  freely.  The  special  part  played  by  the 
nutriment  provided  in  the  form  of  globules  is  that  it  supplies  a 
store  of  sustenance  which  the  Cercaria  takes  with  it  when  it 
leaves  the  Sporocyst  and  sets  out  in  search  of  a  second  host. 

I  can  find  no  mention  in  the  literature  accessible  to  me  of  this 
giving  off  of  globules  by  the  cells  of  the  germinal  epithelium. 
That  the  case  is  not  an  altogether  isolated  one,  however,  is  shewn 
by  the  fact  that  Reuss  in  a  recently  published  preliminary 
notice  (H)  mentions  the  presence  in  the  Sporocysts  of  DidcnmLm 
duplicatum  of  many  "fat  drops."  It  is  at  least  possible  that  the 
phenomenon  is  a  general  one,  and  that  it  is  only  owing  to  the 
colourless  character  of  the  globules  in  most  instances  that  it  has 
hitherto  escaped  notice.  On  the  other  hand,  it  has  to  be  borne 
in  mind  that  in  many  Cercariie  the  digesti^  e  canal  does  not  become 
suflicientiy  well  developed  to  be  functional  until  after  the  larva 
has  escaped  from  the  Redia  or  Sporocj^st.  In  such  cases  the 
formation  of  the  globules,  except  merely  in  order  to  be  subsequently 
dissolved  in  the  nutrient  fluid  of  the  cyst,  would  evidentl}^  be 
superfluous. 

The  occurrence  of  a  pigment  in  the  germinal  epithelium  appears 
to  be  exceptional.  I  have  met  with  only  three  recorded  cases  of 
such  a  thing — by  Wagener  (14)  in  the  Sporocyst  of  Cercaria  cysto- 
phora,  by  Pagenstecher  (10)  in  the  case  of  the  Sporocyst  of  Cercaria 
cotylocerca,  and  by  Heckert  (4)  in  that  of  Lencochhridnwi  para- 
doxiim. 

The  embryos  are  developed  from  a  mass  of  specialised  cells  of 
the  germinal  epithelium — the  ovary  (figs.  5-7).  This  is  constant 
in  position,  at  the  posterior  end,  and  there  is  never  more  than 
one.  It  projects  as  a  free  process  into  the  cavity  of  the  Sporo. 
cyst.  At  its  base  every  gradation  is  to  be  observed  between  the 
ordinary  cells  of  the  posterior  thickening  of  the  germinal  epithe- 
lium and  the  cells  destined  to  give  rise  to  embryos.  Towards  its 
free  extremity  the  ova  undergo  segmentation  which  results  in 


BY    PROFESSOR    W.    A.    HASWELL.  oOl 

the  formation  of  a  spherical  bod}^  containing  a  number  of  nuclei 
with  cell-outlines  ill  defined,  though  sometimes  recognisable. 
These  spherical  embryos  are  still  enclosed  within  a  delicate  mem- 
brane which  invests  the  whole  ovary.  As  they  reach  a  size  of 
0*025  mm.  or  thereabouts,  they  become  free,  and  eventually 
develop  into  the  mature  Cercaria  condition  while  floating  in  the 
fluid  in  the  interior  of  the  Sporocyst. 

In  Ainphistoniuni  subclavaium  according  to  Loos  (9),  in  Leuco- 
chloridiiuii  para<ioxum  according  to  Heckert  (4),  and  inDistomum 
duplicatum  according  to  Reuss  (H),  the  formation  of  the  ova 
originally  takes  place  at  any  point  in  the  wall  of  the  Sporocyst — 
a  definite  ovary  only  becoming  established  at  a  later  stage. 
Whether  this  may  hold  good  in  the  case  of  the  Sporocyst  from 
Mytilus  lalus  with  regard  to  the  primary  Sporocyst  developed 
from  the  Miracidium  there  are  no  data  from  which  to  decide.  I 
have  found  no  Sporocysts,  however  small,  among  the  hundreds 
examined  in  which  the  formation  of  ova  w^as  not  localised  in  a 
single  ovary;  but  I  may  not  have  seen  any  except  those  formed 
secondarily  by  fission  or  endogenous  formation,  or  at  all  events 
may  not  have  happened  to  meet  with  any  early  primary  Sporocysts. 

The  cells  of  the  germinal  epithelium  give  rise,  however,  not 
only  to  embryo  Cercarise,  but  sometimes  also,  though  very  rarely 
so  far  as  m}'  specimens  are  concerned,  to  a  new  generation  of 
Sporocysts.  These  become  set  free  in  the  interior  of  the  parent 
Sporocyst.  From  their  germinal  epithelium,  before  they  escape 
to  the  exterior,  embryos  of  Cercariae  msiy  already  have  become 
formed.  The  young  Sporocysts  (fig.  3)  on  escaping  readily 
multiply  by  binar}^  fission  (fig.  4):  a  constriction  appears  dividing 
the  Sporocyst  into  two  equal  or  slightly  unequal  parts,  and  by 
the  deepening  of  the  constriction  the  two  parts  become  eventually 
completely  severed. 

In  young  Sporocysts  the  germinal  epithelium  is  of  unifornt 
thickness  throughout  ;  but  at  a  very  earl}^  stage  an  anterior 
extremity  capable  of  involution  becomes  differentiated,  after 
which,  the  cyst  increasing  in  length,  the  germinal  epithelium  soon 
comes  to  be   attenuated   except  at   the  anterior   and   posterior 


502  ON    TWO    REMARKABLE    SPOROCYSTS, 

extremities.  In  young  Sporocysts  the  internal  cavity,  owing  to 
the  small  size  of  the  cyst  and  the  thickness  of  the  germinal 
epithelium,  is  extremely  small.  Occasionally  in  sections  of  later 
as  well  as  earlier  stages  a  delicate  reticulated  substance  appears 
to  till  this  cavity  \vholly  or  partially  :  but  there  can  be  no  doubt 
that,  at  least  so  far  as  the  later  stages  are  concerned,  this  is  an 
entirely  artificial  product  formed  as  a  result  of  the  coagulation 
of  the  fluid  contained  in  the  cyst. 

The  Mature  Cercari^. 

The  fully  developed  Cercaria  (fig.  9)  reaches  a  total  length  of 
about  4:  mm.,  the  tail  being  included.  The  head  or  part  of  the 
body  bearing  the  anterior  sucker  is  quite  distinctly  marked  off 
from  the  rest,  being  much  narrower.  On  its  ventral  aspect  is  the 
large  cup-shaped  anterior  sucker,  which  is  capable  of  being  com- 
pletely everted.  Around  the  anterior  margin  of  the  head  on  the 
dorsal  side,  and  usuall}'^  inclined  outwards  and  backwards,  are  a 
series  of  flattened  tooth-like  spines  which  are  capable  of  being 
turned  forwards.  E3'^es  are  not  present.  At  the  sides  of  the 
head  region  are  a  series  of  six  nearly  transverse  ridges  which  do 
not  extend  on  to  the  dorsal  or  ventral  surfaces.  The  remainder 
of  the  body  is  of  oval,  dorso-ventrally  compressed,  form;  in  front 
of  the  middle  is  the  posterior  sucker. 

The  tail  consists  of  a  main  trunk  and  two  branches.  When 
extended  to  its  utmost  the  former  is  over  three  times  the  length 
of  the  body;  the  branches  are  nearly  as  long  as  the  trunk.  The 
latter  expands  proximally  where  it  joins  the  body  into  a  disc,  the 
margin  of  which  is  divided  by  notches  and  fissures  into  a  series 
of  branched  lobes.  I  have  not  seen  this  peculiar  structure  per- 
forming any  function;  but  it  can  hardly  be  an3^thing  else  than  an 
organ  of  adhesion.  It  contracts  and  expands  slowly  at  irregular 
intervals.  The  main  axis  of  the  tail,  when  at  rest,  is  not  in  line 
with  the  body,  but  bent  towards  the  ventral  side.  It  performs 
extremely  rapid  vibrating  movements,  propelling  the  larva  swiftly 
through  the  water.      The  tail  has  a  remarkable  segmented  appear- 


BY    PROFESSOR    W.    A.    HASWELL.  503 

ance,  due  to  the  presence  of  the  regular!}'  arranged  transverse 
bundles  of  muscular  fibres  referred  to  below. 

The  body  is  enclosed  in  a  cuticle  0-003  mm.  in  thickness. 
Beneath  this  is  a  muscular  layer  consisting  of  external,  longitudi- 
nal, and  internal  circular  fibres.  The  space  between  the  body-wall 
and  the  enteric  canal  is  tilled  with  parenchyma,  in  which  are 
numerous  large  cells  about  0*04  mm.  in  diameter  —  the  cystogenic 
cells.  In  young  specimens  these  contain  a  coarse  reticulum. 
In  mature  specimens  they  appear  more  homogeneous.  Their 
ducts  (processes  of  the  cells)  branch,  and  the  branch  ducts  per- 
forate the  cuticle  to  open  over  the  entire  surface  of  the  body. 
There  are  no  cystogenic  cells  in  the  tail.  No  rhabdites  were 
observed  in  any  part. 

The  pharynx  (fig.  10)  is  a  relatively  long  cylinder.  The  region 
of  the  digestive  canal  following  upon  this  behind,  commonly 
known  as  the  oesophagus,  is  very  short.  There  is  nothing  special 
to  be  noted  with  regard  to  the  structure  of  the  various  parts  of 
the  digestive  system.  The  intestine  is  fully  developed  long  before 
the  Cercaria  becomes  mature. 

The  tail  (figs.  11  and  12)  is  a  hollow  cylinder  with  a  thin  wall, 
tensely  filled  with  a  watery  fluid.  It  has  a  thin  cuticle,  beneath 
which  is  a  single  layer  of  circular  muscular  fibres  of  extreme 
fineness.  Internal  to  this  are  four  bundles  of  longitudinal 
muscular  fibres,  two  dorsal  and  two  ventral.  On  each  side  is  a 
fairly  wide  interval — the  lateral  line  as  it  may  conveniently  be 
termed — between  the  dorsal  and  the  ventral  bundle.  Between 
the  dorsal  and  ventral  bundles  on  each  side  run  a  number  of 
vertical  (dorso-ventral)  fibres  for  the  most  part  in  narrow  strands 
occasionally  with  nuclei.  These  are  arranged  at  intervals  along 
the  tail  in  such  a  way  as  to  produce  in  some  cases  an  appearance 
of  segmentation.  Along  each  lateral  line,  immediately  below  the 
cuticle  and  circular  muscle,  runs  a  delicate  filament.  This  gives 
off  branches  which  run  both  dorsally  and  ventrally,  frequently 
dividing  into  finer  threads.  At  several  points  in  its  course  the 
main  filament  presents  a  thickening.  Accompanying  this  is  a 
very  regular  row  of  small,  usually  rounded,  cells  of  a  peculiarly 


504  ON    TWO    REMARKABLE    SPOROCYSTS, 

clear  and  bright  appearance  in  the  living  condition,  so  that  they 
are  very  conspicuous  even  under  a  low  power.  These,  which  are 
separated  from  one  another  by  very  regular  intervals,  are  all 
connected  with  the  lateral  filament,  usually  by  means  of  side- 
branches,  sometimes  (rarely)  directly  with  the  main  filament  itself, 
which  then  seems  to  pass  through  them.  There  is  no  direct 
evidence  of  the  nature  of  these  filaments  of  the  lateral  line;  but 
from  the  way  in  w^hich  they  branch  I  have  very  little  doubt  that 
they  ai-e  nerves.  If  this  should  prove  to  be  correct,  the  regularly- 
arranged  cells  connected  with  them,  though  they  have  no  processes 
projecting  on  the  exterior,  are  probably  sense-cells  of  some  sort. 

Applied  to  the  inner  surfaces  of  the  bundles  of  longitudinal 
muscular  fibres  are  a  number  of  very  large  granular  cells  about 
0-04  mm.  in  diameter,  with  vesicular  nuclei  O'Ol  mm.  in  diameter. 
As  shown  by  their  relations  and  their  developmental  history 
these  are  the  myoblasts  of  the  longitudinal  muscular  fibres. 

The  branches  of  the  tail  resemble  the  tail  proper  in  structure, 
except  that  the  transverse  fibres  are  more  irregularly  arranged  and 
closer  together.  In  the  lobed  disc  at  the  proximal  end  of  the 
trunk  the  bundles  of  longitudinal  fibres  become  spread  out  so 
as  to  be  inserted  into  the  lobes.  The  dorso-ventral  bundles  are 
here  represented  by  thick  columnar  cells. 

Not  much  attention  appears  to  have  been  paid  to  the  structure 
of  the  tail  in  Cercaria;  but,  so  far  as  I  am  able  to  judge,  the 
form  at  present  under  consideration  would  appear  to  be  excep- 
tional in  this  respect,  especially  with  regard  to  the  entire  absence 
of  any  axial  strand.  Thus  Leuckart  states  (6,  p.  512):  "Unterhalb 
der  Cuticula  erkennt  man  deutliche  Rings-  und  Liingsmuskeln 
und  das  nicht  bios  am  Rumpfe,  sondern  auch  am  Schwanze,  nur 
dass  sich  letztere  hier  gewuhnlich  rechts  und  links  zu  breiten 
Bandern  gruppiren,  und  somit  eine  Anordnung  zeigen  die  man 
bei  aufmerksamerBetrachtung  schon  aus  der  entschieden  seitlichen 
Beweoungen  des  Schwanzes  erschliessen  konnte.  Die  Zellen  des 
Achsenstranges  sind  in  eine  glashelle  Bindesubstanz  verw^andelt 
die  nur  noch  einzelne  spindelformige  Korper  erkennen  lasst  und 
nach  Art  eines  (unvollstiindig  erhiirteten)  Skelettes  dazu  dienen 


BY    PROFESSOR    AV.    A.    HASWELL.  505 

mochte,  den  Schwanz  elastisch  zu  machen  und  ihm  einen  gewissen 
Grad  von  Rigiditat  zu  verleihen.  Zwischen  Muskeln  und  Achsen- 
strang  liegt  eine  Schicht  von  grossen  platten  Zellen,  die  bei 
Cercaria  macrocerca  bei  welcher  der  Achsenstrang  zu  eines 
machtigen  Entwickelung  heranwiichst,  sehr  sonderbar  veriistelte 
Formen  haben." 

Here  it  will  be  observed  the  axial  strand  is  described  as  being 
composed  of  cells,  which  have  become  converted  into  a  hyaline 
connective  tissue  acting  as  the  main  support  of  the  organ. 

The  structure  of  the  tail  in  Bucephalus  is  described  b}''  Ziegler 
(18),  but  as  this  is  by  no  means  a  typical  Cercaria  the  statements 
which  he  makes  probably  do  not  hold  good  generally;  they  are 
certainly  entirely  inapplicable  to  the  form  Avhich  I  am  now 
describing. 

Thiry  (13)  states  that  in  Cercaria  macrocerca  the  tail  has  an 
external  circular  layer  of  muscle.  The  longitudinal  fibres  run  in 
two  broad  bands  at  the  sides.  Beneath  this  is  a  mass  which 
anteriorly  consists  of  large  cells,  posteriorly  appears  homogeneous. 

Schwarze's  account  (12)  of  the  structure  of  the  tail  in  Cercaria 
armata  is  also  not  in  any  way  in  agreement  with  what  is  to  be 
observed  in  the  Cercaria  from  Mytilus.  He  describes  the  con- 
tractile substance  of  the  tail  as  forming  an  axial  strand  surrounded 
by  a  layer  of  cj'-stogenic  cells,  and  these  again  by  a  further  layer 
of  contractile  substance,  the  outer  layer  being  connected  with  the 
axial  strand  by  numerous  radiating  bundles. 

Reuss  (11,  p.  378)  states  that  in  the  Sporocyst  of  Distomum 
duplicatum  the  tail  has  a  layer  of  cells  underlying  the  cuticle, 
and  the  interior  is  traversed  by  transverse  muscular  fibres. 

It  does  not  seem  to  be  possible  to  reconcile  these  various  state- 
ments, and,  if  they  are  correct,  there  must  be  considerable  diversity 
in  the  structure  of  the  tail  in  different  Cercariae. 

In  the  posterior  part  of  the  body  opening  near  the  root  of  the 
tail  is  the  excretory  bladder.  This  is  broad  behind,  narrower  in 
front,  where  it  becomes  forked.  Each  of  the  branches  terminates 
anteriorly  in  a  slight  enlargement  into  which  opens  the  corres- 


506  ON    TWO    REMARKABLE    SPOROCYSTS, 

ponding  main  excretory  vessel,  right  or  left  as  the  case  ma}^  be. 
Each  of  these  runs  outwards  and  forwards  to  a  point  near  the 
margin  of  the  body,  where  it  divides  into  an  anterior  and  a 
posterior  longitudinal  trunk.  The  ciliated  funnels  are  0-0075  mm. 
in  diameter.  There  are  always  twelve  of  them  in  mature  Cercariae, 
and  their  arrangement  is  quite  constant.  One  funnel  terminates 
each  of  the  anterior  and  posterior  longitudinal  vessels  ;  the 
remainder  are  situated  at  the  ends  of  longer  or  shorter  side- 
branches  in  the  way  shown  in  fig.  10. 

No  funnels  and  no  vessels  were  observed  in  the  tail.  In  this 
respect  this  Cercaria  differs  apparently  from  other  described 
forms.  Wagener  (17)  states  that  in  the  Cercariee  with  forked  tail 
the  excretory  bladder  extends  into  the  tail  and  becomes  bifurcate, 
each  branch  opening  at  the  end  of  the  corresponding  branch  of 
the  tail;  and  Loos  (7)  describes  and  figures  the  extension  into  the 
tail  of  the  main  excretory  vessel  in  the  Cercaria?  of  Amphistomum 
suhclavatum. 

The  reproductive  system  is  fairly  well  advanced  in  the  mature 
Cercaria.  The  ovary  is  a  rounded  mass  of  cells  situated  a  little 
behind  the  point  of  bifurcation  of  the  intestine.  From  it  the 
oviduct  proceeds  as  a  narrow  tube  which  runs  forwards  and 
towards  the  left  passing  on  the  ventral  side  of  the  left  ramus 
of  the  intestine,  and  bending  in  again  tow^ards  the  middle  line  to 
open  at  the  genital  aperture  immediately  in  front  of  the  ventral 
sucker.  A  rounded  mass  surrounding  the  oviduct  shortly  after 
it  leaves  the  ovary  is  evidently  the  shell-gland.  The  rudiments 
of  the  testes  are  two  small  oval  bodies  situated  between  the  rami 
of  the  intestine  some  distance  behind  the  ovary.  The  vitelline 
glands  are  not  yet  distinguishable.  A  rounded  bod}^  dorsal  to 
the  anterior  part  of  the  ventral  sucker  is  probably  the  vesicula 
seminalis. 

In  the  form  and  arrangement  of  the  spines  around  the  anterior 
end,  the  series  of  lateral  ridges  in  the  region  immediately  follow- 
ing, and  the  form  of  the  phar3mx,  this  Cercaria  resembles  the 
adult  Distomum  fasciatum,  Rud.,  and  probably  is  the  larva  of  a 


BY    PROFESSOR    W.    A,    HASWELL.  507 

form  nearly  related  to  that  species."^  In  the  first  point  it  also 
resembles  D.  acanthocephalam  and  other  members  of  Rudolphi's 
sub-genus  Echinostomum. 

Development  of  the  Cercari^. 

For  what  we  know  of  the  development  of  the  Cercaria  within 
the  Sporocyst  or  Redia  we  are  mainly  indebted  to  Leuckart  (17), 
Thomas  (14),  Heckert,  Schwarze  (12),  and  Loos  (9)-  In  spite  of 
their  labours,  however,  it  must  be  said  that  a  good  many  points 
remain  obscure.  This  holds  good  particularly  with  regard  to  the 
precise  mode  of  origin  of  certain  of  the  organs;  and  it  is  due 
mainly  to  the  almost  complete  absence  of  differentiation  among 
the  cells  (meristem  cells  of  Schwarze  and  others)  of  the  earl}- 
stages,  the  rudiments  of  various  parts  having  the  appearance  of 
becoming  quite  suddenly  crj^stallised  out  from  a  previousl}^  homo- 
geneous mass. 

As  already  stated,  the  embryos  of  the  Cercaria  from  Mytilus 
latns  are  set  free  from  the  ovary  when  they  have  attained  a 
diameter  of  about  0*025  mm.*  At  about  this  stage  (figs.  15  and  16) 
the  embryonic  investment  is  formed.  First  a  single  cell  on  the 
surface  becomes  flattened  out,  then  others  become  similarl}^ 
modified  until  the  spherical  mass  of  cells  becomes  enclosed  in  a 


*  I  infer  this  from  Braun's  reproduction  (3,  taf.  xxi.,  fig,  8)  of  a  figure  by 
Stossich,  the  original  paper  by  that  author  (in  the  Bolletino  della  Soc.  adriat. 
scienze  natur,  Trieste,  vol  ix.,  1885)  not  being  accessible  to  me. 

*  Reuss  (11)  met  frequently  with  a  stage  in  which  one  large  cell  (ovum) 
was  accompanied  by  three  considerably  smaller  cells,  with  nuclei  0-0025  in 
diameter  with  uniformly  distributed,  coarsely  granular  chromatin.  Since  the 
large  cell  next  divides  into  quite  equal  and  similar  cells  he  takes  these 
previously  formed  three  smaller  cells  to  be  of  the  nature  of  polar  bodies,  and 
their  formation  to  be  a  maturation  process.  This  stage  I  have  not  observed. 
But  in  a  large  proportion  of  specimens  there  occur  lying  loose  in  the  Sporo- 
cyst in  the  immediate  neighbourhood  of  the  ovary  a  varying  number  of  cells 
(fig.  14)  which  have  homogeneous  deeply  staining  nuclei  0*002  mm,  in 
diameter.  If  these  are  not  of  the  nature  of  polar  bodies  it  seems  difficult 
to  account  for  them. 


508  ON    TWO    REMARKABLE    SPOROCYSTS, 

thin  layer  composed  of  a  small  number  of  flattened  cells  which 
become  completely  united  at  their  edges.  This  investment 
persists  to  a  comparatively  late  stage  in  the  development  of  the 
Cercaria,  onl}'  disappearing  as  the  cuticle  becomes  formed. 

Schwarze's  account  (13)  of  the  development  of  the  investing 
layer  in  Cercaria  armala  agrees  closely  with  what  I  have  observed. 
I  cannot,  however,  follow  him  in  his  opinion  that  the  embryonic 
investment  gives  rise  to  the  cuticle  of  the  Cercaria.  The  invest- 
ment in  question  is  an  excessively  thin  membrane  which  at  no 
time  contains  more  than  about  half-a-dozen  nuclei.  As  the 
embryo  grows,  this  simph''  becomes  stretched,  and  in  early  tailed 
stages  appears  quite  loosely  connected  with  the  underlying  la3^ers. 
I  have  very  little  doubt  that  it  is  the  laj^er  of  cells  which,  at  the 
period  when  the  tail  is  being  formed,  becomes  arranged  in  a  regular 
manner  beneath  the  original  investing  layer,  that  secretes  the 
cuticle. 

Heckert"^  states  that  in  embryo  Cercarise  of  a  diameter  of 
0-075  mm.  a  second  investment  similar  to  the  first  appears  beneath 
it;  and  Loos  (9)  observed  a  similar  phenomenon  in  the  develop- 
ment of  Rediae  (letter  to  Braun  quoted  in  foot-note,  3,  p.  818). 
There  is  no  appearance  of  this  second  investing  layer  in  the  case 
of  the  Cercaria  from  Mytilus  latus. 

Small  nuclei  with  deeplj^-staining,  apparently  homogeneous,' 
plasma,  similar  to  those  described  by  Schwarze  (10)  in  Cercaria 
armata  as  giving  rise  to  the  rudiment  of  the  reproductive  system, 
appear  before  the  embryo  leaves  the  ovary.  But  these  have  no 
definite  arrangement,  occurring,  frequently  two  together,  here 
and  there,  both  near  the  surface  and  more  deeply,  instead  of 
forming  a  central  mass  as  stated  by  Schwarze.  They  are  most 
probably  merely  phases  in  the  histor}'  of  the  ordinary  meristem 
nuclei,  and  have  no  special  histogenetic  significance. 


*  I  only  know  this  paper  ('  Untersuchungen  iiber  die  Entwickelungs-  und 
Lebensgeschichte  des  Distomum  macrostomum.'  '  Bibliotheca  Zoologica,' 
Leuckart  u.  Chun,  Heft  4),  as  quoted  by  Braun  (3). 


BY    PROFESSOR    W.    A.    HASWELL.  509 

As  the  embryo  increases  in  size,  it  becomes  somewhat  oval,  and 
soon,  when  the  long  diameter  is  about  0-08  mm.,  the  future 
posterior  end  becomes  marked  by  a  slight  depression,  the  anterior 
end  remaining  rounded  (Hg.  17).  In  this  position  a  narrow  cleft 
appears  running  forwards  towards  the  centre  of  the  embryo. 
This  cleft  does  not  open  on  the  exterior,  but  is  covered  over  by 
the  thin  investing  layer  of  cells.  On  either  side  of  it  is  a  mass 
of  cells  from  which  the  tail  buds  out;  in  front  it  terminates  in  a 
slight  transverse  enlargement  which  partly  separates  off  a  dense 
central  mass  of  cells  from  two  lateral  masses.  The  enlargement 
subsequently  becomes  the  bladder  of  the  excretory  sj^stem.  In 
the  lateral  masses  the  rami  of  the  intestine  are  developed  at  a 
later  stage. 

The  central  mass  of  cells  is  distinguishable  (figs.  18  and  19) 
into  an  anterior  and  a  posterior  portion  which  are  not  sharply 
marked  off  from  one  another.  The  posterior  portion  is  the  rudi- 
ment of  the  reproductive  system.  Of  the  anterior  portion  the 
more  ventrally  placed  cells  form  the  rudiment  of  the  pharynx, 
the  more  dorsally  placed  that  of  the  brain.  Of  these  two  organs 
the  pharynx  appears  first  as  a  cylindrical  cord  of  cells  soon 
distinguishable  (fig.  20)  into  an  investing  layer  and  a  central 
core.  The  cells  of  the  investing  layer  are  the  myoblasts  of  the 
wall  of  the  organ.  They  become  enlarged,  and  increase  in 
number,  though  never  becoming  very  numerous.  In  their  plasma 
are  developed  the  radial  muscular  fibres  of  the  pharynx.  The 
enclosed  row  of  cells  do  not  persist.  They  at  first,  fill  the  entire 
lumen,  but  spaces  appear,  and  the  cells  gradually  become  absorbed 
and  eventually  entirely  disappear  (fig.  22).  Whether  they  give 
rise  to  the  thin  cuticle  lining  the  interior  of  the  pharynx  is 
uncertain.  It  is  difficult  to  understand  by  what  other  agency 
that  membrane  could  be  formed. 

At  the  anterior  end,  towards  the  ventral  surface,  when  the 
rudiment  of  the  pharynx  first  becomes  formed,  a  rounded  group 
of  cells  in  all  respects  similar  to  the  rest  becomes  enclosed  by  a 
thin  membrane  formed  of  flattened  cells.  This  is  the  rudiment 
of  the  anterior  sucker. 
34 


510  ON    TWO    REMARKABLE    SPOROCYSTS, 

A  continuation  backwards  of  the  cord  of  cells  which  forms  the 
rudiment  of  the  pharynx  constitutes  the  rudiment  of  the  short 
oesophagus.  This  soon  bifurcates  posteriorly  to  form  the  biramous 
intestine.  In  the  latter  the  central  cells  of  the  cord  become 
enclosed,  as  in  the  case  of  the  pharynx,  in  a  cylinder  formed  of 
the  peripheral  cells.  But  the  latter  layer  remains  very  thin, 
developing  into  the  thin  layer  of  muscle  of  the  wall  of  the 
intestine,  while  the  enclosed  cells,  few  and  irregularl}^  disposed 
at  first  (fig.  23),  multiply  rapidly,  at  the  same  time  becoming 
disposed  as  a  regular  epithelial  layer  (figs.  24-26)  bounding  the 
lumen.  In  the  cesophagus  the  muscular  investing  laj'er  also 
remains  thin,  but  the  enclosed  cells  give  rise  to  a  layer  devoid  of 
nuclei,  but  divided  into  numerous  minute  bodies  somewhat  smaller 
than  the  cells  of  the  epithelium  of  the  intestine. 

In  Cercaria  ao'mata,  Schwarze  does  not  refer  to  the  formation 
of  a  cleft  at  the  anterior  end  preceding  the  development  of  the 
unpaired  part  of  the  enteric  cavity;  but  describes  the  latter  as 
appearing  in  the  form  of  a  solid  process  which  only  develops  a 
lumen  at  a  later  stage  by  the  absorption  of  the  axial  cells.  In 
that  Cercaria  the  full  development  of  the  forked  intestine  only 
takes  place  after  the  Cercaria  has  become  encysted,  and  the  entire 
enteric  canal  can  be  functional  in  the  Cercaria  stage  only  very 
imperfectly,  if  at  all. 

The  rudiment  of  the  posterior  sucker  is  distinguishable  as  a 
rounded  projection  on  the  ventral  surface,  shortly  after  the  tail 
begins  to  be  developed  ;  this  consists,  like  the  anterior  sucker, 
of  a  rounded  mass  of  undifferentiated  cells  separated  internally 
from  the  rest  of   the  body  cells  bj'-  a  thin  membrane. 

When  the  rami  of  the  intestine  first  make  their  appearance,  a 
space — the  origin  of  which  has  been  already  referred  to — is 
developed  between  them.  This  is  the  median  bladder  of  the 
excretory  system.  At  first  it  has  no  definite  wall,  but,  later, 
cells  become  arranged  to  form  a  thin  epithelium  as  in  the  adult. 

When  the  cord  of  cells  destined  to  give  rise  to  the  pharynx 
first  becomes  differentiated,  a  mass  of  cells,  as  already  stated,  lies 
on  the  dorsal  side  of  it.     From  the  central  part  of  this  the  first 


BY    PROFESSOR    W.    A.    HASWELL.  511 

rudiment  of  the  nervous  system  becomes  formed.  This  appears 
as  a  bridge  of  finely  fibrillated  material  (fig.  27)  running  trans- 
versely oyer  the  pharynx  and  surrounded  by  numerous  cells — the 
rudiment  of  the  brain.  This  is  well  established  when  the  pharynx 
is  still  a  narrow  tube  with  a  thin  wall  enclosing  a  core  of  cells. 
From  the  brain  two  narrower  strands  of  similar  material  run 
backwards  at  the  sides,  each  surrounded  by  groups  of  cells;  and 
are  traceable  for  some  distance  through  the  body,  though  not 
extending  at  first  very  far  back.  Before  long  each  longitudinal 
cord  bifurcates  to  form  a  dorsal  and  a  ventral  branch.  About 
the  same  time  a  nerve  is  formed  running  forwards  from  the  brain 
towards  the  anterior  extremity. 

The  tail  (figs.  29-31)  appears  first,  when  the  embryo  is  about 
0-09  mm.  in  length,  as  a  rounded  process  consisting  of  an 
undifferentiated  mass  of  cells  covered  by  a  continuation  of 
the  thin  investing  layer.  While  still  very  short  it  becomes 
bilobed  at  the  extremity.  Before  this  takes  place  internal 
differentiation  has  begun.  A  very  regularly  arranged  line  of 
small  nuclei  becomes  ranged  along  each  side  below  the  investing 
layer;  and  the  rest  of  the  tissue  forms  a  central  core.  As  the  two 
branches  grow  out  at  the  bilobed  extremity,  their  constituent  cells 
take  on  a  similar  arrangement.  The  central  core  becomes  divided 
into  four  longitudinal  rows  of  cells — two  dorsal  and  two  ventral; 
these  are  the  myoblasts.  They  become  enlarged,  and  their  nuclei 
become  vesicular.  In  the  substance  of  their  protoplasm  external 
to  the  nuclei  appear  bright-looking  longitudinal  bands  which 
stain  deeply  with  eosin;  these  are  the  first-formed  muscular  fibres. 
They  soon  become  more  numerous,  and  as  the  tail  becomes  fully 
developed,  take  the  form  of  the  four  longitudinal  bands.  The 
small  cells  of  the  lateral  rows,  at  first  placed  close  together, 
become  separated  from  one  another  by  distinct  intervals  as  the 
tail  elongates,  and  become  the  lateral  series  of  small  cells  referred 
to  above  in  the  account  of  the  tail  of  the  adult. 

The  Sporocysts  and  Cercari^  of  a  Species  of  Gasterostomum. 

Only  one  specimen  of  the  mussel  was  observed  to  be  infested 

with  this  parasite.     It  presented  a  very  remarkable  appearance. 


512  ON    TWO    REMARKABLE    SPOROCYSTS, 

The  mantle-folds  and  the  visceral  mass  in  the  region  between  the 
adductor  muscles  had  the  appearance  of  being  provided  with  a 
ramifying  system  of  vessels  containing  a  blood-red  fluid.  These 
apparent  vessels,  when  examined  under  the  microscope,  were 
found  to  be  narrow  ramifying  Sporocysts,  the  germinal  epithelium 
of  which  contained  a  red  colouring  matter  apparently  identical 
with  that  occurring  in  the  SjDorocyst  described  in  the  first  part 
of  this  paper. "^  In  these  Sporocysts  were  mature  Cercarise  and  all 
stages  in  their  development.  The  Cercari?e  proved  to  belong  to 
the  remarkable  form  known  as  Buceiji]ialus^  v.  Baer,  the  larva  of 
G aster ostomum.  All  of  these  are  developed  in  narrow,  tubular, 
usually  branching  Sporocysts;  but  in  no  case  hitherto  recorded, 
so  far  as  I  am  aware,  is  a  red  colouring  matter  present. 

The  Cercarise  (fig.  32)  are  relatively  small,  being  only  0*25  mm. 
in  length  when  fully  matured.  They  are  sluggish  in  their  move- 
ments, and  the  tail  was  never  observed  to  be  used  as  a  swimming 
appendage,  its  only  movements  being  slow^  waving  ones,  or  such 
as  resulted  in  each  of  the  branches  being  coiled  into  a  spiral  and 
uncoiled  again.  Such  spiral  movements  were  noticed  by  v.  Baer 
in  Bucephalus  p)oIyinorphus.  It  seems  likely  that  the  tail  is  used 
more  for  attachment  than  as  an  organ  of  active  locomotion. 

The  body  is  long  and  narrow.  The  anterior  portion,  lodging  the 
anterior  sucker  or  proboscis,  is  separated  off  from  the  rest  by  a  slight 
constriction.  Its  anterior  end  (figs.  33-36)  varies  in  appearance 
according  to  its  condition.  When  somewhat  contracted  it  usually 
appears  trilobed;  when  it  is  more  extended  the  rounded  aperture 
of  the  sucker  appears  at  its  extremity.  The  anterior  extremity 
is  beset  with  excessively  minute  cuticular  spinules  which  gradually 
decrease  in  size  as  they  pass  backwards,  and  are  no  longer  distin- 
guishable when  the  middle  of  the  body  is  reached.  A  pair  of 
relatively  long  and  slender  cilia,  presumabl}"  sensory,  extend 
forwards  from  the  extreme  anterior  end. 


*  The  nature  of  this  coloaring  matter  has  not  yet  been  investigated.  Its 
spectum  does  not  shew  the  absorption  bands  characteristic  of  that  of  haemo- 
globin. 


BY    PROFESSOR    W.    A.     HAS  WELL.  513 

The  mouth  is  situated  on  the  ventral  surface  considerably 
behind  the  middle.  It  leads  into  a  rounded  pharynx,  from  which 
the  simple  intestine  leads  forwards  as  far  as,  or  further  than,  the 
middle  of  the  length  of  the  body.  No  contents  of  any  kind  were 
ever  observed.  Behind  the  mouth  and  pharynx  are  two  irregular 
masses  of  cells,  forming  a  slightly  uns3niimetrical  pair;  these  are 
apparently  rudiments  of  the  gonads.  Close  to  them  in  the  middle 
is  the  excretory  sac. 

The  tail  consists  of  a  basal  part  and  two  long  branches  or 
appendages.  The  former  is  somewhat  elliptical,  with  the  long 
axis  transverse,  the  anterior  border  nearly  straight,  or  at  least 
less  convex  than  the  posterior.  The  branches  are  attached  to  the 
basal  part  antero-laterally;  they  are  slender,  nearly  twice  the 
length  of  the  body,  and  taper  slightly  at  the  extremity. 

LITERATURE. 

1. — Baer,  K.  E.  von.^ — "  Beitrage  zur  Kenntniss  der  niederen  Thiere. "  Nova 

act.  Caes,  L.  C.  Nat.  Cur.     Vol.  xiii.  1827. 
2. — BiEHRiNGER,  J. — " Beltiage   zur    Anat.    u.   Entw.    der  Trematoden." 

Arb.  Zool.-zoot.  Inst,  in  Wiirzburg,  viii.  1885. 
3. — Braun,  M. — "Vermes"  of  Bronn's  '  Thierreich.' 
4. — Heckert,    G.  —  "  Zur    Naturgeschichte    des  Leucochloridium    para- 

doxum."     Zool.  Anz.  x.  1887. 
5. — Lacaze-Duthiers,  H. — "  Memoire  sur  le  Bucephale  Haime,  helminthe 

parasite    des    huitres  et  des  bucardes."       Ann.   Sci.  nat.  4  ser. 

Tome  i.  1854. 
6. — Leuckart,  R. — "  Die  Menschliche  Parasiten,"  1863. 
7. "  Zur  Entwickelungsgeschichte  des  Leberegels."     Zool. 

Anz.  1881  and  1882. 
8. "  Zur  Entwickelungsgeschichte  des  Lebo'egels."     Arch. 


f.  Naturg.  48,  1882. 
9. — Loos,  A. — "  Ueber  Amphistomum  subclavatum,  Rud.,  und  seine  Ent- 

wickelung."     Festschrift  fiir  Leuckart,  1892. 
10. — Pagenstecher,  a.  H. — "  Untersuchungen  liber  niedere  Seethiere  aus 

Cette."     Zeitschr.  f.  wiss.  Zool.  12,  1868. 
11. — Reuss — "  Beobachtungen  an  der    Sporocyste  und  Cercarie  des  Disto- 

mum  duplicatum,  Baer."     Zool.  Anz.  25,  1902. 
12. — ScHWARZE,  W.  — "Die  postembryonale  Entwickelung  der  Trematoden." 

Zeitschr.  f.  wiss.  Zool.  43,  1886. 


514  ON    TWO    REMARKABLE    SPOROCYSTS, 

13. — Thiky,  L. — "  Beitriige  zur  Kenntniss  der  Cercaria  macrocerca  Filippi." 

Zeitschr.  f.  wiss.  Zool.  10,  1850. 
14. — Thomas,  A.  P. — "  The  life-history  of  the  Liver-fluke."     Quart.  Journ. 

Micro.  Sci.  23,  1883. 
15. — Ulicxy,    J. — "  Helminthologisehe    Beitrage."       Arch,  f.    Naturg.  44 

Jahrg.  Bd.  i.  1878. 
16.— WaCtEner,  G. — "  Helminthologische  Bemerkungen."     Zeitschr.  f.  wiss. 

Zool.  9,  1858. 
17. "  Ueber  Redien  u  Sporocysten."      Arch,  f,  Anat.  u. 

Physiol.  1866. 
18. — ZiEGLEK,    H.    E. — "Bucephalus  und  Gasterostomum."      Zeitschr.  f. 

wiss.  Zool.  39,  1883. 


EXPLANATION  OF  PLATES. 

Reference  Letters. 

6r.,  brain.  c,  cuticle.  ccZ. ,  disc  at  base  of  tail.  f/.v.??i.,  dorso- ventral 
muscles  of  tail,  ex.,  principal  excretory  vessel.  ^.r. &.,  excretory  bladder. 
i.,  intestine.  I.e.,  lateral  cells  of  tail,  l.m.,  longitudinal  muscles  of  tail. 
Z.n.,  lateral  nerve  of  tail,  m.,  mouth,  w./.,  muscular  fibres  developing  in 
myoblasts,  my.,  myoblasts,  o.d.,  oviduct,  ce.,  cesophagus.  ov.,  ovary, 
jp^.,  pharynx,  r.,  rudiment  of  gonads,  .s'.,  anterior  sucker,  s". ,  posterior 
sucker,     sp. ,  spines,     te. ,  testes. 


Plate 


XIX. 


-Sporocyst  from  Mytilus  latus,  magnified. 

-Sporocyst,  entire  preserved  and  stained  specimen,  somewhat  com- 
pressed ( X  75). 
-Young  stage  of  Sporocyst,  compressed  (  x  75). 
-Young  Sporocyst  in  the  act  of  undergoing  fission  (  x  75). 
-Ovary  :  appearance  in  entire  specimen  (  x  1000). 
-Ovary:  section,  showing  ova  and  early  stages  (  x  1000). 
-Ovary  :  section  next  to  that  represented  in  fig.  6. 
-Wall  of  Sporocyst :  vertical  section  of  one  of  the  thickened  regions 
of  the  germinal  epithelium  (  x  1000). 
Fig.    9. — Cercaria,  magnified.     The  tail  is  here  not  so  long  in  proportion  as 

it  is  destined  to  become. 
Fig.  10. — Body  of  Cercaria  (with  base  of  tail),  showing  excretory  and  diges- 
tive systems.     The  posterior  sucker  is  not  shown. 
Fig.  11.  — Transverse  section  of  tail  (  x  750). 

Fig.  12. — Lateral  view  of  a  portion  of  the  tail,  showing  the  lateral  line  and 
its  nerve  (  x  340). 


Fig. 

1.- 

Fig. 

2. 

Fig. 

3.- 

Fig. 

4. 

Fig. 

5. 

Fig. 

6. 

Fig. 

7. 

Fig. 

8. 

BY    PROFESSOR    W.    A.    HASWELL.  515 

Fig.  13. — Two  cystogenic  cells  (  x  950). 

Fig.  14. — Cells  supposed  to  be  possibly  of  the  nature  of  polar  bodies  (  x  1500)* 

Fig.  15,— Early  free  stage  (  x  1000). 

Plate  XX. 

Fig.  16. — Somewhat  later  stage  than  that  represented  in  fig.  15  (  x  950). 

Fig.  17.  —  Stage  with  flattened  posterior  end  (  x  1000). 

Fig.  18.— Stage  with  clefts  and  rudiments  of  pharynx  and  reproductive 
system  :  entire  specimen  in  optical  section. 

Fig.  19. — Longitudinal  section  of  similar  stage  :  combined  from  two  adjoin- 
ing sections  ( x  750). 

Fig.  20. — Transverse  section  of  pharynx  at  the  stage  in  which  its  wall  has 
become  well  established  but  the  musculature  has  not  become 
formed  (x700). 

Fig.  21. — Transverse  section  of  pharynx  at  a  somewhat  later  stage  :  the  cells 
still  intact  in  the  interior. 

Fig.  22. — Somewhat  later  stage  in  which  the  cells  in  the  lumen  have  disap- 
peared and  the  cuticle  has  become  developed  (  x  700). 

Fig.  23. — Transverse  section  of  early  Cercaria  through  the  posterior  region, 
showing  the  intestinal  epithelium  in  the  condition  of  irregularly 
distributed  cells  enclosed  within  the  developing  muscular  layer 
( x  700). 

Fig.  24.  —Transverse  section  of  one  of  the  rami  of  the  intestine  at  the  stage 
in  which  the  internal  cells,  though  still  few  in  number,  have 
become  arranged  in  a  definite  layer  (  x  1500). 

Fig.  25. — Section  similar  to  that  represented  in  fig,  24,  but  of  a  somewhat 
later  stage,  in  which  the  epithelium  has  become  more  fully 
developed  (  x  1500). 

Fig.  26.  — Section  of  a  small  part  of  the  epithelium  of  the  intestine  in  the 
mature  Cercaria,  to  show  the  form  and  arrangement  of  the  cells 
(X  1500). 

Fig.  27.  —Section  through  Cercaria  in  the  region  of  the  brain  at  a  stage  when 
muscular  tissue  is  only  beginning  to  be  formed  (  x  750). 

Fig.  28. — Tail  at  an  early  stage  in  its  development :  viewed  as  a  transparent 
object  (  X  750). 

Fig.  29. — Similar  view  of  later  stage. 

Fig.  30. — Transverse  section  of  tail  at  a  stage  corresponding  to  that  repre- 
sented in  fig.  23  (  x  700). 

Fig.  31. — Transverse  section  of  later  stage  in  which  the  cuticle  has  become 
formed  and  the  formation  of  the  muscular  fibres  has  begun. 

Fig.  32. — Cercaria  stage  {Bucephalus)  of  a  species  of  Echinostomum  (  x  240). 

Figs.  33-36.— Outline  of  head  end  of  living  Bucephalus  in  different  conditions. 


516 


6.— ON  EUCALYPTUS  BICOLOR,  A.  Cunn.* 
By  J.   H.  Maiden. 

E.  bicolor,  A.  Cunn.,  was  first  alluded  to  in  a  published  work 
in  the  following  passage  : — 

"  E.  hicoloi\  A.  Cunn.,  MS.,  a  species  closely  allied  to  E. 
hcemastoma,  Sm.,  but  the  marginal  nerve  is  not  so  close  to  the 
edge  of  the  leaf  (this  is  the  '  Bastard  Box '  of  the  carpenters) " 
(Hooker  in  Mitchell's  '  Journ.  Trop.  Australia,'  390,  1848).  I 
hardly  think  this  can  be  taken,  by  itself,  as  an  adequate  descrip- 
tion for  purposes  of  priority. 

I  have,  however,  examined  the  following  specimens,  and  it 
becomes  a  matter  for  consideration  as  to  how  far  herbarium  labels 
shall  be  taken  as  evidence  of  publication.  I  have  seen  all  the 
specimens,  and  as  regards  those  in  Cunningham's  handwriting,  it 
may  be  pointed  out  that  he  died  in  1839  :  — 

1.  "  Eucalyptus  bicolor"  in  A.  Cunningham's  handwriting  and 
bearing  the  label  "New  Holland,  A.  Cunningham,  Hooker,  1835." 
This  specimen  was  given  b}^  Sir  William  Hooker  to  Bentham. 

2.  '' E.  bicolor,  1846.  Sub-tropical  New  Holland,  Lieut.-Col. 
Sir  T.  L.  Mitchell." 

The  above  are  from  Herb.  Kew. 

A  second  specimen  from  Herb.  Melb.  of  No.  2  labelled  "  No.  446 
of  Nov.  1846." 

There  are  two  specimens  on  one  sheet  in  Herb,  Cant,  ex  herb. 
Lindl,  both  from  sub-tropical  New  Holland,  Lieut.  Col.  Sir  T.  L. 
Mitchell,  and  both  labelled  "  E.  bicolor,  A.  Cunn.,"  by  Cunning- 
ham himself.     One  label  carries  the  additional  information  *'No. 


*  See  also  Deane  and  Maiden,  "  K.  larcjijiorem,  F.v.M.  (Syn.  E.  bicolor 
A.  Cunn.)."     These  Proceedings,  1899,  621. 


BY    J.    H.    MAIDEN.  517 

439,  Nov.  20,  1846,  'Bastard  Box  of  carpenters,'"  and  the  other 
"No.  614,  Nov.  30,  1846,  camp  86." 

Then  comes  Mueller's  very  full  description  of  E.  bicolor,  A. 
Cunn.,  in  Journ.  Linn.  Soc.  iii.  90  (1859),  Mueller  being  then 
ignorant  that  F.  bicolor,  A.  Cunn.,  was  a  synonym  of  E.  largi- 
ftorens,  F.v.M.,  described  in  1855. 

Bentham  accepted  E.  bicolor,  A.  Cunn.,  as  having  priority,  in 
B.  Fl.  iii.  214,  without  comment,  reducing^. /an/i/?ore/is,  F.v.M., 
to  a  synonym. 

Mueller's  own  quotation  of  the  sj^nonymy  is  interesting : — 
('Eucalyptographia,'  under  E.  largijlorens,  F.v.M.). 

"  E.  largifiorens,  F.v.M.  (1854);  Fragmenta,  ii.  58.  E.  peyidula, 
A.  Cunn.,  in  Steudel  (1840);  E.  bicolor,  A.  Cunn.,  in  Mitchell 
(1848)." 

He  proceeds  to  say  : — 

'*  Preference  is  here  given,  in  accordance  with  De  Candolle's 
code,  to  the  name  under  which  this  species  was  tirst  defined,  and 
chosen  as  expressive  of  the  exuberance  of  its  flowers.  Of  neither 
of  the  names  bestowed  by  Allan  Cunningham  on  this  species, 
timely  description  was  given;  the  pendulous  branches  suggesting 
the  one  name  and  perhaps  the  sometimes  but  often  pale  colour  of 
the  filaments*  giving  rise  to  the  other  unless  it  was  derived  from 
the  coloration  of  the  bark." 

I  would  point  out  that  early  descriptions  of  Eucalyptus  and 
other  plants  were  often  vague,  and  we  have  had,  in  some  cases, 
bo  appeal  to  herbarium  specimens  and  other  less  certain,  collateral 
evidence,  to  decide  what  is  intended  as  a  species.  I  frequently 
hear  that  conchologists,  entomologists  and  others  are  in  a  similar 
.situation.  I  think  it  would  place  a  dangerous  power  in  the 
hands  of  any  man  to  enable  him  to  pass  over  these  imperfect 
descriptions,  especially  when  they  are  supported,  as  in  the  present 
case,  by  authentically  named  herbarium  specimens  deposited 
under  proper  safeguards,  in  important  herbaria. 


*  The  late  Dr.  Woolls  wrote  to  me  that,  in  his  opinion,  the  name  was 
given  owing  to  the  variation  in  colour  of  the  filaments. 


518  ox  EUCALYPTUS  B [CO LOR,  A.  CUNN., 

Many  of  Sieber's  names  have  been  accepted  from  herbarium 
labels  only.  Schauer  adopted  Cunningham's  names  and  described 
the  plants,  rectifying  Cunningham's  omission.  It  is  etiquette  not 
to  suppress  a  herbarium  name  when  it  is  proved  to  be  valid,  and 
in  1859  Mueller's  act  in  fully  describing  E.  bicolor,  A.  Cunn., 
shows  that  he  had  no  wish  to  suppress  Cunningham's  name. 

I  therefore,  after  the  most  careful  consideration,  have  decided 
to  follow  Bentham  in  adopting  the  name  £J.  bicolor,  A.  Cunn. 
(B,  Fl.  iii.  214).  It  is  proper  to  state,  however,  that  this  is  a  case 
in  which,  all  evidence  being  available  in  regard  to  the  two  names 
in  question,  it  is  not  absolutely  decisive  in  regard  to  the  adoption 
of  either. 

Synonyms. 

a.  Prefatory  Note  on  E.  bicolor,  Woolls  (also  R,  T.  Baker). 

b.  Note  on  E.  parviflora,  F.v.M. 

1.  E.  bicolor.  Duff  {partim). 

2.  E.  pendula.  Page  (?). 

3.  E.  pendula,  A.  Cunn. 

4.  E,  largijiorens,  F.v.M. 

5.  E.  haniastoma,  Miq. 

a.  E.  bicolor,  Woolls  ('  Contrib.  Flora  of  Australia,'  232);  also 
R.  T.  Baker. 

In  the  WooUsian  herbarium,  which  is  m}'  property,  there  is  a 
specimen  in  Dr.  Woolls'  handwriting  bearing  the  following 
label  :— 

"  Yellow  or  Bastard  Box.  Half-barked  when  young,  nearly 
smooth  when  full  grown.  Hard  wood.  Height  120  feet.  Cab- 
ramatta.      E.  bicolor. '' 

On  another  occasion  Dr.  Woolls  labelled  a  similar  specimen 
from  Cabramatta  ''E.  larcfi/lorens." 

There  is  no  question  as  to  the  identity  of  this  tree,  even  if  his 
specimens  did  not  make  it  quite  clear.  It  is  E.  Bosistoana,  F.v.M., 
is  typical  for  the  species,  as  determined  by  Mueller  himself,  and 
the  assumption  that  Woolls'  determination  of  the  tree  as  E.  bicolor 
was  correct  has  given  rise  to  the  following  ingenious  mistakes. 
(It  will  be  shown  that  E.  i^endxila  is  a  nomen  nudum). 


BY    J.    H.    MAIDEN.  519 

"  In  many  instances  it  is  impossible  to  classify  Eucalypts 
on  the  shape  of  fruits,  anthers,  buds,  and  leaves,  and  in  this  con- 
nection is  mentioned  the  case  of  E.  bicolor*  and  E.  pendulaj  of 
A.  Cunningham.  It  has  been  customar}-  in  recent  times  to 
synonymise  these  species  under  the  name  of  ^.  largijlorens,  F.v.M. 
Now  Cunningham,  who  was  a  field  botanist,  and  who  was  familiar 
with  these  trees,  named  the  bastard  box  of  Cabramatta  E.  hicolor* 
a  tree  with  a  dark  box  bark  on  the  stem,  and  with  clear  ivhite 
limbs,  and  having  a  light  brown-coloured  timber,  whilst  the 
"  Coolabah "  of  the  interior  he  named  E.  jjendula.i  from  its 
drooping  habit.  This  tree  has  a  red-coloured  timber,  and  a  box- 
bark  extending  to  the  ultimate  branches.  The  oils  of  the  two 
trees  are  also  quite  distinct.  The  economic  and  systematic 
materials  of  E.  pendula  have  been  obtained  from  many  parts  of 
the  colony,  and  show  the  usual  constancy  of  specific  characters 
which  the  author  has  found  to  hold  in  almost  all  other  Eucalyptus 
species.  This  also  applies  to  E.  bicolor,  and  on  these  grounds  it 
is  contended  that  the  two  trees  should  be  regarded  as  distinct 
species.  The  only  resemblance  is  the  venation  of  the  lanceolate 
form  of  leaf.  If  placed  under' E.  largiJlo7'ens,l  then  there  would 
be  the  anomaly  of  having  under  one  species  a  tree  with  two  kinds 
of  bark,  two  kinds  of  timber,  two  kinds  of  oil,  and  a  variation  in 
leaves"  (R.  T.  Baker,  Proc.  A.A. A.S.  Melbourne,  1900,  p.  230). 

Again  the  same  writer  states  :  — 

''  I  am  much  indebted  in  this  instance  to  the  writings  of  the 
late  Dr.  Woolls  for  finding  the  particular  tree  of  E,  bicolor.f  In 
his  '  Contributions  to  the  Flora  of  Australia '  (p.  232)  he  gives  the 
locality  Cabramatta,  where  will  be  found  trees  that  exactlj^ 
coincide  with  Cunningham's  description  of  E.  bicolor,  and  in  no 
way  agree  with  E.  largiflnrens,  F.v.M.  {E.  pendula,  A.  Cunn.f), 
of  the  interior.  I  and  others  have  now  seen  both  trees  in  the 
field  and  agree  that  the  two  are  quite  distinct,  and  Cunningham 

''  E.  Bosistoaiia,  F.v.M. 

t  E.  bicolor,  A.  Cunn. 

X  E.  bicolor,  A.  Cunn.,  is  a  synonym  of  E.  largijlorens,  F.v.M. 


520  ON  ECCALYPTUS  B  ICO  LOR,  A.   CUNN., 

was  quite  justified  in  making  two  species,  viz.,  E.  pendida,  'Red 
Box,'  and  E.  bicolor,  '  Bastard  Box.' 

'*  This  latter  species  occurs  all  along  the  banks  of  the  South 
Creek"  (These  Proceedings,  xxv.,  666.) 

b.   Note  on  E.  parvijiora,  F.v.M. 

This  is  a  name  only  given  as  a  synonym  of  E.  hicolor,  A.  Cunn., 
(in  Journ.  Linn.  Soc.  iii.  90)  and  it  is  referred  to  here  in  order 
that  it  may  be  cleared  up.  It  is  the  same  as  E.  bicolor,  A.  Cunn., 
var.  parvifloi'a,  F.v.M.  (B.Fl.  iii.  215),  and  is  E.  populifolia, 
Hook.,  as  noted  by  Mueller  himself  in  Herb.  Melb.  It  is  doubt- 
less the  same  as  "  var.  parvijlora,  Benth."  (should  be  F.v.M. 
'■  Eucalyptographia '  under  E.  largiflorens). 

1.  E.  bicolor,  Duff,  in  '  Catal.  of  N.S.W.  Forestry  Exhibits,' 
Melbourne,  Adelaide,  and  other  Exhibitions,  is  called  "  Slat}'- 
Gum,"  and  its  timber  is  described  as  "  hard,  tough,  strong, 
durable,  and  said  by  experts  to  be  one  of  the  best  hardwoods; 
used  for  fencing,  wheelwrights'  work,  bridges,  railway  sleepers, 
and  house  building;  plentiful.  Hab.  open  forests  south-western 
river  districts,  Blue  Mountains  and  the  Darling  River." 

It  is  evident  that  the  above  partly  refers  to  E.  bicolor,  A. 
Cunn.,  and  to  E.  polyanthemos,  Schauer,  and  it  is  only  referred 
to  on  the  present  occasion  as  the  source  whence  "^.  bicolor,  Slaty 
Cum,"  has  crept  into  numerous  official  reports. 

2.  E.  pendula,  Page. — This  name  was  first  published  (name 
only)  in  "Page's  Prodromus;  as  a  general  nomenclature  of  all  the 
plants  .  .  .  cultivated  in  Southampton  Botanic  Gardens,  by 
William  Bridgewater  Page,  London,  1818."     8vo.  pp.  136. 

In  Steudel's  '  Nomenclator  Botanicus  '  (ed.  ii.  Yol.  i.  p.  600) 
appears  the  simple  entry  "  pendula,  Page,  Nov.  Holl."  I  have 
been  unable  to  ascertain  that  Page's  name  is  more  than  a  nonien 
nudum. 

3.  E.  pendula,  A.  Cunn. — In  B.Fl.  (iii.  215)  this  is  quoted  as 
"A.  Cunn.  in  Steud.  Xom.  Bot.  Ed.  2."  It  is  assumed  to  be  a 
synonym  of  E.  pe^idula,  Page. 


BY    J.    H.    MAIDEN.  521 

The  origin  of  A.  Cunningham's  name  i-)endula  doubtless  arose 
from  the  following  : — 

"  The  '  Weeping  Eucalyptus  '  of  Major  Mitchell's  Expedition, 
1836,  No.  83  of  20th  April."  Specimen  in  Herb.  Cant,  ex  herb. 
Lindl.  This  is  no  doubt  the  plant  referred  to  in  the  '  Three 
Expeditions,'  ii.  45,  in  these  words,  "In  the  woods  I  observed  a 
Eucalyptus,  of  a  graceful  drooping  character,  apparently  related 
to  E,  2)ilularis  and  E.  amygdalina.'^  Mitchell  was  then  on  the 
Lachlan. 

The  specimen  No.  83  has  been  examined  by  me  and  is  identical 
with  E.  bicolor,  A.  Cunn.,  or  E.  largifloreyis,  F.v.M. 

"  The  interior  species,  E.  pendula,  has  a  Box  bark  right  out  to 
the  branchlets  (W.  Biiuerlen  and  R.  H.  Cambage),  a  red  timber, 
the  leaves  being  longer  than  those  of  E.  bicolor  and  glaucous, 
whilst  the  fruits  are  only  half  the  size  of  the  eastern  species.  The 
oils  are  also  quite  different "  (R.  T.  Baker,  these  Proceedings,  xxv. 
666).  The  name  E.  pendula,  A,  Cunn.,  has  no  real  existence, 
and  cannot  stand,  and  the  tree  referred  to  is  E.  bicolor,  A.  Cunn. 
The  E.  bicolor  referred  to  in  this  passage  is,  as  has  been  shown, 
E.  Bosistoana,  F.v.M. 

4.   E.  largiflorens,  F.v.M. 

Arboreous  :  leaves  alternate,  glaucous,  opaque,  oblong-lanceolate,  acute, 
slightly  oblique,  thinly  veined,  hardly  dotted;  umbels  pedunculate,  panicled, 
few-flowered;  flowers  smal]^  on  short  pedicels;  lid  double,  thin,  nearly  even, 
hemispherical,  blunt  or  minutely  apiculate;  tube  of  the  calyx  obconical-bell- 
shaped,  hardly  angular,  twice  as  long  as  the  lower  lid;  fruits  small,  half 
ovate,  short  stalked,  slightly  contracted  at  the  top;  valves  of  the  capsule 
inclosed. 

In  bushy  barren  localities  on  the  Murray,  Avoca,  Wimmera,  and  on  St, 
Vincent's  Gulf.  A  small  tree,  with  persistent  grey-blackish  bark  "  (Trans. 
Vict.  Inst.  i.  34,  1855). 

A  type  specimen  in  Herb.  Melb.  bears  the  following  label  : — 
"  Eucalyptus   largiflorens,  ferd.  MIL  herb.  Stuartii  153.     Nov. 
HoU.  Austral.    1847.     Murra3\      (E.  hsemastoma,  Sm.,  Miquel." 
in  Miquel's  handwriting,  and  also  bearing   the   species   number 

16,  in  his  Ned.  Kruidk.  Arch.  iv.).    "^.  bicolor,  A.C."  (in  Bentham's 
handwriting). 


522  ON  EUCALYPTUS  BICOLOR,  A.  CUNN., 

5.   E.  hcemastovia,  Miq. 

16.  Eucalyptus  haemastoma,  Smith,  Act.  Soc.  Linn.  Lond.  iii.  285  DC. 
Prodr.  l.c.  219,  n.  23.— E.  largiflorens,  Ferd.  Miill.  Herb. 

Locis  humidis  ad  fl.  Murray,  fl.  aestate  (F.M. ).    Van  Diemensland  (Stuart). 

Umbellge  sub-5-floraB,  nunc  superne  paniculato-confertse.  Operculum 
duplex;  exterius  depressum  obsolete  apiculatum,  interius  membranaceum 
convexum  muticum  ;  calycis  tubus  (p.  131)  obconicus  apice  ampliatus  hoc 
paulum,  illo  duplo  amplior,  ambobus  multo  longior  (ex  Miill.  adnot.)" 
Nederl.  Kruidk.  Arch.  iv.  130  (1856). 

This  is  B.  bicolor,  A.  Cunn.,  "as  to  the  Murray  specimens " 
(B.Fl.  iii.  215).  The  Tasmanian  (Van  Diemen's  Land)  specimens 
were  probably  £J.  amygdalina. 

Affinities. 

a.  With  E.  odorata,  Behr. — Mueller  (in  '  Eucalyptographia ') 
says  E.  odorata  perhaps  nearest  approaches  to  E.  largiflorens 
(bicolor),  which  is  best  distinguished  by  its 

1.  Usually  narrow  leaves  of  thinner  consistence. 

2.  With  finer  and  rather  more  spreading  veins. 

3.  By  the  more  extensively  paniculated  flowers. 

4.  Of  smaller  size. 

5.  By  the  shorter  lid,  often  of  less  width  than  the  tube  of  the 
calyx. 

6.  By  the  latter  more  suddenly  meeting  the  stalklet. 

7.  By  smaller,  more  globular  and  less  truncate  anthers. 

8.  By  fruits  of  less  size,  with  more  contracted  orifice  and  valves 
close  to  the  summit. 

This  species  runs  into  odorata  and  in  extreme  forms,  and  I 
doubt  if  it  is  possible  to  separate  them.  Usually  the  leaves  of 
E.  bicolor  are  dull-coloured,  but  this  is  not  an  infallible  guide. 
The  fruits  are  usually  smaller,  sometimes  much  smaller,  and  more 
cylindrical  than  those  of  E.  odorata.  The  fruits  of  E.  bicolor  are 
sub-cylindrical  or  ovoid,  while  those  of  odorata  are  hemispherical. 
E.  bicolor  lias  usually  short  filaments  and  blunt  opercula.  E. 
bicolor  is  usually  found  on  flats  liable  to  floods;  E.  odorata  prefers 
drier  situations. 


BY    J.    H.    MAIDEN.  523 

h.  With  E.  liiicrotheca,  F.v.  M. — The  foliage,  bark  and  timber 
of  the  two  species  are  often  very  similar.  The  fruits  are  very 
different;  the  valves  of  those  of  E.  microtheca  being  exserted. 
Both  timbers  are  red,  but  where  they  grow  together  the  timber 
of  E  microtheca  is  reputed  less  durable,  softer  and  more  faulty 
than  that  of  E.  hicolor. 

c.  With  E.  Behriana,  F.v.M. — See  my  paper  "On  Eucalyptus 
Behriana,  F.v.M.,"  in  Trans.  R.S.S.A.,  1902. 

d.  With  E.  iJopulifoUa,  Hook. — Specimens  from  Cobham  Lake 
in  the  extreme  north-west  of  the  State  (W.  Biiuerlen)  have  verj'- 
broad,  shiny  leaves,  which  resemble  those  of  E.  pajmlifolia  a  good 
deal,  as  well  as  E.  Behriana  (as  far  as  herbarium  specimens  are 
concerned). 

e.  With  E.  hemiphloia,  F.v.M. — The  likeness  of  E.  hicolor  to 
E.  hemiphloia,  F.v.M.,  var.  ynicrocarpa,  Maiden,  when  herbarium 
specimens  are  alone  available,  is  often  very  striking  and  some- 
times deceived  Mueller.  The  timbers  at  once  separate  them,  that 
of  E.  hemiphloia  being  pale  and  that  of  E.  hicolor  being  red. 

In  this  connection  the  following  specimen  is  interesting. 

E.  odorata,  var.Jioribunda,  Benth.  "  Inflorescence  occasionally 
compound,  connecting  the  species  in  some  measure  with  E.  hicolor, 
but  the  foliage  and  shape  of  the  flowers  and  fruits  are  rather 
those  of  E.  odorata.      On  the  Yarra,  Victoria  (B.Fl.  iii.  216)." 

I  have  not  seen  the  fruits,  but  from  the  specimens  I  have  seen 
I  would  suggest  that  it  is  referable  to  E.  hemiphloia.  The  form 
should,  however,  be  further  enquired  into. 

/.  With  E.  crebra,  F.v.M. — The  similarity  of  this  species  to  E. 
hicolor  is  sometimes  marked,  particularly  in  Queensland  specimens. 

With  typical  crehra  in  the  field  the  confusion  could  never  arise 
as  it  is  an  Ironbark,  but  as  the  tropics  are  approached  Ironbarks 
lose  some  of  their  characteristics  and  more  closely  approximate 
to  the  Red  Boxes  such  as  E.  hicolor. 

g.  With  E.  calycogona,  Turcz. — E.  hicolor  shows  affinity  to  E. 
calycogona  through  the  variety  gracilis  of  the  latter. 


524  on  eucalyptus  bicolor,  a.  cunn., 

Kange. 

This  species  prefers  rich  flats  which  are  liable  to  occasional 
submergence. 

South  Australia.— M annum  (W.  Gill). 

Victoria-— Herb.  Melb.  bears  the  following  label : — 

"16.  Eitc.  largiflorens,  ferd.  Mil.  herb.  Stuartii,  153.  Nov. 
Holl.  Austral.  1847.     Murray." 

"  E.  hcemastoma,  Sm.  Miquel "  (in  Miquel's  handwriting). 

"jS'.  bicolor,  A.C."  (in  Bentham's  handwriting). 

8  to  10  m.  N.W.  of  Nhill,  on  somewhat  moist  flats  (St.  Eloy 
D' Alton). 

North-west  of  Lake  Albacutya  (C.  French). 

Wimmera,  with  normal  flowers  (J.  Reader  and  others). 

Wimmera,  1891  (J.  P.  Eckert),  with  red  flowers;  an  instance 
of  two-coloured  flowers  (bicolor). 

Swan  Hill  (Dr.  Griffiths). 

Murray  River  (J.  P.  Eckert). 

Benjerup,  Murray  River  (C.  Walter). 

New  South  Wales.—"  Grey  Box,  the  common  Box  of  Riverina  " 
(T.  G.  Sloane,  Mulwala).     The  fruits  urceolate  when  unripe. 

Nyanda,  Booligal  (J.  O'Brien,  through  F.  B.  Guthrie),  used  for 
feeding  sheep  through  1902  drought. 

"  Black  or  Flooded  Box,"  Deniliquin  (Forester  Wilshire). 

Jerilderie.  Dwarf  Box  (25-30  ft.),  with  a  blackish  persistent 
bark  on  the  stem.  Wood  dark  red  inside,  but  lighter  in  the 
sapwood  (W.  H.  Suttor).  Mr.  W.  H.  Suttor's  specimens  were 
labelled  "  Goborro  "  by  Dr.  Woolls,  and  are  bicolor. 

Ivanhoe  via  Hay  (H.  Deane).     Red  timber. 

"Black  Box,"  "Swamp  Box,"  Hillston.  "A  sure  indication 
of  swampy  country  or  countr}^  flooded  at  times,  and  is  dense  and 
low  growing  "  ( W.  S.  Campbell). 

Lake  Cudgellico  (G.  S.  Home). 

"Drooping  Box,"  Condobolin  Flats  (shiny  leaves)  (J.H.M.). 

Euabalong  Road  (J.H.M.). 

"Swamp  White  Box"  or  "  Coolabah "  of  Lachlan  (F.  R. 
Kids  ton). 


BY    J.    H.    MAIDEiV.  625 

"Sample  taken  from  a  tree  about  15  feet  high  and  1  foot  in 
diameter.  I  selected  a  young  tree,  as  almost  all  the  large  ones 
are  hollow.  It  grew  on  flooded  land  on  the  first  creek  three 
miles  south  of  Condobolin.  I  should  call  it  dwarf  or  stunted  Box 
with  drooping  branches  like  a  '  Willow  tree  ' "  ( W.  H.  Suttor). 

"Grey  Box"  or  "Apple  Box"  or  "Red  Box"'  of  Lachlan 
River,  30  miles  below  Condobolin.  "Rough  grey  bark  on  limbs  " 
(R.  H.  Cambage). 

"White  Box,"  Mt.  Hope  Road  to  Euabalong;  "  White  Box," 
"  Grey  Box,"  "  Apple  Box,"  "  Red  Box."  Persistent  bark  on  the 
branchlets,  wood  redder  than  E.  hemipliloia^  and  not  so  hard; 
bark  not  so  useful  either.     Condobolin  "  (R.  H.  Cambage). 

"  White  or  Grey  Box  "  with  limbs  partly  white  (R.  H.  Cambage). 

Paldrumatta  Bore,  via  Wilcannia,  "Box,"  "  Curra  Curra "  of 
the  aborigines.  "The  onl}^  Eucalypt  growing  in  the  creek  here" 
(P.  Corbett). 

Mt.  Oxley,  Bourke  (E.  Betche). 

(a)  Victorian  Expedition,  1860,  towards  Barrier  Range. 

(6)  Clay  flats,  near  River  Darling,  31st  October,  1860. 

(c)  Victorian  Expedition,  1st  November,  1860,  Bambouroo, 
Dry  Lake,  near  Menindie. 

id)  High  sandy  banks  of  River  Darling. 

(These  four  specimens  were  collected  by  the  Burke  and  Wills' 
Expedition,  and  are  from  Nat.  Herb.  Melbourne). 

Cobham  Lake  (W.  Bauerlen),  No.  263,  20-30',  12".  Bark  per- 
sistent. Dull  leaves.  Venation  strongly  marked.  A  very  broad- 
leaved  form. 

"  River  Box,"  Bourke.  "  Plentiful  on  all  the  low  lands  of  this 
and  adjacent  districts.  Trees  small,  much  resembling  mallee  in 
appearance.  Long  pendulous  branches,  bark  rough,  dark  to  the 
extreme  tips  of  the  branches.  Timber  reddish-brown,  of  superior 
quality.      A  handsome  tree  "  (J.  L.  Boorman).     Leaves  narrow. 

"River  Box,"  Cobar  Road,  near  Bourke.     "Found  in  plenty; 

much  larger  specimens  than  those  growing  in  the  Lignum  Swamps 

35 


526  ON  EUCALYPTUS  BICOLOB,  A.  CUNN. 

on  the  Darling,  near  Bourke  "  (J.  L.  Boorman).  The  leaves  of  this 
specimen  are  mostly  coarser  than  those  of  the  preceding  specimen. 

Bourke  to  Barringun  (W.  S.  Campbell). 

River  Darling  at  Bourke  (J.H.M.).  1 

Tinapagee,  Paroo  River  (E.  Betche). 

"River  Box,"  Belalie  Bore  (H.  V.  Jackson). 

Wilcannia  (H.  V.  Jackson). 

Murtee  Holding  (Stock  Inspector  TuUy). 

"  Black,  narrow-leaved  or  Bogan  River  Box."  Coolabah  (J. 
N.  Peacock). 

Dubbo,  Tomingley  to  Narromine  (J.H.M.). 

"Narrow-leaved  Box,"  on  the  plains  near  Baradine  (Mr. 
Forsyth's  No.  5);  also  Dubbo-Gilgandra  Road,  18  miles  from 
Dubbo  (W.  Forsyth).     Both  with  narrow,  shiny  leaves. 

Castlereagh  River  (W.  Woolls).  These  and  the  Baradine  and 
Gilgandra  specimens  w^ith  very  small  fruits.  The  Castlereagh 
River  specimens  have  narrow  leaves,  but  leaves  of  the  normal 
width,  and  showing  the  characteristic  venation  also. 

"  Narrow-leaved  Box,"  Moree  (W.  S.  Campbell). 

Narrabri  (J.H.M.). 


527 


7. —ON  EUCALYPTUS  POLYANTHEMOS,  Schauer. 
By  J.  H.  Maiden. 

(Plate  xxi.) 

Eucalyptus  polyanthemos,  Schauer,  MSS. 

Arborea  glauco-virens  :  foil,  coriaceis  ovatis  subito  in  petiolum  contractis 
obtusis  apiculatis,  margine  crasso  subrevoluto  cinctis  impunctatis,  utrinq. 
opacis;  paniculis  axillarib.  terminalibq.,  umbellis  3-5-floris,  pedicellis  brevib. 
cupulae  clavato-turbinataB  continuis;  operculo  brevissime  conico  acutiusculo 
vertici  cupulas  exangulatce  imposito.— Foliorum  lamina  2  poll,  circiter  longa, 
2^  poll,  lata,  petiolus  8  lin.  longus,  flores  cum  pedicello  et  operculo  3  lin.  longi, 
operculum  resinoso-punctatum,  stamina  brevissima. — In  Nova?  Cambriae 
australis  interioribus  septentriones  versus  ab  urbe  Bathurst.  —  A.  Cunn., 
Herb.  No.  136,  1822  (Walpers'  'Repertorium  Botanices  Systematicae. '  Tomus 
ii.  p.  924). 

I  have  seen  the  type;  it  is  referred  to  at  p.  533.  I  would 
invite  attention  to  Schauer's  spelling  of  his  specific  name. 

Synonyms. 

1.  E.  polyanthema,  Schauer,  var.  glauca,  R.  T.  Baker. 

2.  E.  Dawsoni,  R.  T.  Baker. 

3.  E.  ovalifolia,  R.  T.  Baker  [partim). 

4.  E.  ovalifolia,  R.  T.  Baker,  var,  lanceolatn,  R.  T.  Baker. 

I  have  given  careful  attention  to  this  species  for  many  years, 
but  could  never  find  any  botanical  differences  sufficient  to  justify 
the  establishment  of  a  second  species  out  of  the  "  Red  Box  or 
Slaty  Gum "  series.  Accompanied  by  a  collector,  Mr.  J.  L. 
Boorman,  I  made  careful  observations.  Then  I  sent  Mr.  Boorman 
alone  to  carefully  investigate  the  trees,  and  he  performed  his 
duty  in  a  most  intelligent  manner.  Although  quite  satisfied  with 
his  reports,  I  went  again  into  the  field  with  him,  and  the  three 
trips  resulted  in  the  accumulation  of  a  large  and  varied  quantity 
of  botanical  material,  including  timber  and  bark.      I  found  people 


528  ON  EUCAl  YPTUS  POLYAXTHEMOS,  SCHAUEK, 

indifferent  a.s  to  the  use  of  the  names  "  Slaty  Gum  or  Red  Box," 
applying  them  indiscriminately  as  a  very  general  rule.  More 
than  one  timber  worker  told  me  they  were  the  same  tree,  and  all 
the  evidence  I  accumulated  points  in  the  same  direction.  As 
the  confusion  in  regard  to  "Red  Box  and  Slaty  Gum"  has  become 
considerable,  I  have  at  pp.  531-535  given  full  notes  made  in  the 
field  by  my  collector  and  by  other  observers,  and  add  the  follow- 
ing report  from  him  which  was  written  in  the  field  : — 

"  With  reference  to  your  instructions  to  collect  all  available 
evidence  in  regard  to  Red  Box  and  Slaty  Gum,  I  respectfully 
report  that  I  went  to  Tallawang  (some  6  miles  from  Gulgong) 
out  towards  Cobborah,  where  it  grows  to  fine  tall  trees,  on  the 
ridgy  ground  near  to  the  Reedy  Creek;  at  Gulgong  it  grows  all 
around  the  district  of  a  much  more  stunted  growth.  At  Lue  it 
grows  (large  trees)  about  5-6  miles  in  a  western  direction  from 
the  railway  line;  the  more  stunted  forms  occur  all  around  the 
immediate  district  of  Lue.  There  again  at  Rylstone,  some  5 
miles  on  the  Ilford  road,  it  again  reaches  to  fine  trees. 

"  From  a  personal  conviction  and  from  reliable  information, 
the  Slaty  Gum  and  Red  Box  are  identical.  The  term  '  Slaty 
Gum'  is  applied  to  the  more  robust  trees  of  the  '  Red  Box'  from 
the  cleaner  stems,  for  these  trees  have  a  greater  tendency  to 
throw  off  the  bark  more  regularly,  falling  in  ribbons,  and  not  in 
'  scales,'  so  to  speak,  as  in  the  case  of  the  less  healthy  trees;  this 
is  not  invariably  the  case,  as  exceptions  do  constantly  occur. 
This  (April)  seemingly  is  the  season  of  the  j^ear  for  shedding  of 
the  bark  A  Mr.  Taylor,  native  of  Rylstone  (a  carpenter),  says 
there  is  no  difference  in  the  timbers  or  in  the  habit  of  the  trees, 
'  Slaty  Gum  '  being  only  larger  than  '  Red  Box.'  Mr.  James 
Holmes,  Gulgong,  who  pointed  out  to  me  the  tree  at  Reedy 
Creek,  calls  '  Red  Gum '  and  '  Slaty  Gum  '  one  and  the  same. 
Mr.  Hatton,  living  at  Coomber,  says  that  '  Slaty  Gum'  is  nothing 
but  '  Red  Box.'  " 

Mr.  Baker  deals  with  the  matter  of  Red  Box  and  Slaty  Gum 
in  these  Proceedings  (xxi.  448,  1896),  and  I  think  the  conclusion 


BY    J.    H.    MAIDEN.  529 

he  then  arrived  at  to  keep  the  forms  to  which  he  referred  under 
E.  polyanthe7nos  was  sound. 

1.  E.  polyanthema,  Schauer,  var.  glauca,  R.  T.  Baker. 

Mr.  Baker  does  not  under  E.  'polyantheraos  (p.  448)  name  a 
variet}^,  but  at  page  431  occurs  the  following  passage: — ."The 
most  valued  timber  is  perhaps  '  Slaty  Gum,'  E.  polyanthema  var. 
glauca,  var.nov.,  and  I  consider  it  a  distinct  gain  to  the  botany 
of  the  colony  to  have  the  correct  botanical  sequence  of  this 
valuable  tree  made  clear." 

Glaucousness  being  a  character  of  the  species,  it  seems  to  me 
that  to  call  one  form  glauca  would  be  inconvenient  and  could  not 
be  carried  out  in  practice. 

2.  E.  DaiDsoni,  R.  T.  Baker,  these  Proceedings,  xxiv.  295, 1899. 
"  Slaty  Gum."     Type  localities,   "  Ridges  on  the  Watershed  of 

the   Goulburn   River   (R.T.B.) ;    across   the    Main    'Divide'    at 
Cassilis,  and  north-west  to  Pilliga  "  (Professor  Warren). 

3.  E.  ovalifolia,  R.  T.  Baker,  these  Proceedings,  xxv.  680, 
1900  (pa7^tim).    . 

"  Red  Box."  Type  localities  given  are  Bathurst,  Rylstone  and 
Camboon  (R.  T.  Baker);  Hargraves,  Mudgee  to  Wellington  (A. 
A.  Suttor);  Gerogery  (L.  Mann). 

Mr.  Baker  (loc.  cit.)  speaks  of  "  the  typical  E.  polyanthema, 
Sch.,  of  Victoria,"  but  the  type  comes  from  Bathurst,  N.S.W., 
like  that  of  E.  ovalifolia.     See  p.  533. 

I  am  of  opinion  that  some  confusion  has  arisen  in  regard  to 
Mr.  Baker's  species.  Some  bud-bearing  twigs  kindly  presented 
by  Mr.  Baker  are  those  of  E.  Baueriana,  Schauer,  var.  conica, 
Maiden  (these  Proceedings,  1902,  p.  216),  or  rather  one  of  those 
intermediate  forms  which  show  that  it  is  impossible  to  separate 
E.  Baueriana  from  its  variety.  Mr.  Smith's  report  on  the  oil 
(these  Proceedings,  xxv.  682)  in  which  he  says  that  "  there  is 
very  little  difference  in  the  constituents  of  this  oil  and  that  of  E. 
Fletcheri,  Baker  "  i^E.  Baueriana,  Schauer),  bears  out  my  view. 


530  ON  EUCALYPTUS  POLYANTHEMOS,  SCHAUEE, 

On  the  other  hand,  Mr.  Baker's  label  saj^s — "  E.  ovalifolia, 
R.T.B.  Red  Box  Hard,  red-coloured  durable  timber;  bark 
rough  at  base."     This  is  a  description  of  E.  jjolyantheinos,  Schauer. 

I  am  therefore  of  opinion  that,  through  inadvertence,  E.  ovali- 
folia,  K.  T.  Baker,  has  been  parti}'-  described  from  E.  j)olyan- 
thenios,  Schauer,  and  partly  from  E.  Bauerania,  Schauer. 

4.   E.  ovalifolia,  R.  T.  Baker,  var.  lanceolata,  R.  T.  Baker. 

A  specimen  of  timber  from  Rylstone  labelled  as  above  by  Mr. 
Baker,  with  the  addition  "  Red  Box,"  and  kindh''  presented  by 
that  gentleman,  is  E.  j^olyanthe^nos,  Schauer. 

Affinities. 

a.  With  E.  Baueriana,  Schauer. — This  species  has  long  been 
confused  with  E.  polyanthemos,  which  is  the  best  evidence  that 
it  is  necessary  to  issue  a  caution  on  the  subject.  As  regards 
herbarium  specimens,  those  of  E.  Baueriana  are  less  glaucous  and 
often  more  slaty-looking  than  E.  j^olyanthemos,  the  opercula  less 
pointed,  and  the  fruits  more  conical  and  with  thinner  rims.  The 
forester  would  not  confuse  the  trees  for  a  moment.  The  bark  of 
E.  Baueriana  is,  woolly  up  to  the  small  branches,. that  of  E.  'poly- 
anthemos being  smooth  or  ribbon3^  The  timber  of  E.  Baueriana 
is  pale  brown  or  brovrn,  while  that  of  E.  jjolyanthemos  is  deep  red. 

I  have  seen  E.  polyanthemos  with  a  true  "  box"  bark,  and  am 
of  opinion  that  the  use  of  the  term  "box"  (other  than  "bastard 
box")  as  applied  to  this  tree,  has  arisen,  in  some  measure,  from 
the  long-continued  confusion  with  E.  Baueriana.  See  also  these 
Proceedings,  1899,  p.  613. 

6.  With  E.  melliodora.,  A  Cunn. — Sometimes  these  two  species 
from  bark  alone  are  hard  to  discriminate.  They  both  may  have 
scaly-box  bark  at  the  butt  for  a  considerable  distance,  thereafter 
ribbony  bark  or  a  "  White  Gum  "  appearance. 

The  bark  of  E.  melliodora  is  more  persistent  than  that  of  E. 
2)olijanthemos;  the  ribbony  bark,  however,  never  descending  so 
low  down  as  in  the  latter.  A  man  might  readily  be  forgiven  if 
he  called  one  polyanthemos  tree  a  Box  and  another  a  Gum,  and 
as  a  matter  of  fact  this  is  commonl}^  done. 


BY    J.    H.    MAIDEN.  531 

E.  melliodora  has  a  very  yellow  inner  bark  and  sapwood,  with 
longer  and  narrower  and  more  pendulous  leaves  and  fewer  flowers. 
E.  polyanthemos  has  a  white  sap  and  red  heart  wood.  E,  'polyan- 
themos  has  comparatively  narrow  leaves  at  the  top  of  the  tree. 

c.  With  E.  tereticornis,  Sm. — Jn  the  field  E.  tereticornis  and 
E.  'polyanthemos  when  fully  grown  might  very  readily  be  con- 
fused. The  smooth,  gnarled  trunk,  with  the  bark  falling  off  in 
patches  or  in  ribbons  is  common  to  both  species,  and  knowing 
them  so  well  I  still  sometimes  considered  it  prudent  to  procure  a 
twig  to  decide  the  identity  of  a  particular  tree.  The  timber  of 
both  trees  is  red,  but  the  ordinary  botanical  characters  of  the 
two  species  are  very  different. 

Range. 

Victoria. — '-''The  Red  Box  (E.  polyanthema)  grows  in  places  all 
over  Victoria.  The  timber  is,  however,  as  a  rule  rather  small, 
the  boles  and  limbs  crooked,  in  some  places  so  much  so,  for 
instance  in  the  Havelock  State  Forest,  as  to  be  of  no  value  but 
for  firewood.  In  parts  of  Gippsland  it  is  larger,  but  there  it 
often  becomes  so  hollow  as  to  be  a  mere  shell.  Although  the 
wood  is  hard  and  durable,  these  defects  cause  it  to  be  of  little 
economic  value"  (A.  W.  Howitt  hi  lilt.). 

"Red  Box"  with  red  timber  and  gnarled  greyish  boxy  bark. 
Euroa;  Lilydale;  Buchan,  Gippsland.  It  grows  on  hillsides 
(A.  W.  Howitt). 

Wangaratta,  handsome,  spreading  trees,  glaucous  all  over 
(J.H.M.).  hence  sometimes  called  Grey  Box.  Mr.  J.  V.  de  Coque 
wrote  to  me  : — "  The  Victorian  Grey  Box,  particularly  about 
Wangaratta,  is  remarkably  durable  and  takes  the  place  of  Iron- 
bark  for  railway  bridges  construction.  I  used  nearly  500  piles 
in  the  Myrtleford  to  Bright  railway,  all  Grey  Box  and  all  pipy; 
when  piles  are  driven  the  practice  is  to  plug  the  pipe  holes." 

"  Hill  Box,"  Mt.  Kosciusko  Range,  red  wood  (Findlay),  pro- 
bably on  the  Victorian  side  (National  Herbarium,  Melbourne). 

"  36.  Eucalyptus  polyanthemos,  Schauer,  I.e.  n.  5.  Beyond 
Mount  Disappointment  (Ferd.  Muell.)"  is  a  note  by  Miquel  in 
Ned.  Kruidk.  Arch. 


532  ON  EUCALYPTUS  POLYANTHEMOS,  SCHAUEE, 

New  South  Wales. — Quiedong,  Bombala.  Bark  persistent,  but 
deciduous  on  top  of  branches.  40-60  feet.  2-4  feet.  Leaves 
and  buds  glaucous  (W.  Bauerlen). 

Tumberumba  (Forester  Taylor;  W.  Forsyth).  "Red  Box," 
"Round  Leaf  Gum,''  "  Red  Mahogany,"  Kyambn,  WaggaWagga 
district  (Forester  Taylor). 

"  Red  Box,"  Wagga  Wagga  district  (W.  Orr).  Mr.  Forester 
Tajdor  has  the  following  note  on  some  other  specimens  :  "Red 
or  round  Shining-leaf  Box.  Yields  a  very  useful  timber  of  a 
dark  red  colour.  The  only  similarity  between  this  and  Bimbil 
(E.  populifolia)  is  the  roundish  shiny  leaves,  which  glisten  in  the 
sun."  Personally  I  have  never  noticed  shining,  glistening  leaves, 
though  in  B.  populifolia  this  is  common  enough. 

"Round-leaf  Gum,"  Tumut  (W.  S.  Campbell). 

Tumut  (H.  Deane).  Mr.  Deane  has  the  following  note : 
"  Deciduous,  smooth  bark;  var.  U.  poli/antheiua.  Same  as  Red 
Box  from  Queanbeyan  apparently." 

"  Round-leaved  Gum,"  Tumut  (W.  W.  Froggatt).  Some  of  the 
leaves  tending  to  lanceolate.  The  fruits  shiny,  and  showing  the 
cracked  appearance  at  the  orifice  common  in  the  siDecies. 

"Red  Box,"  Queanbeyan  (H.  Deane).  In  fruit,  otherwise  not 
to  be  distinguished  from  the  Bombala  specimen.  Fruits  narrow, 
tapering  at  orifice. 

Murrumbidgee  district.  So  glaucous  as  to  be  called  "'  White 
Box"  (J.  Du£e). 

"Red  Box"  is  a  most  durable  timber,  and  is  extensively  used 
for  fencing,  bridge  building,  railway  sleepers,  and  wheelwrights' 
work.  It  is  almost  invariably  found  in  company  with  Ironbark, 
and  in  some  parts  of  my  district  there  is  a  considerable  quantity, 
but  close  to  Grenfell  it  is  not  so  common,  and  is,  therefore,  not 
much  used  (Forester  John  G.  Postlewaite,  Grenfell). 

"  Red  Gum  or  Red  Box,"  Stuart  Town  (A.  Murphy). 

"Red  Box,"  Stuart  Town.  "Plentiful  all  over  the  district; 
trees  of  fair  size,  with  whitish  ribbony  stems  and  glaucous  tint 
throughout,  leaves  and  branches  pendulous,  wood  hard  but 
faulty"  (J.  L.  Boorman). 


BY    J.    H.    MAIDEN.  533 

''Red  Box,"  Apsley,  Bathurst  (R.  H.  Cambage). 

"  Red  Box,"  Cow  Flat,  Bathurst.  Typical  for  E.  ovalifolia,  R. 
T.  Baker  (partiTn,  see  p.  529). 

"Interior  of  Southern  Australia,  N.  from  Bathurst."  A.  Cun- 
ningham (Herb.  Heward,  then  Herb.  Lemann.  Now  Herb.  Cant.). 
Type  of  U.  polyanthemos,  Schauer.  Examined  by  me,  and 
absolutely  identical  with  the  Stuart  Town  and  Bathurst  specimens. 

The  following  entry  in  Allan  Cunningham's  manuscript  journal 
refers  to  E.  polyanthemos  : — 

"20th  April,  1817.  18  miles  W.  of  Bathurst.  A  species  of 
Eucalyptus  (conferta)  rising  about  20  feet  with  obovate  leaves, 
inflorescence  umbelled  and  terminal,  is  at  this  period  just  expand- 
ing its  flowers  on  the  sides  of  the  hills." 

"  Red  Box,"  Reedy  Creek,  near  Gulgong.  "  Narrow  leaves; 
big  tree  glaucous  all  over  "  (J.  L.  Boorman). 

"  Slaty  Gum,"  Reedy  Creek.  "  Fine  large  trees  growing  on 
slight  ridges  awa}^  from  the  creek;  at  this  period  (April)  of  the 
year  the  stems  are  pure  white  with  irregular  patches  of  green; 
the  bark  is  \  inch  thick,  falling  away  in  long  ribbons;  the  leaves 
and  tips  of  branches  are  of  a  mealy  whiteness.  Sapwood  pale 
yellow,  centre  red  "  (J.  L.  Boorman). 

"  Red  Box,"  Gulgong.  "The  whole  of  the  tree  of  a  silvery 
whiteness." 

A  second  specimen  with  leaves  from  lanceolate  to  ovate  (J.  L. 
Boorman). 

"Red  Box  or  Slaty  Gum,"  rarely  called  "Bastard  Box." 
Gulgong.  "Leaves  on  shoots  from  nearly  orbicular  to  nearly 
linear  lanceolate"  (J.H.M.  and  J.  L.  Boorman). 

"Slaty  Gum,"  on  ridges,  Mudgee.  "  Smooth  bark,  good  wood 
for  weatherboards,  fencing  and  other  wheelwrights'  work,  bad 
for  burning,  does  not  crack  in  the  sun  "  ( W.  Woolls). 

Note  by  Dr.  Woolls  on  another  specimen: — "Leaves  vary, 
more  lanceolate  in  the  large  trees.     Slaty  Gum,  E.  polyanthemaJ' 

" '  Slaty  Gum'  is  considered  the  best  in  the  district  (Mudgee), 
but  unfortunately  the  supply  is  not  equal  to  the  demand.  Grows 
in  the  district  west  of  Reedy  Creek.     Used  for  all  work  where 


534         ON  EUCALYPTUS  POLVAXTHEMOS,   SCHAUEE, 

toughness  is  required.  Differs  from  Ironbark,  inasmuch  as  the 
log  when  drying  at  the  butt  cracks  in  rings,  while  Ironbark 
radiates  from  the  centre"  (Forest  Ranger  Marriott).  Mr. 
Marriott  called  this  timber  "  F.  bicolor,"  an  error  which  has  been 
referred  to  at  p.  520. 

Mr.  A.  G.  Hamilton  (these  Proceedings  [2],  ii.,  277)  states 
that  the  "  Slat}'  Gum "  is  common  on  the  ridges  at  Mudgee, 
where  it  does  not  attain  a  great  size,  but  on  the  flats  at  Tallewang 
it  runs  up  into  splendid  trunks,  which  are  much  used  in  bridge 
building,  &c.,  and  the  timber  is  considered  very  durable. 

"Slaty  Gum  or  Red  Box,"  Lue,  Mudgee  Line.  "Fine  large 
trees,  40  to  60  feet  high,  patchy  or  at  times  quite  smooth,  white 
with  patches  of  grey  bark;  this  is  more  applicable  to  the  large 
trees;  the  smaller  are  more  of  a  scaly  nature.  The  latter  desig- 
nated '  Red  Box  ';  the  larger  '  Slaty  '  or  '  Spotted  Gum  '  "  (J.  L. 
Boorman). 

"Slaty  Gum,"  Rylstone.  "  Large  tall  trees,  clean  stems,  bark 
falling  away  in  long  ribbons,  leaving  a  clean  white  stem  with 
patches  of  dark  green,  the  tips  of  the  branches  of  a  mealy, 
powdery  whiteness,  sap  timber  pale  3"ellow,  centre  red  "  (J.  L. 
Boorman). 

"  Red  Box,"  Rylstone.  "  Trees  not  so  large  as  those  known 
as  '  Slaty  Gum,'  and  the  bark  is  more  scaly,  but  in  wood  and 
every  other  respect  identical  with  Slaty  Gum  "  (J.  L.  Boorman). 

"Red  Box,"  typical  for  E.  ovalifolia,  R.  T.  Baker,  Rylstone 
(R.  T.  Baker).  See  H.  G.  Smith  (Proc.  R.S.N.S.W.  xxxiv. 
290).  Labelled  indifferently  "  Slaty  Gum,  Red  Gum,  Red  Box," 
Capertee.  "  Some  trees  large  and  entirely  smooth-barked;  otliers 
small  and  stunted,  with  a  ribbony  bark.  A  large  tree  which  was 
felled  showed  narrow  leaves  at  the  top "  (J.H.M.  and  J.  L. 
Boorman). 

Following  are  some  field  notes  : — 

"  Cullen  Bullen  to  Capertee.  This  species  reminds  one  of 
Yellow  Box  [E.  melliodora)  slightly — a  ribbony  gum  on  young 
trees,  scaly  bark  when  older,  at  least  as  far  as  the  first  fork  and 
often  beyond.     Handsome  trees;  wood  red;  we  then  found  a  tree 


BY    J.    H.    MAIDKN.  535 

3'  ill  diameter,  and  reminding  one  a  good  deal  of  E.  tereticornis, 
the  smooth  part  of  a  3'ollow  cast.  The  buds  of  a  yellowish  cast 
and  often  a  little  glaucous.  Little  conical  opercula.  Wood  red. 
Very  free  fiowerer.      Fruits  conoid  "  (J.H.M.). 

"Slaty  Gum,"  Bylong  Creek,  typical  for  F.  Dawsoni,  R.  T. 
Baker. 

"Red  Box."  "  Mountain  Gum.  Murrumbo,  Goulburn  River; 
Camboon ;  Bylong  (R.  T.  Baker).  Three  specimens  showing 
transition  from  lanceolate  leaves  to  ovate." 

Baerami,  Goulburn  River  ?  (H.  Deane). 

"  Red  Box,"  near  Taree.  "  Timber  \'ery  durable,  of  red  colour. 
Bark  persistent,  and  like  that  on  the  trunk  of  Grey  Box  (hemi- 
2)hloia)  (A.  Rudder)." 

"Box,"  Cundletown.  "Wood  dark  red.  Height  120  feet; 
diam.  2-3  feet"  (A.  Rudder).  This  brings  the  range  of  the 
species  to  the  Lower  Manning  River,  These  specimens  have 
lanceolate  leaves  of  thinner  texture  than  the  mountain  forms  of 
E.  ijolyanthenios. 


EXPLANATION  OF  PLATE  XXL 

Flowering  twig  of  Eucalyptus  polyanthemos,  Schauer. 

Drawn  from  the  type  (see  p.  533). 

(The  two  leaves  below  the  principal  figure  are  shown  separately  as  if 
removed  from  their  attachments  on  the  left  side.  This  was  done  to  bring 
the  drawing  within  the  space  necessary  for  a  single  plate). 


536 


CONTRIBUTIONS    TO    A    KNOWLEDGE    OF    THE 
FLORA    OF    AUSTRALIA. 

Part  IV. 

By  R.  T.  Baker,  F.L.S.,  Curator  and  Economic  Botanist, 
Technological  Museum,  Sydney. 

(Plates  xxii.-xxiii.) 

CELASTRINE^. 

El^odendron  australe,  Ve7it. — This  species  occurs  as  far  west 
as  Narrabri  (W.  Bauerlen),  thus  showing  that  it  belongs  to  the 
arid  country  flora  as  well  as  that  of  the  mountain  and  coast. 

RUTACE^. 

AcRONYCHiA    L.EVis,  FoTst. — Mt.   McKcnzie,  Tenterfield  (W. 

Bauerlen). 

HHAMNE^. 

Pomaderris  ferruginea,  Sieb. — Previously  recorded  from  Port 

Jackson  to  Victoria,  but  it  extends  as  far  north  as  Tenterfield 

(W.  Bauerlen). 

ANACARDIACEJE. 

EuROSCHiNUS  FALCATUS,  Hook.  f. — -Specimens  of  this  species 
have  recently  been  forwarded  to  me  from  Stockton,  near  New- 
castle, thus  showing  it  to  have  a  much  more  southern  distribution 
than  previously  suspected.  In  that  locality  the  tree  is  known  as 
"  Rosewood  "  or  "  Brown  Beech,"  and  grows  in  poor  sandy  soil 
adjacent  to  salt  water.  At  one  time  it  was  very  plentiful,  but 
owing  to  its  proximity  to  settlement  it  has  been  much  thinned 
out.  The  timber  has  a  very  decided  scent  when  freshly  cut,  and 
is  specifically  light,  porous,  and  pale-coloured  with  sometimes  a 
tinge  of  yellow.  It  is  a  splendid  working  timber,  suitable  for 
cabinet  work,  and  is  worthy  of  forest  conservation. 


f 


BY    R.    T.    BAKER.  537 

LEGUMINOS^. 

OxYLOBiUM  TRiLOBATUM  var.  iLiciFOLiUM,  Maiden  &  Betche. — 
Drake  (W.  Bauerlen).  Only  previously  recorded  from  the  Tweed 
River. 

Daviesia  latifolia,  R.Br. — Extends  as  far  north  as  Tenter- 
field  (W.  Bauerlen). 

PuLTENiEA  viLLOSA,  WUld. — Tliis  species  extends  into  the 
interior  of  New  South  Wales,  as  it  is  found  to  occur  at  Grenfell 
(W.  Bauerlen). 

Acacia  leprosa,  Sieh. — Mudgee  is  the  most  northern  locality 
previously  recorded  for  this  species.  It,  however,  extends  as  far 
north  as  Tenterfield  (W.  Bauerlen). 

A.  LANiGERA,  A.  Citnu. — Tenterfield,  which  is  now  its  most 
northern  locality  in  New  South  Wales. 

A.  VERNiciFLUA,  A.  Cunn. — Grenfell  (W.  Bauerlen). 

A.  PENNiNERVis,  Sieh. — Tliis  species  has  a  range  much  further 
west  than  previously  recorded,  as  it  is  found  on  Mount  Bolton 
near  Moura,  Parkes,  where  also  occurs 

A.  MELANOXYLON,  R.Br..,  the  most  westerly  record  for  that 
species. 

A.  PYCNOSTACHYA,  F.v.M. — Narrabri  (W.  Bauerlen).  This  is 
the  most  southern  and  western  point  recorded  for  this  species. 

HALORAGE^. 

Haloragis  serra,  Brongn. — Occurs  as  far  south  now  as  Cowra 
(W,  Bauerlen)  and  Gundagai  (H.  Deane). 

MYRTACE^. 

KuNZEA  OPPOSiTA,  F.V.M. — Timbarra  is  the  only  locality  so 
far  recorded  for  this  species.  It,  however,  occurs  as  far  south  as 
Tenterfield  (W.  Bauerlen). 

Eucalyptus  Smithii,  R.  T.  Baker. — This  species,  now  famous 
for  the  excellence  of  its  oil,  is  found  to  occur  in  almost  all  the 
gullies  around  Mittagong  (H.  G.  Smith),  where,  however,  it  must 


538  FLORA    OF    AUSTRALIA, 

not  be  confounded  with  the  rough-barked  variety  of  E.  viminalis, 
LabilL,  var.  jyedicellaris,  of  Deane  (fc  Maiden. 

E.  Bridgesiana,  B.  T.  Baker. — This  is  now  known  to  extend 
as  far  west  as  Moura,  Grenfell  (W.  Bauerlen). 

E.  viTREA,  R.  T.  Baker. — Extends  as  far  south  as  Delegate, 
and  probably  into  Victoria  (W.  Bauerlen). 

E.  OVALIFOLIA,  R.  T.  Baker. — This  "Red  Box"  tree  occurs  as 
far  west  as  Moura,  near  Parkes  (W.  Bauerlen),  where  it  differs 
in  no  respect  from  the  type  which  was  described  from  Rj^stone 
specimens.  To  further  prove  its  claims  to  specific  rank  as  dis- 
tinct from  E.  polyantheyna,  Schau.,  it  may  be  mentioned  that  the 
former  is  found  at  Linley  Creek  near  Goulburn,  being  known  also 
as  "  Red  Box,"  where  the  latter  species  also  occurs  and  is  dis- 
criminated by  settlers. 

E.  Fletcheri,  R.  T.  Baker.— Victon  (R.  T.  Baker).  This 
species  is  synonymised  by  Mr.  Maiden  in  the  Proceedings  of  this 
Society  (1902,  p.  216)  with  E.  Baueriana,  Schau.,  the  t3qDe  of 
which  from  the  Vienna  herbarium  was  exhibited  at  the  June 
meeting.  This  specimen  is  only  in  flower  and  bud,  whilst  the 
description  of  it  (Walp.  Repert.  ii.  924,  Suppl.  i.  1845)  is  practi- 
cally useless  for  the  purposes  of  Eucalyptus  determination.  E. 
Fletcheri  as  described  places  beyond  dispute  the  tree  referred  to, 
whilst  the  fragmentary  herbarium  material  now  brought  forward 
as  its  type,  and  which  may  do  duty  for  any  of  the  "  Boxes  " 
placed  in  the  past  with  E.  jwlyarithemos,  Schau.,  will  only  lead 
to  still  further  confusion  in  the  species  of  the  genus,  and  cause 
doubts  in  the  minds  of  future  workers. 

From  the  Vienna  type  material  and  its  published  description 
I  fail  to  see  the  connection  between  these  two  species,  much  less 
E.  conica,  Deane  &  Maiden,  which  was  also  synonymised  at  the 
same  time  under  E.  Baueriana,  Schau.,  a  species  recorded  by 
Bentham  as  tropical. 

E.  OVALIFOLIA,  R.  T.  Baker,  var.  lanceolata,  R.  T.  Baker  et 
H.  G.  Smith. — This  "  Box  ''  tree  occurs  at  Putty,  midway  between 
Richmond  and  Singleton  (A.  C.  Barwick).  The  only  previous 
recorded  locality  for  this  species  is  Camboon,  Rylstone. 


BY    R.    T.    BAKER.  539 

E.  DIVES,  Schau. — This  species  is  known  at  Adelong  as  "Mess- 
mate," which  district  is  now  its  most  western  limit. 

E.  OCHROPHLOIA,  F.v.M. — This  species  is  found  as  far  east  as 
Goondiwindi,  Queensland  (C.  N.  Chapman),  under  the  local  name 
of  "  Yuppunyah." 

E.  MACRORHYNCHA,  F.V.M. — This  species  occurs  at  Gramen, 
N.W.  of  Glen  Innes  (Henry  Hurst).  This  locality  greatly 
extends  the  northern  limit  of  this  important  economic  Eucalypt, 
which  will  probably  now  be  found  to  extend  into  Queensland. 

E.  TERETICORNIS,  Sm.,  var.  LINEARIS,  R.  T.  Baker  et  //.  G.  Smith. 
— This  variety  has  now  been  found  at  Stroud,  where  the  timber 
is  regarded  as  much  superior  to  that  of  the  type,  as  posts  made 
from  it  have  stood  for  50  years  in  the  ground  and  still  remain 
perfectly  sound.  Since  this  var,  was  recorded  (' Eucalypts  and 
Essential  Oils,'  by  Baker  and  Smith)  further  investigation  has 
shown  that  its  differences  from  the  type  are  much  greater  than 
was  originally  supposed;  in  fact,  they  are  of  such  a  nature  as 
to  fully  justify  its  being  raised  to  specific  rank. 

E.  MoRRisii,  R.  T.  Baker. — Occurs  as  far  west  as  IMt.  Drysdale, 
on  rocky  ridges,  and  is  known  as  "  Scrub  Gum  "  (G.  Wardrop). 

E.  DEALBATA,  A.  Cunn. — The  most  easterly  known  locality  for 
this  species  is  now  to  be  recorded,  namely,  Putt}^,  south-west  of 
Singleton,  where  it  grows  to  a  height  of  from  40  to  50  feet,  with 
a  smooth,  dark-coloured  bark  falling  off  in  patches,  giving  the 
tree  a  spotted  appearance  (A.  C.  Barwick). 

In  the  Cassilis  district  there  appears  to  be  a  form  of  this 
species  with  a  very  short  blunt  operculum;  further  investigation 
is  being  taken  to  prove  whether  it  is  really  a  variety  or  a  new 
species. 

E.  SQUAMOSA,  Deane  <k  Maiden. — This  tree,  only  previously 
recorded  from  National  Park  and  Richmond,  is  now  found  to 
extend  as  far  north  as  Putty,  50  miles  south  of  Singleton,  where 
it  is  known  as  "  Blood-tree."  Mr.  A.  C.  Barwick,  who  collected 
the  specimen,  states  that  he  has  never  found  the  tree  to  attain  a 
height  of  over  30  feet. 


540  FLORA    OF    AUSTRALIA, 

E.  QUADRANGULATA,  Dea7ie  <k  Maiden. — This  rather  rare  species 
has  recently  been  found  at  Putty,  south  of  Singleton  (A.  C. 
Barwick),  so  far  its  northern  limit. 

E.  RADiATA,  Sieb. — Occurs  at  Putt}^  via  Singleton,  on  the  banks 
of  creeks,  where  it  is  known  as  "Peppermint,"  "  Water  Gum," 
and  "  Box  "  (A.  C.  Barwick). 

ONAGREiE. 

Epilobium  glabellum,  Forst.  — Another  remedy,  it  appears,  is 
now  to  be  added  to  those  already  recorded  for  this  species,  as 
Mr.  J.  A.  Daley  reports  that  it  is  greatly  used  at  Bowna,  Albury, 
in  cases  of  dysenterj^,  and  with  marked  success  in  chronic  cases. 

COMPOSITE. 

Olearia  glutescens,  F.v.M. — Girilambone  (W.  Bauerlen). 
Previously  only  recorded  from  the  southern  coast  district  and 
southern  interior. 

O.  alpicola,  F.v.M. — Previously  recorded  from  the  Southern 
Dividing  Range,  at  high  elevations;  but  it  has  recently  been 
found  at  Mount  Mackenzie,  Tenter jfield  (W.  Bauerlen). 

O.  MYRSTNOiDES,  F.V.M. — This  southern  species  also  occurs  at 
the  same  place  (W.  Bauerlen). 

Brachycome  scapigera,  DC. — This  hitherto  reported  southern 
species  is  now  found  to  occur  as  far  north  as  Tenterfield  (W. 
Bauerlen). 

Erechtites  mixta,  DC. — This  species  has  an  extensive  range 

along  the  coast  district  of  New  South  Wales,  having  been  collected 

at  Tenterfield  in  the  north  and  Clyde  Mountain  in  the  south  (W. 

Bauerlen). 

ARALIACE^. 

Astrotricha  ledifolia,  DC. — The  previously  known  range, 
Port  Jackson  to  Victoria,  of  this  species  is  now  much  extended, 
as  it  is  now  found  to  occur  as  far  north  as  Tenterfield  and  west 
to  Narrabri  (W.  Bauerlen). 


BY    R.    T.    BAKER.  541 


L0RANTHACE5:. 


LoRANTHUS  BiDWiLLi,  Benth. — The  only  locality  in  New  South 
Wales  previously  recorded  for  this  species  is  Murrumbo  (R.  T. 
Baker);  it  has,  however,  recently  been  obtained  at  Narrabri  on 
Casuarina  Luehmanni,  R.  T.  Baker  (W.  Bauerlen). 

RUBIACE^. 

Knoxia  corymbosa,  Willd. — Woodbarn  is  now  its  most  southern 

locality  in  New  South  Wales  (W,  Bauerlen). 

ASCLEPIADE^. 

Lyons  I A  straminea,  R.Br.  —Putty,  south-west  of  Singleton,  its 
northern  limit  so  far  (A.  C.  Barwick). 

ILLECOBRACE^. 

ScLERANTHUs  BiFLORUS,  Hook.  f. — Mount  Mackenzie,  Tenter- 
field  (W.  Bauerlen). 

POLYGONE^. 

Emex  australis,  Steinb. — This  maritime  plant  has  been  found 
at  New  Angledool  (A.  Paddison)  in  the  northern  interior,  where- 
it  is  locally  known  as  "  Wild  Rhubarb." 

In  Western  Australia  it  is  known  as  "  Doublegee." 
MuEHLENBECKiA  ADPRESSA,   Meissii. — The  Nepean  River    has 
been  regarded   as  the   usual   northern   limit   of   this  species;    it 
extends,  however,  as  far  north  as  Tenterfield  (W.  Bauerlen), 

EUPHORBIACE^. 

Phyllanthus  thymoides,  Sieh. — A  rather  interesting  red 
variet}"  of  this  plant  occurs  at  Bateman's  Bay  (W.  Bauerlen). 

P.  GuNNii,  Hook./. — Only  previously  recorded  from  the  Two- 
fold Bay  district,  but  it  has  now  been  found  in  the  dry  interior 
at  Palesthan,  Condobolin  (M.  A.  C). 

CASUARINE^. 

Casuarina  Luehmanni,  R.  T.  Baker. — Under  the  local  name 
of  "  Bull-Oak"  (W.  Bauerlen)  this  tree  occurs  at  Narrabri,  which 
is  thus  its  nearest  recorded  eastern  limit. 
36 


542  FLORA    OF    AUSTRALIA, 

LYCOPODIACE^. 

Lycopodujm  ceknuum,  Linn. — E\'ans  River,  between  the  Rich- 
mond and  Clarence  Rivers  (W.  Baiierlen).  This  is  its  most 
southern  locality  recorded. 

FILICES. 

*PoLYPODiUM  AUSTRALE,  Metten.,  var.  alpicola,  F.v.M.  —  Bull- 
rock  Mountain,  Snowy  Mountains  (W.  Biiuerlen). 

Lomaria  capensis,  Willd.,  var.  Baeuerleni,  Bailey. — This 
variety  was  first  discovered  by  Mr.  Bauerlen  at  Milton  in  1884, 
but  so  far  it  has  not  been  recorded  for  New  South  Wales.  It 
was  described  by  Bailey  in  his  Supplement  to  the  '  Fern  World 
of  Australia'  (p.  4).  Baron  von  Mueller  in  writing  to  the  finder 
stated  that  if  it  should  prove  to  be  not  merely  local  it  might  be 
looked  upon  as  a  distinct  species. 

FUNGI. 

^Agaricus  (Pleurotus)  lividulus,  B.  c6  C. — Tantawanglo 
Mountain  (W.  Bauerlen). 

*A  DiVERSiPES,  Berk. — Brown's  Camp,  Delegate  (W.  Bauerlen). 

*BoLETUS  gbanulatus,  Linn. — Tingiringi  Mt.,  Delegate  (W. 
Bauerlen). 

*PoLYPORUS  SQUAMOSUS,  Huds. — Tingiringi  Mt.  (W.  Bauerlen). 

POLYPORUS    MYLITT^,   Cke.  d'  MasS. 

(Plates  xxii.-xxiii.) 

The  sclerotium  of  this  species  is  of  common  occurrence  in  the 
eastern  States  of  Australia  as  well  as  in  Tasmania  under  the 
name  of  "  Black  Fellows'  Bread,"  but  onl}^  one  specimen  in  fruc- 
tification has  so  far  been  recorded,  and  that  was  obtained  from 
South  Australia. 

A  good  specimen  of  the  "  bread  "  weighing  8  lbs.,  and  measur- 
ing 8  inches  long  and  5  in  diameter,  was  recently  received  by  me 

*  New  for  New  South  Wales. 


BY    R.    T.    BAKER.  543 

from  Mr.  A.  O.  Black,  who  procured  it  from  Jerula,  Black 
Springs,  Lithgow.  Unfortunately,  or  perhaps  fortunately  as 
results  proved,  it  had  been  cut  in  halves.  After  keeping  it  in 
the  herbarium  for  a  week  without  moisture,  the  exposed  septa  of 
one  half  began  to  throw  out  innumerable  hyphae  which  eventually 
covered  the  whole  surface,  those  of  the  other  half  remaining 
quiescent  during  this  period. 

At  the  end  of  seven  days  portion  of  the  hyphse  appeared  to  gravi- 
tate towards  one  end  of  tlie  edge  and  to  accumulate  into  a  mass 
which  assumed  a  pufF-ball  shape,  and  four  days  later  developed 
a  pileus,  the  upper  surface  of  which  was  at  first  of  an  orange- 
brown  colour  in  the  centre  with  a  mauve  band  of  four  lines 
broad  slightly  removed  from  the  edge,  the  under  surface  showing 
the  pores  of  a  Polyporus.  It  gradually  grew  till  it  attained  a 
diameter  of  3  inches,  when  it  began  to  harden  and  eventually 
lost  its  colour.  As  the  process  of  induration  proceeded,  the 
edges  gradually  recurved,  and  eventually  in  its  final  stage  the 
whole  much  resembled  P.  ovinus,  Schseff.  The  pores  were 
characteristic  of  the  genus,  but  unfortunately  no  spores  were 
detected. 

The  second  half  was  then  taken  in  hand  and  placed  on  soil 
and  kept  moist.  The  hyphse  soon  became  active,  and  at  the  end 
of  a  week  the  whole  surface  of  the  gelatinous  cells  exposed  in  the 
section  was  covered  with  a  perfectly  white  mass  of  mycelium. 
The  convex  side  was  then  buried  in  soil,  except  a  small  part  of 
one  end  which  showed  signs  of  fructification,  and  well  watered. 
From  the  exposed  end  protuberances  began  to  form,  and  eventu- 
ally three  developed,  the  largest  being  a  fine  specimen 
measuring  over  4  inches  in  diameter  and  with  exactly  the  same 
markings  and  colour  as  the  first  obtained  and  described  above, 
and  like  it  resembling  a  poached  egg  in  colour  and  form  when 
viewed  from  above.  The  stipes  was,  however,  more  fan-shaped 
than  in  the  previous  specimen.  The  hyphse  next  developed  into 
a  thick,  porous,  foamy  mass  covering  almost  the  whole  surface 
and  eventually  spreading  over  and  into  the  adjacent  soil  opposite 
to  the  exposed  end.     Two  small,  cup-shaped  specimens  were  the 


544  FLORA    OF    AUSTRALIA. 

last  to  appear  on  the  cut  surface  of  this  half  of  the  "  bread." 
At  the  end  of  a  month  it  formed  a  very  interesting  specimen. 
Attempts  to  detect  spores  were  again  fruitless,  the  spore-bearing 
stage  appearing  not  to  be  reached,  probabl}^  owing  to  the  artificial 
method  of  propagation. 

■^PoLYSTiCTUS  TRIZONATUS,  Cooke. — Snowy  River  ( W.Bauerlen). 

*Hexagonia  decipiens,  Berk. — Snowy  River  (W.  Bauerlen). 

*HYMEN0CHiETA  MouGEOTTii,  FHes. — Tantawanglo  Mt. 

jPatellaria  (Patixella)  filicicola. — Cambewarra  (W.  Bau- 
erlen). 

LICHENES. 

*Parmelia   limbata,   Laurer. — Bangley  Creek    (Shoalhaven); 

(W.  Bauerlen). 

*Parmelia  perforata,  Achorius. — Cambewarra  (W.  Bauerlen). 

*Sph^rophoron    tenerum,    Laurer. — Sugar    Loaf    Mt.    (W. 

Bauerlen). 

MUSCI. 

•^Brentelina    Baeuerlexi,    Cke.   6c    Mass. — The    Clyde    (W. 
Bauerlen). 


EXPLANATION  OF  PLATES  XXII.-XXIIL 

PolyporiLS  mylittce,  Cke.  &  Mass. 

Plate  xxii. 

Exposed  end  of  one  section  showing  fructification-stage  developing  from 
the  outer  coating  of  the  sclerotium. 

Plate  xxiii. 
Half  of  the  sclerotium  with  the  fructification  developing  from  the  cut 
surface. 


*  New  for  New  South  Wales, 
t  Not  previously  recorded  for  Australia. 


t 


545 


ON  THE  MAMMALIAN  AND  REPTILIAN  VOMERINE 

BONES. 

By  R.  Broom,  M.D.,  B.Sc,  C.M.Z.8.,  Corresponding  Member. 

(Plates  xxiv.-xxvi.) 

Eor  a  hundred  years  and  more  almost  every  anatomist  of  note 
has  agreed  in  regarding  the  paired  elements  which  lie  immediately 
behind  the  premaxillaries  in  the  reptilian  palate  as  together 
homologous  with  the  mammalian  vomer ;  and  the  opinion  of 
authorities  has  been  so  unanimous  that  it  is  naturally  with  some 
hesitation  that  one  ventures  to  suggest  that  the  currently  accepted 
view  is  probably  incorrect.  It  is  quite  likely  that  this  view  might 
have  long  continued  to  be  almost  universally  accepted  had  not 
the  difficulty  of  explaining  the  homology  of  the  dumbbell-shaped 
bone  of  Urnithorhynchus  forced  us  to  a  reconsideration  of  the 
position.  This  little  bone,  whose  anatomical  relations  and 
development  are  now  so  well  known,  lies  beneath  the  nasal  septum 
and  clasps  the  cartilages  of  Jacobson.  At  no  stage  in  its 
development  is  it  connected  with  either  the  vomer  behind  or  the 
premaxillaries  in  front.  The  earlier  anatomists,  Rudolphi, 
Meckel,  and  Owen,  very  naturall}^  agreed  in  finding  the  homologue 
of  this  little  bone,  which  lies  in  front  of  the  vomer,  in  the  palatine 
processes  of  the  premaxillae  of  the  higher  forms;  and  in  more 
recent  times  this  view  has  been  supported  by  Albrecht,  Turner, 
Symington,  and  myself.  But  if  this  view  be  correct,  it  becomes 
necessary  to  regard  the  premaxillary  as  a  complex  bone,  and  as 
possibly  representing  more  than  the  premaxillary  of  the  reptilian 
or  amphibian  ancestors.  Before  looking  more  closely  into  the 
problem  it  may  be  well  to  briefly  review  one  or  two  recent  papers 
dealing  with  the  subject. 


546  MAMMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

In  1879  Albrecht  (1)  published  a  paper  dealing  with  the 
ossification  of  the  premaxillary  bone,  and  advanced  some  reasons 
for  believing  that  this  bone  is  a  complex  element,  its  palatine 
process  being  frequently  formed  from  a  distinct  centre.  In  a 
later  paper  he  advocated  the  homology  of  the  dumbbell-shaped 
bone  of  Omithorhynchus  with  the  palatine  process  of  the  pre- 
maxillary in  the  higher  mammals. 

In  1884  Bland  Sutton  (2)  communicated  a  paper  to  the  Zoolo- 
gical Society  of  London  mainly  on  the  question  under  consideration. 
In  it  he  not  only  held  that  the  palatine  process  is  a  distinct 
morphological  element,  but  that  it  is  the  homologue  of  the 
so-called  "vomer"  of  the  Ichthyopsida,  and,  further,  that  the 
mammalian  vomer  has  its  homologue  in  the  "  parasphenoid  "  of 
the  lower  forms. 

In  1885  Turner  (3),  writing  on  the  homology  of  the  dumbbell 
bone  of  Omithorhynchus,  came  to  the  conclusion  that  it  is  the 
homologue  of  the  palatine  process  of  the  premaxillary. 

In  the  same  year  Kitchen  Parker  (4)  published  his  magnificent 
monographs  on  the  development  of  the  skull  in  the  Insectivora 
and  Edentata.  In  a  number  of  forms  he  found  evidence  of  the 
palatine  process  being  developed  as  a  distinct  element.  To  this 
element  he  gave  the  name  of  "anterior  paired  vomer,"  and 
recognised  it  to  be  the  homologue  of  the  paired  "vomer"  of  the 
Lacertilia  and  Ophidia.  In  many  animals  he  also  recognised  the 
existence  of  a  pair  of  posterior  or  lateral  vomers.  Of  Rhynchocyon 
he  says : — "There  are  here  five  vomerine  bones,  as  in  the  Hedgehog 
and  Armadillo.  .  .  ,  The  two  antero-lateral  vomers  are  the 
largest  I  have  seen  as  yet,  reminding  one  of  their  large  cupped 
counterparts  in  the  Ophidia  and  Lacertilia.  .  .  .  The  narrow 
hind  part  of  the  main  vomer  is  partly  hidden  by  the  two  large 
postero-lateral  vomers."  The  Marsupials,  Parker  considers,  have 
"many  separate  vomers" — the  genus  Cuscus  having  as  many  as 
ten.  It  may  be  remarked,  however,  that  these  observations  of 
Parker's  require  confirmation. 

In  1891  Howes  (6)  read  a  paper  before  the  Zoological  Society 
of  London  on  the  peculiar  condition  of  the  vomer  in  the  Caiman. 


BY    R.    BROOM.  547 

He  showed  that  in  this  Crocodilian  the  vomer  has  its  anterior  end 
greatly  developed,  and  that  in  connection  with  this  anterior 
development  there  is  a  cartilaginous  sac,  which  he  concluded  was 
most  probably  the  cartilaginous  capsule  of  Jacobson's  organ.  He 
discusses  at  some  length  the  question  of  the  homology  of  the 
bony  supports  of  Jacobson's  organ  in  mammals  and  lizards;  and 
concludes  that  what  he  calls  the  prepalatine  lobe  of  the  vomer  in 
the  Caiman  is  homologous  with  the  "anterior  paired  vomer"  of 
Parker  in  mammals.  These  paired  vomers  he  regards  as  morpho- 
logicalty  portions  of  the  vomer  proper  which  have  become  separated 
by  absorption.  According  to  this  view  the  lacertilian  "  vomer  " 
would  be  morphologically  equivalent  to  the  mammalian  vomer 
and  anterior  vomers  combined. 

In  1894  Wilson  (6),  of  Sydney,  published  a  paper  giving  a  very 
full  description  of  the  dumbbell-shaped  bone  of  OrnitJiorhynchiis, 
and  discussing  at  some  length  the  homologies  of  the  bone.  He 
was  specially  impressed  by  the  fact  of  its  relationship  to  the 
vomer  being  apparently  much  more  important  than  its  relation- 
ship to  the  premaxillaries;  and  was  led  to  conclude  that  the 
dumbbell  bone  is  no  part  of  the  premaxillary,  but  a  true  anterior 
vomer  homologous  with  the  small  anterior  vomers  discovered  by 
Parker  in  the  early  stages  of  a  number  of  long-nosed  mammals.  He 
seems  inclined  to  the  opinion,  though  he  does  not  explicitly  state 
it,  that  the  dumbbell  bone  is  also  the  homologue  of  the  lacertilian 
so-called  "vomer." 

In  the  following  year,  having  been  for  some  time  engaged  in 
the  study  of  the  comparative  anatomy  of  Jacobson's  organ,  and 
having  come  across  one  or  two  new  facts  bearing  on  the  subject, 
I  (7)  published  a  short  paper  on  the  homology  of  the  palatine 
process  of  the  mammalian  premaxillary.  In  the  Marsupials  which 
I  had  studied,  I  had  found  the  palatine  process  to  be  invariably 
developed  along  with  the  premaxillary,  and  with  no  trace  of  a 
distinct  anterior  vomer;  but  as  the  palatine  process  invariably 
formed  a  splint  to  the  cartilage  of  Jacobson,  similar  to  that 
formed  by  the  dumbbell  bone  in  Ornithorhynchus,  and  having 
relations   very  similar    to    those    of    the    so-called    "vomer"    in 


548  MAMxMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

lizards,  I  arrived  at  the  conclusion  that  the  process  is  morpho- 
logically an  anterior  vomer  even  when  not  formed  from  a  distinct 
centre,  and  is  the  homologue  of  the  dumbbell  bone  of  Orniiho- 
rhynchus  and  of  the  so-called  "vomer"  in  at  least  lizards  and 
snakes.  For  this  element  I  proposed  the  name  of  pre  vomer  as 
more  in  harmony  with  the  general  terminology  than  "anterior 
vomer." 

In  1896  Symington  (8),  who  had  some  years  previously  described 
the  dumbbell  bone  of  Ornithorhynchus,  replied  to  Wilson,  and 
showed  that  the  dumbbell  bone  in  most  of  its  relations  agrees 
with  the  palatine  process  of  the  premaxillary  with  which  he 
regards  it  as  homologous.  He  is  not  inclined  to  regard  the 
palatine  process  as  vomerine,  since  it  is  only  exceptionally  formed 
from  a  distinct  centre. 

This  year  Wilson  (9)  has  published  a  most  interesting  paper  on 
the  anatomy  of  the  snout  of  the  young  Ornithorhynchus  and 
Echidna,  of  which  a  preliminary  notice  was  issued  two  years  ago. 
In  this  paper  he  confirms  the  observation  of  Newton  Parker  that 
the  dumbbell  bone  is  developed  as  two  splints  to  Jacobson's 
cartilages,  and  further  shows  that  in  the  early  young  the  pre- 
maxillaries  not  only  meet  in  the  middle  line  but  send  back 
palatine  processes.  These  processes  are  quite  distinct  from  the 
dumbbell  bone  ending  some  distance  in  front  of  its  developing 
splints. 

Van  Bemmelen  (10)  also  last  year  published  an  important  paper 
on  the  Monotreme  skull  in  which  he  supports  the  pre  vomerine 
theory  of  the  dumbbell  bone.  I  regret,  however,  that  I  have  not 
yet  seen  the  paper. 

The  Development  of  the  Mammalian  Premaxillary. 

The  large  majority  of  anatomists  have  always  regarded  the 
premaxillary  bone  with  its  palatine  process  as  a  single  element, 
and  I  think  it  may  be  regarded  as  an  undoubted  fact  that  in  the 
large  majority  of  mammals  the  palatine  process  is  developed  along 
with  the  premaxillary,  and  that  only  in  comparatively  few  forms 


BY    R.    BROOM.  549 

is  the  process,  or  even  a  part  of  it,  ever  formed  from  a  distinct 
centre. 

In  Marsupials,  so  far  as  they  have  been  examined,  the  palatine 
process  is  never  distinct. 

If  the  results  of  Kitchen  Parker's  work  on  the  development  of 
the  skull  in  the  Edentata  and  Insectivora  were  fully  confirmed 
there  would  be  no  longer  any  doubt  that  in  a  number  of  forms 
at  least  the  palatine  process  is  a  distinct  element  which  at  only 
a  comparatively  late  stage  in  development  unites  with  the  pre- 
maxillary.  Parker,  in  fact,  seems  to  consider  this  to  be  the 
normal  condition,  since  he  states  that  the  "  anterior  paired 
vomers  are  very  constant  in  Mammals."  He  describes  distinct 
"anterior  vomers  "  in  the  following  forms  : — Tatusia  hybrida,  T. 
peha,  Dasypus  villosus,  ChoIo2?us  didactylus,  Manis  brevicaudata, 
Orycteropus  capensis,  Eriyiaceus  europceus,  and  Bhynchocyon 
cernei.  In  Centetes^  Hemicentetes,  and  GaleopitltecvSj  Parker 
apparently  satisfied  himself  that  there  are  no  distinct  "anterior 
vomers."  In  the  mole  and  shrew  the  "  anterior  paired  vomers  " 
are  stated  to  be  "probably  connate  with  the  premaxillary." 

When  one  considers  what  a  small  structure  the  palatine  process 
is  even  in  the  adult  animal,  and  that  the  greater  part  of  Parker's 
results  were  obtained  by  the  dissection  of  small  embryos,  one 
hesitates  in  accepting  as  final  the  results  of  even  such  a  marvellous 
minute  anatomist  as  Parker.  And  the  hesitation  becomes  greater 
when  one  finds  Parker  on  one  page  contradicting  a  statement 
made  only  a  few  pages  before.  Thus,  on  page  166,  we  are  told 
that  the  anterior  paired  vomers  of  the  mole  "  are  very  slightly 
separated  from  the  palatine  processes  of  the  premaxillaries,"  and 
on  page  179  that  they  have  a  "doubtful  existence  independent 
of  these  processes." 

I  regret  that  I  have  not  had  at  my  disposal  embryos  of  any  of 
the  Edentates  studied  by  Parker  ;  but  in  young  Hedgehogs, 
corresponding  to  Parker's  4th  and  5th  stages,  I  have  been  unable 
to  find  any  distinct  "  anterior  vomers."  Recently  I  made  a  series 
of  sections  of  the  snout  of  a  foetal  Macroscelides  proboscideus 
(head  length  15*5  mm.) — a  form  closely  allied  to  Rhynchocyon, 


550  MAMMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

and  only  slightly  less  developed  than  the  foetus  studied  by  Parker. 
The  premaxillaries  were  well  developed,  and  had  large  palatine 
processes  which  ran  far  back  by  the  sides  of  Jacobson's  cartilages, 
but  there  was  no  trace  of  a  distinct  "  anterior  vomer,"  and  in  the 
gradual  thinning  of  the  plates  as  they  pass  backwards  there  was 
prett}'-  conclusive  proof  that  the  plates  had  been  ossitied  from  the 
premaxillaries. 

It  seems  probable  to  me  that -further  research  will  show  that 
in  none  of  the  Insectivora  are  there  any  distinct  "anterior 
vomers,"  but  on  the  other  hand  it  is  quite  likely  that  Parker  is 
right  in  finding  distinct  "anterior  vomers"  in  most  of  the 
Edentates. 

In  only  a  very  few  of  the  higher  mammals  have  we  any  record 
of  the  palatine  process  being  formed  from  a  distinct  centre. 
Howes  records  it  in  the  case  of  the  rabbit,  and  Schwink  in  the 
fcetal  sheep.     Probably  some  other  instances  will  be  recorded. 

In  the  Chiroptera  we  find  a  distinct  palatine  process  very 
generally  absent,  owing,  no  doubt,  to  the  fact  that  Jacobson's  organ 
is  as  a  rule  wanting,  and  also  to  the  fact  that  the  premaxillaries 
frequently  do  not  meet  in  the  middle  line.  In  Miniopteras,  one 
of  the  few  bats  in  which  the  organ  of  Jacobson  is  known  to  occur, 
we  have  the  cartilage  of  the  organ  supported  by  a  median  bone 
almost  exactly  after  the  manner  of  the  dumbbell  bone  in  Orni- 
thorhynchus.  As  in  Ornithorhynchus,  too,  this  median  bone  is 
formed  by  the  anchylosis  of  a  pair  of  bony  splints.  And,  though 
in  adult  life  such  extensive  anchylosis  takes  place  that  even  the 
vomer  becomes  anchjdosed  to  the  maxillaries,  the  little  median 
bone  remains  quite  free. 

It  will  thus  be  seen  that  while  in  most  mammals  the  cartilage 
of  Jacobson's  organ  is  supported  by  a  bony  process  from  the 
premaxillary,  in  a  few  forms  {Orniihorhynchus^  Minioj^terus),  the 
cartilage  is  supported  by  a  distinct  element ;  while  in  others 
apparently  we  have  an  intermediate  condition  in  that  the  element 
which  supports  the  cartilage  is  at  first  independent,  and  then 
anchyloses  with  the  premaxillary  to  form  its  palatine  process. 


BY    R.    BROOM.  551 

The  question  now  arises  whether  the  independent  element  in, 
say,  07'7iithorhynchu>i,  is  homologous  with  the  palatine  process  in 
the  majority  of  mammals.  Unquestionably  both  occupy  corre- 
sponding morphological  tracts,  and  by  the  large  majority  of 
anatomists  they  have  thus  been  regarded  as  homologous,  and 
such,  until  recently,  was  my  own  opinion. 

The  skeleton  in  mammals  differs  from  that  of  their  Theriodont 
ancestors  mainly  in  the  reduction  of  the  number  of  skeletal 
elements.  In  very  many  instances  we  find  a  certain  region  in 
the  Theriodont  occupied  by  two  elements,  and  in  the  mammal 
the  corresponding  morphological  region  occupied  by  only  one. 
Where  this  occurs  we  have  in  most  cases  direct  evidence  that 
what  has  happened  is  that  one  of  the  two  elements  has  become 
steadily  reduced  in  size,  and  ultimately  completely  replaced  by 
the  larger  element,  and  not  that  the  single  element  represents  a 
fusion  of  the  original  two.  And  we  have  further  evidence  that 
the  element  which  replaces  the  other  has  the  form  of  the  newly 
developed  part  determined  by  the  element  which  it  replaces. 

Now,  with  regard  to  the  palatine  process  of  the  premaxillary, 
it  seems  much  more  satisfactory  to  regard  it  as  a  true  portion  of 
the  premaxillary  which  has  replaced  an  anterior  vomerine  element, 
such  as  the  dumbbell  bone  of  Ornitliorliynchus,  than  to  regard  it 
as  itself  the  homologue  of  that  bone.  We  know  that  in  the 
higher  Theriodonts  (H)  {e.g.,  Gomij^liognaihus)  the  tract  occupied 
by  the  palatine  process  in  mammals  is  occupied  by  a  pair  of 
apparently  distinct  small  bones.  The  dumbbell  bone  of  Ornitho- 
rhynchus  and  the  little  median  bone  of  MiniojyteA-ics  represent 
these  bones  of  Gomi)hognaihiis  anchylosed  together.  If  Parker's 
researches  are  confirmed  we  shall  have  in  some  of  the  Edentates 
at  least  the  condition  where  a  rudimentary  anterior  vomerine 
element  is  being  replaced  by  a  palatine  process  from  the  pre- 
maxillary, and  to  which  in  the  adult  it  is  anchylosed.  In  the 
large  majority  of  higher  mammals  the  anterior  vomer  is  completely 
aborted,  and  its  place  and  functions  taken  by  the  replacing 
palatine  process  of  the  premaxillary. 


00 2  mammalian  and  reptilian  vomerine  bones, 

The  Lacertilian  "Yomer." 

The  bones  in  the  lizard  which  have  usually  been  regarded  as 
vomers  agree  with  the  mammalian  vomer  in  lying  for  the  most 
part  in  the  anterior  half  of  the  skull,  and  in  articulating  with  the 
premaxillaries  and  the  palatines.  But  here  the  resemblance 
ceases;  and  when  we  compare  the  intimate  relations  of  the  bones 
we  tind  it  impossible  to  regard  them  as  homologous.  The 
mammalian  vomer  is  a  median  element  developed  in  connection 
with  that  part  of  the  basicranial  axis  which  is  formed  by  the 
united  trabeculse;  the  lacertilian  "  vomer  "  is  a  paired  element 
which  is  formed  in  connection  with  the  nasal  capsules. 

Though  many  figures  of  sections  of  the  nasal  capsules  of  lizards 
and  snakes  have  been  published  by  Born,  Parker,  Seydel,  and 
others,  perhaps  the  best  illustration  is  a  figure  by  Seydel  (12)  of 
a  reconstructed  model  of  the  nasal  capsule  of  Lacerta,  in  his 
paper  on  the  organ  of  Jacobson  of  Chelonia.  From  this  figure, 
which  I  have  reproduced,  it  will  be  seen  that  the  nasal  capsule 
consists  of  an  irregular  cartilaginous  sac  with  three  openings,  one 
for  the  anterior  and  one  for  the  posterior  nasal  opening,  and  a 
third  opening  towards  the  brain  to  admit  the  nasal  nerves.  The 
external  wall  is  much  folded,  and  the  inner  wall  is  to  some  extent 
formed  by  the  nasal  septum.  How  much  of  the  septum  belongs 
to  the  nasal  capsule,  and  how  much  is  trabecular,  is  difficult  to 
decide.  By  the  side  of  the  base  of  the  septum  is  a  long  narrow 
cartilaginous  band  which  is  united  to  the  nasal  floor  proper  in 
front,  and  to  the  posterior  wall  of  the  capsule  behind,  and  which 
forms  to  some  extent  the  lower  and  inner  wall  of  the  capsule. 
This  is  the  paraseptal  cartilage.  If  this  lacertilian  capsule  be 
compared  with  the  nasal  capsule  of  the  marsupial  it  will  be  seen 
that  the  two  are  essentially  similar  in  structure — the  marsupial 
capsule  only  differing  in  being  somewhat  simpler.  In  the 
marsupial — at  least  in  the  fcetal  condition — the  paraseptal 
cartilage  stretches  from  the  nasal  floor  in  front  to  the  hinder  part 
of  the  nasal  capsule  exactly  as  in  the  lizard.  B}^  Parker  the 
paraseptal   cartilage    was    regarded    as    a    development    of    the 


BY    R.    BROOM.  553 

trabecular  cornu,  and  any  one  who  had  only  studied  it  in  the 
rudimentary  condition  in  which  it  is  found  in  the  higher  mammals 
might  readily  come  to  that  conclusion.  The  posterior  attachment 
of  the  cartilage,  however,  in  the  lizard  and  marsupial  proves,  I 
believe,  that  the  paraseptal  is  a  true  portion  of  the  nasal  capsule. 

Whatever,  however,  be  the  morphological  significance  of  the 
paraseptal  cartilage  there  is  no  doubt  that  it  is  in  connection 
with  this  cartilage  that  the  so-called  "vomer  "of  the  lizard  is 
formed.  On  transverse  section  the  cartilage  is  seen  passing 
downward  and  a  little  outwards  from  near  the  base  of  the  septum, 
and  it  is  as  a  splint  on  the  inner  side  of  this  cartilage  that  the 
"  vomer "  first  develops.  It  soon,  however,  passes  outwards 
round  the  lower  border  of  the  cartilage,  and  forms  to  some  extent 
a  floor  to  the  nasal  capsule;  but  though  it  extends  laterally 
beyond  the  limits  of  the  cartilage  its  length  is  determined  b}^  the 
length  of  the  paraseptal. 

In  Ornithorhynchus  and  Minioi^teriis  the  paraseptals,  being 
retained,  as  in  most  mammals,  only  as  supports  to  Jacobson's 
organ,  have  their  length  determined  by  the  length  of  the  organ, 
and  are  hence  much  shorter  than  the  nasal  capsule.  On  the 
inner  side  of  each  cartilage  a  splint  bone  develops  exactly  as  in 
the  lizard,  and,  as  in  the  lizard,  the  length  of  these  splints  is 
determined  by  the  length  of  the  cartilages.  In  both  Ornitho- 
rhynchus and  Miniopterus  these  splints  in  their  later  development 
become  anchylosed,  in  the  one  case  to  form  the  dumbbell  bone, 
and  in  the  other  the  little  median  bone  which  lies  in  front  of  the 
vomer. 

As  the  paraseptal  cartilage  in  the  marsupial  is  undoubtedly 
the  true  homologue  of  the  paraseptal  cartilage  in  the  lizard,  the 
paraseptal  in  Ornithorhynchus,  even  though  somewhat  rudimen- 
tary, must  also  be  homologous.  It  is,  therefore,  impossible  to 
doubt  that  the  splint  bones  which  form  on  their  inner  sides,  and 
whose  lengths  are  determined  by  the  cartilages,  must  also  be 
homologous.  As  the  so-called  "  vomer  "  of  the  lizard  is  thus  seen 
to  be  the  homologue  of  the  dumbbell  bone  of  Ornithorhynchus,  it 
can  no  longer  be  regarded  as  a  true  vomer,  since  the  dumbbell 


554  MAMMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

bone  is  not  the  vomer  of  OrnithorliyncJms^  and  must  in  future  be 
referred  to  b}^  a  new  name.  The  name — Prevomer — which  I 
originally  suggested  (7)  seems  the  most  appropriate. 

In  the  large  majority  of  the  reptilian  orders  the  so-called 
''  vomers  "  are  undoubtedly  homologous  with  the  prevomers  of  the 
lizard.  This  is  the  case  in  the  Ophidia,  Rhynchocephalia, 
Plesiosauria,  Ichthyosauria,  Pelycosauria,  Dinosauria,  and  Pareia- 
sauria.  In  the  Theriodontia  and  Anomodontia  the  bone  which 
has  been  referred  to  as  the  vomer  is  the  true  homologue  of  the 
mammalian  vomer,  and  this  is  almost  certainl}'-  also  the  case  in 
the  Chelonia. 

On  the  "Paraspiienoid  "  and  the  Mammalian  Vomer. 

While  the  pair  of  bones  in  the  front  of  the  palate  in  the 
Amphibia  and  other  forms  were  regarded  as  the  homologues  of 
the  mammalian  vomer,  it  became  necessar}^  to  find  another  name 
for  the  large  median  membrane  bone  that  supports  the  basicranial 
axis,  and  the  name  "  paraspiienoid  "  proposed  by  Huxley  has  been 
almost  universally  adopted. 

This  bone  is  greatly  developed  in  the  Amphibia,  where,  as  a 
rule,  it  forms  a  very  complete  floor  to  the  cartilaginous  cranium, 
extending  from  the  basioccipital  region  to  near  the  anterior  part 
of  the  palate.  It  is  invariably  a  median  bone,  and  is  ossified 
from  a  single  centre.  According  to  Parker,  the  "paraspiienoid" 
first  appears  in  the  frog  as  "  a  parostosis  beneath  the  intertrabe- 
cular  space." 

Among  reptiles  the  "  parasphenoid,"  though  almost  alwaj^s 
present,  is  as  a  rale  feebly  developed.  In  the  primitive  reptile 
Sphenodon  its  development  has  recentl}^  been  carefull}'  worked 
out  by  Howes  and  Swinnerton  (13),  according  to  whom  "  the 
paraspiienoid  first  appears  in  the  form  of  a  minute  bony  element 
.  lying  freety  beneath  the  antero-median  border  of  the 
post-pituitary  plate."  The}^  add  that  "  with  advancing  develop- 
ment it  assumes  an  arrow-head  shape." 

In  the  lizard,  Parker  (14)  figures  the  first  appearance  of  the 
"  parasphenoid  "  as  a  minute  median  ossification  lying  just  in 
front  of  the  pituitary  space. 


BY    R.    BROOxM.  555 

In  the  snake  the  "  parasphenoid  "  is  moderately  well  developed, 
and  extends  far  forwards.  In  its  earliest  stage  of  development 
it  is  described  by  Parker  (15)  as  "a  lx)ng  styloid  tract  of  granular 
tissue  .  .  .  wedged  between  the  trabeculag "  and  reaching 
*'  the  pituitary  body  by  its  broad  hinder  end." 

It  will  thus  be  seen  that  in  both  amphibians  and  reptiles  the 
'' parasphenoid "  is  a  median  ossification  which  develops  in  the 
neighbourhood  of  the  pituitary  space — occasionally  arising  behind 
the  space  as  in  Sjyhenodon,  and  at  other  times,  as  in  the  snake, 
having  its  entire  development  in  front  of  the  space. 

Now,  the  mammalian  vomer  is  developed  as  a  median  splint 
bone  along  that  part  of  the  basicranial  axis  which  in  later 
development  becomes  the  presphenoid  and  mesethmoid  regions; 
and  as  this  is  exactl}^  the  region  in  which  the  median  splint  bone 
of  the  snake  is  developed,  we  are  forced  to  conclude  that  the 
"parasphenoid"  of  the  snake  is  really  the  homologue  of  the 
mammalian  vomer.  In  the  reptiles,  no  doubt  the  exact  position 
of  the  median  bone  varies  considerably;  but  so  in  mammals  does 
the  vomer  vary  greatly  as  regards  its  anterior  and  posterior 
developments.  In  most  mammals  the  vomer  extends  far  forwards, 
and  in  the  Cetaceans  extends  practically  to  the  end  of  the  long 
rostrum;  but  in  Echidna,  on  the  other  hand,  the  anterior  extremity 
of  the  vomer  is  found  in  the  posterior  nasal  region;  and  while  in 
many  Rodents  we  find  the  vomer  rudimentary,  and  posteriorly  not 
even  reaching  the  presphenoid,  in  some  Cetaceans  the  vomer 
extends  back  even  to  the  basioccipital. 

The  Phylogeny  of  the  Vomerine  Bones. 

Having  shown  that  the  mammalian  vomer  has  its  homologue 
in  the  "  parasphenoid  "  of  the  reptiles  and  amphibians,  and  that 
the  so-called  "  vomers  "  of  these  lower  forms  are  really  homologous 
to  the  dumbbell  bone  of  Ornithorhynchus,  it  is  now  only  necessary 
to  consider  briefl}^  the  steps  b}'  which  the  vomerine  bones  of  the 
higher  forms  have  been  derived  from  those  of  their  Labyrinthodont 
ancestors. 


556  MAMMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

In  the  extinct  Labyrinthodonts,  as  in  their  degenerate  descen- 
dants, the  Amphibia  of  to-day,  we  find  the  basicranial  axis 
supported  by  a  powerful  median  vomer  (parasphenoid),  and  in 
front  we  have  a  pair  of  prev^omers  supporting  the  nasal  capsules. 

In  the  most  primitive  reptilian  order  with  which  we  are 
acquainted — the  Cotylosauria — we  find  that  the  palate  differs 
from  the  ancestral  Labyrinthodont  condition  mainly  in  the  greatly 
increased  development  of  the  pterygoids,  with  a  corresponding 
decrease  in  the  size  of  the  median  vomer.  Howes  has  already 
recently  suggested  that  the  reduction  of  the  "  parasphenoid  "  in 
the  ascending  reptilian  series  has  been  brought  about  by  the 
increased  development  of  the  pterygoids;  and  it  seems  moderately 
certain  that  this  is  the  true  explanation.  It  is  more  difficult  to 
assign  a  cause  for  the  increased  development  of  the  pterygoids. 
But  it  seems  probable  that  as  the  descendants  of  the  Labyrintho- 
donts took  to  terrestrial  habits  a  stronger  palate  than  could  be 
formed  by  the  median  vomer  became  advantageous,  and  was 
formed  by  the  pterygoids,  which,  besides  becoming  greatly 
increased  in  size,  became  dentigerous. 

In  the  majority  of  more  recent  reptiles,  we  have  a  modification 
of  the  condition  found  in  Sphenodon  where  there  is  a  small 
median  vomer  and  two  moderately  large  prevomers. 

In  the  Theriodonts,  however,  an  interesting  modification  arises. 
With  the  formation  of  a  secondary  palate  a  new  function  is  given 
to  the  median  vomer,  which  becomes  again  well  developed  as  a 
support  to  the  new  palate.  The  prevomers,  on  the  other  hand, 
being  now  no  longer  required  to  protect  the  nasal  capsules,  become 
greatly  reduced  in  size,  and  are  retained  only  as  supports  to  the 
cartilages  of  Jacobson, 

In  the  Dicynodonts  the  true  vomer  is  greatly  developed,  while 
the  prevomers  are  completely  lost  owing  to  the  enormous  develop- 
ment of  the  premaxillaries. 

In  the  Chelonians  we  have  a  condition  somewhat  resembling 
that  of  the  Dicynodonts,  and  probably  brought  about  by  a 
parallel  development.  The  median  vomer  is  developed  as  a 
support  to  the  palate,  and  the  prevomers  are  lost. 


BY    K.    BROOM. 


557 


In  the  early  mammals  the  condition  was  probably  very  similar 
to  that  of  the  Theriodonts,  and  this  condition  is  still  retained  in 
a  few  mammals  {Ornithorhynclmis,  Miniopterus).  In  the  large 
majority  of  recent  mammals  the  small  prevomers  have  been 
replaced  by  the  palatine  processes  of  the  premaxillaries. 

The  principal  modifications  of  the  vomer  and  prevomer  will  be 
readilv  seen  from  the  adjoining  table  : — 


Primitice  Orders — 
Labyrinthodontia 
Cotylosauria 

Peevomers. 

Vomer. 

Large. 
Large. 

Large, 
Small. 

RhynchocephaUan 
Phylum — 
Pelycosauria            ! 
Rhynchocephalia 
Lacertilia 

Large. 
Large. 

Large,  small,  or  rudimen- 
tary. 

Small. 
Small. 
Small  or  rudimentary. 

Aberrant  Order  — 
Chelonia 

Lost. 

Large. 

Tlieriodont  Phylum  — 
Theriodontia 
Anomodontia 
Monotremata 
Marsupialia 
Eutheria 

Small. 
Lost. 

Small  or  lost. 
Replaced  by  premaxilla. 
Small,  or  replaced  by  pre- 
maxilla. 

Large. 
Large. 

Large. 
Large. 
Large. 

References  to  Literature. 
1. — Albrecht,  p. 
2. — Sutton,  J.  Bland — "Observations  on  the  Parasphenoid,  the  Vomer 

and  the  Palatopterygoid  Arcade. "     P.Z.S.,  1884. 
3. — Turner,  W. — "  The  Dumb-bell-shaped  Bone  in  the  Palate  of  Ornitho- 
rhynchus  compared  with  the  Prenasal  Bone  in  the  Pig."      J.  Anat. 
and  Phys.,  Vol.  xix. 
4. — Parker,  W.  K.  —"On  the  Structure  and  Development  of  the  Skull  in  the 
Mammalia.      Pt.  ii.,  Edentata;  Pt.  ii'i.,  Lisectivora. "      Phil.  Trans., 
1885. 
5.— Howes,  G.  B. — "On  the  probable  Existence  of  a  Jacobson's  Organ 
among  the  Crocodilia,  &c."     P.Z.S.,  1891. 
37 


558  MAMMALIAN  AND  REPTILIAN  VOMERINE  BONES, 

6. — Wilson,  J.  T. — "  Observations  upon  the  Anatomy  and  Kelations  of  the 

Dumbbell-shaped  Bone  in  Ornithorhynchus,  &c."     Proc.  Linn.  Soc. 

N.S.W.,  1894. 
7. —Broom,  R.  —  "  On  the  Homology  of  the  Palatine  Process  of  the  Mamma- 
lian Premaxillary."     Proc.  Linn.  Soc.  N.S.W.,  1895. 
8. — Symington,  J. — "On  the  Homology  of  the  Dumbbell-shaped  Bone  in 

the  Ornithorhynchus."     J.  Anat.  and  Phys.,  1896. 
9. — Wilson,  J.  T. — "  On  the  Skeleton  of  the  Snout  of  the  Mammary  Foetus 

of  Monotremes."     Proc.  Linn.  Soc.  N.S.W.,  1901. 
10. — Van  Bemmelen,  J.  F.  —  "Der  Schadelbau  der  Monotremen."     Semon's 

Zoolog.  Forschungsreisen,  iii.  Band,  4  Lieferung,  1901. 
11. — Broom,  R  — '*  On  the  Occurrence  of  an  Apparently  Distinct  Prevomer 

in  Gomphognathus."     J.  Anat.  and  Phys.,  Vol.  xxxi.,  1896. 
12. — Seydel,  0. — "  Ueber  die  Nasenhohle  und  das  Jacobson'sche  Organ  der 

Land-  und   Sumpfschildkroten."     Festschrift  f.  C.  Gegenbaur,  1896. 
13. — Howes,  G.  B.,  and  Swinnerton,  H.  H. — "  On  the  Development  of  the 

Skeleton  of  the  Tuatara,  Sphenodon  inuictatus,  &c. "     Trans.   Zool. 

Soc.  Vol.  xvi.,  1891,  p.  52. 
14. — Parker,  W.  K. — "On  the  Structure  and  Development  of  the  Skull  in 

the  Lacertilia.     Pt.  1."     Phil.  Trans.,  1879. 
15. "  On  the  Structure  and  Development  of  the  Skull  in 

the  Common  Snake  (Tropidonotus  natrix)."     Phil.  Trans.,  1879. 


References  to  Plates, 

a.n.,  anterior  nares  ;  J.o.,  Jacobson's  organ;  mx.,  maxillary  ;  n.,  nasal; 
71./.  c,  nasal  floor  cartilage  ;  ??..§.,  nasal  septum;  pal.,  palatine;  p.c,  para- 
septal  cartilage  ;  ^.wa;.,  premaxillary  ;  pf.,  pterygoid;  P.  ro.,  prevomer ;  qu., 
quadrate;  r.c,  recurrent  cartilage;  s.mx.,  septomaxillary ;  t.p.,  trans- 
palatine;  ^y. ,  tympanic  ;  Fo.,  vomer. 


EXPLANATION  OF  PLATES  XXIV. -XXVI. 

Plate  xxiv. 

Fig.     1. — Nasal  capsule  of  embryo  of  Lacerta  agilis,  from  the  underside 

(after  Seydel).     Reconstructed  from  sections.     Magnified. 
Fig.    2. — Nasal  capsule  of  10  mm.  foetus  of  Trichositrus  vulpecula,  from  the 

underside.     Reconstructed  from  sections.     Magnified. 
Fig.     3. — Transverse  section  of  snout  of  young  Gecko  (Heteronota  hinoei). 

Showing  the  prevomers  as  splints  to  the  paraseptal  cartilages. 

Magnified. 


BY    R.    BROOM.  559 

Fig.  4. — Transverse  section  of  snout  of  ripe  foetus  of  Miniopterus  schreiberdi. 
Showing  the  prevomers  developing  in  connection  with  Jaeobson's 
cartilage  (the  modified  paraseptal).  [In  the  adult  the  two 
prevomers  are  found  anchylosed  together  to  form  a  median  bone 
which  reaches  from  the  one  cartilage  of  Jacobson  to  the  other.] 

Plate  XXV. 
Fig.    o.— Under  view  of  skull  of  IchthyopMs  glutinosus.    Showing  the  large 
median  vomer  (parasphenoid)  and  the  large  paired  prevomers 

(vomers).     Enlarged. 
Fig.     6. — Under  view  of  skull  of  Amblystovia  (after  Parker),  showing  the 

very  large  median  vomer  (parasphenoid)    and    the    modified 

paired  prevomers  (vomers). 
Fig.     7— Under  view  of  skull  of  ripe  embryo  of  Sphenodon  punctatm  (after 

Howes  and  Swinnerton).     Showing  the  small  median  vomer 

(parasphenoid)  and  the  large  prevomers  (vomers).     Enlarged. 
Fig.     8. — Semidiagrammatic  view  of  under  side  of  skull  of  Plesiosaunis  (after 

Andrews).     Showing  the  small  median  vomer  and  the  paired 

prevomers.     Keduced. 
Fig.     9. — Under  view  of  skull  of  nearly  ripe  embryo  of  Zootoca  (after  Parker). 

The  median  true  vomer  is  very  small,  and  the  paired  prevomers 

large.     Enlarged. 
Fig.  10.— Under  view  of  skull  of  young  Tropidonotus  (after  Parker).     The 

median  true  vomer  is  almost  entirely  situated  in  the  anterior 

half  of  the  skull. 

Plate  xxvi. 
Fig.  11. — Under  view  of  skull  of    Gomphognathus  (partly  restored  behind). 

Showing  the  median  true  vomer  and  the  paired   prevomers. 

The  dotted  lines  indicate  the   probable  anterior  limit  of  the 

vomer.     In  a  fragmentary  specimen  of  the  snout  Seeley  figures 

a  bone  which  he  calls  the  vomer  considerably  further  forward, 

but  the  bone  which  he    so    regards  is    almost    certainly  the 

ethmoid.      The    posterior    extensions   of   the    prevomers    are 

assumed.     Esduced. 
Fig.  12. — Under  view  of  skull  of  Ornithorhynchus.     Showing  the  prevomers 

united  to  form  the  dumbbell-shaped  bone.     Slightly  reduced. 
Fig.  1,3. — Under  view  of  skull  of  Ptyclwgnatlius.     Here  only  the  true  median 

vomer  is  developed,  the  great  palatal  development  of  the  pre- 

maxillary  having  led  to  the  loss  of  the  prevomers.     The  hinder 

part  of  the  skull  is  restored.     Reduced. 
Fig.  14. — Under  view   of  skull   of    Testudo.     Only  the   median   vomer   is 

developed.     The  loss  of  the  prevomer  may  be  due  to  the  fact 

that  the   organs   of   Jacobson   are  imperfectly   differentiated. 

Nat.  size. 


560  MAMMALIAN    AND    REPTILIAN    VOMERINE    BONES. 

Addendum    (added  14th  Felruary,  1903). 

Since  the  above  paper  was  sent  off  I  have  discovered  the 
presence  of  a  pair  of  large  prevomers  in  Titanosuchus;  and  also 
a  well  developed  pair  of  prevomers  in  a  moderately  large  jDrimi- 
tive  Theriodont,  allied  to  Lycosuclius,  but  as  yet  undescribed. 
As  a  large  pair  of  prevomers  are  known  to  occur  in  Tcipino- 
ceplialus,  the  various  stages  in  the  evolution  of  the  prevomer 
may  now  be  regarded  as  at  least  approximately  known.  From  a 
form  like  PareiasauruSy  with  a  palate  formed  on  the  Rhj^ncho- 
cephalian  t3^pe,  we  get  derived  the  Prototheriodonts  Tainno- 
cephalus  and  Titanosuchus — unfortunately  as  yet  very  imperfectly 
known.  In  the  next  stage  we  find  the  true  primitive  Theriodonts 
of  the  type  of  Lycosuclius  (Proc.  S.  Af.  Phil.  Soc.  1902)  with  a 
pair  of  moderately  developed  and  quite  distinct  prevomers  And 
lastly,  we  have  the  higher  Theriodont  type  represented  by  Gom- 
phognathus  and  closely  resembling  the  condition  in  Ornitlio- 
rliynclms.  In  Gorgonops — which  though  veiy  imperfectly  known, 
probably  belongs  to  the  Prototheriodonts, — we  have  the  pre- 
vomers anchylosed  as  in  Ornitliorliynclius  and  JSLiniopterus  (R.B., 
7th  January,  1903). 


561 


NOTES  ON  THE  BOTANY  OF  THE  INTERIOR  OF 
NEW  SOUTH  WALES. 

By  R.  H.  Cambage. 

Part  VII. — From  Forbes  to  Bathurst. 

(Plate  xxvii.) 

The  notes  in  this  paper  refer  more  particularly  to  the  vegetation 
on  the  western  slopes  than  to  that  of  the  interior,  beginning  with 
the  botany  of  the  low  country  near  the  Lachlan  and  terminating 
with  that  of  the  western  elevation  of  the  Blue  Mountains.  It 
will  be  noticed  that  there  is  a  complete  change  in  the  flora  between 
Forbes  and  the  hills  around  Bathurst,  caused  chiefly  by  the 
difference  in  altitude  of  the  two  districts.  This  difference  amounts 
to  over  2,000  feet,  for  while  Forbes  is  only  about  800  feet  above 
sea  level,  many  of  the  hills  around  Bathurst  and  Orange  exceed 
3,000  feet.  Much  of  the  country  referred  to,  especially  that  to 
the  westward  of  Orange,  is  open  box  forest,  considerably  cleared, 
and  the  shrubs  are  therefore  not  numerous. 

From  Forbes  to  Eugowra  is  about  24  miles  in  a  general  easterly 
direction,  the  ascent  being  very  slight,  and  various  trees  and 
shrubs  noticed  were  : — Callitris  rohusta  {y^hitQ  or  Cypress  Pine), 
Pittosporum  phillyrceoides,  Casua7'ina  Luehma7ini  (Bull  Oak),  C. 
Cunninghamiana  (River  Oak),  and  Acacia  decora.  On  the  ridges 
a  few  miles  back  from  the  river  the  shrubs  are  more  plentiful. 

The  Eucalypts  passed  were  : — E.  7nelliodora  (Yellow  Box),  E. 
Woollsiana  (Black  Box),  E.  hemiphloia  var.  alhens  (White  Box), 
E.  conica  (Apple  Box),  E.  rosti-ata  (River  Red  Gum),  E.  tereti- 
cornis  (Forest  Red  Gum),  and  var.  dealhata  (Mountain  Gum). 

Mr.  J.  H.  Maiden  in  a  recent  paper  on  E.  Behriana  (Trans.  R. 
Soc.  S.  Aust.  1901,  p.  11)  includes  E.  Woollsiana  under  E.  herai- 
phloia  as  a  variety  which  he  names  microcarpa. 


562  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

Reference  h  made  in  these  papers  (Part  ii.  p.  714)  to  the 
affinities  between  £.  hemij^hloia  and  B.  WooUsia7ia. 

Some  fine  trees  of  E.  tereticornis  were  seen  in  places  on  the 
flats  along  the  road,  var.  deaJbata  being  plentiful  on  the  granite 
hills  towards  Eugowra,  while  E.  rostrata  was  confined  to  the  river 
and  large  creeks.  On  questioning  a  saw-mill  proprietor  as  to  the 
relative  merits  of  E.  rostrata  and  E.  tereticornis^  he  stated  that 
the  River  Red  Gum  was  more  liable  to  warp  in  exposed  situa- 
tions than  the  Forest  Red  Gum,  and  he  had  seen  an  instance  in 
which  the  former  had  been  used  for  window  frames,  but  after  a 
few  years  the  wood  had  w^arped  to  such  an  extent  that  it  became 
necessary  to  replace  it.  A  resident  of  Riverina,  however,  who 
has  had  considerable  experience  with  the  timber  of  River  Red 
Gum,  states  that  it  gives  little  or  no  trouble  from  warping. 

From  Eugowra  to  Cudal  is  about  30  miles  north-easterly,  the 
ascent  in  the  latter  part  of  the  road  being  considerable.  Various 
trees  and  shrubs  noted  along  the  roadside  were  : — Callitris 
robusta,  C.  cal  car  ata  {Mountain,  Black  or  Green  Pine)  only  on  ridges, 
Casuarina  Cunninghamiana,  C.  Luehnianni,  Stercidia  diversifolia 
(Currajong),  Acacia  doratoxylon  (Currawong),  on  ridges  only,  A. 
homaJophyUa  (Yarran),  and  A.  obtusatoj,  Sieb,  (Black  Wattle). 
This  latter  species  was  noticed  at  3  miles  from  Eugowra  growing 
close  to  the  large  granite  rocks  known  as  the  Bushrangers'  Rocks, 
behind  which  the  bushranger  Gardiner  and  party  waited  when 
they  stuck  up  the  gold  escort  in  1862.  It  was  not  found  in  the 
districts  to  the  westward. 

The  Yarran  seen  at  about  6  miles  from  Eugowra  was  the.  last 
noticed  in  coming  from  the  west,  and  this  spot  therefore  marks  a 
spot  on  the  eastern  limit  of  its  habitat.  A.  homalojjhyUa  is  not 
plentiful  anywhere  to  the  eastward  of  a  line  joining  Junee,  Gren- 
fell  and  Wellington,  though  to  the  north  of  Gulgong  it  comes 
eastward  to  the  Hunter  River,  and  again  southward  to  the  east 
of  Rylstone  (R.  T.  Baker)  and  near  Glen  Alice  east  of  Capertee 
(J.  E.  Carne).  A  sister  tree,  A.  pendula,  (Myall  or  Boree)  has 
its  eastern  margin,  roughly  speaking,  about  20  or  30  miles  to  the 


BY    R.    H.    CAMBAGE.  563 

westward  of  that  of  the  Yarran,  except  on  the  plains  to  the  west 
of  Grenfell,  where  the  eastern  limits  of  the  two  species  nearly 
coincide.  A.  pendiila  does  not,  so  far  as  I  am  aware,  come  east 
of  the  Great  Dividing  Range  at  any  point,  though  it  may  possibly 
just  do  so  in  the  region  of  the  Liverpool  Range.  It  is  not 
uncommon  in  the  western  districts  to  hear  of  Ringed  or  Curly 
Yarran,  and  by  some  this  tree  is  thought  to  be  different  from 
ordinary  Yarran.  They  are  the  same  species,  viz.,  A.  homalo- 
2)hylla  occurring  in  two  forms,  one  with  the  wood  having  a  wavy 
grain  and  the  other  a  straight  grain.  From  considerable  obser- 
vation it  appears  to  me  that  Curly  Yarran  is  not  nearly  so  plenti- 
ful as  that  with  the  straight  grain,  nor  could  I  ever  find  any 
outward  sign  by  which  to  recognise  a  difference  in  the  trees,  but 
an  axe  cut  will  at  once  settle  the  question.  Curly  Yarran  pos- 
sesses a  very  ornamental  figure  owing  to  the  presence  of  this 
waving  in  the  grain  of  the  wood. 

Another  western  tree  having  a  similar  feature  is  A.  Cambagei, 
Baker,  the  Gidgea  of  the  Bourke  district,  but  in  this  case  the 
curly  form  seems  to  be  the  more  general  one,  hence  it  is  quite 
common  to  find  Ringed  Gidgea.  The  cause  of  the  rings  in  the 
Gidgea  is  the  same  as  in  the  Yarran.  One  of  the  ordinary  uses 
of  these  timbers,  especially  the  former,  is  for  stockwhip  handles, 
and  this  tapering  form  assists  considerably  in  developing  the 
rings  in  the  figure.  The  same  result  is  obtained  when  the  timber 
is  worked  up  into  walking  sticks  or  otherwise  used  in  turnery. 
Among  stockmen's  lore  I  have  heard  it  stated  that  in  order  to 
produce  rings  it  was  necessary  that  the  stem  of  the  trees  should 
grow  in  concentric  rings  of  different  colours,  and  that  these  were 
seen  alternately  in  the  length  of  the  tapering  whip  handle.  I 
was  never  able  to  find  anything  of  this  kind,  and  have  no  doubt 
that  the  undulation  in  the  grain  is  always  sufficient  to  account 
for  the  feature.  In  mature  trees  of  Yarran  the  timber  often  has 
flaws  inside  owing  to  the  operations  of  some  boring  insect,  and 
this  is  the  more  annoying  as  these  affected  spots  are  often  undis- 
covered till  the  working  up  of  the  wood  is  well  in  progress. 
Owing  to  this  blemish  the  usefulness  of  the  tree  is  considerably 


564  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

lessened.  This  species  suffers  in  many  western  localities  from 
the  attack  of  a  mistletoe,  usually  Loranthus  'pendidus. 

Near  Eugowra  this  mistletoe  was  found  growing  on  Eucalyptus 
Gonica,  and  L.  linojjhyllus,  Fenzl.,  on  Casuarina  Luehma7ini. 

Along  the  roadside,  betw^een  Eugow^ra  and  Cudal,  the  following 
Eucalj'^pts  were  noticed  : — E.  Woollsiana,  E.  conica,  E.  rostrata, 
E.  melliodora,  E.  tereticornis,  and  its  var.  dealbata,  the  latter  on 
hills  of  Devonian  sandstone,  E.  hejiiiphloia,  var.  alhens,  E. 
Bridgesiana  (Apple,  near  Murga),  and  E.  macrorhyncha  (Stringy- 
bark)  on  a  ridge  of  Devonian  sandstone. 

E.  Bridgesiana  was  mentioned  in  Part  vi.  (p.  202)  as  the  New 
South  Wales  apple  tree  which  has  been  recognised  for  so  many 
years  as  E.  Stitartiana. 

E,  conica  ceases  on  this  road  near  Toogong,  the  elevation 
beyond  this  being  too  great  for  it. 

E  rostrata  was  not  noticed  after  Cudal  was  passed,  it  also 
preferring  a  warmer  climate  than  is  to  be  found  near  Orange. 

From  Cudal  to  Orange,  a  distance  of  25  miles,  the  ascent  is  so 
great  that  all  the  typical  interior  trees  are  left  behind,  and 
replaced  by  others  which  are  usually  found  in  cold  regions,  the 
effect  of  climatic  influence  being  very  pronounced.  At  about 
half-way  Casuarina  Cunning hamiana  is  seen  for  the  last  time. 
Sterculia  diversifolia  is  noticed  at  intervals,  and  along  the  last 
10  miles  of  the  road  Acacia  dealhata^  the  Silver  Wattle,  becomes 
common.  From  the  notes  which  follow  it  may  be  seen  that  this 
is  a  very  common  "Wattle"  on  the  highlands,  and  when  flowering 
in  September  it  is  a  very  beautiful  tree. 

As  most  of  the  country  along  the  Cudal  to  Orange  road  has 
been  cleared,  the  flora  is  chiefly  represented  by  the  Eucalypts, 
those  passed  being  E.  nielliodora,  E.  Bridgesiana,  E.  Woollsiana, 
E.  hemijyhloia  var.  albens,  E.  tereticornis,  E.  viminalis,  Labill. 
(White  Gum),  at  about  17  miles  from  Orange,  E.  Cambagei 
(Bastard  Box,  Apple  or  Bundy),  near  the  10-mile  post,  E.  macro- 
rhyncha (7-mile  post),  E.  dives,  Schauer  (Peppermint),  (6-m.)  E. 
stellulata,  Sieb.    (Sally,   Black  Sally,   Sallow  or   Sally  Butt),  E. 


BY    R.    H.    CAMBAGE.  565 

macrorhyncha,  E.  vimiyialis,  E.  dives  (-i-m.),  E.  melliodora,  E. 
Bridgesiana  and  E.  viminalis. 

E.  Woollsiana  ceases  at  about  20  miles  from  Orange,  while  E. 
hemiphloia  var.  albens  continues  to  within  about  10  miles,  their 
eastern  margins  thus  being  only  10  miles  apart.  This  is  owing 
to  the  great  difference  in  elevation  which  takes  place  in  the 
comparatively  short  distance.  Had  the  ascent  been  gradual,  as 
from  Cootamundra  to  Harden,  the  difference  might  have  amounted 
to  -iO  or  50  miles  instead  of  only  10.  As  var.  albens  does  not 
extend  much  to  the  westward  of  Forbes,  it  may  be  noticed  that 
it  is  neither  an  interior  nor  a  mountain  species,  but  occupies  that 
area  which  may  be  designated  as  the  western  slopes.  It  continues 
much  in  this  position  north  and  south  throughout  New  South 
Wales. 

E.  viminalis  is  perhaps  one  of  the  best  known  white  gum  trees 
of  the  cold  regions.  It  generally  grows  as  a  somewhat  spreading 
tree  with  rather  drooping  foliage,  and  may  often  be  found  on 
flats  near  large  creeks,  as  well  as  on  basaltic  tablelands.  Its 
habitat  seems  to  be  regulated  chiefly  by  climatic  conditions,  for 
while  around  Orange  and  Bathurst  it  is  seldom  seen  below  an 
altitude  of  2,000  feet,  yet,  as  southern  latitudes  are  approached, 
it  is  found  at  much  lower  levels,  till  in  Tasmania  and 
Southern  Victoria  it  is  common  right  down  to  sea-level. 
It  may  be  pointed  out  that  so  far  as  is  at  present 
known,  none  of  the  Eucalypts  mentioned  in  previous  papers 
occur  in  Tasmania.  E.  viminalis  is  not  a  coast  species,  but 
comes  to  within  20  or  30  miles  of  the  ocean  in  the  district 
south  of  Moss  Vale.  It  also  occurs  on  a  flat  a  mile  or  two  above 
the  town  of  Picton.  Though  often  a  white  tree  of  the  ribbony 
gum  type,  it  not  infrequently  has  a  hard  black  bark  near  the 
butt,  arranged  in  very  strong  thick  scales,  and  this  rough  bark 
extends  varying  distances  up  the  barrel,  from  a  few  feet  in  some 
cases,  to  the  whole  length  of  the  trunk  in  others,  and  in  rare 
instances  even  on  the  branches.  Still  above  the  rough  bark  the 
general  appearance  of  the  tree  remains  the  same,  and  it  is  to  the 
upper  part  of  a  tree  that  the  bushman's  eye  wanders  when  in 


566  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

doubt  about  its  identity.  The  timber  of  this  species  is  considered 
to  be  ver}^  inferior,  both  in  or  out  of  the  ground,  and  has  a  porous 
appearance,  though  I  have  heard  of  an  instance  where  it  has 
lasted  very  well  in  a  wet  place,  A  reference  to  the  inflorescence 
shows  that  the  flowers  are  usually  arranged  in  threes,  though  this 
is  by  no  means  a  constant  feature,  for  even  where  at  first  sight 
there  appears  to  be  no  departure  from  this  arrangement,  a  careful 
inspection  will  often  reveal  umbels  of  fours  and  fives,  or  even 
more."^  On  the  other  hand,  there  are  some  trees  of  this  species 
which  show  scarcely  any  threes,  but  these  I  should  say  are  greatly 
in  the  minority.  I  have  collected  specimens  from  a  tree  near 
Orange  with  buds  arranged  in  threes,  but  a  little  lower  on  the 
same  branch  with  as  many  as  seven  mature  fruits  in  the  umbel, 
thus  showing  that  the  same  tree  does  not  necessarily  produce  an 
equal  number  every  season.  E.  viminalis  is  one  of  the  gum  trees 
from  which  manna  is  often  collected,  and  has  been  referred  to 
under  the  name  E.  mannifera,  A.  Cunn. 

E.  dives  seems  to  be  known  everywhere  as  Peppermint,  and 
this  name  is  suggested  from  the  odour  emitted  by  the  leaves  when 
bruised.  It  is  confined  to  the  cold  parts  of  New  South  Wales, 
and  may  be  generally  separated  from  other  Peppermints  by  its 
broad  stem-sucker f  leaves.  Usually  it  grows  on  ridges,  and  seems 
to  avoid  alluvial  flats.  It  has  fibrous  grey  bark  covering  the 
trunk  and  sometimes  the  larger  branches,  the  older  bark  being 
honey-combed  in  some  cases,  giving  it  a  lace-like  appearance. 
The  timber  of  this  tree  is  not  much  prized. 


*  Vide  remarks  by  Messrs.  Deane  and  Maiden.     These  Proceedings,  1901, 
Part  i.,  139. 

t  As  the  term  "sucker"  is  strictly  confined  in  botany  to  young  plants 
formed  on  underground  rootstocks,  while  in  Australia  the  same  term  is' 
popularly  applied  to  adventitious  growths  on  various  parts  of  the  stem  or 
branches  caused  chiefly  by  that  particular  part  of  the  tree  being  either  cut 
or  bruised,  it  has  been  thought  expedient  in  future  to  refer  in  these  papers 
to  the  latter  form  as  "  stem-suckers  "  by  way  of  distinction,  and  to  the  former 
as  either  "  suckers"  or  "  root-suckers." 


BY    R.    H.    CAMBAGE.  567 

E.  stellulata  is  a  well  known  tree  on  the  highlands  both  north 
and  south,  being  usually  found  along  damp  flats  and  watercourses 
in  open  grazing  countr}^  The  upper  parts  of  the  trunk  and 
branches  are  covered  with  a  smooth  greenish  to  slate-coloured 
bark,  while  near  the  butt  the  bark  is  in  rough  thick  flakes,  often 
deeply  furrowed,  and  not  unlike  that  sometimes  found  on  the 
lower  part  of  E.  viminalis.  Its  chief  flowering  time  in  the  Orange 
district  is  about  March,  and  its  flowers  are  arranged  in  numerous 
star-like  clusters.  The  venation  of  the  leaves  is  of  interest,  the 
veins  being  arranged  longitudinally  and  almost  parallel  to  the 
midrib.  The  timber  of  this  species  is  soft  like  that  of  most  of  the 
cold  country  trees,  and  somewhat  brittle. 

From  Orange  to  Ophir  is  a  distance  of  about  16  miles  north- 
easterly, and  the  country  falls  gradually  all  the  way.  Along  the 
latter  part  of  the  road  some  of  the  low  country  forms  reappear. 
Various  trees  and  shrubs  noticed  along  the  roadside  were : — 
Acacia  dealhata  most  of  the  way,  A.  vestita,  Ker,  A.  falcata  (?), 
Willd.,  A.  linifolia  (?),  Willd.,  A.  armata,  Cassia  Sojjhera,  L., 
Casuarina  Cunninghamiana,  Poinade7'ris  'phillyrmoides^  Sieb., 
Grevillea  7'amosissima,  Meissn.,  Sterculia  diversifolia,  Disca^'ia 
australis,  Hk.,  Callistemon  salig7ius,  DC,  var.  angusti/olia,  along 
the  creek,  and  Exocaiyus  cupressiformis  (Native  Cherry). 

Acacia  vestita  was  found  only  near  Ophir.  It  grows  in  a  very 
graceful,  spreading  and  pendulous  form. 

The  Eucalypts  noticed  along  this  road  were : — E.  viminalis^  E, 
Bridgesiana,  E.  [coriacea,  A.  Cunn.,  E.  stellulata,  E.  melliodoray 
E.  tereticornis,  E.  rubidcc,  Deane  and  Maiden,  E.  dives,  E.  macro- 
rhyncha,  E.  Cambagei,  E.  hcemastoma,  Sm.,  var.  micrantha,  E. 
macidosa,  Baker,  E.  polyanthema,  Schau.,  and  E.  hemiphloia  var. 
alhens. 

E.  coriacea  is  another  of  the  white  gums  of  the  cold  country, 
and  although  it  may  often  be  found  in  company  with  E.  viminalis 
it  leaves  that  species  behind  in  some  of  its  ascents  on  our  highest 
mountains.  This  is  the  Eucalypt  which  may  be  found  growing  at 
a  higher  elevation  than  any  other  Eucalypt  in  New  South  Wales, 
or  probably  for  that  matter  in  the  world  {vide  Mr.  J.  H.  Maiden's 


568     BOTANY  OF  THE  INTKRIOR  OF  NEW  SOUTH  WALES, 

^' Flora  of  Mount  Kosciusko").  It  is  common  along  the  highlands 
of  New  South  Wales,  both  north  and  south,  extending  also  into 
Victoria  and  Tasmania.  It  is  the  Snow  Gum  of  the  Kiandraand 
Kosciusko  districts,  where  it  often  grows  in  a  stunted  form.  A 
conspicuous  feature  in  the  general  appearance  of  this  tree  is  its 
large  coarse,  longitudinally  veined,  and  somewhat  shining  leaves, 
which  in  times  of  drought  are  considered  to  be  moderately  good 
fodder,  and  this  in  some  places  has  earned  for  the  tree  the  name 
of  Cattle-gum.  Its  fruits  are  fairly  large,  and  in  the  Oberon 
district  the  seeds  are  eaten  by  the  Gang  Gang  Cockatoo  {Callo- 
cephalon  galeatiim^  Latham).  The  bark  of  this  tree  is  usually 
smooth  and  white  to  the  ground,  also  thin,  and  when  growing 
with  E.  viminalis  bushmen  can  separate  the  two  species  on  sight 
by  the  bark  and  foliage.  The  timber  of  E.  coriacea  is  one  of  the 
best  of  the  cold  country  gums,  and  is  often  used  for  fence  posts. 
This  species  is  not  restricted  to  any  particular  geological  formation, 
but  seems  to  have  a  preference  for  igneous  over  sedimentary,  and 
is  usually  found  growing  in  open  forests.  It  shares  wdth  some 
other  trees  the  names  of  White  Gum,  Cabbage  Gum,  and  Scribbly 
Gum.  The  latter  name  is  suggested  by  certain  scribble  marks 
on  the  bark,  and  these  are  made  by  a  small  grub  eating  its  way 
along  the  outside  of  the  inner  bark,  the  eiFect  being  that  the 
"  scribble  "  is  left  thereon;  also  on  the  inside  of  the  outer  bark, 
the  outside  of  which  was  perhaps  previously  operated  upon  while 
it  was  yet  inner  bark,  so  that  a  piece  of  bark  just  falling  off  will 
often  have  scribble  marks  on  both  sides,  but  in  no  w^ay  connected 
with  each  other.  A  casual  examination  once  made  of  this  little 
grub  in  the  Oberon  district  showed  it  to  measure  only  two. lines 
in  length.  E.  lutmastoma  is  another  white  gum  tree  which  also 
has  has  scribble  marks  on  the  bark. 

E.  rubida  is  also  one  of  the  high  country  gums,  and  is  some- 
times found  associated  with  E.  viminalis,  with  which  species  it 
is  often  confused  by  bushmen.  Of  the  two  species,  E.  ruhida 
generally  takes  the  drier  land,  being  found  on  hillsides,  while  E. 
viminalis  is  more  plentiful  in  the  valleys;  but  it  is  also  quite 
common  to  see   them  mingle   on  elevated  land.     As  a  rule  E. 


BY    R.    H.    CAMBAGE.  569 

ruhida  is  neither  so  spreading  nor  so  pendulous  as  E.  vhninalis; 
and  its  identification  is  further  assisted  by  the  glaucous  appear- 
ance of  the  foliage  and  fresh  bark,  as  well  as  some  red  patches 
on  the  outer  bark  just  before  it  begins  to  peel  off.  The  greatest 
diiference  is  perhaps  in  the  stem-sucker  leaves,  those  of  F. 
vimiiialis  being  generall}^  narrow,  while  those  of  E.  ruhida  are 
nearly  round,  even  on  the  young  shoots  high  up  among  the 
branches.  The  flowers  are  usually  arranged  in  threes  like  those 
of  E.  viminalis,  and  in  my  collecting  I  have  been  unable  to  find 
more  than  four  in  the  uml^el,  and  that  rarely,  though  this  number 
is  recorded  by  other  collectors  as  being  exceeded  at  times.  The 
timber  of  U.  ruhida  is  regarded  as  very  inferior,  and  resembles 
that  of  E.  viminalis.  The  species  has  a  considerable  range  in  the 
highlands,  and  extends  into  Victoria.  It  may  often  be  found 
growing  on  Silurian  slate  formation,  though  not  usually  in  very 
rocky  situations,  but  it  is  also  common  on  soil  formed  from 
decomposed  igneous  rocks.  This  is  another  tree  from  which 
manna  has  been  collected. 

E.  hmmastoma  var.  micrantha  is  a  well  known  white  gum  tree 
both  along  the  coast  and  on  the  highlands.  Around  Bathurst 
and  Orange  it  is  known  variously  as  Brittle  Gum,  Brittle  Jack, 
Cabbage  Gum,  Scribbly  Gum,  White  Gum  and  Red  Cabbage 
Gum.  Along  the  coast  the  timber  of  this  tree  is  not  much  valued, 
partly  because  it  does  not  split  well,  and  partly  owing  to  the  fact 
that  there  is  an  abundance  of  other  much  finer  timber  available. 
Still,  on  the  highlands,  it  is  often  in  request  for  fence  posts,  as  it 
has  been  proved  to  be  durable  in  the  ground.  It  is  also  con- 
sidered to  be  moderately  good  fuel,  but  the  wood  of  the  cold 
country  is  much  inferior  in  this  respect  to  that  of  the  western 
districts.  Its  timber  when  freshly  cut  is  of  a  pale  red  colour, 
with  a  very  short  grain.  The  bark,  which  is  white  and  rather 
thin,  sheds  off  in  short  patches.  The  short  or  long  grain  in  the 
timber  of  the  Eucalypts  is  usually  manifested  by  a  corresponding 
feature  in  the  bark,  and  there  is  consequently  much  truth  in  the 
lines  of  Henry  Kendall,  where  he  writes  of  Jem  the  Splitter  : — 
His  knowledge  is  this — he  can  tell  in  the  dark, 
What  timber  will  split  by  the  feel  of  the  bark. 


570     BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

The  fruits  of  the  Bathurst  and  Orange  representatives  of  E. 
hcemastoma  are  usually  small  and  hemispherical,  and,  therefore, 
more  properly  belong  to  var.  micrantha,  but  they  have  the  reddish 
rim  which  gave  rise  to  the  botanical  name  of  the  species.  In 
shape,  the  fruits,  with  their  rounded  base,  have  often  reminded 
me  of  miniature  wine  glasses,  and  the  similarit}"^  is  increased  by 
the  slenderness  of  the  pedicels  which  support  the  fruit.  In  some 
of  the  coast  forms,  where  the  fruits  are  larger,  they  are  also  more 
conical,  perhaps  approaching  those  of  E.  Sieheriana,  F.v.M.,  the 
Mountain  Ash. 

E.  hcemastoma  var.  micrantha  is  generally  found  growing  on 
poor  soil,  and  undoubtedly  prefers  a  geological  formation  of  sedi- 
mentary origin  to  that  of  an  igneous,  and  will  thrive  better  on 
the  top  of  a  Silurian  slate  ridge  among  the  rocks  than  on  the 
surrounding  flats  where  the  soil  is  infinitely  better. 

E.  'maculosa  is  another  white  orum  tree  ofenerallv  found  associated 
with  E.  hcemastoma  var.  micrantha,  and  in  habit  as  well  as  out- 
ward appearance  so  much  resembles  it  that  by  the  casual  observer 
the  two  species  are  often  confused.  Both  appear  to  flourish  on 
Silurian  slate  ridges,  and  may  also  be  found  on  the  Triassic 
sandstone  around  Mount  Victoria,  but  I  have  never  seen  the 
former  on  the  low  coastal  area  between  Wollongong  and  New- 
castle where  the  latter  is  quite  common.  For  a  further  comparison 
of  these  two  trees,  see  Mr,  Maiden's  remarks  in  these  Proceedings 
for  1901  (p.  582). 

E.  maculosa  is  generally  a  small  tree,  but  in  rare  instances  I 
have  seen  it  with  a  diameter  of  three  feet  six  inches,  though  not 
proportionately  tall.  In  some  parts  on  the  highlands  a  feature 
of  the  bark  is  that  when  young  it  often  becomes  covered  with  a 
white  powder,  but  this  disappears  from  the  old  bark  which  is 
shed  in  patches,  thereby  suggesting  the  name  of  Spotted  Gum  for 
the  tree.  This  is  the  name  by  which  the  trees  are  known  between 
Orange  and  Ophir,  though  at  AViseman's  Creek  they  are  called 
White  Cabbage  Gum  to  distinguish  them  from  E.  hcemastoma 
var.  micra?itha,  which  is  there  called  Red  Cabbage  Gum,  the 
wood  of  the  former  being  somewhat  paler  than  that  of  the  latter, 


BY    R.    H.    CAMBAGE.  571 

not  SO  brittle,  nor,  so  far  as  I  could  learn,  is  it  considered  so 
durable.  E.  maculosa  may  be  readily  separated  from  E.  hcema- 
stoma  var.  micrantha  by  its  coarser  buds,  thicker  pedicels,  and 
slightly  domed  fruits."^ 

E.  poJyanthema  is  the  Red  Box  of  the  Bathurst  district,  and  is 
usually  found  on  elevated  land,  showing  a  decided  preference  for 
sedimentary  formation.  Although  this  species  grows  in  fairly 
cold  country,  its  regular  home  in  the  Bathurst  to  Goulburn 
district,  appears  to  be  approximately  in  a  zone  between 
1,500  and  2,500  feet  above  sea  level,  though  I  have  seen 
it  both  above  and  below  these  elevations.  It,  therefore, 
occasionally  comes  low  enough  to  associate  with  E.  hemijMoia 
var.  a/bens,  and  is  only  at  times  found  high  enough  to  occupy  the 
same  localities  as  E.  coriacea  or  E.  dives.  It  has  a  fairly  smooth 
gum  tree  bark  except  near  the  base,  where  for  several  feet  up  the 
barrel  it  is  of  a  rough  flaky  nature.  The  name  of  Box  is 
suggested  by  the  hardness  and  texture  of  the  wood,  and  perhaps 
by  a  little  colouring  of  the  bark  near  the  base.  Judging  the 
trees  by  the  bark  alone,  they  may  at  times  be  alternately  confused 
with  Yellow  Box  (E.  melliodora)  or  Forest  Red  Gum  (E.  terefi- 
cornis).  The  oval  and  sometimes  glaucous  leaves,  however,  will 
generally  determine  the  species  when  seen  growing  in  the  districts 
where  it  is  known.  R.ed  Box  is  prized  most  for  its  strong  red 
timber  which  is  very  durable  in  or  out  of  the  ground.  Un. 
fortunately  around  Bathurst  it  does  not  grow  to  very  stately 
dimensions,  and  in  addition  to  having  only  a  comparatively  short 
trunk,  the  wood  has  generally  a  very  irregular  grain,  and  is, 
therefore,  diflacult  to  split.  The  curly  nature  of  the  timber  is 
well  indicated  by  similar  windings  in  the  bark. 

This  tree  has  been  recently  named  E.  ovalifolia  by  Mr.  R.  T. 
Baker  (these  Proceedings,  1900,  p.  680)  in  order  to  separate  it 
from  the  Victorian  Red  Box.  So  far  as  I  have  been  able  to 
study  the  latter  tree,  I  find  that  it  has  timber  similar  to  the  Red 


*  Messrs.  Baker  and  Smith,  in  their  ''Eesearch  on  the  Eucalypts,"  have 
given  this  variety  micrantha  specific  rank  under  the  name  of  E.  Rossii. 


572     BOTANY  OF  THE  INTERIOR  OF  XEW  SOUTH  WALES, 

Box  of  Bathurst,  but  appears  constantly  to  have  larger  and  more 
orbicular  leaves,  as  well  as  more  of  a  box  bark  covering  the  whole 
of  the  trunk.  The  fruits  and  buds  of  these  two  trees  approach 
each  other  considerably  in  shape.  The  Bathurst  tree  continues 
southward  along  the  western  slopes,  past  Goulburn,  to  the 
Gundagai  and  Tumut  district,  showing  practically  no  variation, 
and  is  recorded  by  Mr.  Baker  from  Gerogery.  Before  Albury 
is  reached,  the  Victorian  tree  may  be  seen,  and  it  extends 
southerly  right  across  that  State,  varying  very  little.  As  seen 
from  the  railway  line  near  Albury  the  southern  Red  Box  appears 
quite  dissimilar  to  the  Bathurst  tree,  chiefly  owing  to  the  distinc- 
tive characters  of  the  bark  and  foliage. 

On  the  road  from  Orange  to  Ophir  a  splendid  spot  from  which 
to  study  the  local  Eucalypts  is  in  the  paddock  opposite  Bosedale 
House,  about  five  miles  from  Orange.  Of  the  fourteen  Eucalypts 
seen  along  this  road,  twelve  may  be  found  in  this  paddock,  though 
they  occur  in  various  other  places  on  the  way,  the  two  which  are 
absent  being  E.  hemvphloia  var.  aJhens  and  E.  tereticoryiis,  though 
the  latter  may  be  found  quite  near.  E.  jyolyanthema  is  scarce  at 
this  spot. 

In  the  creek  at  Ophir  is  the  site  of  the  first  discovery  of  pay- 
able gold  in  Australia,  and  the  locality  is  well  illustrated  in  "The 
Mineral  Resources  of  New  South  Wales,"  by  Mr.  E.  F.  Pittman, 
A.R.S.M.  The  trees  growing  along  the  bank  of  the  creek  are 
River  Oak  (Casuarina  Cunninghamiana),  while  among  others 
growing  around  are  E.  polyanthema,  E.  mxacrorhyndia^  E.  Cam- 
bagei,  and  E.  maculosa,  all  of  them  typical  gold-field  trees  on  the 
elevated  Silurian  slate  areas. 

In  speaking  of  the  various  zones  of  elevation  at  which  different 
trees  grow,  it  may  be  pointed  out  that  over  the  area  referred  to 
in  these  papers  the  Eucalypt  that  shows  most  disregard  for 
climatic  conditions  is  E.  vielliodoj'a  [Yellow  Box  or  Yellow  Jacket). 
It  is  mentioned  in  Part  iii.  as  having  been  noticed  down  the 
Lachlan  at  Euabalong.  I  have  also  collected  it  just  above  Dan- 
daloo  on  the  Bogan,  both  localities  being  well  beyond  the  western 
limit  of  E.  hemiphloia  var.  alhens.     Coming  eastward  it  is  found 


BY    R.    H.    CAMBAGE.  573 

ascending  the  western  slopes,  passing  right  through  the  belt  and 
beyond  the  eastern  limit  of  var.  albeiis,  and  continuing  upwards 
to  within  a  few  miles  of  Orange,  though  above  an  altitude  of 
3,000  feet  it  seems  rare.  This  distribution  is  important  in  view 
of  the  probable  future  of  this  tree  in  regard  to  its  bearing  on  the 
Eucalyptus  oil  industry. 

In  going  from  Ophir  to  Mullion  Creek  Railway  Station  the 
country  again  rises,  and  E.  hemiphloia  var.  albens  is  soon  left 
behind.  Other  Eucalypts  growing  on  the  elevated  parts  are  E. 
viminaUs,  E.  Bridyesiana,  E.  mdliodora^  E.  tereticoruis,  E.  maci'o- 
7'hyncha,  E.  maculosa,  E.  Cambagei,  E.  hcemastoma  var.  Tnicrantha, 
E.  polyanthema,  E.  dives,  E.  amygdaJina,  Labill.  (Peppermint),  E. 
rubida  and  E.  coriacea. 

Other  plants  within  sight  of  the  road  are  Acacia  dealbata,  A. 
tindidifolia,  A.  lanigera,  A.  Cunn.,  Cassinia  Theodori,  Daviesia^^. 

Two  other  plants  found  in  the  district  were  Pidtencea  ternata, 
F.V.M.,  and  F.  styphelioides,  A.  Cunn. 

Eucalyptus  amyydalina  is  usualh^  known  on  the  western  high- 
lands as  Peppermint,  but  towards  the  coast  where  the  trees  have 
greater  proportions  they  are  very  often  called  Messmate.  The 
first  name  is  suggested  from  the  strong  smell  of  the  crushed  leaves, 
while  the  appellation  of  Messmate  infers  that  these  trees  "  mess- 
mate "  with  or  partake  of  the  characters  of  other  trees.  The 
trees  showing  outward  affinities  to  E.  aniygdalina,  and  which 
suggest  the  name  of  Messmate  are  probably  E.  dives  (Peppermint) 
and  one  of  the  Stringybarks  growing  in  the  locality. 

E.  amygdalina  usually  grows  with  a  grey  bark  not  unlike  that 
of  E.  dives,  but  perhaps  a  little  more  fibrous  in  large  trees.  The 
two  species  are  easily  separated  on  sight  by  the  leaves,  those  of 
the  former  being  narrow,  while  those  of  the  latter  are  fairly 
broad,  especially  on  the  young  shoots.  The  fruits  of  E.  dives  are 
slightly  the  larger.  The  timber  of  E.  amygdalina  is  not  con- 
sidered to  be  of  much  value,  being  soft  and  likely  to  decay  when 
used  i]i  exposed  situations.  This  species  is  not  very  plentiful  in 
the  Orange  and  southern  Bathurst  districts,  and  Mullion  Creek 
marks  the  most  north-v/esterly  locality  in  which  I  have  ever  found 


574  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

it.  This  is  one  of  those  species  which  seem  to  slightly  prefer  a 
geological  formation  of  sedimentary  origin  to  that  of  an  igneous, 
but  apparently  it  is  more  particular  as  a  first  consideration  in 
having  a  fairly  cool  climate,  and  ma}^  be  found  both  north  and 
south  on  elevated  Silurian  slate  areas,  as  well  as  on  the  Triassic 
sandstone  around  Mount  Victoria  and  elsewhere.  From  Goul- 
burn  and  Moss  Vale  it  continues  eastward  to  the  top  of  the 
Illawarra  Range  at  Macquarie  Pass  overlooking  Albion  Park,  but 
I  have  never  found  it  down  in  the  Illawarra  district. 

E.  radiata,  Sieb.,  is,  however,  found  along  man}'  of  the  rivers 
and  creeks  of  the  coastal  area,  though  not  absolutely  confined  to 
these  situations,  and  is  known  sometimes  as  Peppermint  but  often 
as  Ribbony  Gum  from  the  appearance  of  the  streamers  of  decor- 
ticating bark  as  they  hang  from  the  upper  parts  of  the  trees. 
The  trunks  of  E.  radiata  are  covered  at  the  base  and  for  ^arj'ing 
distances  upwards  with  a  grey  bark  very  similar  to  that  of  E. 
amygdalina^hMt  the  upper  part  is  usuall}^  clean  and  fairly  white. 
In  general  appearance  this  is  the  chief  difference  noticed  between 
the  two  species  except,  of  course,  in  habit,  E.  radiata  showing  a 
preference  for  growing  along  creek  banks  and  on  flats.  Still  the 
latter  trees  may  be  found  on  the  hillsides  in  many  localities,  and 
although  they  appear  to  grow  at  lower  levels  in  Xew  South  AVales 
than  E.  ainygdalina,  they  may  sometimes  be  found  at  elevations 
at  least  exceeding  2,000  feet  above  sea  level.  The  fruits  of  E. 
radiata  have  appeared  to  me  to  be  uniformly  rather  more  con- 
stricted at  the  rim  than  are  those  of  E.  amygdalina,  but  undoubt- 
edly the  two  species  approach  each  other  ver}'  closely,  and  there 
are  instances  where  it  is  extremely  difficult  to  separate  them. 

E.  amyydaJiiia  has  an  extensive  range  on  the  highlands  in  New 
South  Wales  and  Victoria,  and  is  common  in  Tasmania,  ]3ut  under 
the  influence  of  southern  latitudes  may  there  be  found  practically 
down  to  the  sea  level,  although  also  at  considerable  elevations, 
and  apparently  heedless  of  whether  the  formation  is  igneous  or 
sedimentary,  being  common  on  the  former  near  Launceston  and 
on  the  latter  around  Hobart.  In  southern  Victoria,  also,  it 
appears  to  show  that  a  suitable  climate  is  its  chief  desideratum, 


BY    R.    H.    CAMUAGE.  575 

as  it  grows  there  at  a  much  lower  level  than  in  the  latitude  of 
Sydney. 

At  about  10  miles  south-westerly  from  Orange  there  is  a  high 
mountain  of  volcanic  origin,  exceeding  4,000  feet  above  sea  level 
and  known  as  the  Canoblas.  It  is  of  intense  geological  interest,  for 
in  addition  to  the  petrological  study  it  affords  of  the  various  forms 
of  volcanic  rock  of  which  it  is  composed,  and  the  probable  remains 
of  an  ancient  crater  on  the  top,  just  south  of  the  Trig,  station, 
the  original  lava  flow  has  covered  and  thereby  protected  older 
sedimentary  rocks  of  the  Devonian  and  Silurian  periods,  and 
these  ma}^  be  seen  in  places  protruding  from  under  the  base  of 
the  mountain.  To  the  eastward,  near  Forest  Reefs,  Tertiary 
fossil  fruits  may  be  found  buried  under  a  great  thickness  of  basalt 
which  was  possibly  emitted  from  or  contemporaneous  with  the 
outflow^s  from  the  Canoblas.  Some  rocks  containing  graptolites 
of  probably  Lower  Silurian  origin  may  be  found  towards  Diggers' 
Creek  near  Cadia. 

The  road  from  Orange  to  Cargo  passes  the  base  of  the  mountain 
on  the  western  side  at  about  7  miles  from  Orange,  and  the 
Eucalypts  noticed  between  Orange  and  this  point  are  E.  dives, 
E.  viminalis,  E.  Br-idgesiana,  E.  r)iacrorhyncha  and  E.  coriaceaj 

On  ascending  the  mountain  from  about  the  7-miIe  post  the 
following  plants  are  noticed  : — Eucalyptus  macrorhyncha,  E. 
melliodoi'a,  E.  Bridyesiana,  E.  viminalis,  E.  Cambagei,  E.  dives, 
Acacia  dealbata,  A.  melanoxylon,  ^.^v.,  Exocarpus  ciqjressiformis, 
Thysanotus  fube?-osus,  R.Br.  (Fringed  Violet),  Veronica  perfolio- 
lata,  R.Br.,  Calythrix  tetragona,  Stella7'ia  2Junge7is,  Brong.,  Ery- 
fh?rea  australis,  R.Br.,  Helichrysur)i  apicidatum,  DC,  and  Lepto- 
spermum  lanigevam  (?),  Sm. 

Erythr(pa  australis  is  a  small  plant  which  has  been  seen  extend- 
ing over  most  of  the  area  described  in  these  papers,  and  is  to  be 
found  in  the  coastal  area  as  well.  It  was  used  by  the  aborigines 
for  medicinal  purposes,  and  is  not  confined  to  this  State. 

Acacia  melanoxylon  is  the  same  as  the  famous  Tasmanian 
Blackwood,  but  in  New  South  Wales  it  does  not  grow  into  such 
fine  trees  as  may  be  found  in  Victoria  and  Tasmania.     It  is  often 


576     BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

known  as  Hickory  from  the  toughness  of  the  wood.  Along  parts 
of  the  North  Coast  and  Table  Lands  it  is  called  Mudgerabah  and 
has  various  other  local  names.  The  species  is  very  likely  to  be 
confused  b}^  the  casual  observer  with  A.  implexa  owing  to  a 
general  similarity  in  the  bark  and  venation  of  the  leaves.  The 
simplest  test,  if  ripe  pods  are  available,  is  to  examine  the  funicle 
which  connects  the  seed  to  the  pod,  that  of  A.  meJanoxylon  being 
red  and  carried  round  the  seed  twice,  while  that  of  A.  imijlexa  is 
paler  and  folded  at  the  hilum  end  of  the  seed. 

The  only  Eucalj^pts  growing  on  the  top  of  the  Canoblas  are 
E.  coriacea  and  E.  rubida,  the  others  ceasing  at  about  500  feet 
below  the  summit.  Mr.  J.  H.  Maiden  has  shown  that  this  par- 
ticular form  of  E.  ruhida  growing  on  the  top  of  the  Canoblas  is 
ver}^  little  removed  from  typical  E.  Guniiii,  Hook.,  of  Tasmania 
(these  Proceedings,  1901,  p.  581). 

On  the  roadside  between  Canoblas  and  Cargo  the  following 
Eucalypts  were  noticed  : — E.  riminalis,  E.  Eridgesiana,  E.  dives, 
E.  fereficoniis,  E.  meUiodora,  E.  macrorhyncha,  E.  Camhagei  (18 
mile  post),  and  E.  hemiphloia  var.  alhens. 

Between  Orange  and  Cargo,  a  distance  of  23  miles,  the  country 
falls  almost  continuously,  and  the  climatic  effect  on  the  flora  is 
again  very  noticeable.  E.  coriacea  is  left  behind  at  the  7-mile 
post.  Near  the  15-mile  post  the  last  is  seen  of  the  Peppermint, 
E.  dives  and  of  E.  viminalis.  At  the  20-mile  post  E.  macrorhyn- 
cha  ceases,  the  country  below  this  being  too  hot  for  these  species. 
On  the  other  hand,  E.  hemiphloia  var.  alhens  does  not  appear  till 
the  18-mile  post  is  reached,  the  country  above  that  being  too  cold 
for  it. 

Between  the  15-  and  16-mile  posts  and  on  the  left  hand  side  of 
the  road,  specimens  w^ere  collected  from  a  tree  ^vhich  about  the 
greater  part  of  the  trunk  considerably  resembled  E.  Bridgesiana, 
but  the  upper  part  and  branches  were  clean,  giving  the  tree  the 
appearance  of  E.  viminalis.  The  fruits  and  buds  most  resemble 
those  of  the  multiflowered  form  of  E.  viminalis,  and  in  the 
absence  of   stem-sucker  leaves  have  been  provisionally  identified 


BY    R.    H.    CAMBAGE.  577 

as  belonging  to  that  species.  One  or  two  other  similar  looking 
trees  were  seen  in  the  locality. 

From  Cargo  to  Cowra  via  Canowindra  the  country  still  falls, 
and  the  vegetation  gradually  assumes  the  western  forms.  On 
the  hills  at  Cargo  is  rather  a  glaucous  form  of  E.  Cambcu/ei,  and 
this  is  the  last  seen  of  the  species  in  travelling  towards  the 
Lachlan.  It  is  here  associated  with  JE.  hemiphloia  var.  alhens,  a 
coalition  which  only  takes  place  near  the  western  or  lowest  level 
of  the  former,  and  the  eastern  or  highest  part  of  the  latter. 
Growing  with  them  is  E.  te7-eticornis  var.  dealbata. 

The  country  is  generally  clear  of  shrubs,  but  the  larger  trees 
noticed  between  Cargo  and  Canowindra  were  : — Eucalyptus  melli- 
odora,  E.  fereticornis,  E.  hemiphloia  var.  albens,  E.  Woollsiana, 
Callitris  7'obusta,  and  Sterculia  diversifolia.  The  re-appearance  of 
Eucalyptus  Woollsia^ia  and  Callitris  robusta  denotes  a  return  to 
the  low  country. 

Sterculia  diversifolia  (Currajong)  is  the  well  known  fodder  tree, 
and  there  are  few  plants  having  a  wider  range  over  this  State, 
although  it  rarely  grows  in  clusters  but  as  isolated  trees.  It  is 
found  over  the  whole  of  the  area  described  in  these  papers,  and 
is  dotted  nearly  all  along  the  coastal  districts.  It  grows  on 
various  geological  formations,  but  if  it  shows  any  partiality  it  is 
first  for  limestone  and  then  igneous  over  slate  and  sandstone. 
Near  Parramatta,  at  the  Pennant  Hills  quarry,  it  it  found  on 
basalt,  while  at  Newcastle  and  West  Dapto  it  grows  on  Permo- 
Carboniferous  formation.  At  Tillowrie  near  Milton  it  may  be 
found  in  limited  quantities  on  an  igneous  rock  extending  only  a 
short  distance  among  the  sandstone,  and  was  known  as 
Curranimai  by  the  aborigines  who  used  the  young  roots  as 
an  article  of  food.  The  aborigines  of  the  district  had  a 
curious  belief  that  to  cut  one  of  these  trees  would  produce 
rain,  consequently  they  were  often  spoken  of  as  Rain 
trees.  But  they  must  not  be  confused  with  two  trees  of  Euca- 
lyptus macidata  (Spotted  Gum)  which  grow  on  the  roadside  a  few 
miles  to  the  southward  and  are  known  to  residents  and  regular 
travellers  as  the  "  Water  Trees,"  owing  to  the  fact  that  they  have 


578  BOTANY    OF    THE    INTERIOR    OF   NEW    SOUTH    WALES, 

small  hollows  in  the  base  of  the  trunks  which  hold  water  nearly 
all  the  year  round.  One  in  particular  has  been  the  means  of 
quenching  many  a  thirst  on  this  particularly  dr}^  piece  of  road. 

The  name  of  Currajong  or  Kurrajong  appears  to  be  a  settler's 
name  for  the  various  trees  which  have  a  bark  easily  stripped  and 
containing  a  strong  fibre,  as  is  the  case  with  the  young  shoots  of 
this  species.  In  the  Ulladulla  district,  sjDecies  of  Commersonia 
and  Pimelea  are  better  known  as  Currajong  than  the  Sterculia, 
while  in  other  places  some  of  the  Hibiscus  family  bear  this  name. 
For  other  "Kurrajongs,"  see  Mr.  Maiden's  'Useful  Native  Plants 
of  Australia.' 

In  the  western  districts  the  young  leaves  are  often  divided  into 

three  long  lobes,  and  it  is  quite  common  to  see  divided  leaves  on 

large  trees,  but  this  feature  is  much   more  unusual   along   the 

coast.     Seeds  of  the  Currajong  collected  in  the  western  districts, 

and  planted  on  the  coast,  produce  plants  with  leaves  more  divided 

than  are  those  of  the  indigenous  coastal  trees.     A  plant  grown 

at  Burwood  from  seed  collected  on  the  Upper  Macleay  in  1 889, 

seldom  has  any  divided  leaves,  and  the  divisions  were  very  slight 

on  some  young  shoots  recently  produced  by  cutting  back  one  of 

the  older  branches.     This  particular  tree,  though  now  13  years 

old,  has  never   flowered.     A  tree  growing  beside  it  from  seed 

collected  on  the  Lachlan  has  the  much  divided  leaves.     Another 

slight  difference  noticed  between  the  coastal  and  interior  Curra- 

jongs  is  that  the  coast  tree  will  sometimes  shed  all  its  leaves, 

though  this  is  not  a  constant  feature,  and  is  a  matter  which 

requires  further  investigation,  possibly  only  applying  to  old  trees. 

A  tree  at  Milton  was  leafless  during  most  of  January,  1901.     In 

the  interior  I  have  noticed  them  partially  shed  their  leaves,  but , 

never  wholly.     During  the  year  of  heaviest  flowering  the  Illawarra 

Flame-tree,  Sterculia  aceri/olia,  A.  Cunn.,  is  almost  leafless  just 

before  blooming  in  November  and  December,  so  that  the  feature 

is   not  restricted   to  one    species   of    this    genus.     These    slight 

differences  are  not  sufficient  to  suggest  a  separation  of  the  species, 

but  they  may  point  to  a  possibilit}^  of  some  slow  evolution  being 

in  progress,  perhaps  the  result  of  climatic  influence. 


BY    R.    H.    CAMBAGE.  579 

Recentl}''  some  prominence  has  been  given  to  the  fact  that  the 
soft  wood  of  the  Queensland  Bottle  Tree,  Sterculia  rupestris, 
Benth.,  after  being  sawn  and  put  through  a  chaff  cutter,  is  useful 
as  a  fodder.  The  branches  of  the  Currajong  are  little  more  than 
bark  and  wood  of  a  very  pithy  nature,  and  may  yet  be  found 
useful  for  the  same  purpose  if  treated  in  a  similar  manner.  The 
tree  is  a  most  valuable  one  to  pastoralists,  and  its  conservation 
should  receive  every  consideration  at  their  hands.  Near  the  29- 
mile  post,  on  the  Orange  to  Canowindra  road,  may  be  seen  a 
luxuriant  growth  of  these  trees  on  a  hillside  composed  almost 
wholly  of  limestone. 

Along  the  creek  banks  and  flats,  near  Canowindra,  are  Casua- 
rina  C iinninghamiana,  Eucalyptus  rostrata,  and  E.  Bridgesiana. 
After  that  the  following  may  be  noticed  distributed  along  the 
road  to  Cowra  : — E.  tereticoruis  (not  plentiful),  E.  melliodora,  E. 
hemiphloia  var.  cdhens,  E.  Woollsiana,  Ccdlitris  7'obitsta,  Acacia 
implexa  (Hickory),  and  Sterculia  diver sif alia,  while  Eucalyptus 
conica  (Woolly-butt)  may  be  seen  on  the  flats  near  Cowra. 

Morongle  Creek  is  situated  some  few  miles  above  Cowra,  and 
along  its  banks,  as  well  as  on  the  flats  near  the  Laehlan,  E. 
conica  is  fairly  plentiful.  It  is  locally  known  under  the  names 
of  Box,  Apple,  and  Woolly-butt,  but  there  are  strong  grounds  for 
believing  that  it  was  known  to  the  aborigines  as  Morongle.  The 
latter  name,  however,  appears  to  have  fallen  into  disuse,  and  I 
could  only  hear  of  one  person  who  had  heard  this  name  from  an 
aborigine. 

From  Cowra  to  Grenfell  is  about  34  miles  westerly,  and  the 
Eucal3^pts  noticed  along  the  roadside  were : — E.  rostrata,  E. 
melliodora,  E.  conica,  E.  Woollsiana,  E.  hemijMoia  var.  alhens, 
E.  tereticoruis,  and  var.  dealhata,  E.  sideroxylon  (Ironbark),  E. 
affinis  (White  Ironbark  or  Ironbark  Box),  E.  macrorhyncha,  E. 
polyanthema  and  E.  Bridgesiana.  These  were  all  seen  before  the 
15-mile  post  was  reached,  and  each  species  was  again  noticed 
between  that  point  and  Grenfell. 

Various  shrubs  and  trees  passed  between  Cowra  and  Grenfell 
were  : — Callitris    robusta,    C.    calcarata,    Sterculia    diver sifolia, 


580  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

Macrozamia  secunda,  C.  Moore  (a  dwarf  form  of  Cycad  found  in 
various  western  areas  chief!}"  on  sandstone  ridges),  Casitarina 
quadrifcdvis,  Acacia  diffusa  (sometimes  called  Pin  Bush  from  the 
shape  of  the  rigid  pointed  leaves),  Exocmyus  cujiressiformi^,  and 
Pterostylis  rpjiexa,  R.Br. 

Going  north  from  Grenfell  along  the  Goolagong  road  for  about 
nine  miles,  and  then  turning  to  the  right  for  three  miles,  the 
following  trees  and  shrubs  were  noted  : — Eucalyptus  hemfiiphloia 
var.  alhens^  E.  tereticornis  and  var.  dealhata,  E.  raeUiodora^  E. 
Woollsiana,  E.  macrorhyncha,  E.  sideroxylon,  E.  Bridgesiana,  E. 
couica,  E.  rostrata,  E.  affinis,  E.  polyanthema^  Callitrls  calcarata, 
C.  robusta,  Acacia  doratoxylon  (Currawong),  A.  dealhata  (green 
variety),  A.  hakeoides,  Casuarina  quadrivalvis  (She  Oak  or 
Mountain  Oak),  C.  Luehmanni  (Bull  Oak),  Lpptospermurti 
scojmrium,  Forst.  (Tea  Tree,  the  form  with  narrow  pointed  leaves), 
StercuUa  diversifolia,  Pittosporum  phillyroioides,  Eusamis  acumi- 
natus  (Quandong),  Exocarpus  cupressiformis ,  Eriostemon  myopo- 
rides,  DC,  and  Cassinia  sp. 

The  rabbits  had  eaten  the  bark  off  the  roots  of  the  Currawong 
trees  in  many  cases. 

About  four  miles  north-easterly  from  Grenfell,  on  the  Newgrove 
Road,  are  Eucalyptus  hcemastoma  var.  micrantha,  and  Acacia 
obtusata  (I). 

Between  Orange  and  Cadia,  a  distance  of  about  15  miles,  the 
country  is  basaltic  most  of  the  way  and  for  the  greater  part 
elevated.  The  Eucalypts  growing  near  the  road  w^ere  : — E.  vimi- 
7ialis,  E.  rubida,  E.  Bridgesiana  (3-m.),  E.  coi-iacea,  E.  stelhdata 
(4-m.),  E.  aggy^egata,  Deane  and  Maiden  (5-m.),  E.  dives  (8-m.),  E. 
vimiiialis,  E.  coriacea,  E.  macrorhyncha,  E.  stelhdata  (12-m.),  E. 
Cambagei,  E.  melliodora  and  E.  dives. 

Other  trees  and  shrubs  were  : — Acacia  dealhata  (Silver  Wattle), 
A.  melauoxylon  [T\  (7-m.),  Banksia  inargriiata,  Cav.  (Honeysuckle), 
Exocaiyus  c2ip7'essiJ'o7'inis,  E.  striata,  R.  Br.,  Leptospermum  sp., 
and  Casuarina  Cunniughamiana,  the  latter  growing  along  the 
banks  of  the  creek  at  Cadia.  These  River  Oaks  extend  for  some 
miles  along  the  creek  above  Cadia,  as  well  as  downwards  to  the 


BY    R.    H.    CAMBAGE.  581 

Lachlaii  River,  and  this  must  be  one  of  the  locaHties  where  they 
are  found  at  their  greatest  elevation  above  sea-level. 

Eucalyptus  agyreyata  follows  alluvial  flats  or  damp  places  on 
the  tablelands,  and  is  found  in  various  localities  south  of  Bathurst, 
between  Burraga  and  Oberon.  It  may  also  be  seen  from  the 
railway  line  on  the  flats  near  Wallerawang.  There  is  ver}^  little 
of  it  in  the  Orange  district,  but  in  addition  to  the  above  locality 
on  the  Cadia  road  it  is  growing  between  the  6-  and  7-mile  posts 
on  the  road  from  Orange  to  Forest  Reefs,  probably  a  continuation 
of  the  same  patch.  It  appears  to  grow  in  exactly  similar  situa- 
tions to  the  Camden  Woolly-butt  {E.  2Iacarthuri,  Deane  and 
Maiden),  found  in  the  Moss  Vale  district.  It  has  a  fairly  rough 
bark  extending  right  up  among  the  branches,  in  texture  being 
something  between  a  flbrous  and  a  flaky  bark.  At  Hobby's 
Yards,  south  of  Newbridge,  the  tree  is  know^n  as  Woolly-butt, 
while  in  the  district  south  of  Oberon  it  is  often  called  Flooded 
Gum  from  the  fact  that  it  follows  near  the  banks  of  the  creeks. 
Still  it  is  very  distinct  from  the  various  Woolly-butts  and  Flooded 
Gum  of  the  coast  districts.  The  leaves  are  fairly  narrow,  the 
stem-sucker  foliage  being  somewhat  broader,  but  at  the  same 
time  not  remarkably  broad.  The  timber,  which  is  pale  and 
rather  light,  does  not  seem  to  have  any  reputation  for  durability. 
The  flowers  are  numerous,  the  fruits  small  and  clustered  in  a 
manner  which  suggested  the  botanical  name  for  the  species.  It  is 
recorded  by  the  authors  from  around  the  Goulburn  district,  north 
and  south,  but  may  possibly  have  a  considerable  range  on  the 
tablelands  of  this  State. 

Between  Cadia  and  Mandurama  the  country,  which  is  generally 
clear,  is  of  more  geological  than  botanical  interest.  The  rocks 
are  chiefly  igneous,  and  amongst  others  is  an  interesting  horn- 
blende-andesite  near  Burnt  Yards.  The  trees  growing  along  the 
roadside  are  '.—Eucalyi)tus  viininalis^  E.  Bridgesiana^  E.  tereti- 
cor7iis,  E.inacror'hyncha,  E.  Cambagei,  E.  'polyantheina^  E.  melli- 
odora,  E.  hemvphloia  var.  aJbens  (which  ceases  5  miles  before 
Mandurama  is  reached),  E.  dives,  Casuarina  Ciinninyhamimia, 
Acacia  diffusa,  A.  dealhata,  and  A.  melanoxylon  (?). 


582     BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

From  Mandurama  to  Hobby's  Yards  the  country  gradually 
rises,  and  the  following  species  ma}^  be  seen  : — Acacia  dealbata, 
Eucalyptus  mellioclora,  E.  Bridyesiaua,  E.  Camhayei  (not  plenti- 
ful), E.  vwiinalis,  E.  macrorhyncha^  E.  dives,  E.  tereticoriiis  (not 
plentiful  on  the  highest  parts),  E.  stelhdata,  E.  coriacea  (on  the 
highest  parts),  E.  ruhida,  and  E.  agyr-egata,  the  latter  being  on 
the  flats  near  Neville  and  again  at  Hobby's  Yards  on  the  road  9 
miles  from  Newbridge  towards  Trunkey. 

From  Hobby's  Yards  to  Trunkey,  a  distance  southerly  of  about 
9  miles,  the  country  falls  and  the  following  trees  were  passed  : — 
Acacia  dealbata,  Eucalyptus  aggregata,  E.  coriacea,  E.  stellulata, 
E.  rubida,  E.  Bridgesiana,  E.  macrorhyncha,  E.  dives  (14  miles 
from  Newbridge),  E.  melliodora,  E.  maculosa,  E.  polyanthema,  E. 
Car/ibagei,  and  E.  hcemastoma  var.  micrantha.  The  latter  being 
a  few  hundred  yards  to  the  westward  and  not  visible  from  the 
road.  Gonipholobium  Huegelii,  Benth.,  was  found  a  few  miles 
east  of  Hobby's  Yards. 

Continuing  from  Trunkey  to  Tuena,  the  country  still  falls  to 
the  Abercrombie  River,  and  within  the  first  mile  the  trees  passed 
were  : — Eucalyptus  polyanthema,  E.  Cambagei,  E.  maculosa,  E. 
macrorhyncha,  E.  Bridgesiana,  E.  melliodo7'a,  E.  tereticoriiis,  E. 
viminalis,  and  E.  rubida.  Thence  to  the  Abercrombie  River,  at 
12  miles  from  Trunkey,  are  E.  hemiphloia  var.  albens,  E.  polyan- 
thema, E.  melliodora,  E.  Cambagei,  and  E.  macrorhyncha. 

It  may  be  noticed  that  during  the  last  1 1  miles  all  the  tj^pical 
cold  country  gums  have  been  left  behind  owing  to  the  increased 
warmth  of  the  somewhat  lower  country.  Still  an  occasional  tree 
of  E.  viminalis  may  be  found  even  below  this  level,  there  being 
one  solitary  tree  of  this  species  just  below  the  junction  of  the 
Burro wa  and  Lachlan  Rivers,  although  it  looks  quite  out  of  place 
amidst  its  surroundings. 

Near  the  Abercrombie  River  and  towards  Tuena  are  Casuarina 
C unninghamiana,  E.  hemiphloia  var.  albens,  E.  polyanthema,  E. 
melliodora,  E.  tereticornis,  E.  Bridgesiana,  E.  macrorhyncha,  and 
a  small  patch  of  E.  hcemastoma  var.  micrantha,  near  Tuena  and 
close  to  the  36-mile  post  from  Newbridge. 


BY    R.    H.    CAMBAGE.  583 

Many  of  these  notes  were  made  for  purposes  of  personal  refer- 
ence only  and  not  with  the  object  of  publication.  The  conditions 
also  under  which  they  were  taken  did  not  permit  of  a  thorough 
investigation  throughout  a  district,  consequently  they  are  not  at 
all  exhaustive.  The  genus  Eucalyptus  is  the  only  one  which  may 
be  considered  approximately  complete.  So  far  as  the  Bathurst 
district  is  concerned,  a  much  more  detailed  list  of  plants  may  be 
found  in  the  '  Notes  on  the  Flora  of  Bathurst '  of  Mr.  W.  J. 
Clunes  Ross,  B.Sc,  F.G.S.  (Kept.  Assoc.  Adv.  Sci.  vii.,  1898,  467). 

From  Bathurst  to  Limekilns  is  northerly  about  20  miles,  and 
the  conspicuous  trees  growing  along  the  roadside  are  Eucalyptus 
tereticornis,  E.  Bridgesicina,  E.  melliodora,  E.  viminalis,  E.  ruhida 
(9-m.),  E.  Camhagei,  E.  macrorhyncha,  E.  polyanthema^  E.  inacu- 
losa,  E.  pidverulenta^  Sims,  Casuarina  Cunni7ighamiana,  Ango- 
jjhora  intermedia,  DC,  (Apple),  Acacia  vestita,  A.  dealbata,  A, 
diffusa,  A.  lanigera,  A.  linifolia,  Willd.,  and  Kunzea  peduncular-is, 
F.v.M. 

Typical  Devonian  fossils  may  be  found  in  some  of  the  creeks 
along  this  road,  having  been  washed  down  from  the  hills  to  the 
eastward. 

The  trees  of  AngopJiora  intermedia  are  first  met  with  just 
beyond  the  6-mile  post,  and  this  is  the  only  locality  within  the 
area  described  in  these  papers  where  a  species  of  Angophora  has 
been  noticed. 

A.  intermedia  is  a  very  common  "Apple  Tree"  along  the  coast 
and  north-westerly  districts,  at  least  as  far  out  as  Gilgandra,  but 
over  the  Great  Dividing  Range  it  keeps  practically  to  the  northern 
side  of  the  Great  Western  Railway  Line.  Its  timber  is  soft  but 
useful  for  some  kinds  of  wheelwrights'  work.  Asa  fuel  it  burns 
well,  and  a  dead  tree  lying  on  the  ground  will,  after  being  lighted, 
quietly  burn  right  away,  leaving  in  white  ashes  the  imprint  of 
both  trunk  and  branches.  South  of  Bathurst  (as  intimated  in 
Part  vi.)  Eucalyptus  Bridgesiana  is  usually  known  as  Apple,  but 
when  found  growing  in  company  with  an  Angophora  it  appears 
to  be  either  WooUybutt  or  Peppermint,  though  in  Gippsland  it 
bears  the  native  name  of  But  But  according  to  Mr.  A.  W.  Howitt. 


584:  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

The  trees  growing  near  the  20-mile  post  at  Limekihis  and 
mentioned  as  Eucalyi^tiis  jnilverulenta,  Sims,  are  the  same  as  those 
found  in  the  Goulburn  to  Marulan  district  and  known  as  Argyle 
Apple.  It  has  a  fibrous  short  stringy  bark  and  pale  glaucous 
leaves  exactly  similar  to  the  Goulburn  trees,  while  its  timber  has 
the  same  reddish  tint.  In  going  southwards  from  Limekilns 
towards  Goulburn  there  seems  to  be  a  break  of  at  least  70  or  80 
miles  before  the  species  is  again  found.  The  question  once  more 
comes  up  as  to  whether  in  such  cases  as  this  the  growth  was 
formerly  continuous,  and  has  partly  perished  from  climatic  or 
other  causes,  or  whether  the  seeds  were  originally  distributed 
only  at  these  wide  intervals.  A  careful  detailed  geological  survey 
of  the  intervening  area  would  be  one  necessary  part  of  the 
research  required  to  assist  in  the  determination  of  this  interesting 
question. 

In  going  from  Bathurst  southwards  to  Rockley,  a  distance  of 
about  22  miles,  various  trees  and  shrubs  noticed  near  the  roadside 
were  : — Eucaly2)tus  viminalis,  E.  Bridyesiana^  E.  j^olyanfhema^  E. 
melliodora,  E.  Camhagei,  E.  hcemastoma  var.  micrantha,  E.  macro- 
rhyncha,  E.  tereticornis,  E.  cordata,  Labill.,  var.,  E.  ruhida,  E. 
coriacea,  E.  stellulata,  E.  dives,  Acacia  dealbata,  A.  lanigera,  A. 
discolor,  A.  diffusa,  A.  ohtusata,  A.  armata,  A.  lunata,  Sieb., 
Casuarina 2)aludosa{X),  Calythrix  fetragona,  and  Oia:r  stricta,^.^Y. 

For  about  10  miles  the  formation  is  granite,  and  the  land  being 
cultivated,  it  follows  that  the  trees  are  scarce.  But  about  the 
11-  and  12-mile  posts  are  hills  of  micaceous  schist,  slate  and  lime- 
stone, w^here  the  flora  changes  entirely  and  the  plants  are  fairly 
numerous. 

On  a  continuation  of  these  hills  towards  Wiseman's  Creek,  the 
Flannel  Flower,  Actinotus  Helianthi,  Labill.,  may  be  found,  and 
this  is  its  most  western  locality  in  the  Bathurst  district  known 
to  me. 

In  their  *  Research  on  the  Eucalypts '  Messrs.  Baker  &  Smith 
express  the  opinion  that  the  t3^pical  E.  pulveridenta,  Sims,  which 
was  described  in  1819  from  a  seedling  plant  grown  in  a  nursery 
in  England,  was  not  the  Argyle  Apple  as  commonly  supposed. 


BY    R.    H.    CAM13AGE.  585 

but  a  tree  growing  at  Cow  Flat  near  Bathurst  and  closely  allied  to 
the  Tasmanian  E.  cordata,  Labill.  From  the  drawings  and  the 
description  of  the  drooping  stems,  the  similarit}'  also  seems  to 
me  greater  to  what  I  shall  term  in  these  papers  the  Bathurst  E. 
cordata  or  E.  cordata  var.  than  to  the  Argyle  Apple.  The  first 
difficulty  that  presents  itself  is  to  settle  upon  the  locality  where 
the  original  seeds  were  collected  from  which  the  plants  in  Eng- 
land were  grown.  The  locality  is  stated  as  New  Holland,  and  if 
this  could  at  those  early  dates  have  sometimes  erroneously 
included  Tasmania,  it  would  seem  possible  the  seeds  were  collected 
there  and  were  perhaps  typical  E.  cordata,  for  it  must  be  remem- 
bered that  the  species  {E.  pidvericleuta)  was  described  from  culti- 
vated specimens. 

In  the  year  1822  Allan  Cunningham  collected  near  Cox's  River 
a  shrubby  species,  which  he  named  E.  pidvlge7'a,  and  from  his 
general  description  this  appears  to  be  the  same  as  the  Bathurst 
E.  cordata.  Curiously  this  form  does  not  appear  to  have  been 
again  collected  till  February  of  1900,  when  I  found  it  while 
surveying  on  the  hills  near  Cow  Flat,  opposite  the  old  Apsley 
Copper  Mine,  at  about  one  mile  west  of  the  11-mile  post  on  the 
road  from  Bathurst  to  Rockley.  It  was  not  until  the  year  1813 
that  the  Blue  Mountains  were  crossed,  so  that  if  the  seeds  of  the 
typical  E.  pulvei'identa  were  collected  beyond  that  point,  no  time 
was  lost  in  having  them  bearing  fruit  in  England  by  1819,  when 
the  description  was  published.  Still  a  flowering  plant  of  Euca- 
lyptus glohidus,  Labill.,  said  to  be  only  two  years  old,  was 
exhibited  at  the  last  meeting  of  the  Australasian  Association  for 
the  Advancement  of  Science  in  Hobart,  but  this  would  seem  to 
be  exceptionally  young. 

As  there  is  some  doubt  from  the  information  at  present  avail- 
able as  to  which  is  the  typical  E.  indveridenta,  I  shall  continue 
to  refer  it  in  these  papers  to  the  Argyle  Apple.  "^ 


*  Mr.  J.  H.  Maiden,  F.L.S.,  &c,,  refers  to  this  matter  in  a  paper  on  E. 
pulverulenta  (these  Proceedings,  1901,  p.  547). 


586     BOTANY  OF  THE  INTERIOR  OF  NEW  SOUTH  WALES, 

The  question  as  to  whether  the  Bathurst  E.  cordata  differs  from 
the  Tasmania!!  one  sufficiently  to  be  regarded  as  a  separate  species 
will  probably  remain  to  some  extent  a  matter  of  opinion.  There 
is  a  noticeable  difference  in  the  shape  and  texture  of  the  leaves, 
those  of  the  Bathurst  trees  being  thicker  and  more  leathery  than 
are  those  of  the  Tasmanian  species,  while  the  latter  are  often 
crenulate  and  more  acuminate.  The  amount  of  variation  in  the 
length  of  peduncle  appears  to  be  about  the  same  in  both  cases. 
Some  difference  may  also  be  noticed  in  the  shape  of  the  fruits,  an 
important  point  in  botany,  those  of  the  Tasmanian  trees  having 
a  globular  tendency.  In  both  cases  the  fruits  are  sessile  and 
a.rranged  in  three-j.  The  Bathurst  trees  are  growing  near  the 
top  of  a  rock}^  hill  of  micaceous  schist  formation,  seldom  exceed- 
ing more  than  20  feet  high,  and  little  more  than  3  inches  in 
diameter,  with  the  heads  of  the  trees  bending  right  over.  The 
buds  commonly  have  a  partially  second  operculum.  The  wood  is 
pale,  the  bark  white  and  inclined  to  be  ribbony.  The  whole  of 
the  trees  to  be  found  cover  very  little  more  than  an  acre  of 
ground,  and  no  others  of  the  same  species  being  known  on  the 
mainland  of  Australia,  it  is  difficult  from  this  small  area  to  make 
a  satisfactory  comparison  with  the  Tasmanian  trees. 

To  the  south  of  Ilobart  E.  cordata  m^j  be  found  on  hills  of 
volcanic  rock.  From  a  bushman's  point  of  view,  they  are  very 
like  the  Bathurst  trees,  except  that  they  grow  larger  and  are 
usually  much  more  erect,  only  two  or  three  instances  being 
noticed  on  one  hill  where  the  heads  showed  a  tendency  to  bend 
over,  though  not  to  the  extent  which  may  be  seen  at  Bathurst. 
In  bark  and  timber  they  appear  identical.  The  Bathurst  trees 
are  the  toughest  to  be  found  among  the  hills  on  which  they  grow, 
and  the  same  may  be  said  of  these  Hobart  trees,  except  it  be 
some  stunted  forms  of  E.  cocc{fcra,  Hook.,  growing  on  Mount 
Wellington.  The  Tasmanian  or  typical  E.  cordata  also  has  the 
partially  double  operculum,  though  this  feature  may  be  found  at 
times  on  other  species  of  Eucalyptus.  Considering  the  widely 
different  conditions,  both  geological  and  climatic,  under  which 
the  two  trees  grow,  some  difference  at  least  in  general  appearance 


BY    R.    H.    CAMBAGE.  587 

might  be  expected.  Still,  in  my  experience,  the  differences  found 
among  various  trees  in  scattered  localities  have  not  always  been 
so  great  as  expected,  except  perhaps  those  affected  by  extreme 
cold,  or  by  boisterous  weather  along  the  sea  shore.  In  the 
absence  of  other  causes,  the  better  soil  in  which  the  Hobart  trees 
are  growing  might,  however,  be  sufficient  to  account  for  the 
larger  growths  found  there. 

Altogether,  the  differences  in  herbarium  specimens  of  these 
two  trees,  so  far  as  I  know  them,  seem  sufficient  to  enable  a 
botanist  to  separate  them,  while  at  the  same  time  they  have  so 
many  points  in  common  that  I  am  disposed  for  the  jDresent  to 
regard  one  as  a  variety  of  the  other. 

In  view  of  the  scarcity  and  the  dwarfed  form  of  the  Bathurst 
trees,  together  with  the  fact  that  they  are  growing  on  some  of 
our  oldest  rocks  (a  fact  in  itself  not  sufficient  to  prove  much),  I 
am  inclined  to  believe  that  the  species  was  once  more  plentiful, 
and  that  this  patch  is  only  a  remnant  of  what  previousl}^  extended 
over  a  very  much  larger  area,  the  limits  of  which  it  is  impossible 
to  determine. 

Oberon  is  situated  about  32  miles  south-easterly  from  Bathurst, 
and,  being  higher,  has  a  slightly  different  flora.  The  most  con- 
picuous  difference,  however,  is  in  the  size  of  the  trees,  those  in 
the  Oberon  district  being  often  very  tall,  reminding  one  in  this 
respect  of  many  spots  along  the  coastal  area. 

In  going  easterly  towards  the  Duckmaloi  Creek  for  about  seven 
miles,  partly  along  the  Hampton  Road,  the  following  species  may 
be  noticed  : — Eucalyptus  viminalis,  E.  aggregata,  E.  stellulata,  E. 
coriacea,  E.  rubida,  E.  dives,  E.  amygdalina,  E.  vitrea,  Baker, 
Exocarpus  cupressiformis,  Banksia  marginata  and  Acacia  penni- 
nervis,  Sieb.  (Black  Wattle). 

The  trees  of  E.  vitrea  were  noticed  between  the  5-  and  6-mile 
posts  on  both  sides  of  the  road.  They  have  the  general  appear- 
ance of  the  Peppermint  group,  but  are  distinguishable  from  E. 
dices  and  E.  amggdalina,  both  of  which  may  be  found  before  the 
4-mile  post  is  reached.  In  this  locality  E.  vitrea  has  fairly  rough 
grey  peppermint  bark  on  the  trunk  for  a  distance  upwards  of 


588  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH    WALES, 

about  15  or  20  feet  on  a  tree  50  feet  high.  Above  20  feet  the 
bark  is  smooth  and  a  dirty  white.  The  leaves,  which  have  the 
lateral  veins  almost  parallel  to  the  midrib,  are  somewhat  shining, 
and,  even  when  dried,  have  a  glassy  appearance.  The  fruits  seem 
slightly  larger  than  those  of  E.  dives,  which  are  again  usually 
larger  than  those  of  E.  amygdalina. 

The  species  had  never  been  noticed  by  me  before  these  Oberon 
trees  were  found  in  February,  1900,  at  which  time  it  was  unde- 
scribed,  but  shortly  afterwards  I  recognised  the  same  species  on 
the  roadside  between  Marulan  and  Moss  Vale.  In  both  localities 
the  trees  seemed  to  average  about  50-60  feet  high.  In  some  forms, 
owing  to  its  shining  leaves  and  the  smoothness  of  the  upper  part 
of  the  trunk  and  branches,  it  somewhat  approaches  E.  coriacea 
in  general  appearance,  but  its  closest  affinities  seem  to  be  the 
Peppermints. 

In  going  southerly  for  18  miles  from  Oberon  towards  Swatch- 
field  the  following  Eucalypts  may  be  seen  : — E.  steUidata,  E. 
Bridgesiana,  E.  coriacea,  E.  viminalis,  E.  fastiyata,  Deane  and 
Maiden  (5-m.),  E.  aygregata  (7-m.),  E.  amygdalina,  E.  dives 
(14-m.),  E.  coriacea,  E,  stellulata,  E.  aggregata,  E.  viminalis,  E. 
fastigata,  and  E.  amygdalina. 

Other  trees  are: — Acacia  dealbata,  Banksia  marginata.  Acacia 
melanoxylon,  and  Goodenia  ovata,  Sm.  (?),  a  shrub  locally  known 
as  Wild  Hops. 

Eucalyptus  aggregata  is  fairly  common  on  the  southern  side  of 
Oberon. 

E.  fastigata  has  something  the  appearance  of  the  coast  Black- 
butt  {E.  pilularis),  but  the  rough  bark  generally  extends  at  least 
up  to  and  sometimes  on  the  branches,  though  its  terminal  point 
is  variable.  It  occurs  plentifully  in  the  southern  part  of  the 
Oberon  district  and  towards  the  Jenolan  Caves,  and  appears  to 
be  known  throughout  as  Blackbutt,  this  name  liaving  probably 
been  applied  to  it  by  the  early  settlers  because  of  its  general 
resemblance  to  the  coast  trees  which  bear  that  name.  The  trees 
are   among   the   largest  in  the  Oberon  district,  and  though  the 


BY    R.    H.    CAMBAGE.  589 

fissile  timber  is  freely  used  it  is  not  considered  to  have  any 
special  value. 

Messrs.  Deane  and  Maiden  have  expressed  the  opinion  (these 
Proceedings,  1901,  p.  123)  that  E.  vitrea  is  a  form  of  E.fastigata. 
The  fruits  of  E.  vitrea  are  usually  rounder  than  those  of  E. 
fastigata,  which  are  somewhat  pear-shaped  and  slightly  domed. 
Speaking  with  a  somewhat  limited  acquaintance  with  these  trees 
they  appear  to  me  to  be  separable. 

There  appears  to  be  some  doubt  as  to  whether  E.  fastiyata  is 
sufficiently  distinct  from  E.  regnans,  F.v.  M.,  as  to  warrant  its 
having  specific  rank.  My  own  observations  on  the  matter  made 
from  specimens  collected  near  Dandenong  in  Victoria,  west  of 
Albion  Park,  west  of  Milton  and  at  Oberon  in  jS'ew  South  Wales, 
lead  to  the  conclusion  that  the  two  trees  are  ver}'  closely  allied,  and 
it  may  yet  be  found  that  E.fastigata  is  only  a  variety  or  form  of  E. 
regnans.  Certainly  the  bark  of  E.  regnans  in  Victoria  is  usually 
smooth  to  perhaps  within  10  or  15  feet  from  the  ground  or  some- 
times less,  while  in  N.S.  Wales  that  of  E.  fastigata  is  rough  up 
to  and  often  on  the  branches,  but  this  character  in  the  variability 
of  bark  is  v^ery  deceptive.  Again,  the  fruits  of  E.  regnans  which 
I  collected  at  Dandenong  are  chiefly  truncate,  while  at  Oberon 
those  of  E,  fastigata  are  chiefly  domed,  with  the  valves  often 
slightly  exserted,  but  those  from  Albion  Park  show  forms  both 
truncate  and  slightly  domed,  though  the  rough  somewhat  stringy 
bark  on  the  trees  (locally  called  Messmate)  at  the  latter  place 
extends  on  to  the  branches.  In  all  three  cases  the  peduncles 
may  be  found  in  pairs  in  the  axils,  as  well  as  solitary.  So  far 
as  the  rough-  and  smooth-barked  trees  are  concerned  those 
characters  are  sufficient  to  enable  the  two  to  be  separated,  but 
there  are  probably  some  forms  that  would  present  considerable 
difficulty. 

The  following  is  a  list  of  Eucalypts  passed  between  Forbes  and 
Oberon  : — E.  77ieUiodora,  E.  Woollsiana,  E.  hemiphloia  var.  alhens, 
E.  Gonica,  E.  rostrata,  E.  tereticornis  and  var.  dealhata,  E.  Bridge- 
siana,  E.  macrorhyncka,  E.  virninalis,  E.  Cambagei,  E.  dives,  E. 
stelhdata,  E.  coriacea,  E.  ruhida,  E.  hcemastoma  var.  micranthay 
39 


590  BOTANY    OF    THE    INTERIOR    OF    NEW    SOUTH     WALES, 

E.  maculosa,  E.  polyanthema,  E.  amygdalina,  E.  sideroxyhn 
(only  near  Grenfell),  E.  affinis  (only  near  Grenfell),  E.  aggregata, 
E.  pulverulenta  (only  near  Limekilns),  E.  cordata  var.  (only  near 
Cow  Flat),  E.  vitrea  and  E.  fastigata. 

About  eighteen  years  ago  the  Native  Bears  (Phascola)  cfus 
cinereus)  were  fairly  plentiful  along  parts  of  the  road  between 
Bathurst  and  Rockley,  and  I  noticed  that  although  they  might 
at  times  be  found  on  most  kinds  of  Eucalypts,  they  seemed  to 
show  a  preference  for  E.  melliodora,  the  Yellow  Box.  In  AYestern 
lllawarra,  about  twenty  years  ago,  their  favourite  trees  were  the 
White  Box,  since  named  E.  quadrangulata,  Deane  and  Maiden. 
Around  Bolivia,  and  towards  Tenterfield,  about  fifteen  years  ago 
some  rather  small  Box-trees  were  constantly  inhabited  by  the 
Koala.  Evidently  they  had  some  jDreference  for  the  Box  group, 
and  it  was  often  remarked  that  they  w^ere  rarely  seen  in  an 
Angophora,  and  also  were  usually  more  numerous  among  the 
Gums  than  the  Stringybarks. 

From  the  above  remarks,  and  as  the  Native  Bears  live  on  the 
leaves  of  trees,  it  may  be  gathered  that  the  chemistry  of  our 
Eucalypts  had  been  investigated  long  before  that  branch  of  study 
was  taken  up  by  man. 

Many  things  to  be  met  with  in  Nature  command  some  affection, 
but  the  amount  varies  according  to  different  conditions.  Our 
vegetation,  however,  when  it  presents  its  greatest  beauty,  is 
admired  rather  than  loved.  This  is  particularly  the  case  with 
our  coastal  scenery,  which  has  much  more  to  commend  it  than 
the  less  favoured  flora  of  the  interior.  Still  there  are  many 
evidences  of  admiration,  and  even  veneration  toward  members  of 
the  floral  family  in  Western  New  South  Wales,  being  partly 
brought  about  because  of  their  usefulness  and  partly  from  their 
beauty.  The  result  of  some  observations  made  on  the  matter 
leads  me  to  suggest  that  the  species  which  commands  the  greatest] 
affection  over  the  area  described  in  these  papers  is  Acaeia  pendiday 
the  Myall  or  Boree. 

This  paper  concludes  for  the  present  the  series  of  "  Notes  on 
the  Botany  of  the  Interior." 


BY    R.    H.    CAMBAGE.  591 

I  wish  to  acknowledge  assistance  in  various  ways  from  Mr.  J. 
H.  Maiden,  F.L.S.,  who  has  been  ever  ready  to  discuss  with  me 
any  knotty  points  which  have  arisen  in  regard  to  the  identification 
of  plants,  and,  in  addition  to  naming  many  of  the  species,  to  give 
any  information  that  might  add  to  the  usefulness  of  these  papers. 

My  thanks  are  also  due  to  Mr.  R.  T.  Baker,  F.L.S.,  who  has 
also  contributed  in  an  exactly  similar  manner. 

I  also  desire  to  acknowledge  services  rendered  by  Mr.  E. 
Betche,  Botanic  Gardens,  Mr,  W.  S.  Dun,  Palaeontologist  Mines 
Department,  Mr.  J.  J.  Fletcher,  and  Mr.  H.  G.  Smith,  F.C.S. 


EXPLANATION  OF  PLATE  XXVII. 

Map  of  New  South  Wales  showing  Author's  Koute 

Part 

i. 

— Bourke  to  Cobar. 

Part 

ii. 

— Cobar  to  Muclall. 

Part 

ill. 

— Mudall  to  Euabalong, 

Part 

iv. 

— Mt.  Hope  to  Parkes. 

Part 

V. 

— Parkes  to  Marsden. 

Part 

vi. 

— Marsden  to  Narrandera. 

Part 

vii. 

—Forbes  to  Bathurst. 

592 


NOTES    AND    EXHIBITS. 


Professor  Haswell  showed  under  the  microscope  preparations 
of  the  Sporocysts  described  in  his  paper. 

Mr.  Froggatt  exhibited  specimens  of  the  curious  coccid,  Frenchia 
casuarince,  Mask.,  recently  found  by  him  on  casuarinas,  near 
Condobolin,  N.S.W.;  the  species  was  originally  described  from  the 
Wimmera  district,  Victoria,  and  is  now  recorded  for  the  first 
time  from  New  South  Wales.  Also  specimens  of  the  larvae  of 
the  pine-scrub  beetle  (Diadoxus  erythrurus)  recently  collected 
from  dead  or  dying  Currawong  bushes  (Acacia  doratoxyloii)  on 
the  ranges  about  the  Lachlan  River  beyond  Condobolin.  As 
living  trees  are  to  be  found  side  hy  side  with  dead  or  dying  ones, 
and  as  some  of  the  latter  may  yield  as  many  as  half  a  dozen  larvae, 
Mr.  Froggatt  expressed  his  belief  that  in  the  locality  mentioned 
the  destruction  of  the  trees  was  attributable  to  the  insects,  and 
not  to  the  drought. 

Mr.  Fred.  Turner  exhibited  specimens  of  {\)  Plantago  coronopus, 
Linn.,  collected  by  him  in  the  north  reserve  at  Coogee  Bay,  a 
species  not  hitherto  found  in  New  South  Wales,  he  believed, 
though  it  has  been  recorded  from  the  Southern  and  Western 
States;  (2)  Lobelia  pur pni^ascens,  R.Br.,  in  a  state  of  malformation, 
a  condition  not  previously  observed  by  him. 

Mr.  Baker  exhibited  two  sections  of  "  Blackfellow's  bread," 
Polyporus  mylittce,  Cooke  and  Massee,  showing  the  fructification  of 
this  fungus — one  in  the  growing,  and  the  other  in  the  indurated 
stage;  timber  of  Euroschinus  falcatus,  Hook,  f.;  a  particularly 
fine  specimen  of  the  flannel  flower,  Actinotus  Helianthi,  Labill., 
showing  fasciation  of  the  stem;  and  the  foliage  and  bark  of  a 
probably  now  species  of  phyllodineous  Acacia  from  Kingstown 
viaUralla;  alsoa  "Box"seedlingof  Eucalyptus populifolia^  Hook.f., 
from  Mount  Drysdale,  to  illustrate  the  contention  that  the  so-called 
•'  Box"  suckers  of  the  western  plains  are  not  suckers  in  the  true 
botanical  sense,  but  seedlings;  and  in  support  of  the  view  that 


NOTES    AND    EXHIBITS.  593 

Eucalypts  do  not  sucker  at  all.  The  term  "  sucker  "  as  used  in 
Australia  is  a  colloquialism  applied  to  adventitious  growths  from 
stems  and  branches  of  Eucalypts. 

Mr.  Cambage  exhibited  herbarium  and  timber  specimens  in 
illustration  of  his  paper.  He  also  showed  fruiting  branches  of 
the  form  of  Pitto>^porum  undulatum,  Yent.,  which  bears  flowers 
with  long  stamens,  from  AVaratah  and  Burwood.  In  the  neigh- 
bourhood of  Sydney,  however,  it  is  quite  a  common  occurrence  to 
find  the  species  in  a  dio3cious  condition,  only  the  form  with  rudi- 
mentary stamens  setting  any  fruits. 

Mr.  Maiden  exhibited  a  type  specimen  of  Eucalyptus  hicolor, 
A.  Cunn.,  which  was  originally  presented  by  Cunningham  to  Sir 
W.  J.  Hooker  and  presented  by  Sir  William  Thiselton  Dyer  to 
the  exhibitor.  He  also  showed  a  drawing  of  a  properly  authen- 
ticated type  specimen  of  E.  polyanthemos,  Schauer,  from  the  her- 
barium of  the  University  of  Cambridge.  Also  a  specimen  of 
Distegia  Ledehouri%  Greene,  from  California,  belonging  to  an 
anomalous  genus  of  the  Compositse. 


WEDNESDAY,  NOVEMBER  26th,   1902. 


The  last  Ordinary  Monthly  Meeting  of  the  Society  for  the 
Session  was  held  in  the  Linnean  Hall,  Ithaca  Road,  Elizabeth 
Bay,  on  Wednesday  evening,  November  26th,  1902. 

Mr.  J.  H.  Maiden,  F.L.S.,  &c..  President,  in  the  Chair. 

Messrs.  Ebenezer  Campling,  Superior  Public  School,  Wing- 
ham,  N.S.W.,  and  Harald  C.  Dannevig,  Department  of 
Fisheries,  Sydney,. were  elected  Members  of  the  Society. 

The  Donations  and  Exchanges  for  the  month,  amounting  to  5 
Vols.,  26  Parts  or  Nos.,  5  Reports,  1  Pamphlet,  received  from  26 
Societies,  (fee,  were  laid  upon  the  table. 


594 


ON  A  NEW  SPECIES  OF  SY2IPL0C0S  FROM  NEW 
SOUTH  WHALES. 

By  R.  T.  Baker,  F.L.S.,  Curator,  Technological  Museum. 

Sydney. 

(Plate  xxviii.) 

Symplocos  Baeuerleni,  sp.nov. 

A  small,  glabrous,  delicate  shrub,  about  6  to  9  feet  high. 

Leaves  ovate,  acuminate,  or  lanceolate,  terminating  in  a  sharp 
point,  glabrous,  pale  on  the  underside,  membranous  or  chartaceous, 
irregularly  toothed  or  entire,  slightly  shining  on  the  upper  surface, 
about  2  inches  long  and  under  9  lines  wide  ;  principal  veins 
distant  and  more  distinct  on  the  underside,  pale-coloured,  slightly 
oblique,  not  always  parallel;  petiole  about  1  line  long. 

Flowers  small,  solitary  in  the  axils  or  3  to  5  in  an  exceedingly 
short  raceme;  petiole  slightly  under  a  line  long.  Bracts  few, 
small,  minutely  hirsute.  Calyx  glabrous,  lobes  exceedingly  short 
and  broad.  Petals  glabrous,  imbricate,  broad,  about  1  line  long. 
Stamens  united  at  the  base  of  the  petals,  indefinite,  numbering 
from  20  or  more.  Pistil  about  as  long  as  the  stamens.  Stigma 
slightly  bifurcated. 

Fruit  ovoid,  bluish-black,  under  4  lines  long,  2  lines  in  diameter, 
contracted  at  the  top  and  crowned  by  the  calyx  lobes. 

Hah. — Tumbulgum,  Murwillumbah, Tweed  River  (W.  Bauerlen). 

This  shrub  differs  considerably  in  its  foliage  from  S.  spicata, 
Roxb.,  S.  Thwaitesii,  F.v.M.,  and  S.  paucistamineics,  F.v.M. — the 
three  species  at  present  recorded  for  Australia  ;  in  fact,  the 
difference  is  so  great  in  this  feature  that  one  would  never  at  first 
sight  take  this  plant  to  be  a  Symjyhcos.  The  floral  and  carpellary 
characters,  however,  show  it  conclusively  to  be  such.  The 
inflorescence  of    the  above  three  species  of  Si/mplocos  described 


BY    R.    T.    BAKER.  595 

for  Australia,  is  in  the  form  of  spikes  or  panicles  with  numerous, 
flowers,  whilst  in  this  species  the  flowers  are  either  solitary  or  few 
(from  3  to  5)  in  an  exceedingly  short  raceme.  The  individual 
flowers,  as  well  as  the  leaves  and  fruits,  are  all  much  smaller  than 
in  any  of  the  above  species. 

It  differs  from  -S'.  spicata,  Roxb.,  also  in  its  small,  delicate,  thin 
leaves  and  smaller  and  pedicellate  flowers.  S.  Thwaitesii,  F.v.M. 
and  S.  paucistamineus  have  firm,  comparatively  large,  coriaceous 
leaves,  and  numerous  spikes  of  over  an  inch  long.  The  fruits 
are  also  smaller  than  those  of  any  of  these  species. 

It  also  appears  to  be  quite  distinct  from  any  Symplocos  des- 
cribed in  the  '  Flora  of  British  India  '  (Hooker). 

It  occurs  in  the  rich  alluvial  gullies  of  Tweed  River,  growing 
amongst  the  recently  described  new  plants  of  that  district, 
obtained  by  Mr.  W.  Bauerlen. 


EXPLA.NATION  OF  PLATE  XXVIII. 

Symplocos  Bduerleni,  sp.n. 

Fig.  1. — Flowering  twig. 
Fig.  2. — Fruiting  twig. 

Fig.  3.  — Slightly  expanded  bud  showing  imbricate  petals. 
Fig.  4. — Individual  flower  showing  disposition  of  petals  and  stamens. 
Fig.  5.  —Individual  flower  showing  disposition  of  sepals  and  petals. 
(Figs.  3,  4,  and  5  are  enlarged). 


596 


STUDIES  ON  AUSTRALIAN  MOLLUSC  A. 

Part  YII. 

By  C.  Hedley,  F.L.S. 

(Plates  xxix.,  xxxi.,  xxxii.,  xxxiii.) 

(Continued  from  p.  29.) 

Chione  lagupus,  Lamarck. 
Vemi^s  lagopus,  Lamarck,  An.  s.  Vert,  v.,  1818,  p.  591. 

The  original  locality  for  this  species  is  King  George's  Sound, 
W.A.  It  was  characterised  by  its  author  as  40  mm.  long,  exter- 
nally white  splashed  with  chestnut,  within  tinged  with  rose; 
remarkable  for  its  close  longitudinal  furrows,  which  are  crenulated 
on  their  upper  sides,  anteriorly  the  intervening  ridges  become 
almost  lamellate.  He  classed  it  next  but  one  to  Venus  yallinula, 
Lamk.  Probably  his  type  passed  with  Delessert's  collection  to 
the  Geneva  Museum.* 

The  species  was  recognised  from  West  Australia  by  Menke,t 
who  had  proposed  for  it  the  name  of  Venus  amyydalum.  Except 
that  Tate  (Trans.  Hoy.  See.  S.A.  ix.,  1887,  p.  91)  has  reduced  it 
to  a  synonym  of  Chione  yallinula^  Lamarck's  name  has  apparently 
dropped  out  of  notice  of  conchological  writers. 

Yet  his  description  applies  exactly  and  exclusively  to  the 
Venus  australis,  Sowerby,  I  which  name  it  should  supersede.  The 
name  Venus  australis  was  in  the  same  year  given  to  another 
species  by  Quoy  k,  Gaimard.     But  in  1790  it  had  already  (S3\st. 

*  Crosse,  Journ.  de  Conch,  xvii.,  1S69,  p.  208. 

t  Menke,  Spin.  Moll.  Nov.  Holl.  1843,  p.  43. 

X  Sowerby,  P.Z.S.,  1835,  p.  22. 


BY    C.    HEDLEY.  597 

Nat.  xiii.,  p.  3282)  been  preoccupied  by  Gmelin  for  yet  another 
species. 

Mactra  abbreviata,  Lamarck. 
(Plate  xxix.,  figs.  1-3.) 

This  species  has  been  mentioned  as  occurring  in  Port  Jack- 
son."^ The  description  of  it  suggested  to  me  that  it  might  perhaps 
be  identical  with  Mactra  pusilla,  A.  Adams.  I,  therefore,  sent 
examples  of  J/,  pusilla  to  Paris,  with  a  request  for  a  comparison 
between  it  and  J/,  ahhreviata. 

Mr.  J.  Mabille  very  kindly  replied  tomy  inquiries  as  follows:  — 
"  La  Mactra  abbreviata  est  representee  dans  les  collections  du 
Museum  d'histoire  naturelle  de  Paris  par  2  exemplaires  en  fort 
bon  etat  de  conservation,  ces  2  exemplaires  constituent  le  type 
meme  de  Lamarck.  lis  sont  completment  distincts  de  la  M. 
pusilla.  Cette  Mactra  abbreviata  est  une  coquille  blanche,  un 
peu  brilliante  ornee  de  s tries  d'accroissement  bien  visibles,  un  peu 
larges  et  comme  aplaties;  de  forme  ovale  triangulaire,  renflee, 
surtout  dans  la  region  umbonale;  a  I'etat  tres  frais  elle  est  revetue 
d'un  epiderme  jaunatre,  ecailleuse,  caduc,  ainsi  qu'en  temoigne 
I'un  des  individus,  qu'en  possede  encore  une  bande  au  bord  ventral. 

"  La  lunule  et  le  corselet  sont  grands,  bien  delimites  par  deux 
carenes  plus  accuses  que  celles  qui  existent  chez  le  Mactra  pusilla; 
les  crochets  sont  plus  fort  que  ceux  du  Mactra  pusilla;  cette 
derniere  est  tres  deprim^e,  beaucoup  plus  transverse;  les  cotes  qui 
ornent  la  lunule  et  le  corselet  sont  identiques  dans  les  deux 
especes. 

"  La  M.  abbreviata  mesure  en  largeur  33  a  36  millimetres;  en 
hauteur  29  a  30  et  en  epaisseur  20  a  24  millimetres.  Elle  se 
rapproche  assez  de  la  Mactra  pura,  Deshayes,  d'Australie,  mais 
cette  derniere  egalement  blanche  comme  I'abbreviata,  est  plus 
grande,  plus  deprimee,  a  lunule  et  a  corselet  depourveux  de  sillons." 

Seeing  that  the  unknown  J/,  abbreviata  proved  not  to  be  M. 
pusilla,  I  obtained,  through  the  courtesy  of  Dr.  J.  C.  Verco,  a 

*  Lamarck,  An.  s.  Vert,  v.,  1818,  p.  477. 


598  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

series  of  the  shell  known  to  South  Australian  conchologists  as  M. 
p^ii'a,  and  transmitted  them  to  Mr.  J.  Mabille  with  a  request  that 
they  be  compared  with  Mactra  ahhreviata,  Lamarck. 

Mr.  Mabille  replied  (30/x./02)  : — "  Les  coquilles  que  vous 
m'avez  envoyes  ont  des  crochets  moins  inclines  en  avant  que  ceux 
du  type :  par  leur  reunion  le  bord  des  valves  dans  toutel'etendue 
de  la  lunule  et  du  corselet  present  chez  le  type  de  Lamarck  une 
saillie  tres  accusee  dans  I'une  des  exemplaires,  moins  dans  I'autre: 
cette  saillie  manque  dans  vos  specimens;  enfin  le  caractere  le  plus 
serieux,  celiii  par  lequel  Lamarck  a  suffisamment  caracterise  son 
espece  (ano  vulvaque  eleganter  plicatis)  sont  presque  absent,  au 
moins  sur  le  corselet.  Dans  le  type  la  lunule  et  le  corselet  sont 
ornes  de  plis  ou  pour  mieux  dire  de  lamelles  tres  reguli^res,  egale- 
ment  espacees,  partant  des  crochets,  couvrant  entierement  ces 
deux  parties,  surdirigeant  sur  le  centre  des  valves  en  s'afFaiblissant 
au  sortir  de  la  lunule  et  du  corselet,  de  telle  sorte  que  sur  la  valve 
meme  elles  ne  forment  que  des  stries  d'accroissement. 

"  Les  exemplaires  marques  de  la  lettre  B  [other  individuals  of 
my  parcel. — C.H.]  sont  trop  transverses^  trop  deprimes  pour 
pouvoir  se  rapporter  a  I'espece  de  Lamarck,  au  moins  dans  I'etat 
actuel  de  nos  connaissances.  Les  jeunes  coquilles  en  raison  de  leur 
depression  se  rapportent  certainement  a  la  forme  B. 

"  Ces  different  exemplaires  de  Mactra  ne  constitnent-ils  que 
des  varietes  de  la  Mactra  ahhreviata,  c'est  possible  mais  j'en  doute 
fort,  je  crois  plutot  a  une  espece  voisine  de  I'abbreviata,  carparmi 
tous  les  exemplaires  communiques,  je  n'en  vois  pas  un  seul  qui 
presente  les  caracteres  distinctifs  de  I'espece  en  question,  a  savoir 
la  presence  des  plis  ornant  la  lunule  et  le  corselet  de  la  M. 
abbreviata,  ce  qui  ne  permet  pas  de  trouver  une  forme  inter- 
mediaire  reliant  entre  elles  ces  diverses  Mactres." 

My  own  conclusions  from  the  information  so  kindly  imparted 
by  Mr.  Mabille  are  as  follows  : — Firstly,  the  record  of  Mactra 
ahhreviata  from  Port  Jackson  had  best  be  regarded  as  an  error 
and  the  species  expunged  from  the  fauna  of  N.  S.  Wales. 
Secondly,  the  identification   of    a  South  Australian  bivalve  by 


BY    C.    HEDLEY.  599 


ra 


Tate  as  Jlactra  picra,  Deshayes,"^  is  erroneous.  The  real  Mad 
pura,oi  which  I  gathered  an  example  on  the  beach  at  Green  Island, 
off  Cairns,  Queensland,  is  a  larger  and  flatter  species,  with  smooth 
dorsal  areas.  Thirdly,  after  comparing  examples  of  the  so-called 
Mactra  jnira  from  South  Australia  with  the  types  of  Mactra 
ahbreviata,  Lamk.,  Mr.  Mabille  finds  that  the}^  differ  in  minor 
characters  and  especially  by  the  want  of  lamellae  on  the  lunule 
and  escutcheon.  He  considers  that  the}^  may  be  a  variety  of 
Lamarck's  J/,  ahbreviata.  but  prefers  to  regard  them  as  a  related 
but  distinct  species. 

Under  these  circumstances  I  refrain  from  describing  it  as  a 
new  species,  but  as  a  temporary  expedient  will  call  it  M. 
ahbreviata  var.  Since  the  shell  has  never  been  figured  I  submit 
drawings  of  the  hinge  and  exterior  of  a  specimen  40  mm.  in 
length,  33  mm.  in  height,  and  23  mm.  in  breadth  of  conjoined 
valves.  It  appears  to  be  related  to  Mactra  rvfescens,  Lamk. 
Collectors  more  fortunate  than  myself  who  have  access  to  the 
species  are  invited  to  ascertain  whether  the  range  of  variation 
includes  Lamarck's  type  and  to  search  Peron's  collecting  grounds, 
especially  King  Island,  for  the  type  form. 

Purpura  pseudamygdala,  n.sp. 

Buccinam  amy(jdala,  Reeve,  Conch.  Icon.,  1846,  PI.  viii.,  sp.  60. 

Purpura  (Cronia)  amygdala,  Angas,  Proc.  Zool.  Soc.  1867, 
p.  191;  Brazier,  Proc.  Linn.  Soc.  N.S.W.  i.,  1876,  p.  200;  Smith, 
Alert  Coll.  1884,  p.  51  ;  Watson,  Rep.  Zool.  Chall.  xv.,  1886, 
p.  172;  Melvill  &  Standen,  Journ.  Linn.  Soc.  Zool.  xxvii.,  1899, 
p.  163.  (Not  Purpura  amygdala,  Kiener,  Coq.  Viv.,  Purpura,  1836, 
p.  39,  pi.  x.,  f.  26;  and  Chenu,  Man.  de  Conch,  i.,  1859,  p.  167, 
f.  807). 

(Plate  xxix.,  figs.  4,  5.) 

Shell  ovate-conic,  angled  at  the  shoulder,  concave  below  the 
suture,  solid,  everywhere  roughened  by  small  scales.  Whorls 
about  six.      Colour  pale  brown,  with  four  spiral  buff  bands  upon 


Tate,  Trans.  Roy.  Soc.  S.A.,  ix.,  1887,  p. 


600  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

the  prominent  lyne,  light  orange  within  the  aperture.  Sculpture : 
longitudinal  ribs,  about  ten  to  a  whorl,  mount  the  spire  obliquely, 
are  comparatively  stronger  on  the  earlier  whorls,  undulate  the 
suture,  attain  sudden  prominence  at  the  shoulder  and  vanish 
towards  the  base.  Crossing  these,  four  prominent  lyrse,  two  of 
which  ascend  the  spire,  are  produced  into  tubercles.  Between 
the  lyrae  are  fine,  densely  squamose  threads.  An  elevated  fasciolei 
winds  round  the  canal.  Protoconch  of  the  sinusigera  type,  broad 
with  rounded  whorls,  brown,  smooth  and  glossy.  Aperture  ovate, 
the  columella  expanding  laterally  to  a  free  edge  and  spreading 
above  into  a  broad  sheet  of  callus.  Inner  lip  with  a  few  den- 
ticules,  variable  in  number  and  development.  Length  33,  breadth 
16  mm. 

Hah. — The  species  ranges  from  Sydney  to  Torres  Straits;  the 
adult  shell  here  figured  is  from  Sydne}^  Harbour;  the  larva  (fig.  5) 
was  collected  at  Caloundra,  Queensland,  by  H.  L   Kesteven. 

Type. — To  be  presented  to  the  Australian  Museum. 

On  a  priori  grounds  it  is  improbable  that  Kiener  knew  the 
shell  here  described  as  P.  pseudamt/gdah',  for  he  was  unacquainted 
with  the  Queensland  fauna,  but  was  well  provided  with  material 
from  North-west  Australia.  Kiener  did  not  know^  the  locality  of 
his  P.  amygdala:  his  figure  and  description  do  not  suit  the  East 
coast  shell  with  which  the  London  writers  have  identified  it.  On 
the  contrary,  Kiener's  account  agrees  well  with  a  West  Australian 
species  related  to,  jjossibl}^  even  a  variety  of,  Buccnuwi  avellana, 
Reeve  (Conch.  Icon,  iii.,  Buccinum,  1846,  PL  viii.,  sp  52),  and 
which  has  been  obtained  by  Mr.  A.  U.  Henn  at  Geraldton.  W.A. 
Purpura  amygdala,  as  I  recognise  it,  is  easily  distinguished  from 
the  species  above  described  by  wanting  the  rough  scabrous 
sculpture,  by  being  shorter  and  relativel}'  broader,  and  liy  having 
fewer  and  stouter  lyr?e  within  the  lip. 

Capulus  austhalis,  Lamarck. 

Patella  australis,  Lamarck,  An.  s.  Vert,  vi.,  1^19,  pt.  i.,  p.  335; 
Delessert,  Recueil,  1841,  pi,  xxiii.,  f.  ll-a,  6,  c.  (Not  Hipponyx 
australis  of  Quoy  ct  Gaimard,  and  subsequent  authors). 


BY    C.    HEDLEY.  601 

Confusion  early  enveloped  this  species.  Deshayes  was  the  first 
to  transfer  it  from  Patella  to  Hifponyx.'*'  Under  the  name  of 
Hipponyx  aust7'alis,  Quoy  &,  Gaimard  described  an  Australian 
shell  for  which  they  say  they  kept  Lamarck's  name  but  amended 
his  description  to  agree  with  their  specimens,! 

It  has  been  remarked  that  Lamarck's  description  differs  in 
important  details  from  that  of  Quoy  &  Gaimard.  Menke 
endeavoured  to  harmonise  the  discrejDancy  by  supposing  that 
Lamarck's  characters  '■^  testa  tenui,  seinipellucida,"  applied  to  the 
young  shell,  and  that  Quoy  &  Gaimard's  phrase  " cras.^a "  properly 
described  the  adult.;  As  the  size  (one  inch)  specified  by  the 
earlier  writer  equals  that  given  by  the  later  authors,  Menke's 
explanation  is  untenable. 

To  avoid  doulbt  several  writers  have  ignored  Lamarck  and 
have  quoted  the  species  as  of  Deshayes,  or  of  Quoy  ife  Gaimard. 
Angas  considered  that  the  species  illustrated  in  the  '  Voyage  of 
the  Astrolabe '  is  probably  identical  with  the  earlier  Amalthea 
Gonica,  Schumacher.§  Tryon  suggests  that  Patella  cassida, 
Dillwyn,||  embraces  both,  which  is  likely  to  be  the  correct  view. 

Granting  the  argument  that  Lamarck  dealt  with  one  shell  and 
Quoy  &  Gaimard  with  another,  then  to  what  species  should 
Lamarck's  name  apply  1  The  answer  I  would  give  is  that  what 
Delessert  figured  and  Lamarck  described,  is  the  shell  now  wrongly 
known  to  Australian  conchologists  as  Capidus  danieli.  The  mis- 
appropriation of  this  name  to  the  Australian  shell  is  due  to 
Angas.  IT 

Capidus  danieli,  Crosse,  was  described  from  New  Caledonia, 
and  it  is  improbable  that  it  should  also  occur  in  South  Australia. 
Watson,  who  examined   the  type  of    C.  australis  yet  failed  to 

*  Deshayes,  Encycl.  M(^th.  Vers,  ii.,  IS.SO,  p.  274. 

t  Quoy  &  Gaimard,  Voy.  Astrolabe,  iii.,  p.  434,  PI.  Ixxii.,  f.  25-34. 

Z  Menke,  Zeit.  f.  Malak.,  1844,  p.  61. 

§  Angas,  Proc.  Zool.  Soc.  1865,  p.  175. 

Ij  Dillwyn,  Descr.  Cat.  Rec,  Shells,  ii.,  1817,  p.  1037,  based  on  Martini, 

Conch.  Cab.  i.,  pi.  xii.,  f.  116. 

^  Angas.  Proc.  Zool.  Soc.  1865,  p.  175. 


602  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

comprehend  that  two  species  and  genera  were  involved  in  his 
synonomy,  remarked  that  the  apices  of  the  two  species  turn  in 
opposite  directions,*  an  observation  which  I  have  verified.  Dall 
remarks  "that  Hipponyx  danieli,Qvo^%e,  is  ahnost  certainly  Pa^e/^a 
calyptra  of  Martyn,  described  in  the  last  century.  A  magnificent 
specimen  from  the  Fiji  Islands  is  in  the  National  Museum.  It  is 
a  Capulus  and  not  a  Hipponyx.^' j 

Serpulus  sipho,  Lamarck. 

This  common  and  widely  distributed  Australian  shell  was 
figured  from  King  George's  Sound,  W.A.,  by  Quoy  &l  Gaimard 
in  the  '  Voyage  of  the  Astrolabe,'  under  the  name  of  Vermetus 
arenarius,  Lamarck.  Tracing  back  the  citation  to  Lamarck  we 
find  that  he  in  turn  based  his  account  on  a  species  described  by 
Linne  in  the  Systema  ISTatura  (xii.,  1758,  p.  1266).  According  to 
Hanleyi  the  original  specimen  used  by  Linne,  still  preserved  in 
the  Museum  of  the  Linnean  Society  of  London,  is  the  Mediter- 
ranean species  otherwise  called  Vermetus  gigas,  Bivona.§ 

Consequently  the  name  which  Quoy  &  Gaimard  attach  to  their 
drawing  must  be  rejected.  Tate  ct  May,  to  cite  the  latest  state- 
ment, replace  it  by  "  Thylacodes  sulcatus,  Lamk." 

But  Vaillant,||  after  examining  the  types  of  Lamarck  andi 
of  Quo}^  &  Gaimard  in  the  Paris  Museum,  remarks  that  the  namej 
of  Serpula  sulcata,  Lamk.,  covered  tw^o  species;  the  first  a  fossil 
from  Touraine,  the  second  a  recent  Australian  shell.  He  restricts  i 
the  name  S.  sulcata  to  the  former,  and  unites  the  latter  with' 
Vermetus  arenarius,  Q.  &  G.  (not  Lamk.),  and  V.  dentiferus^\ 
Q.  &  G.  (not  Lamk.),  to  Serpula  sipho,  Lamk.  (Anim.  s.  Vert. 
1818,  p.  626),  under  the  name  of   Serpulorhis  sipho.     The  genusl 


*  WatsoH,  Chall.  Exped.  Zool.  xv.,  p.  457- 

t  Dall,  Bull.  Mus.  Comp.  Zool.  Harv.  Coll.  xviii.,  18S9,  p.  287. 

X  Hanley,  Ips.  Linn.  Conch.  1855,  p.  447. 

§  Therefore  the  shell  discussed  in  these  Proceedings,  (2  ,  Vol.  ix.,  p.  465,  agj 

Ktiphus  arenarius  should  take  the  name  of  Kuplins  polythalamins,  Linn. 

II  Vaillant,  Nouv.  Archiv.  Mus.  vii.,  1871,  pp.  193,  197. 


BY    C.    HEDLEY.  603 

Serpulorhis,  Sassi  (1827),  is,  however,  of  later  date  thsiU  Serpulus 
proposed  by  Moiitfort  (Conch.  Syst.  ii.,  1810,  p.  27). 

The  species  under  discussion  has  been  entered  in  local  lists 
under  the  name  of  Thylacodes  decussaUis,  Gmel.  (Angas,  Proc. 
Zool.  Soc.  1867,  p.  211). 

ASSIMINEA    PACiODELLA,   n.sp. 

(Plate  xxix.,  fig.  6.) 

Shell  perforate,  ovate,  rather  solid.  Colour  reddish-brown. 
Whorls  four,  rapidly  increasing,  wound  obliquely,  spire  pagodi- 
f orm.  The  straight  slope  of  the  upper  part  of  the  last  two  whorls 
ends  in  a  sharp  projecting  rim,  beneath  which  the  perpendicular 
periphery  bears  several  small  spiral  threads  diminishing  succes- 
sively. Base  rounded.  Umbilicus  a  narrow  oblique  perforation. 
Aperture  ovate,  oblique,  columella  expanded  and  reflected,  outer 
lip  simple.     Height  1*96  mm.,  breadth  1*2  mm. 

Hab. — Manly  Lagoon,  in  brackish  water  (H.  L.  Kesteven). 

Ty  p  e  to  be  presented  to  the  Australian  Museum. 

Cecum  lilianum,  n.sp. 
(Plate  xxix.,  fig.  7.) 

Shell  curved,  tapering,  pale  brown,  with  about  twenty  broad, 
projecting,  regular,  rounded,  transverse  rings  about  their  own 
breadth  apart.  Lip  formed  of  a  ring  more  massive  than  the  rest. 
Septum  flat.  Length  2-32,  breadth  at  aperture  0*44,  at  truncated 
end  0-28  mm. 

ffab.  —Long  Bay,  near  Sydney;  one  specimen,  in  shell  sand; 
collected  by  Miss  L.  Parkes. 

T  y  p  e.  —  Miss  Lily  Parkes  has  generously  presented  the  unique 
specimen  to  the  Australian  Museum. 

This  specimen  resembles  C.  vertebrale,  Hedley,  from  the  Ellice 
Islands,  than  which  it  is  narrower,  more  curved  and  more  tapering 

Hydatina  tasmanica,  Beddome. 

This  species  was  figured  in  Part  iii.  of  these  Studies.  I  am 
now  able  to  add  it  to  the  fauna  of    New  South  Wales,  having 


604  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

found  a  specimen  (15/vi./02)  in  a  stranded  bottle  on  the  rocks 
at  Middle  Head. 

Cylindrobulla  fischeri,  a   Ad.  &  Ang. 

C ylindrohiilla  fischeri^  A.  Ad.  &  Ang.,  Proc.  Zool.  Soc  1864, 
p.  37;  Angas,  op.  cit.,  1871,  p.  98. 

(Plate  xxix.,  tigs.  8,  9.) 

This  species  has  never  been  tigured.  My  illustration  is  based 
on  authentic  material  obtained  by  Brazier  in  the  Lane  Cove. 

Helix  subangulata,  Pfeiffer. 

Those  interested  in  the  land  shells  of  Tasmania  have  for  long 
been  troubled  by  two  large  and  conspicuous  land  shells  falsely 
ascribed  to  their  fauna  by  Pfeiffer.^  These  are  Helix  bi sulcata  and 
H.  subangulata.  The  former  of  these,  as  Mr.  Brazier  has  sug- 
gested to  me,  is  evidently  Rhytida  be r audi,  Gassies.f  from  JST.E. 
New  Caledonia. 

To  the  kindness  of  Mr.  J.  Ponsonby,  of  London,  lam  indebted 
for  a  solution  of  the  second  puzzle.  At  my  request  he  and  Mr. 
E.  A.  Smith  searched  for  Pfeiffer's  type  of  H.  subangulata  and 
succeeded  in  identifying  it  as  the  3^ouug  of  Flanispira  zonalis, 
Ferussac,  from  Halmaheira.  Mr.  Smith  regards  this  determina- 
tion as  beyond  doubt.  This  Moluccan  species  can,  therefore,  be 
erased  from  future  Tasmanian  lists. 

Endodonta  melbournensis,  Cox. 

Helix  melbournensis,  Cox,  Mon.  Austr.  Land  Shells,  1868,  p.  22, 

Pl.xii,  f.  10. 

(Plate  xxxi.,  figs    16,  17.) 

As  the  figure  quoted  is  inexact,  I  supph'  another  from  authentic 
material  kindly  lent  by  Dr.  Cox.  The  specimen  drawn  is  6  mm. 
major  diam.,  5-1  mm.  minor  diam.,  height  3-3  mm. 


*  Legrand,  Journ.  of  Conch,  ii.,  1879,  p.  95. 
t  Helix  beraudi,  Gassies,  Journ.  de  Conch,  vii.,  1858,  p.  68. 


BY    C.    HEDLEY.  605 

Endodonta  subdepressa,  Brazier. 

Helix  subdepressa,  Brazier,  Proc.  Zool.  Soc.  1871,  p.  641. 
Helix  dandenongensis,  Petterd,  Journ.  of  Conch,  ii.,  1879,  p.  356; 
Tate,  Proc.  Roy.  Soc.  S.A.  iv.,  1882,  p.  75. 

(Plate  xxxi.,  figs.  13,  14,  15.) 

This  species  has  not  yet  been  illustrated.  The  specimen  drawn 
is  part  of  Brazier's  original  series;  it  is  4*1  mm.  in  major  diameter, 
3*3  mm.  in  minor  diameter,  and  1-8  mm.  in  height.  Prof.  Tate  is 
responsible  for  the  above  synonomy. 

Endodonta  otwayensis,  Petterd. 

Helix  otwayensis,  Petterd,  Mon,  Land  Shells  Tas.,  April,  1879, 
p.  39;  Journ.  of  Conch,  ii.  Dec.  1879,  p.  356;  Johnston,  Proc.  Roy. 
Soc.  Tas.  1879  (1880),  p.  24. 

(Plate  xxxi.,  figs.  10,  11,  12.) 

The  specimen  drawn  was  kindly  identified  for  me  by  Mr.  W. 
F.  Petterd;  it  measures  2*3  mm.  in  major  diameter,  2  mm.  in  minor 
diameter,  and  1  -5  mm.  in  height.  It  was  presented  to  me  by 
Mr.  T.  S.  Hall,  who  collected  it  at  Fern  Tree  Gully,  Victoria. 

Endodonta  tamarensis,  Petterd. 

Helix  tamarensis,  Petterd,  Mon.  Land  Shells  Tas.,  April,  1879, 
p.  30.  Endodonta  tamarensis,  Hedley,  Records  Aus.  Mus.  ii., 
1896,  p.  104.  Helix  rosacea,  Petterd,  Journ.  of  Conch,  ii.,  July, 
1879,  p.  213  (not  Helix  rosacea,  Muller,  1774). 

(Plate  xxxi.,  figs.  18,  19,  20.) 

The  present  is  the  first  illustration  presented  of  this  species;  it 
is  derived  from  an  authentic  specimen  3*6  mm.  in  major  diameter, 
3  mm.  in  minor  diameter,  and  2  mm,  in  height. 

The  drawings  of   this  and  the  last  three  species  were  made 

about  seven  years  ago,  but  their  publication  has  been  delayed  by 

constant  pressure  of  work.     I  owe  apologies  for  their  tardy  issue 

to  many  friends  who  have  kindly  contributed  material  for  study. 

40 


606  studies  on  australian  mollusca, 

The  Thiphop.id.e  of  New  South  Wales. 

Investigation  proves  that  the  well  known  name  of  Triforis  can 
no  longer  be  used  at  the  current  valuation  If  maintained  at  all, 
it  must  be  restricted  to  a  minor  group  typified  by  the  eocene 
fossil,  T.  plicata,  Deshayes. 

It  has  been  generally  understood  (for  example,  by  Fischer, 
Man.  de  Conch.,  1884,  p.  678)  that  the  genus  in  question  was 
first  proposed  in  1824  as  Triforis  by  Deshayes  in  '  Descrip. 
Coquilles  Fossiles  des  Environs  de  Paris '  (ii.  p.  429).  Though 
the  title-page  of  this  volume  is  dated  "1824,"  the  notice  in  ques- 
tion did  not  appear  till  much  later.  Thus  Menke,  writing  in 
1830,  knew  nothing  of  "  Triforis,''^  and  failed  to  include  the  name 
in  his  '  Synopsis  Methodica  Molluscorum.'  Xewton  states*  that 
pages  427  to  498  of  Deshayes'  work  were  issued  in  1834.  This, 
then,  is  the  date  to  be  assigned  to  '^Triforis"  of  Deshayes. 
There  are  vague  hints  in  literature  that  Basterot  referred  to 
^^Tri/oris"  in  1825.  A  paper  by  Basterot  is  contained  in  the  first 
volume  of  the  Memoires  Soc.  Hist.  Nat.  Paris  (1825,  pp.  1-100). 
This  I  have  been  unable  to  consult,  but  since  no  one  has  claimed 
that  Basterot  established  the  genus,  I  shall  presume  that  the 
reference  was  a  mere  allusion,  not  of  s3'stematic  importance. 

Blainville  mentioned  the  genus  (Man.  de  Malac.  i.,  1825,  p.  404) 
as  a  division  of  Cerithium  under  the  French  popular  names 
"  Triphore  ou  Tristome."  Certain  authors  have  wrongly  received 
this  as  a  Latin  generic  name  Tristoma,  which,  however,  would 
fall  before  Cuvier's  Tristoma  (1817). 

Later  Blainville  regularly  characterised  the  genus  under  the 
name  Triphora  (Diet.  Sci.  Nat.  Iv.,  1828,  p.  344)  with  T.  gemma- 
turn,  Blainville  (  =  T.  tristoma,  Blainville)  as  type.  This  name 
Triphora  was  noted  in  the  following  year  by  Menke,  and  in  the 
year  after  by  Rang. 

Later  Deshayes  published,  in  1832,  the  name  Trlphoris  (Encyc. 
Meth.  ii.,  p.  1053).  Finally,  as  alread}^  stated,  Deshaj^es  described 
Triforis  in  1834. 

*  K.  B.  Newton,  Brit.  Mus.  Cat.  Eocene  Moll.,  1891,  p.  309. 


BY    C     HEDLEY.  607 

It  seems  clear  that  Triphoro.,  Blainville  (1828),  demands 
recognition  as  the  prior  genus  of  the  family.  Certain  reconstruc- 
tion follows  this  deduction.  The  type  of  Triphora,  T.  tristoma, 
is  mentioned  by  Hervier  (Journ.  de  Conch.,  xlv.  1898,  p.  289)  as 
belonging  to  Inella,  Bayle  ( =  Ino,  Hinds).  If  this  be  right, 
Tnella  should  disappear  as  a  synonym  of  Triphora.  I  should, 
however,  have  regarded  T.  tristoma  as  referable  to  Jousseaume's 
genus  Mastoni(mtrmis  (Bull.  Soc.  Malac.  France,  i.,  1884,  p.  239). 

The  family  Triphoridae  is  divided  by  Jousseaume  into  eleven 
genera.  Probabl}-  none  of  the  species  I  record  from  this  State 
fall  into  Triphora,  sensu  stricto.  Though  some,  if  not  all,  of 
Jousseaume's  genera  are  natural,  they  are  so  vaguely  defined  that 
I  prefer  to  wait  before  allotting  the  Australian  species.  Probably 
at  least  a  hundred  species  exist  in  Australian  seas,  and  a  better 
knowledge  of  these  is  desirable  before  framing  a  classification, 
especially  as  opinions  are  divided  as  to  whether  these  eleven 
sections  are  of  generic  or  subgeneric  value. 

Passing  from  the  consideration  of  genera  to  that  of  species,  a 
far  larger  number  of  Triphora  are  known  to  Australian  collectors 
than  are  named  in  literature.  Some  of  these  might  already 
have  been  dealt  with,  but  a  necessary  preliminary  to  the  descrip- 
tion of  new  species  is  the  distinction  of  old  species,  and  here  all 
workers  have  encountered  an  insuperable  obstacle. 

Several  new  Australian  Triphora  unluckil}'-  fell  into  the  incom- 
petent hands  of  Arthur  Adams,  who,  in  naming  them,  deliberately 
omitted  all  measurements,  neglected  to  figure  species,  and  gave 
scanty  descriptions.  Probably  he  never  used  a  microscope,  and 
Smith  has  observed  that  he  was  colour  blind.*  In  conchology 
his  work  is  of  the  same  grade  as  that  of  his  contemporary  Francis 
Walker  in  entomology. 

Though  enjoying  a  monopoly  of  the  types,  London  writers  have 
made  no  effort  to  rehabilitate  these  species.  In  monographing 
"  Triforis,^^  Tryon  simply  excluded  these  species  of  Adams  as 
unintelligible.     To  ignore  names  which  any  day  may  be  resur- 

*  Smith,  Chall.  Rep.  Zool.  xiii.,  1885,  p.  61. 


608  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

rected  by  those  having  access  to  the  types,  or  to  go  by  guess,  is 
the  dilemma  of  the  student  of  our  Triphora.  Electing  the  latter 
course,  I  note  that  some  of  Adams'  species  were  reported  with 
corroborative  detail  as  from  Sydne3^  Assuming  that  these 
species  are  contained  in  the  large  collection  before  me,  I  have 
applied  his  names  to  the  species  which  best  fit  the  descriptions. 

The  material  dealt  with  was  gathered  around  Sydney  by  the 
late  Mrs.  Starkey,  Messrs.  J.  Brazier,  H.  L.  Kesteven  and  mj^self. 

I  have  to  gratefullj^  acknowledge  kind  assistance  from  Dr.  J. 
C.  Yerco,  Mr.  A.  Morton  and  the  Rev.  H.  D.  Atkinson. 

This  opportunity  is  taken  of  adding  that  the  shell  I  described 
from  Funafuti  as  Triforis  torquatus,  Hedley,  is  probably  identical 
with  the  prior  T.  ustidatus,  Hervier,  (Journ.  de  Conch,  xlvi.,  1898, 
p.  298,  pi.  xvii.,  f.  4),  whose  memoir  did  not  reach  me  till  my 
account  had  appeared. 

Triphora  regina,  n.sp. 
(Plate  xxxii.,  fig.  21.) 

Shell  narrowly  conical,  solid.  Colour  white,  a  narrow  orange 
line  along  the  lowest  gemmule  row  of  each  whorl  and  an  orange 
tip  to  the  canal.  Whorls  about  twelve.  Sculpture  :  on  the  base 
are  three  simple  keels,  on  the  last  whorl  three  equal  sized 
gemmule  rows,  of  which  the  median  diminishes  on  the  penulti- 
mate and  gradually  fades  away  as  it  ascends  the  spire.  Length 
5  mm.,  breadth  1*5  mm. 

Hah. — Balmoral  Beach,  Middle  Harbour;  one  specimen  (self). 

Type. — To  be  presented  to  the  Australian  Museum. 

I  venture  to  describe  this  species  from  a  single  specimen, 
mutilated  at  each  extremity,  because  the  orange  thread  on  each 
whorl  will  render  possible  the  recognition  of  any  fragment.  The 
contour  and  sculpture  are  also  sufficiently  distinct  from  any 
species  known  from  this  coast. 

Triphora  innotabilis,  n.sp. 
(Plate  xxxii.,  figs.  23,  24,  25.) 
Shell  small,  slender  and  very  narrow.     Colour  brown,  gem  mules 
dull  white.     Whorls  ten,  plus  a  five-whorled  protoconch.     Sculp- 


BY    C.    HEDLEV.  609 

ture  :  on  the  base  are  two  plain  spiral  keels;  the  latter  whorls 
with  three  gemmule  rows,  the  median  being  much  the  smaller;  as 
it  mounts  the  spire  it  diminishes,  fading  out  about  half  way  up. 
The  gemmules  are  about  18  to  a  whorl,  large,  prominent,  and  set 
about  half  their  diameter  apart.  A  raised  thread  winds  along 
the  suture.  First  whorl  of  the  protoconch  smooth,  the  rest  with 
a  central  keel  and  transverse  bars.  Aperture  subquadrate,  and 
notch  simple,  spur  of  the  lip  crossing  the  pillar.  Canal  short, 
straight.     Length  4*8;  breadth  1-4  mm. 

Hab. — Sydney  Harbour;  one  specimen  (H.  L.  Kesteven). 

Type. — To  be  presented  to  the  Australian  Museum. 

Distinguishing  features  of  this  species  are  its  small  and  slender 
form,  median  row  of  small  gemmules  between  two  rows  of  large 
ones  and  the  five-whorled  protoconch.  A  second  specimen  from 
Mrs.  Starkey's  collection,  of  broader  proportions,  is  also  figured 
(fig.  25). 

Triphora  albovittata,  n.sp. 

(Plate  xxxii.,  figs.  26,  27.) 

Shell  conical,  moderately  broad.  Whorls  nine,  plus  a  four- 
whorled  protoconch,  parted  by  deep  sutures.  Colour  pale  yellow; 
upper  row  of  gemmules  on  each  whorl  white,  beneath  them  a 
narrow  line  of  ochre,  base  and  protoconch  ochreous.  Sculpture  : 
on  the  base  are  two  plain  spiral  keels,  above  which  are  four  rows 
of  gemmules;  the  four  preceding  whorls  have  three  rows  each, 
and  those  above  again  two  each.  The  gemmules  are  set  their 
own  diameter  apart,  about  twenty  to  a  whorl,  joined  by  a  string 
to  their  fellows  in  the  same  and  their  neighbour  in  the  next  row. 
Protoconch  :  first  and  second  whorls  smooth,  third  and  fourth 
with  a  median  keel  and  transverse  bars.  Aperture  imperfect  in 
the  examples  seen.     Length  4-8;  breadth  1-5  mm. 

Hah. — Balmoral  Beach;  in  shell  sand,  rare. 

Type. —To  be  presented  to  the  Australian  Museum. 

The  white  subsutural  gemmules  underlined  by  ochre,  aff'ord  a 
recognition  mark  for  this  species,  which  is  nearly  allied  to  T. 
gi^anifera,  Brazier,  and  may  indeed  ultimately  prove  only  a  variety 
of  that. 


610  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

Triphora  granifera,  Brazier. 

Triforis  graniferus^  Brazier,  Proc.  Linn.  Soc.  N.S.W.  (2)  ix., 
1894,  p.  173,  pl.xix.,  f.  10;  1  T.  fasciata,  Tate  &  May,  op.  cit., 
1901,  pi.  xxiii.,  ff.  10-11. 

(Plate  xxxiii.,  figs.  28,  29.) 

This  species  is  one  of  the  commonest  in  Sydney  Harbour,  and 
extends  to  Tasmania  and  South  Australia.  As  the  original 
description  was  rather  brief,  I  add  the  following  : — 

Shell  small,  regularly  conical.  Whorls  eight,  plus  a  four- 
whorled  protoconch,  parted  by  deep  sutures.  Colour  light  brown, 
upper  gemmule  row  paler,  base  and  sutures  chocolate,  lip  dull 
white.  Sculpture  :  on  the  base  two  plain  spiral  keels,  on  the 
periphery  a  keel  with  incipient  gemmules  and  above  three  rows 
of  closely  packed  gemmules,  set  about  25  to  a  row,  less  than  their 
own  breadth  apart.  On  the  lower  whorls  the  gemmules  of  every 
row  are  equal  in  size;  above  the  median  row  has  smaller  gemmules. 
A  nodose  thread  runs  along  the  sutures.  Aperture  subquadrate, 
spur  of  the  lip  not  reaching  the  pillar;  anal  notch  simple,  shallow; 
canal  short  and  straight.     Length  4-45;  breadth  Lo  mm. 

Triphoka  angasi,  Crosse  k  Fischer. 

Tripho7'is  angasi,  Crosse  k  Fischer,  Journ.  de  Conch,  xiii.  1865, 
p.  46,  pi.  i.,  ff.  12-13. 

Shell  of  moderate  size,  tapering  slowl)',  glossy,  solid.  Colour 
buff,  with  a  chocolate  base  and  sutural  band,  gemmules  white. 
Whorls  about  thirteen,  parted  by  distinct  sutures.  Sculpture  : 
on  the  earlier  adult  whorls  are  two  gemmule  rows,  on  the  seventh 
whorl  a  minute  intermediate  row  is  intercalated  which  gradually 
attains  the  size  of  the  other  rows.  The  additional  row  is  nearer 
to  the  upper  than  to  the  lower  row.  Along  the  suture  winds  a 
small  plain  thread,  which  on  the  last  whorl  appears  as  a  fourth 
row  of  gemmules.  The  base  has  one  plain  spiral  thread.  In  the 
immature  stage  the  species  has  a  flat  base.  The  gemmules  are 
set  their  own  breadth  apart,  about   22  to  a  whorl,  conspicuous 


BY    C.    HEDLEY.  611 

against  the  yellow  and  brown  interstices.  Aperture  subquadrate, 
anal  sinus  simple,  spur  of  lip  crossing  the  callus  of  the  pillar, 
canal  shorter  than  the  aperture.     Length  7;  breadth  2  mm. 

The  Sydney  shell  above  described  has  a  general  resemblance  to 
T.  cinerea,  but  is  more  cylindrical,  with  flatter  whorls  and  a 
different  colour-pattern.  The  gemmule  rows  appear  narrower 
and  sharper,  but  this  may  be  an  optical  illusion,  due  to  contrast 
of  colour.  No  specimens  examined  afforded  an  opportunity  for 
describing  the  protoconch. 

I  have  identified  Sydney  specimens  by  comparison  with 
examples  from  South  Australia,  the  type-locality,  kindly  furnished 
by  Dr.  J.  C.  Yerco.  Our  shells  are  considerably  larger  and  pro- 
portionately a  little  broader,  but  correspond  in  all  details  of 
sculpture  and  structure. 

Triphora  nigrofusca,  a.  Adams. 

(Plate  xxxiii.,  figs.  34,  35.) 

This  species  was  originally  described  (Proc.  Zool.  Soc.  1851, 
p.  278)  in  the  following  words  : — "  T.  testa  pyramidali,  nigro- 
fusca; anfractibus  planis,triseriatimgranulatis,  granulis  aequalibus, 
confertis,  anfractuum  suturis  impressis,  basi  convexa.  Sydney, 
low  water,  under  stones  (Mr.  Strange).  A  black-brown  species, 
with  three  rows  of  regular,  equal-sized  granules  on  each  whorl." 

To  this  Angas  added  (oi).  cit.  1867,  p.  208)  the  length — 4 
lines. 

This  species  has  not  again  been  recognised,  and  Tryon  includes 
it  in  his  list  of  rejected  species.  Among  ihe  shells  collected  by 
the  late  Mrs.  Starkey  in  Sydney  Harbour  is  a  single  specimen 
which  fairly  suits  the  brief  account  quoted  above,  and  which  I 
regard,  though  not  with  certainty,  as  this  species.  I  note  the 
following  characters  : — 

Shell  rather  long  and  narrow,  small,  apex  blunt,  aperture 
unusually  expanded.  Whorls  ten,  including  the  protoconch, 
constricted  at  the  sutures.  Colour  chocolate,  darker  at  the  base, 
gradually  paler  towards  the  apex.  Sculpture  :  on  the  base  are 
two  plain  spiral  ikeels,  the  upper  coiled  along  the  suture  in  pre- 


612  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

ceding  whorls;  three  equally  spaced  smooth  spiral  cords  ornament 
each  whorl,  broken  into  gemmules  by  a  series  of  longitudinal 
ridges;  the  interspaces  longitudinally  striated.  Protoconch  of 
three  whorls,  first  smooth,  second  and  third  bicarinate,  crossed 
by  numerous  fine  bars.  Aperture  subquadrate,  exserted,  anal 
notch  simple,  spur  almost  but  not  quite  reaching  the  pillar. 
Length  6;  breadth  1-8  mm. 

Among  other  faults,  Adams  gave  the  specific  names  of  Triforis 
a  masculine  termination.  As,  however,  that  generic  name  is 
compounded  from  the  Latin  feminine  substantive  foris,  the 
specific  name  should  have  taken  a  feminine  termination. 

Triphora  tasmanica,  T.  Woods. 

T.  Woods,  Proc.  Roy.  Soc.  Tas.,  1875  (1876),  p.  28. 

(Plate  xxxii.,  fig.  22.) 

In  general  appearance  this  species  is  like  T.  angasi  and  T. 
cinerea,  bearing  on  each  of  the  older  whorls  three  equal  gemmule 
rows.  There  is  a  shell  with  a  lilac  apex,  occurring  in  Tasmania 
and  South  Australia,  wdiich  appears  to  be  a  variety  of  this.  A 
useful  recognition  mark  is  the  brown  dot  between  each  gemmule 
of  the  lower  tier.  But  from  every  member  of  the  genus  T.  tas- 
manica is  distinguished  by  its  remarkable  apex — I  hesitate  to 
write  protoconch.  The  shell  commences  with  two  rows  of  elevated 
rounded  gemmules,  between  which  on  the  4-5th  whorl  a  median 
row  is  intercalated.  It  is  possible  that  a  caducous  larval  shell 
ma}^  have  preceded  the  stage  described. 

A  shell  of  six  whorls,  2*2  mm.  long,  dredged  by  Mr.  G.  H. 
Halligan  and  myself  in  100  fathoms,  16  miles  east  of  Wollongong, 
is  here  figured.  It  is  the  only  record  of  the  species  from  New 
South  Wales,  and  has  been  determined  by  a  careful  comparison 
with  the  type  specimen  kindly  lent  me  by  Mr.  A.  Morton. 

Triphora  cinerea,  n.sp. 

(Plate  xxxiii.,  tigs.  36,  37.) 

Shell  of  moderate  size,  narrow,  conical  with  straight  sides. 
Colour  :  base  dark  chocolate,  spire  marbled  with  ash  and  different 


BY    C.     HEDLEY.  613 

shades  of  brown,  or  brown  with  ash  gem  mules,  an  ashy  tint  pre- 
dominating. Whorls  ten,  plus  a  three-whorled  protoconch. 
Sculpture :  across  each  whorl  are  about  twenty-one  vertical 
rounded  ribs,  separated  by  equal  and  deep  interstices;  these  ribs 
cease  on  either  side  of  the  sutures,  giving  definition  to  the  whorls. 
Three  equal  and  evenly  spaced  spiral  cords  override  both  ribs 
and  interstices,  producing  gemmules  on  the  former  and  partition- 
ing the  latter  into  deep  square  pits.  An  extra  gemmule  row 
runs  along  the  basal  angle  of  the  last  whorl  and  appears  on  the 
spire  as  a  sutural  thread.  On  the  base  is  a  plain  spiral  cord. 
The  first  whorl  of  the  protoconch  is  white,  the  second  and  third 
coloured,  each  with  two  spiral  keels,  whose  interspaces  on  the 
second  and  third  are  latticed  by  wide-set  oblique  bars,  forming 
rhomboidal  meshes.  Aperture  subquadrate,  anal  notch  simple, 
spur  of  the  lip  crossing  the  pillar,  canal  produced,  straight. 
Length  8;  breadth  2  mm. 

Hah.  —  Middle  Harbour  beaches;  common  as  dead  shells  on  the 
sands. 

Type. — To  be  presented  to  the  Australian  Museum. 

Possibly  this  is  the  species  recorded  by  Angas"^"  as  Trijoris 
granulatus,  Ad.  k  Reeve,  from  Botany  Bay. 

Triphora  nocturna,  n.sp, 

(Plate  xxxii.,  figs.  30,  31.) 

Shell  large,  tall  and  slender,  conical.  Colour  uniform  dark 
purple-brown.  Whorls  fourteen,  plus  a  four-whorled  protoconch. 
Sculpture  :  on  the  base  are  three  plain  spiral  keels.  Last  whorl 
with  three  gemmule  rows,  the  uppermost  the  largest;  on  the 
penultimate  whorl  the  median  row  is  reduced  to  a  nodose  thread; 
above  it  disappears,  leaving  two  rows  to  mount  the  spire.  The 
whorls  are  longitudinally  fluted;  upon  the  folds  are  set  the 
gemmules,  about  20  to  a  whorl,  polished,  much  elevated,  distant 
from  each  other,  their  interstices  roughened  and  dulled  by  longi- 
tudinal wrinkles.      Protoconch  :  first  two  whorls  smooth,  third 

*  Angas,  Proc.  Zool.  Soc,  1871,  p.  91. 


614  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

and  fourth  with  a  median  keel  and  transverse  bars.  Aperture 
subquadrate,  spur  of  the  lip  crossing  the  pillar;  anal  notch  simple, 
rather  deep.     Canal  short,  straight.     Length  11;  breadth  3  mm. 

Hah. — Pearl  Bay,  Middle  Harbour;  alive  under  stones  at  low 
tide,  taken  by  myself. 

Type. — To  be  prosented  to  the  Australian  Museum. 

Among  local  species  this  is  distinguished  by  its  large  size, 
slender  shape  and  uniform  purple  colour. 

Triphora  MACULOSA,  n.sp. 
(Plate  xxxii.,  figs.  32,  33.) 

Shell  conical,  rather  broad  and  solid.  Whorls  ten,  parted  by 
distinct  and  deep  sutures.  Colour  :  irregularly  variegated  with 
opaque  white,  buff  and  chocolate,  usually  articulated  with  choco- 
late and  white  beneath  the  sutures,  base  chocolate  and  apex 
white.  Sculpture  :  the  base  has  two  plain  spiral  keels,  above 
which  are  four  rows  of  gemmules,  the  penultimate  whorl  and 
those  immediately  above  have  each  three  rows  of  gemmules;  on 
the  seventh  the  median  row  diminishes  and  on  the  whorls  above 
gradually  disappears,  leaving  two  rows  only  on  the  upper  whorls. 
The  gemmules  are  round  hemispherical  bosses,  set  their  own 
diameter  apart,  about  25  to  a  whorl,  linked  by  a  string  to  their 
fellows  in  the  same  and  their  neighbours  in  the  next  row.  Apex 
rather  styliform,  the  second  whorl  has  one  keel,  the  third  two, 
after  which  the  keels  develop  gemmules  and  assume  the  adult 
sculpture.  Aperture  perpendicular,  subquadrate,  anal  notch  pro- 
duced on  the  right  into  a  spur  which  crosses  the  pillar;  anal 
notch  simple,  shallow.  Canal  short  and  straight.  Length  6*7; 
breadth  2*1 4mm. 

Hah. — Balmoral  Beach,  Middle  Harbour,  Sydney;  common  as 
dead  shells  on  the  beach. 

Type. — To  be  presented  to  the  Australian  Museum. 

Arthur  Adams,  the  evil  genius  of  Australian  conchology, 
applied  the  MS.  name  of  Triphoris  maculosus  to  a  Sydney  species. 
An  unfortunate  entry  of  this  name  in  the  Catalogue  of  Angas 
(P.Z.S.  1877,  p.  184)  is  the  sole  record  concerning  it.     As  local 


BY    C.    HEDLEY.  615 

tradition  has  tlius  identified  it,  and  as  the  adjective  maculosvs 
suits  it,  I  conclude  that  it  was  to  the  species  above  described  that 
Adams  gave  the  name.  But  I  beg  to  call  attention  to  the  fact 
that  I  am  not  naming  the  shell  2\  maculosa  (A.  Ad.,  MS.),  that 
being,  as  E.  R.  Sykes  remarks,  a  complicated  and  cumbersome 
process  of  nomenclature. 

Triphora  AxMpulla,  n.sp. 

(Plate  xxxiii.,  figs.  38,  39.) 

Shell  small,  narrowly  conical,  acuminate.  Whorls  nine,  plus  a 
four-whorled  protoconch,  parted  by  distinct  sutures.  Colour  : 
each  whorl  is  chequered  by  alternate  squares  of  white  and  choco- 
late; apex  white,  base  chocolate.  Sculpture  :  on  the  tirst  four 
adult  whorls  are  two  gemmule  rows,  on  the  tifth  a  small  median 
row  is  intercalated,  which  increases  till  on  the  last  whorl  it  equals 
the  others;  the  gemmules  are  about  18  to  a  whorl,  large  and 
closely  set,  linked  within  the  row,  but  not  from  row  to  row;  on 
the  base  are  two  plain  spiral  keels.  Protoconch  :  first  whorl 
smooth,  remainder  bicarinate,  crossed  by  numerous  fine  bars. 
Aperture  subquadrate,  spur  of  the  lip  not  reaching  the  pillar, 
anal  notch  simple,  shallow;  canal  short,  straight.  Length  5; 
breadth  1'6  mm. 

Hab. — Watson's  Bay;  under  stones  (J.  Brazier). 

Type. — To  be  presented  to  the  Australian  Museum. 

In  colour  and  shape  this  species  is  like  T.  maculosa  and  T. 
festiva.  Than  the  former  it  is  far  smaller,  than  the  latter  it  is 
larger  and  narrower. 

Triphora  fasciata,  T.  Woods. 

Triforis  fasciata,  T.  Woods,  Proc.  Roy.  Soc.  Tas.  1878  (1879), 
p.  34. 

(Plate  xxxiii.,  figs.  40,  41.) 

This  species  has  been  generally  misunderstood.  I  believe  that 
all  references  to  it  in  literature  subsequent  to  the  original  descrip- 
tion are  erroneous.  The  late  M  r.  C.  E.  Beddome,  whose  eyesight 
was  not  good  in  his  later  years,  distributed  other  species,  especi- 


616  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

ally  a  form  of  T.  granifera^  under  the  name  of  T.  fasciata.  Thus 
Brazier  was  misled  into  identifying  (these  Proceedings  (2)  ix., 
1894,  p.  694)  T.  scitulus  as  from  Sydney  and  as  identical  with 
T.Jasciata.  Tate  tfe  May  have  remarked  that  the  two  species 
cannot  be  the  same  (these  Proceedings,  1901,  p.  457). 

To  digress  an  instant,  I  would  hazard  the  conjecture  that 
Triphoris  scihdus,  A.  Ad.,  (Proc.  Zool.  Soc.  1851,  p.  278)  from 
Port  Lincoln,  S.A.,  is  probably  identical  with  T.  pfeifferi,  Crosse 
and  Fischer  (Journ.  de  Conch.  1865,  p.  47,  pi.  i.,  ff.  14,  15).  It  is 
significant  that  T.  2?feifferi  but  not  T.  scitulus  appeared  in  the 
first  South  Australian  list  published  by  Angas,  though  he  included 
the  latter  in  his  supplement  (Proc.  Zool.  Soc.  1878,  p.  ©67). 

By  its  ornament  T.  fasciata  recalls  T.  pfeifferi,  Crosse  and 
Fischer,  a  longer,  narrower  shell  in  which  the  inequality  of  the 
gemmule  rows  is  further  exaggerated.  Indeed  Tasmanian  col- 
lectors have  sent  me  T.  fasciata  labelled  T.  pfeifferi;  and  accord- 
ing to  Woods'  own  shell  in  the  Tasmanian  Museum  (kindly  lent 
to  me  for  the  purpose  of  this  article  by  Mr.  A.  Morton)  T.  pfeifferi 
itself  was  distinguished  by  Tenison- Woods  as  "7".  tasmanica  var. 

It  is  only  through  the  great  kindness  of  the  Rev.  H.  D. 
Atkinson,  who  has  lent  to  me  part  of  the  original  series  studied 
by  Tenison  Woods,  that  I  am  now  enabled  to  recognise  and 
rehabilitate  the  species.  This  authentic  material  shows  that  the 
original  diagnosis  was  obscure  and  confusing.  I  therefore  add 
here  the  figure  and  description  I  had  prepared  from  material  col- 
lected in  Sydney  Harbour  by  the  late  Mrs.  Starkey  and  by  H.  L. 
Kesteven,  an  account  I  composed  under  the  impression  that  I  was 
dealing  with  a  new  species. 

Shell  small,  narrow,  translucent,  glossy.  Colour  white,  base 
and  supersutural  thread  orange-brown.  Whorls  ten,  plus  a  pro- 
toconch  of  five  and  one-half  whorls.  Sculpture  :  base  smooth,  a 
plain  supersutural  thread,  followed  by  two  rows  of  gemmules,  the 

*  Ten.  Woods,  Proc.  Eoy.  Soc.  Tas.  1S77  (lS7S),p.36;  ?  T.tasmanicav&x. 
op.  cit.  1876  (1877),  p.  lol. 


BY  c.  hedlp:y.  617 

lower  much  the  hirger.  between  which  are  intercalated  on  the 
lower  whorls  a  row  of  minute  gemmules.  The  gemmules  are  15 
to  a  whorl;  in  transverse  order  they  mount  the  spire  obliquely; 
those  of  the  principal  row  are  large,  prominent,  hemispherical 
bosses.  Protoconch  of  5|  whorls  with  a  median  keel  and  trans- 
verse bars.  Aperture  subquadrate,  anal  notch  simple,  spur  of  lip 
not  reaching  the  pillai-.  Canal  short,  straight.  Length  5-5  ; 
breadth  1*5  mm. 

Triphora  labiata,  a.  Adams. 
(Plate  xxxiii.,  figs.  42,  43,  44.) 

This  species  was  originally  described  (Proc.  Zool.  Soc.  1851, 
p.  279)  as  follows: — "T.  testa  subulato-pyramidali,  nigro-fusca, 
in  medio  tumida,  spira  apice  obtuso;  anfractibus  10,  planulatis, 
triseriatim  granuloso-carinatis,  suturis  concavo-impressis;  labro 
reflexo,  dilato,  albido;  canali  brevi,  subrecurvo.  Sydney,  under 
stones,  low  water  (Mr.  Strange).  A  small  nearly  black  shell,  with 
the  outer  lip  dirty  white  or  pale  fuscous."  To  this  Angas  added 
(op.  cit.  1867,  p.  209)  the  important  item,  "Length  3  lines." 

Tryon  naturally  included  this  in  a  list  of  lost  species  when  he 
monographed  the  genus.  Though  such  scanty  data  are  insufficient 
to  distinguish  this  among  the  host  of  known  Trijohora,  it  is  enough 
to  pick  it  out  from  the  Triphora  of  Sydney  Harbour.  The  only 
Sydney  shell  which  suits  the  description  of  Adams,  and  one 
which  has  not  hitherto  been  locally  recognised,  is  here  figured. 
I  have  found  it  at  Cape  Banks  alive  at  low  water  under  stones. 
I  add  the  following  description  : — 

Shell  glossy,  short,  broad,  swollen,  apex  slightly  m.ammillate. 
Whorls  ten,  separated  by  deep  sutures.  Colour  :  the  lower 
whorls  are  entirely  dark  chocolate;  towards  the  apex  the  lower 
half  of  each  whorl  becomes  gradually  paler,  reaching  a  buff  colour 
at  the  apex,  the  chocolate  persisting  above  as  a  narrow  sutural 
thread;  outer  lip  dull  white.  Sculpture  :  on  the  base  are  three 
plain  spiral  keels.  The  ultimate  and  penultimate  whorls  have 
three  equally  spaced  and  sized  keels  bearing  numerous  (more  than 
twenty)    small,    close-set,   ill-defined    gemmules.      In    transverse 


618  STUDIES    ON    AUSTRALIAN    MOLLUSCA, 

order  the  gemmules  are  set  in  perpendicular  rows.  On  the  ante- 
penultimate the  median  gemmules  shrink  in  size,  and  a  few 
whorls  higher  up  the  median  row  pinches  out.  First  whorl 
dome-shaped;  secondMnflated,  with  two  small,  sharp,  plain  spiral 
keels;  on  the  third  the  keels  commence  to  resolve  into  gemmules, 
after  which  the  adult  sculpture  is  assumed.  Aperture  rounded, 
closed  on  the  right  by  a  spur  crossing  the  pillar  lip;  anal  notch 
a  deep  subcircular  orifice.  Canal  extremely  short,  recurved. 
Length  4 "2;  breadth  1-5  mm. 

In  colour  and  contour  this  resembles  T.  ohesula,  Jousseaume, 
from  which  the  protoconch  and  anal  notch  immediately  separate  it. 

Triphora  kesteveni,  n.sp. 
(Plate  xxxiii.,  fig.  45.) 

Shell  acicular.  Colour  pale  pink.  Whorls  tw^elve.  Sculpture: 
on  the  last  whorl  four,  on  the  others  three,  smooth  spiral  ribs,  the 
uppermost  the  largest;  interstices  punctate.     Length  6-5  mm. 

Hah. — Ladies  Hall  Beach,  South  Head,  Sydney;  one  specimen 
(H.  L   Kesteven). 

X  y  p  e.  —  To  be  presented  to  the  Australian  Museum. 

This  species  so  differs  in  form  and  sculpture  from  others  known 
from  this  coast  that  I  have  ventured  to  describe  it  from  a  single 
worn  and  decollate  example. 


EXPLANATION  OF  PLATES. 
Plate  xxix. 

Figs.  1,  2,  3. — Exterior  and  hinge  of  Mactra  ahhreviata,  Lanik. ,  var. 

Figs.  4,  5. — Adult,  magnified  sculpture  and  larval  shell  of  Purpura  pseud- 
amygdala,  Hedley. 

Fig.  6.—Assiminea  pagodel/a,  Hedley. 

Fig.  7. — Cacum  lilianum,  Hedley. 

Figs.  8,9. — 6'y^inr^?'o6u^/a^/?sc/ifri,  Adams  and  Angas,  viewed  from  above  and 
from  the  side. 

Figs.  10,  11,  12.  — Various  aspects  of  Endodonta  otwayensis,  Petterd. 


BY    C.    HEDLEY.  619 

Plate  xxxi. 

Figs.  13,  14,  15. — Various  aspects  of  Endodonta  suhdepressa,  Brazier. 
Figs.  16,  17. — Endodonta  nielbournensis,  Cox,  viewed  from  the  side  and  from 

beneath. 
Figs.  18,  19,  20. — Various  aspects  of  Endodonta  tamarensis,  Petterd. 

Plate  xxxii. 
Fig.  21. — Triphora  7^egina,'H.ed\ej. 

Fig.  22. — Young  shell  of  Triphora  tasmanica,  Tenison  Woods. 
Figs.  23,  24,  25. — Aperture,  apex,  adult  and  variety  of  Triphora  innotahilis, 

Hedley. 
Figs.  26,  27. — Ba.ck  view  of  shell  and  apex  of  Triphora  albovittata,  Hedley. 
Figs.  28,  29. — Adult  and  apex  of  Triphora  granifera,  Brazier. 
Figs.  30,  31. — Shell,  apex  and  aperture  of  Triphora  nocturna,  Hedley. 
Figs.  32,  33.— Shell,  apex  and  aperture  of  Triphora  maculosa,  Hedley. 

Plate  xxxiii. 

Figs.  34,  35. — Shell,  apex  and  aperture  of  Triphora  nigrofusca,  A.  Adams. 
Figs.  36,  37. — Shell,  apex  and  aperture  of  Triphora  cinerea,  Hedley. 
Figs.  38,  39.  — Shell,  apex  and  aperture  of  Triphora  amptdla,  Hedley. 
Figs.  40,  41. — Shell,  apex  and  aperture  of  Triphora  fasciata,  Tenison  Woods. 
Figs.  42,  43,  44. — Shell,  apex  and   aperture  from  the   front  and   side   of 

Triphora  lahiata,  A.  Adams. 
Fig.  45. — Shell  of  Triphora  kesteveni,  Hedley. 


620 


NOTES  ON  PROSOBRANCHIATA. 
No.  W.—LITTORINACEA. 
By  H.  Leighton  Kesteven. 

(Plate  XXX.) 

When  studying  the  affinities  of  Fossarina  (9),  I  found  that 
Risella  differed  in  anatomic  characters  from  Littorina,  the  type 
genus  of  the  family,  to  which,  in  the  past,  it  has  been  assigned. 
I  have  since  studied  its  anatomy  from  actual  dissections  and 
from  sets  of  serial  sections,  kindly  prepared  for  me  by  Messrs. 
J.  P.  Hill  and  R.  Greig  Smith.  My  thanks  are  also  due  to  Mr. 
C.  Hedley  for  many  useful  suggestions.  I  have  also  dissected 
Tectarius  and  Littorina.  As  a  result  of  comparing  the  anatomy 
of  these  with  other  Tsenioglossa,  I  have  found  it  advisable  to 
reclassify  the  Littorine  groups  thereof.  It  will  be  as  well  to 
state  my  conclusions  first  and  discuss  them  in  detail  afterwards. 

I  have  taken  as  my  base  Fischer's  classification  (4,  pp.  707-711), 
and  have  altered  it  in  the  following  manner  : — 

Superfamily  LITTORINACEA. 

Family  LITTORINID^.* 

Containing  those  genera  enumerated  hereunder  by  Fischer, 
with  the  exception  of  Fossarina  (which  in  my  paper  above 
referred  to  has  been  shown  to  be  a  Trochid),  Risella  and  Echinella. 


*  In  defining  Littorinidae  thus  I  have  had  to  take  a  good  deal  for  granted, 
since  there  have  been  no  dissections  of  several  of  the  genera  published;  nor 
have  I  been  able  to  study  the  soft  parts  of  any  but  Australian  members. 
I  have  been  unable  to  obtain  even  the  shells  of  some  of  the  genera.  It  is 
probable  that,  as  our  knowledge  of  Prosobranchiate  anatomy  increases, 
several  of  them  will  have  to  be  removed. 


I 


BY    H.    LEIGHTON    KESTEVEN.  621 

Family  RISELLID^. 

Genera  Risella,  Gray,  and  Bissellojjsis,  Kesteven. 

Family  MODULID^. 

Genera  Modulus,  Gray,  Echindla,  Swainson,  and  Peasiella, 

Kevill. 

Family  NASSOPSID^. 

Genus  I^assopsis,  Smith. 
Reviewing  the  superfamily  as  a  whole,  its  anatomic  characters 
are— 1,  a  much  aborted  osphradium;   2,  simple  branchiae;  3,  a 
rather  large  devolopment  of  the  mucous  gland;  4,  small  salivary 
glands;  5,   a  large  buccal  mass;  and  6,   a  long  and  practically 
uniform  radula.     To  these  may  be  added  a  general  similarity  of 
digestive  tract  and  nervous  system,  and  the  absence  of  an  intro- 
vert.    Their  habitat  has   been  the   indicator  which   led  to  the 
inclusion  of  these  families   under  one  designation.     It  seemed 
reasonable  to  expect   that  a  station  such  as   that  occupied   by 
Littorina  should  have  had  a  modifying  effect  on  some  of  the  organs. 
It  is  to  this  that  I  would  assign  the  degenerate  osphradia  and 
branchiae,  and  perhaps  also  the  large  development  of  the  mucous 
gland.    Living  so  much  out  of  water,  the  first  of  these  organs  would 
be  of  little  use  to  the  mollusc,  and  has  accordingly  become  aborted. 
In    Tectarius   nodulosus,   Gmel.   (which    in    New   South   Wales 
lives  beyond  the  reach  of  even  spray,  except  in  rough  weather), 
the  osphradium  is  so  simple  that,  functionally,  it  must  be  almost 
impotent.     The  simplicity  of  the  branchia?  is,  I  would  suggest, 
due  to  the  same  cause.     The  large  supply  of  mucus  may  possibly, 
by  being  deposited    around    the   operculum,    serve   to   seal  the 
closure  of  the  shell,  and  thereby  prevent  evaporation. 

These  three  characters  are  constant  throughout  the  super- 
family;  yet,  if  the  extent  of  their  development  or  abortion  may 
be  accounted  for  by  "environment,"  we  are  at  once  presented 
with  the  question,  are  they  characters  of  systematic  importance  ? 
That  these  characters  are  the  result  of  environment  is  a  matter 
3f  opinion,  but  in  the  present  instance  they  may  bear  weight  as 
41 

i 

A     : 

/<'^\'-'  /  . 

if.-  ^'  .^^  ^<^ 


-'-nk 


622  NOTES    ON    PROSOBRANCHIATA, 

forming  part  of  a  general  similarity  of  organisation.  Returning 
to  the  osphradium,  Pelseneer  (16)  has  said  : — "  II  constitute  alors 
(dans  les  Tenioglosses  les  plus  archaique,  exemple  Paludina^ 
Littoriyia,  Cyclostoma,  Vermetics,  etc.),  un  bourrelet  epithelial 
filiforme,  sur  un  nerf  ou  sur  un  ganglion."  While  not  contesting 
that  this  type  of  osphradium  is  archaic,  I  would  point  out  that 
in  some  instances  it  may  be  a  degenerate  form.  For  instance, 
concomitant  with  a  habitat  such  as  that  affected  by  Littorina, 
Tectarius,  etc.,  there  would  necessaril}^  be  a  disuse  of  this  organ, 
a  condition  favourable  to  degeneration.  The  presence  of  an 
osphradium  in  a  mollusc  endowed  with  a  pulmonary  cavity  would 
seem  to  mark  such  as  an  evolving  Pulmoiiate  rather  than  an 
archaic  Tsenioglossan. 

Although  somewhat  foreign  to  the  present  paper,  a  short 
speculation  on  the  use  of  the  "  glande  pedieuse  "  of  Vermetus  may 
perhaps  be  allowable  here.  In  the  words  of  Lacaze  Duthiers  (10), 
the  osphradium  is  here  "  reduit  a  un  filet."  The  species  of  this 
genus  are  almost  always  covered  by  water,  and,  being  fixed 
permanently  in  one  position,  it  must  be  a  vital  necessity  to  test 
the  quality  of  the  water  they  inhale;  and  yet  the  osphradium, 
although  strongly  innervated,  is  very  simple.  It  has  occurred 
to  the  writer  that  it  may  be  that  the  so-called  pedal  gland  is  a 
highly  specialised  olfactory  organ;  its  free  communication  with 
the  water  would  seem  to  support  this  view.  It  may  be  worthy 
of  note  that  the  analogue  of  such  a  pedal  olfactory  organ  is  to 
be  found  among  the  Pulmonata.  That  it  is  not  a  mucous  gland 
seems  certain  from  Lacaze  Duthier's  paragraph  (10,265): — "Une 
particularite  bien  digne  d'interet  s'est  presentee.  La  glande  placee 
dans  la  cavite  du  corps,  qui  s'ouvre  entre  le  pied  et  la  tete  est, 
tres  probablement  I'intermediaire  entre  la  cavite  generale  du 
corps  et  I'exterieur.  Si  done  il  n'a  pas  ete  possible  de  decouvrir 
ailleurs  I'orifice  exterieur  de  la  circulation,  on  pourrait  le  con- 
siderer  comme  existant  a  la  fa,ce  inferieure  de  la  glande."  This 
explanation  of  its  function  seems  untenable,  since  Vermetus  is 
not  capable  of  much  expansion,  and  one  would  not,  therefore,  look 
to  find  a  highly  specialised  aqueo-vascular  orifice  developed. 


BY    H.    LEIGHTON    KESTEVEN.  623 

I  would  not  be  understood  to  advance  this  theory  as  a  convic- 
tion; it  is  but  a  speculation  of  one  who  has  not  been  able  to 
examine  the  organ  for  himself;  moreover,  the  osphradium  cannot 
be  regarded  as  functionless,  for,  as  already  mentioned,  it  is 
highly  innervated. 

LlTTORINID^. 

The  family  Littoriaidce  has,  at  my  hands,  suffered  only  restric- 
tion. The  reasons  for  removing  the  various  genera  therefrom 
will  be  found   under  the  families  in  which  I  have  placed  them. 

RiSELLID.E. 

The  family  Risellidce  has  been  formed  for  the  reception  of  the 
two  unitypical  genera  Risella  and  Risellopsis. 

I  proceed  at  once  to  describe  the  anatomy  of  Risella  inelano- 
stoma,  Gmelin. 

External  features : — The  mollusc  extruded  from  its  shell  has 
been  figured  by  Quoy  and  Gaimard  (18),  as  also  the  operculum. 
The  ground  colour  of  the  body  is  white,  reticulated  with  black 
markings,  which  in  places  become  confluent.  Foot  white,  tinged 
with  yellow.  Operculum  paucispiral,  corneous,  oval.  The  snout 
is  not  as  long  as  it  is  represented  in  the  figure  referred  to. 
Tentacles  of  medium  length,  carrying  the  eyes  on  prominences 
at  their  bases.  When  compared  with  Littorina  or  Lotorium,  it 
is  seen  that  Risella  is  twisted  half  round  in  its  shell.  In  the 
former  genera  the  dorsal  aspect  of  the  mollusc  is  that  opposite 
to  the  axial,  whilst  in  the  latter,  and  most  probably  all  genera 
possessing  trochiform  shells,  the  dorsal  aspect  is  that  which  is 
uppermost  when  the  shell  is  standing  on  its  base.  To  make  this 
more  plain,  when  Littorina  is  set  perpendicularly,  with  the  spire 
up,  the  dorsal  face  of  the  mollusc  is  on  the  outside  of  the  coils; 
with  the  shell  of  Risella  in  the  same  position  the  mollusc  is  lying 
with  its  dorsal  face  up.  This  twisting  of  the  mollusc  in  the  shell 
has  resulted  in  the  descending  of  the  columellar  muscle  on  to  the 
base  of  the  shell.     In  the  following  description  I  have  spoken  of 


624  NOTES    ON    PROSOBRANCHIATA, 

the  dorsal  as  the  superior,  and  the  ventral  as  the  inferior  aspects 
of  the  mollusc. 

The  mantle  is  thin  and  transparent;  through  it  may  be  seen, 
beginning  from  the  left,  the  osphradium,  branchiae,  mucous  gland, 
rectum,  and  the  uterus  or  vas  deferens.  Immediately  posterior 
to  the  pallia!  cavity,  and  slightly  to  the  left  lies  the  nephridium. 
Below,  posterior  to  the  columellar  muscle,  the  posterior  oesophagus 
and  anterior  aorta  may  be  seen  through  the  thin  connective  wall, 
between  the  body  and  visceral  coil.  The  genital  gland  lies  over 
the  superior  surface  of  the  visceral  coil,  covering  the  liver  which 
constitutes  the  greater  part  of  the  coil;  inferior!}'  the  stomach 
may  be  seen.  The  branches  from  the  aorta  visceralis  reticulate 
over  both  surfaces,  but  more  profusely  on  the  superior.  Along  the 
axis  of  the  coil,  for  about  half  the  length  thereof,  the  posterior 
oesophagus  may  be  seen;  the  main  trunk  of  the  aorta  visceralis  is 
also  situated  here,  but  extends  to  the  end. 

The  osphradium  (PI.  xxx.,  fig.  2,  osph.)  is  extremely  simple;  a 
very  narrow  thread  of  epithelial  tissue,  nearly  as  long  as  the 
branchiae,  without  any  pectinations  whatsoever;  as  already  stated, 
it  is  but  slightly  innervated. 

The  branchice  (fig.  2,  cten.,  and  fig.  5)  are  more  simple  than  in 
Littori^ia,  the  filaments  being  shorter  and  narrower.  The  tine 
thread-like  prolongations  found  in  that  genus,  Lotorimn,  Natica, 
etc.,  are  here  absent.  These  prolongations  should,  I  think,  be 
regarded  as  venous  sinuses  rather  than  as  part  of  the  ctenidium. 
In  the  species  under  consideration  a  large  lacuna  (fig.  5,  /.)  may 
be  observed  defining  the  anterior  distal  half  of  the  organ.  This 
is  provided  with  such  definite  walls  that  it  might  well  be  regarded 
as  a  vein. 

The  mucous  gland  (fig.  1,  m.gl.)  is  normally  large;  it  lies  along- 
side of  the  rectum,  to  the  left,  and  starting  a  little  behind  the 
anus  continues  back  to  the  end  of  the  pallial  cavity.  It  is  of  a 
grey  colour,  and  consists  of  large  cells  of  concentric  structure. 
During  fecundity  the  uterus,  the  walls  of  which  secrete  a  great 
deal  of  mucus,  seems  to  be  developed  somewhat  at  the  expense 
of  this  organ. 


BY  H.  LEIGHTON  KESTEVEN.  625 

The  digestive  tract  (fig.  1)  is  esentially  the  same  as  in 
Littorina.  The  buccal  mass  (figs.  1  and  6,  hue.)  is  large;  the 
cartilaginous  cushions  on  which  the  radula  rides  are  shaped 
like  the  quarters  of  an  orange,  the  thin  or  lower  end  of  the  large 
one  on  the  right  folding  inside  that  on  the  left.  There  being  no 
introvert,  the  anterior  oesophagus  (fig.  6,  ant.  oes.)  is  very  short. 
It  is  provided  with  appendages  (fig.  6,  ajop.)  similar  to  those 
figured  by  Souleyet  in  Littorina  (21).  The  crop  (figs.  1  and  6, 
cro-p)  is  the  largest  organ  in  the  body  cavity;  it  is  of  a  brown 
colour,  and,  as  in  Littorina^  the  lumen  is  along  its  lower  side; 
the  greater  part  of  it  is  traversed  and  divided  by  thin  partitions, 
the  attachment  of  which  may  be  seen  on  the  outside  as  slight 
constrictions.  The  posterior  oesophagus  (figs.  1  and  6,  j^ost.cts.) 
passes  along  the  axis  of  the  visceral  coil  for  about  half  its  length, 
when  it  enters  the  large,  muscular-walled  stomach  (fig.  1,  st.). 
This  latter  is  shaped  like  a  segment  of  a  circle  an tero- posteriorly, 
and  flattened  dorso-ventrally;  its  distal  end  is  a  little  posterior 
to  the  entrance  of  the  oesophagus;  anteriorly  it  tapers  away  to 
the  thin  intestine.  The  intestine  (fig.  1,  int.)  immediately  rises 
towards  the  superior  face  of  the  liver,  and,  having  reached  the 
surface,  traverses  that  face  of  it  to  which  the  nephridium  is 
attached,  in  the  serpentine  manner  depicted  in  my  figure;  thence 
the  rectum  passes  along  the  right  side  of  the  pallial  cavity,  the 
anus  (fig.  1,  an.)  being  situated  well  towards  the  end  thereof. 
The  liver  is  dark  green,  and,  as  is  usual,  is  the  main  constituent 
of  the  visceral  coil  (fig.  1,  liv.).  The  hepato-pancreatic  ducts  (fig. 
1,  h.-p.d.)  are  three  in  number — one  at  the  extreme  posterior  end 
of  the  stomach,  one  emptying  into  the  centre  on  the  superior 
face,  and  the  third  situated  well  towards  the  anterior  end,  on  the 
axial  side.  The  position  of  this  last  may  sometimes  be  seen 
from  the  outside.  The  salivary  glands  (figs.  1  and  6,  sal.gl.)  are 
small;  their  ducts  enter  the  buccal  mass  on  either  side  of  and 
just  anterior  to  the  oesophagus.  The  radula  (fig.  1,  rad.)  is 
typically  littorinoid,  long  (46  mm.)  and  narrow  ;  it  leaves  the 
buccal  mass  on  the  underside  and  passes  to  the  right  of  the  body 
cavity,  where  it  is  coiled  up.     The  rachidian,  as  is  stated  in  the 


626  NOTES    ON    PROSOBRANCHIATA, 

revised  edition  (1871)  is  incorrectly  figured  by  Wilton  in  Wood- 
ward's 'Manual';  my  figure  (fig.  1)  supplies  this  deficiency.  I 
have  counted  over  three  hundred  rows  of  teeth  on  the  ribbon. 

The  nephridiuni  (figs.  1,  2,  3,  -i,  neph.)  is  situated  just  behind 
the  end  of  the  pallial  cavity,  of  which  it,  in  part,  forms  the 
posterior  boundary.  It  is  slightly  more  on  the  left  hand  side 
than  on  the  right.  In  the  figures  it  has  been  detached  from  the 
liver,  and  laid  back  on  the  right  hand  side.  The  reno-pallial 
orifice  is  on  the  left  hand  side;  its  position  is  shown  in  figure 
(tig.  1,  r.h.o.).  The  natural  position  of  the  nephridium  is  shown 
in  the  figures  by  the  dotted  line,  but  the  line  of  the  anterior 
attachment  is  shown  only  on  tig.  4,  by  the  double  dotted  line; 
from  this  it  will  be  seen  that  there  is  a  pocket  of  the  organ  over- 
lying the  pericardium.  The  reno-pericardial  orifice  is  very  small; 
its  position  is  under  the  pocket  just  mentioned  (fig.  4,  r.p.o.). 

Circulatory  system  (tig.  2)  : — The  heart  (fig.  2,  heart;  tig.  5)  lies 
in  the  pericardium,  immediately  behind  the  ctenidium  and  in 
front  of  the  nephridium,  a  pocket  of  which,  as  already  mentioned, 
overlies  it.  It  consists  of  a  thin- walled  auricle  (fig.  5,  aur. ),  well 
divided  from  a  larger  and  thick-walled  ventricle  (fig.  5,  vent.). 
The  arterial  system  consists  of  a  very  short  aortic  trunk,  which 
gives  rise  to  the  anterior  and  posterior  aortae.  The  former 
(fig.  2,  ant.aor.)  enters  the  body  cavity  alongside  and  slightly  below 
the  oesophagus,  underneath  which  it  passes,  apparently  without 
giving  off  any  branches  till  the  centre  of  the  crop  is  reached, 
where  a  small  branch,  which  bifurcates  at  once  on  the  underside  of 
that  organ  is  to  be  seen.  Continuing  forward  it  suddenly 
diminishes  just  posterior  to  the  pedal  commissures  and  ganglia; 
the  main  branch  here  enters  the  foot.  A  smaller  branch  continues 
forward  and  supplies  the  head  and  buccal  mass.  The  posterior 
aorta  (aorta  visceralis)  almost  at  its  inception,  sends  a  branch 
to  supply  the  liver,  under  the  nephridium  (fig.  2,  a.).  Further 
back  another  and  rather  larger  branch  arises  which  supplies 
the  rectum,  uterus,  and  right  side  of  the  mantle  (tig.  2,  b.). 
From  this  second  branch  the  aorta  passes  through  the  liver, 
so  far  as  I  can  ascertain,  without  branching  till  it  reaches  the 


BY  H.  LEIGHTON  KESTEVEN.  627 

surface  on  the  axial  side,  a  little  distance  posterior  to  the 
nephridium  (fig.  2,  c).  From  here  it  may,  without  dissection, 
be  followed  along  the  axis  of  the  coil  to  the  end,  giving  off 
branches  all  the  way  (fig.  2,  c.',  c",  c.'")  ;  those  given  off 
superiorly  are  much  the  largest,  and  in  some  cases  almost  embrace 
the  coil;  the  finer  branches  of  these  reticulate  throughout  the 
genital  gland  and  liver. 

There  are  only  two  veins,  properly  so  called — the  branchial 
(fig.  2,  fig.  5,  br.v.)  and  nephridial  (fig.  2,  neph.v.);  the  former  is 
the  larger,  and  enters  the  auricle  almost  immediately  after  leaving 
the  branchiae.  The  latter,  although  the  smaller,  is  perhaps  not 
less  important;  its  branches  reticulate  over,  and  collect  the  blood 
absorbed  through  the  wall  of  the  nephridium,  to  discharge  itself 
into  the  branchial  vein  just  outside  the  pericardium.  It  seems 
probable  that  this  vein  supplies  the  new  blood  to  the  system. 
My  sections  show  venous  sinuses  between  the  inner  and  outer 
wall  of  the  mantle.  Their  presence  was  to  be  anticipated,  con- 
sidering the  size  of  the  artery-  supplying  the  right  side  of  the 
manile.  From  the  ventricle  through  the  right  pallial  artery,  and 
pallial  sinuses  to  the  branchi^,  and  back  to  the  auricle,  is  then 
the  shortest  circuit.  It  was  this  that  led  me  to  suggest  that  the 
so-called  "filiforme  prolongations"  of  some  branchiae  are  really 
venous  sinuses. 

Nervous  systeiti  (figs.  12,  13)  : — The  main  ganglia  and  their 
commissures  onl}^  are  here  described  and  figured.  So  far  as  my 
dissections  have  gone,  the  system  is  essentially  the  same  as  in 
Littorina.  It  is  hoped  in  a  future  paper  to  describe  this  part  of 
the  anatomy  of  Risella  in  more  detail,  and  to  compare  it  with  the 
nervous  systems  of  several  of  the  other  genera  in  the  superfamily. 
The  difficulty  attendant  on  getting  material  for  such  a  comparison 
has  rendered  it  impossible  to  incorporate  it  in  the  present  essay. 
The  cerebral  ganglia  (figs.  12,  13,  e.g.)  and  their  commissure 
are  not  at  right  angles  to  the  antero-posterior  axis  of  the 
mollusc,  but  at  an  angle  to  it  of  about  45°,  the  right  ganglion 
being  the  more  anterior  of  the  two.  Anteriorly  both  ganglia 
give  off  three  or  four  nerves,  but  I  have  been  unable  to  definitely 


628  NOTES    ON    PROSOBRANCHIATA, 

identify  any  of  them,  the  nervous  system  not  being  decipherable 
in  my  serial  sections.  The  two  inner  ones  {c.h.con.)  are  believed 
to  be  the  cerebro-buccal  connectives.  It  is  also  believed  that  the 
labial  circle  described  for  Nassojysis  by  Moore  (13)  exists  in  this 
genus  also,  but  this  is  entirely  uncertain.  A  nerve  rising  from 
the  posterior  and  inferior  face  of  the  right  cerebral  ganglion 
(n.gen.)  often  comes  away  with  the  ganglion.  After  running  a 
rather  short  course  it  splits  up  into  several  fine  branches.  It  has 
not  been  seen  in  situ,  but  possibly  innervates  the  genitalia  on 
the  body  wall.  Inferiorly  both  cerebral  ganglia  are  connected 
by  their  respective  commissures  (c.pd.con.)  to  the  pedal  ganglia. 
Posteriorly  the  cerebro-pleural  commissures  (c. pi. con.)  are  given 
oflf,  and  connect  the  cerebral  and  pleural  ganglia.  The  left 
cerebro-pleural  connective  is  longer  than  the  right.  The  pleural 
ganglia  (pl.g.)  are  also  connected  directly  with  the  pedal  ganglia, 
the  pleuro-pedal  connectives  (pl.pd.con.)  being  stouter  than  the 
cerebro-pedal  connectives. 

The  left  pleural  ganglion  is  connected  directly  wdth  the  sub- 
intestinal  ganglion  {sh.iiit.g.).  Two  other  nerves  arise  from  the 
left  pleural  ganglion;  one  of  these  {dialxon.)  passes  just  under 
the  floor  of  the  body  cavity  in  the  direction  of  the  left  visceral 
nerve  {n.v.'),  with  which,  although  it  has  not  been  traced  so  far, 
it  possibly  connects,  causing  a  condition  of  dialoneury  on  the  left 
side.  The  third  nerve  rising  from  this  ganglion  is  probably  the 
columellar  nerve  (n.coL).  The  right  pleural  ganglion,  besides 
being  connected  to  its  fellow  pedal  and  cerebral  ganglia,  is  also 
connected  by  the  supra-intestinal  commissure  (sp. int. con.)  to  the 
ganglion  of  that  name  (sp.int.g.).  The  supra-intestinal  commissure 
is  longer  than  is  the  subintestinal  commissure.  There  are  no 
nerves  taking  rise  from  the  right  pleural  ganglion,  so  that  there 
is  no  connection  between  that  and  the  subintestinal  ganglion. 
Both  supra-  and  subintestinal  ganglia  give  rise  to  the  respective 
pallial  (n.pal.)  and  visceral  nerves  (n.v.).  The  pedal  ganglia 
(pcl.g.),  as  in  Littorina,  are  of  a  large  size,  and  give  rise  inferiorly 
to  numerous  nerves,  none  of  which  have  I  been  able  to  identify. 
A    peculiar   ganglionic   knot   (p.g.k.),   partly   divided    from   the 


BY    H.    LEIGHTON    KESTEVEN.  629 

inward  inferior  face  of  each  ganglion,  and  giving  rise  to  two  stout 
and  two  fine  nerves,  is  worthy  of  note;  they  occur  in  Littoi-ina, 
and  are  perhaps  represented  by  slight  swellings  in  the  same 
situation  on  the  pedal  ganglia  of  jS^assopsis  (vide  Moore,  13,  pi.  20, 
f.  7).    _ 

Genitalia  {^)  (fig.  3).  The  gland  (fig.  3,  gl.)  is  situated  on  the 
superior  face  of  the  visceral  coil;  its  lobules  are  arranged  around 
several  centres,  each  centre  apparently  communicating  with  the 
vas  deferens  through  one  main  canal.  The  vas  deferens  (fig.  3, 
v.d.)  is  situate  on  the  axis  of  the  coil,  and  is  very  much  convoluted. 
Arrived  at  the  anterior  end  of  the  liver  it  proceeds,  parallel  with 
the  rectum,  straight  along  the  mantle;  a  little  posterior  to  the 
anus  it  takes  a  sharp  turn  in  towards  the  body,  thence  it  con- 
tinues, still  as  a  closed  vas  deferens,  along  the  bod}'-  to  the  penis 
(fig.  3,  p.).  This  is  situated  on  the  right  side,  almost  under 
the  snout.  The  seminal  products  pass  through  it,  not  along  a 
groove  on  its  side,  as  in  Littorina  (fig.  8). 

Genitalia  (9)  (fig-  ^)  '■ — The  gland,  macroscopically,  is  the  same 
as  that  of  the  male,  except  that  it  is  larger  and  the  centres  of 
arrangement  are  not  so  distinguishable;  some  of  the  lobes  extend 
round  the  outer  side  onto  the  inferior  surface.  The  ovules 
(tig.  9)  may  be  seen  to  be  arranged  around  the  edges  of  the 
lobules.  The  oviduct  (fig.  4,  ovid.)  occupies  the  same  position  as 
the  vas  deferens,  and  is  convoluted  as  much.  The  uterus  (fig.  4, 
ut.)  is  placed  on  the  mantle  between  the  rectum  and  body;  during 
fecundity  it  is  much  enlarged,  and  at  such  times  the  rectum  lies 
in  a  groove  on  its  inward  side;  at  the  posterior  end,  at  the 
junction  of  the  oviduct,  there  is  a  single  convolution.  Its  walls 
are  strengthened  by  muscular  bands  which  may  be  seen  from 
outside  to  form  slight  constrictions.  From  its  anterior  end,  at 
the  same  position  at  which  the  vas  deferens  turns  into  the  body, 
the  uterus  may  be  seen  to  give  off  a  short  tube,  which  opens  into 
a  groove  (fig.  4,  ovip.)  running  along  the  body;  this  groove 
terminates  in  the  same  situation  as  the  penis,  by  its  two  walls 
dividing  and  becoming  enlarged  into  flat  attached  lobes.  Except 
during  the  period  of  fecundity,  the  whole  of  both  the  male  and 


630  NOTES    ON    PROSOBRANCHIATA, 

female  reproductive  complex  is  very  much  shrunken,  and  the 
parts  are  very  difficult  to  dissect  out.  At  such  times  the  glands 
are  restricted  to  the  main  branches  of  the  visceral  arterial  system, 
these  branches  forming  the  centres  around  which  the  lobes  are 
grouped  when  fully  developed.  The  extensions  of  the  female 
gland  around  the  outer  side  and  on  to  the  inferior  face  are 
invariably  along  the  course  of  an  artery.  This  variation  in  the 
size  of  the  gland  is  noted  by  R.  J.  Harvey  Gibson  (6)  in  Patella 
vidgata  ;  it  doubtless  occurs  in  all  molluscs.  I  have  myself 
noticed  it  in  Lotoriura  and  Purpura. 

The  peculiar  form  of  the  genitalia  is  the  character  on  which  it 
has  been  thought  fit  to  found  this  family.  In  the  male  there  is 
no  sperm-groove  as  in  Liltorina,  its  place  being  taken  by  an 
anterior  prolongation  of  the  vas  deferens  ;  and,  further,  the 
seminal  fluid  passes  through  the  penis,  not  along  a  groove  on  its 
side;  nor  are  any  glands  to  be  detected  on  the  penis.  In  the 
female  we  have  an  equally  important  difference — namely,  the 
existence  of  an  ovipositor  C?)  similar  to  that  found  in  Stromhus 
and  Pterocera.  The  former  of  these  differences  is  one  of  the 
most  important  anatomic  differences  between  the  Lotoriidce  and 
Muricidai. 

Moore  has  stated  (12,  p.  162)  that  he  has  found  this 
ovipositor  (?)  slightly  developed  in  Littorina.  This  is  an  observa- 
tion that  needs  contirmatiun.  The  most  careful  examination 
of  innumerable  fresh  and  spirit  specimens  of  L.  scabra,  Linn., 
(some  of  large  size),  and  L.  maurittaiia,  Lamk.,  has  proved  to  the 
writer  that  it  does  not  exist  in  either  of  these  two  species. 
Moreover,  Souleyet  (21),  in  his  excellent  account  of  the  anatomy 
of  L.  littorale,  Linn.,  does  not  mention  its  presence.  If,  therefore, 
it  does  exist  in  one  species  of  so-called  Littorina^  I  would  suggest 
that  species  of  two  genera  are  grouped  under  one  name. 

Tenison- Woods  (23)  has  stated  that  although  the  species  of 
Riselia  are  organically  hemaphrodite,  functionally  the  sexes  are 
separate,  and  may  be  distinguished  by  the  form  of  the  shell. 
Both  these  statements  are  at  variance  with  facts;  it  is  probable 
that  he  mistook  the  ovipositor  (i)  for  the  male  organ;  of  twenty 


BY    H.    LEIGHTOX    KESTEVEN.  631 

specimens,  all  exactly  similar,  of  the  form  locally  kuown  as  R. 
plana,  Q.  ct  G.,  nine  were  males  and  the  remainder  females. 

Tryon's  reference  (24)  of  PlesiotrocJnis,  Fischer  (3),  and  Limno- 
trochtcs,  Smith  (20)^  to  Risella  as  subgenera  was  altogether  wide 
of  the  mark.  Fischer's  reference  (4,  p.  687)  of  the  former  to  the 
Planaxidte  is  much  more  likely  to  be  correct.  The  systematic 
position  of  the  latter  has  lately  been  ably  discussed  by  Lettice 
Digby  (2). 

Risellopsis,  Kesteven  (9),  is  included  here  almost  entirely  on 
conchological  characters,  nothing  beyond  the  operculum  and 
dentition  being  known  of  its  anatomy;  amd  on  these  it  might 
equally  be  referred  to  the  Littorinidse. 

The  genus  appeals  to  the  writer  as  a  perpetuation  of  the 
immature  condition  of  Risella  (vide  fig.  11,  pi.  xxx.). 

MODULID.E. 

To  this  family,  hitherto  having  but  one  genus — Modulus — I 
propose  to  add  Echiiiella,  Swainson,  and  Peasiella,  Nevill  (14). 
Echinellawd.^  proposed  by  Swainson  (22)  in  1840  for  the  reception 
of  two  species — Tl.  granidata,  Swains.,  and  E.  coronaria,  Lamk. 
The  former  of  these  is  apparently  a  nomen  nuduvi,  and  has  never, 
so  far  as  I  can  ascertain,  been  identified.  The  latter  has,  there- 
fore, been  regarded  as  the  type.  Tryon  (25,  p.  231),  sub  voce 
Tectarius  says  : — "  I  am  also  compelled  to  unite  with  this  group, 
as  synonyms,  Nina,  Gray,  1850,  and  Echinella,  Swainson,  1840, 
their  characters  being  very  variable,  and  shading  into  those  of 
the  type."  Fischer  (4)  treated  the  genus  as  valid,  placing  it  after 
Tectarius,  in  the  Littorinid^e.  He  thus  describes  it : — "  Coquille 
imperforee,  granuleuse,  epaisse;  spire  elevee,  pyramidale;  overture 
striee  a  I'interieure;  base  de  le  columelle  inuni  d\ine  dent; 
opercule  poly  gyre,  d  nucleus  central.^''     The  italics  are  mine. 

In  arriving  at  the  above  conclusion,  Trj^on  could  only  have 
studied  the  tooth  on  the  base  of  the  columella  as  the  recognition 
mark  of  Echinella.  This  he  would  have  found  unreliable,  since 
it  is  slightly  developed  in  Tectarius  pagodas,  Linn.  The  more 
important   character  of   the   multispiral   operculum   was,   in  all 


632  NOTES    ON    PROSOBRANCHIATA, 

probability,  entirely  overlooked  by  him.  Adams  Brothers  d) 
enumerated  three  species  of  Echinella,  viz.  : — E.  coronaria,  Lamk. 
E.  granulata,  Swains.,  and  E.  cumingii,  Phil.  Although  devoid 
of  the  tooth  on  the  columella,  Tectarius  spinulosus,  Philippi  (17), 
has  the  multispiral  operculum,  and  is,  therefore,  an  Echinella. 
Tectarius  bullatus,  Martyn,  and  T.  tectum-persicum,  Linn.,  are 
probably  referable  here  also.  Tryon  regards  E.  coronaria,  Lamk., 
as  a  variety  of  the  former  of  these  two  (25,  p.  257),  but  it  is 
doubtful  whether  this  is  correct;  T.  spinulosus,  Phil.,  which  he 
has  stated  to  be  the  young  of  T.  bullatus,  Martyn,  is  certainly  a 
distinct  species.  As  stated  by  Hedley  (8),  Tectarius  inontrouzieri , 
Fischer,  is  a  Turcia. 

E.  granulata,  Swains.,  never  having  been  identified,  we  have 
three  species  certainly  belonging  to  the  genus  Echinella,  with  the 
probability  of  two  others.  E.  coronaria,  Lamk.,  doubtless  has  a 
polygyral  operculum,  and  from  personal  knowledge  the  writer  can 
say  the  other  two  have  also.  The  opercula  of  several  species  of 
Tectarius  have  also  been  examined  b}'-  the  writer,  and  in  no 
instance  can  he  find  an  intermediate  between  the  typically 
polygyral  one  of  E.  cumingii  and  that  of  T.  pagodus,  Linn., 
which  is  typically  littorinoid. 

In  discussing  the  affinities  of  Echinella^  the  writer  is  handi- 
capped in  having  been  unable  to  obtain  specimens  for  dissection. 
The  operculum,  however,  happily  affords  us  an  important  clue  to 
its  systematic  position.  Although  undoubtedly  bearing  great 
conchological  resemblance  to  Tectarius,  the  difference  between 
their  opercula  debars  their  inclusion  in  the  same  family.  There 
are  only  five  or  six  families  in  the  T^enioglossa  in  which  the 
multispiral  operculum  occurs;  of  these  the  Modulida?  is  the  only 
one  to  which  Echinella  can  be  referred.  The  tooth  on  the  base 
of  the  columella,  possessed  by  some  of  the  species,  strengthens 
this  classification,  which,  in  view  of  our  ignorance  of  the  soft 
parts,  is  more  advisable  than  would  be  the  addition  of  another  to 
the  already  large  number  of  families  in  the  suborder. 

Peasidla  was  proposed  by  Nevill  (14)  as  a  subgenus  of  Eisella 
for  the  reception  of  Trochus  tantillus,  Gould  (6).      His  treatment 


BY    H.    LEIGHTON    KESTEVEN. 


633 


of  it  in  this  manner  was  apparently  in  deference  to  the  opinion 
of  E.  A.  Smith,  who,  in  dealing  with  the  type,  had  said  : — "The 
interior  of  the  aperture  is  not  pearly  as  in  the  Trochidse;  and  yet 
the  operculum  truly  appertains  to  that  family,  being  concentric 
and  multispiral;  and  although,  therefore,  differing  from  that  of 
Eisella,  the  shell  itself  appears  to  suggest  the  joropriety  of  its 
being  located  with  that  genus  rather  than  with  the  Trochidse" 
(19).  Disregarding  the  peculiarly  contradictory  phrasing  of  this 
statement,  the  conclusion  seems  to  be  a  good  deal  at  fault.  The 
resemblance  of  the  shell  to  Risella  is  very  slight,  and,  as  stated, 
the  opercula — characters  of  infinitely  more  importance — are  of 
two  distinct  types.     The  dentition  (Text  fig.  1)  resembles  that  of 


Fig.  1 — Operculum  and  dentition  of  Peasiella  tantillus,  Gould. 

Littorina  more  nearly  than  any  other;  but  that  of  Modulus,  with 
which  I  would  place  Peasiella  as  a  full  and  valid  genus,  is  of  the 
same  type.     The  operculum  also  favours  this  allocation. 

Hedley  (8)  has  drawn  attention  to  the  fact  that  Echinella 
gaidii,  Montrouzier  (H),  is  a  synonym  of  Trochus  conoidalis, 
Pease  (15).  Montrouzier  describes  the  operculum  as  normal — 
that  is,  corneous  and  multispiral,  since  such  is  the  operculum  of 
the  genus  to  which  he  referred  it.  The  species  is,  therefore,  a 
Peasiella;  but  whether  the  other  species  which  have  been  placed 
here  (-u^'c^e  Tr^^on,  25,  pp-  263-264)  belong  to  this  or  distinct  genera 


634  NOTES    ON    PROSOBRANCHIATA, 

remains  to  be  ascertained,  so  insufficient  have  purely  conchological 
characters  proven. 

Nassopsid.e. 

The  family  here  proposed  contains  but  one  species,  Ji^assopsis 
nassa,  Woodward  (86, 27),  one  of  the  unique  "Halolimnic"  mollusca 
of  Lake  Tanganj^ika.  The  anatomy  has  been  very  thoroughly 
worked  out  by  J.  E.  S.  Moore  (13),  but  this  writer  has  viewed  the 
characters  of  the  species  of  the  Halolimnic  fauna  from  so  prejudiced 
a  standpoint  that,  instead  of  discussing  their  affinities,  he  has 
exerted  all  his  ability  to  prove  that  they  are  surviving  Jurassic 
archetypes.  To  such  lengths  is  this  conviction  carried  that  the 
arguments  used  tend,  in  some  instances,  to  weaken  the  undoubt- 
edly correct  theor}^  that  the  lake  has  been  an  arm  of  the  sea  in 
early  geologic  times. 

In  the  present  instance  no  definite  opinion  as  to  the  relation- 
ship of  the  genus  is  given,  though  it  is  vaguely  compared  with 
the  Strombidae,  Paludina  and  Littorina.  A  careful  weighing 
of  its  characters,  however,  it  is  here  contended,  shows  its  relation- 
ship to  be  with  the  Littorinidae. 


BiBLIOGEAPHY. 

1.  —Adams,  H.  &  A.— Genera  of  Recent  Mollusca,  i.,  1854,  316. 

2.— DiGBY,  Lettice. — "  On  the  Structure  and  Affinities  of  the  Tanganyika 

Gasteropods,   Chytra  and  Limnotrochus"    Journ.  Linn.  Soc.  Lond. 

Zool.,  xxviii.,  1902,  434,  ct  seq. 
3. — Fischer.— Journal  de  Conchyliologie,  sxvi.,  1S78,  212;  xxvii.,  1879,  29, 

pi.  iii.,  fig.  4. 

4. Manuel  de  Conchyliologie,  1887,  709. 

5_ — Gibson,  R.  J.  Hakvey.  — "  Anatomy  and  Physiology  of  Patella  valgata  " 

Trans.  Roy.  Soc.  Edin.,  xxxii.,  1885,  GOl-638. 
6._GouLD.— Proc.  Boston  Soc,  iii.,  1849,  118.     United  States  Exploring 

Expedition  (Wilkes),  xii.,  1852,  184,  figs.  215,  215a,  2156. 
7.— Hedley,  C. — Mem.  Aust.   Mus.,  iii.,  pt.  7;  "Mollusca  of  Funafuti, 

1899,  424. 
8. "A  Revision  of  the  Types  of  the  Marine  Shells  of  the 

Chevert  Expedition."  Records  Aust.  Mus.,  iv,,  1901,  121. 


BY    H.    LEIGHTON    KESTEVEN.  635 

9. — Kesteven,  H.  Leighton. — The  systematic  position  of  Fossarina,  Ad.  & 

Ang.,  and  of  Fossarina  varia,  Hutton.  Kecords  Aust.  Mus.,  iv.,  1902, 

317-322. 
10. — Lacaze  Duthiees.  —  "  L'Anatomie  et  1'  embriogenie  des  Vermets."  Ann. 

Sci.  Nat.  Zool.  4e  ser.  xiii.  1860,  230,  pi.  4,  tig.  6. 
11.— Montrouzieb. — Jour,  de  Conch,  xxvii.,  1878,  26,  pi.  iii.,  tigs.  3,  3a. 
12. — Moore,  J.  E.  S  — "  The  Molluscs  of  the  great  African  Lakes.  No.  iii. 

Tangdnyikia  and  Spekia.\  Quart.  Journ.  Micr.  Sci.  xlii.,  1899. 
13. Op.  cit.  No.  iv.  Nassopsis  and  BytJwceras.  Loc.  cit. 

1899,  187,  et  seq. 
14. — Nevill. — Hand  List  of  the  Mollusca  in  the  Indian  Museum,  1884,  159. 
15. — Pease.— American  Jour.  Conch.,  iii.,  1867,  287;  pi.  12,  fig.  8. 
16. — Pelseneer. — Introduction  a  I'etude  des  Mollusques,  1894,  57. 
17. — Philippi. — Abbildungen  Conchylien.      Band  iii.,   1847,   17,  Tab.  vi., 

fig.  24. 
18. — QuoY  &  Gaimard.-    Voyage  de  I'Astrolobe.   Zool.  Tome  iii,,  1855,  271. 

Atlas  Moll.,  pi.  62,  figs.  8-11. 
19.— Smith,  E.  A.— Journ.  Linn.  Soc.  Lond.  Zool.  xii.,  1876,  552. 

20. Ann.  and  Mag.  Nat.  Hist.,  ser.  5,  vi.,  1880,  425. 

21— SouLEYET.— Zool.  de  la  Bonite,  ii.,  1853,  551,  pi.  33. 

22. — SwAiNsoN. — A  Treatise  on  Malacology,  1840,  352. 

23.— Tenison- Woods.  — Proe.  Linn.  Soc.  N.S.  Wales,  i.,  1876,  242-249. 

24.— Tryon.— Structural  and  Systematic  Conchology,  ii.,  1883,  242. 

25. Manual  of  Conchology,  ix.,  1887. 

26.— Woodward.— Proc.  Zool.  Soc.  Lond.,  1859,  349,  pi.  47,  figs.  4,  4a,  4&. 
27.— Smith.— Ann.  and  Mag.  Nat.  Hist.,  ser.  6,  vi.,  1890,  93. 


EXPLANATION  OF  PLATE  XXX. 

Anatomy  of  Risella. 

Fig   1.— Digestive  system. — buc,  buccal  mass,     rad.,  radula.     an.,  anus. 

m.gl.,  mucous    gland,      post. as.,  posterior   oesophagus,      neph., 

nephridium.     liv.,  liver,     sal.gl.,  salivary  glands.      crop,  crop. 

r.b.o.,   reno-pallial   orifice.       int.,   intestine.       h.-p.d.,   hepato- 

pancreatic  ducts. 
Fig.  2.— Circulatory    system. — cten.,   ctenidium.      br.v.,    branchial    vein. 

osph.,  osphradium.       nepli.v.,  nephridial    vein.       a.c.c.'c."c."' , 

branches  of  the  posterior  aorta,     b. ,  right  pallial  artery,     ant.aor., 

anterior  aorta,     nepli.^  nephridium. 
Fig.  3. — Genitalia  (  $  )• — A-.,  a spermatozooid.    p.,  penis,    v.d.,  vas deferens. 

neph.,  nephridium.     gl.,  testes. 


636  NOTES    ON    PROSOBRANCHIATA. 

Fig.  4. — Genitalia  (  $  ). — ovij).,  ovipositor  (?).  ut.,  uterus,  ovid.,  oviduct, 
jiej:*/^ ,  nephridium.     gl.,  ovary,     r.j^.o.,  reno-pericardial  orifice. 

Fig.  5. — Ctenidium. — b.,  venous  lacuna,  nejyh.v.,  nephridial  vein,  vent., 
ventricle.     rt?rr.,  auricle,     fe?'.!?. ,  branchial  vein. 

Fig.  6. — Anterior  portion  of  the  digestive  tract. — buc,  buccal  mass,  sal.gl.y 
left  salivary  gland,  app.,  left  appendage,  ant.ces.,  anterior 
oesophagus.  crop,  crop,  post.aes,,  posterior  oesophagus.  I., 
lumen,     rad.,  radula. 

Fig.  7. — Rachidian  tooth. 

Fig.  8. — Penis  showing  sphincter  muscles,  and  seminal  canal  squeezed  out 
between  lines  of  contraction. 

Fig.  9. — One  lobule  of  ovary  showing  ovules  arranged  round  the  edge. 

Fig.  10.— Crop. — I.,  lumen. 

Fig.  11. — Young  shell. 

Fig.  12. — Nervous  system  dissected  out  and  drawn  with  the  camera  lucida 
as  seen  floating  in  water. — e.g.',  left  cerebral  ganglion,  e.g.", 
right  cerebral  ganglion.  pl.g.\  left  pleural  ganglion,  pl.g.", 
right  pleural  ganglion,  pd.g. ',  pd.g." ,  left  and  right  pedal  ganglia, 
sb.int.g.,  subintestinal  ganglion.  sp.int.g.,  supra-intestinal 
ganglion,  sp.int.con.,  supra-intestinal  connective,  sb.int.con., 
subintestinal  connective,  c. 2)1. con.,  cerebropleural  connectives. 
c.pd.con.,  cerebro-pedal  connectives,  pl.pd.con.,  pleuropedal 
connectives,  dial. con.,  dialoneurous  connection  (?)  c.b.con., 
cerebro-buccal  connectives  (?)  n.gen.,  genitalia  nerve  (?)  nv.\ 
n.v." ,  left  and  right  visceral  nerves,  n.pal.',  n.pal." ,  left  and 
right  pallial nerves,  n.col.,  columellar  nerve  (?)  p.g.k.,  ganglionic 
knots. 

Fig.  13. — Nervous  system  seen  from  above.     Lettering  as  in  fig.  12. 


63^ 


NGARRABUL  AND  OTHER  ABORIGINAL  TRIBES. 

Part  I. — Medical  and  Surgical  Practice. 

By  John  MacPherson,  M.A.,  B.Sc,  M.B.,  Ch.M. 

While  engaged  in  professional  work  in  New  England,  I  made 
the  close  acquaintance  of  the  aboriginals  inhabiting  that  district 
and  those  passing  through  it  in  the  course  of  their  wanderings. 
From  them  I  learned  some  details  of  their  methods  in  medicine 
and  surgery,  such  as  were  practised  while  the  tribes  still  preserved 
their  integrity.  The  district  I  speak  of  was  occupied  by  many 
tribes,  known  by  the  names  of  the  languages  spoken  by  their 
members.  Thus,  the  Yukumhul  language  was  spoken  at  Inverell 
and  Boggy  Camp.  JSfgarrahul  was  used  at  Bolivia,  Glen  Innes, 
Stonehenge,  Wellingrove,  Emmaville,  and  the  neighbouring 
localities.  Ea-nee-ivin  obtained  at  Oban  and  its  vicinity.  The 
Bundel  or  Bundela  tongue  prevailed  at  Yugilbar  (upon  the  Upper 
Clarence)  and  Gordon  Brook  (a  tributary  of  the  same  stream). 
In  the  following  account,  unless  otherwise  specified,  the  facts 
relate  to  the  customs  of  the  Ngarrabul  Blacks. 

The  education  of  students  in  medicine  took  place  at  the  great 
Bora  Initiation  ceremonies  (Bull-bah),  the  old  doctors  and  warriors 
imparting  the  information.  Many  youths  from  different  tribes 
received  their  instruction  together.  The  old  people  withdrew, 
and  the  doctors  (in  number  from  three  to  six)  took  the  lads  apart 
to  teach  them  the  mysteries  of  the  healing  art.  From  time  to 
time  their  proficiency  was  tested;  they  were  sent  to  observe  and 
diagnose  cases  of  illness — being  reprimanded  for  their  mistakes — 
until  the  required  degree  of  competency  was  gained.  When  at 
length  the  young  man  (for  such  he  had  now  become)  was  deemed 
42 


638  NGARRABUL    AND    OTHER    ABORIGINAL    TRIBKS, 

fully  qualitied  to  practise,  he  must  needs  comport  himself  with 
circumspection,  else  trouble  awaited  him.  If  his  erstwhile 
instructors  considered  him  unwarrantably  zealous  in  prosecuting 
the  work  of  his  profession,  they  reported  the  matter  to  the  tribal 
chief  {Doorttngd  ov  yyumbaivnh),  who,  if  he  found  the  accusation 
sustained,  ordered  the  culprit  to  be  summarily  put  to  death. 
Such  drastic  measures  were  deemed  necessary,  for  the  doctor 
would,  when  opportunity  favoured,  subject  a  patient,  against 
whom  he  harboured  resentment,  to  a  course  of  treatment  unneces- 
sarily rigorous.  In  the  Yukumbul  tribe,  to  guard  against  the 
possibility  of  such  secret  enmity  being  visited  upon  the  sick  man 
{Dfhikkae^)  by  any  doctor  {Xoonwae-hah),  he  (the  patient)  might 
insist  upon  several  medical  men  being  called  in  consultation  upon 
his  case.  In  Bundel  the  dusky  physician  was  known  as  Xgull- 
loo-hiiyal  —a,  layman  or  ordinar}^  tribesman  being  BiyynL 

Before  the  advent  of  the  white  settler,  the  natives  informed 
me,  there  was  but  little  sickness  or  disease  of  any  kind.  Tumours 
or  neoplasms  were  rare  or  unknown.  The  w^ork  of  the  doctors 
was  mainly  surgical,  and  consisted  of  the  treatment  of  wounds — 
the  result  of  accident  or  sustained  in  warfare.  Sorcer}^  and  witch- 
craft occupied  a  prominent  place  in  their  dealings  with  the  sick. 
As  narrated  to  me  by  the  natives,  their  practices  were,  in  the 
main,  similar  to  those  recorded  by  many  other  observers  through- 
out Australia  —  blowing,  manipulating  and  sucking  being,  as 
elsewhere,  conspicuous  features.  In  the  presence  of  the  doctor, 
the  patient  lay  down  and  was  subjected  to  a  ])h3^sical  examination 
that  the  seat  of  disease  might  be  ascertained.  After  feeling  his 
head,  body  and  limbs,  the  ph3^sician  took  some  bushes  and  bit  off 
a  piece  with  leaves.  Upon  these  he  blew,  and  over  them  spread 
a  mysterious  powder  which  he  kept  in  a  bag.  The  leafy  twig  he 
then  pressed  well  into  the  skin  over  the  supposed  diseased  part, 
or,  calling  to  his .  assistance  one  or  more  professional  colleagues, 
they  all  pressed  vigorously  upon  the  potent  charm.     Thus,  they 

*  A  word  of  doubtful  authenticity. 


BY    JOHN    MACPHERSON. 


639 


said,  the  drug  rapidly  pervaded  the  system  and  soon  effected  a 
cure. 

Suction  was  one  of  the  most  universal  of  practices  amongst  the 
Australian  Medicine  Men.  Dr.  Clutterbuck"^  refers  to  a  similar 
custom  among  the  Victorian  natives.  I  was  surprised  to  observe 
the  evident  faith  in  which,  even  at  the  present  time,  this  manner  of 
treatment  is  held  by  the  civilised  aborigines.  Of  course,  in  some 
cases,  it  would  be  a  simple  method  of  dry-cupping  and  would 
have  the  same  applications.  Again,  it  is  the  most  rational  manner 
of  treating  the  stings  and  bites  of  venomous  animals.  Otherwise 
it  was  purely  "  make-believe"  charlatanism,  or  part  of  dexterous 
sleight-of-hand  performances  by  the  astute  physician.  In  the 
first  category  we  might  include  sucking  the  integument  over  the 
affected  parts  in  various  pains  or  headaches,  "  colds "  (nasal 
catarrh),  &c.  ( Yukumbul).  An  old  Yugilbar  native,  whose  grand- 
child had  an  attack  of  acuie  bronchitis  with  cough  and  dyspnoea, 
sucked  the  infant's  mouth  and  nose,  after  the  fashion  of  the 
doctors  of  his  tribe,  as  an  emergency  measure  before  seeking  my 
professional  advice.  If  a  person  felt  sore  (boo-boo)  anywhere,  the 
doctor  inserted  a  cigar-shaped  needle  into  the  tender  place.  This 
manoeuvre,  I  was  assured,  was  almost  painless.  Through  the 
needle  the  wily  sorcerer  affected  to  extract  by  sucking  the 
materies  morbi  (in  the  shape  of  a  pebble  or  similar  object  pre- 
viously adroitly  concealed  in  his  mouth).  This  he  spat  out,  to 
the  great  satisfaction  of  the  sufferer.  In  the  Yukumbul  tribe, 
similarly,  splinters  or  barbs  of  spears  were  supposed  to  be  removed 
by  sucking.  An  Inverell  native  assured  me  that  if  a  spear-head 
penetrated  the  flesh  and  broke  off,  native  doctors  (but  not  Euro- 
pean surgeons)  could,  by  these  means,  extract  the  buried  fragment. 
Of  course  a  piece  of  spear-head  was,  beforehand,  deftly  hidden 
about  the  performer's  person  to  be  produced  at  an  opportune 
moment. 

Their  materia  medica  comprised  herbs,  plentiful  in  the  Clarence 
River   scrubs,   but    scarce    in    the    Glen    Innes    neighbourhood. 


'  Port  Phillip  in  1849,'  p.  54. 


640  NGARRABUL    AND    OTHER    ABORIGINAL    TRIBES, 

Marvellous  properties  in  the  curing  of  disease  were  ascribed  to 
them.  They  were  more  potent  for  good,  the  Blacks  informed  me, 
than  any  of  the  remedies  at  my  command.  Dr.  Clutterbuck 
spoke  with  disdain  of  the  innocent  herbs  swallowed  in  decoction 
by  the  indigenous  tribes  of  Victoria.  It  is,  however,  highly 
probable  that  a  systematic  investigation  of  their  drugs  would  in 
some  instances  reveal  properties  of  which  advantage  might  well 
be  taken  in  the  practice  of  medicine.  I  need  refer  as  an  example 
only  to  the  Corkwood  tree  of  Eastern  Australia  (Duboisia  myopo- 
roides,  R.Br.)  which  was  employed  by  the  natives  for  its  toxic 
properties'^  and  which  yields  Duboisine,  a  drug  whose  use  in 
medical  practice  has  now  become  world-wide.  The  Bundela 
women  prepared  a  medicine  from  a  small  plant  about  nine  inches 
in  height  confined  to  the  warm  climate  of  the  coastal  regions.  A 
fire  having  been  made,  stones  were  placed  upon  it  until  they 
became  very  hot.  Over  these  stones  was  stood  a  kooliman  or 
wooden  bowl  (Noomie)  filled  with  water.  The  plant  was  put  in 
the  water,  which  was  then  boiled  and  poured  into  a  strong  wooden 
medicine  bucket  (Goolong-gooloo).  The  steam  was  confined  for  a 
while  with  tea-tree  bark.  The  drug  thus  prepared  was  of  sour 
taste  and  great  potency,  being  given  with  caution  and  onl}^  in 
very  small  doses.  The  Yukumbul  natives  used  herbs  found  grow- 
ing upon  the  plains  for  internal  disorders  caused  by  drinking 
contaminated  water.  Unfortunately  I  was  unable  to  procure 
specimens  of  any  of  these  plants. 

Two  species  of  the  genus  Exccecaria  are  found  in  Australia  and 
are  remarkable  for  the  possession  of  a  milky  sap  with  intensely 
virulent  and  acrid  properties  and  capable  of  inflicting,  by  contact, 
the  direst  mischief  upon  the  organs  of  vision.  These  are  the 
Gutta  Percha  tree  of  Queensland  and  Northern  Australia  {E. 
parvifolia,  Muell.  Arg.)  and  the  Milky  Mangrove  {E.  agallocha, 
Linn.)  ranging  from  Northern  Australia  to  the  north  coastal  area 
of  New  South  Wales.     The  former  species  was  studied  by  Dr. 

*  F.   M.   Bailey,  Proc.   Linn.   Soc.   N.S.W.,  Vol.  v.,  ISSO,  pp.  1  et  seq. 
(quoting  Rev.  W.  Woolls). 


I 


BY    JOHN    MACPHEHSON.  641 

T.  L.  Bancroft,  and  Mr.  Edward  Palmer*  speaks  of  its  bark  being 
used  by  the  Gulf  of  Carpentaria  natives,  broken  up  in  water,  as 
a  lotion  for  the  relief  of  pain.  The  juice  of  the  Milky  Mangrove, 
according  to  Mr.  J.  H.  Maiden,!  was  used  by  the  aborigines  of 
Eastern  Australia,  New  Guinea  and  India  to  cure  chronic  ulcers. 
In  Fiji  the  patient  was  fumigated  wdth  the  smoke  of  the  burning 
wood.  In  the  Clarence  River  scrubs  grows  a  tree  perhaps 
identical  with  this  Milky  Mangrove  and  whose  bark  and  sap 
were  used  by  the  blacks  in  the  treatment  of  burns.  When 
cutting  the  bark  it  was  necessary  to  avert  the  face,  else  the 
juice  spurting  into  the  eyes  might  destroy  the  sight.  This  juice 
was  collected  in  a  medicine  bucket.  In  treating  burns,  snake's 
fat  was  first  smeared  upon  the  injured  surface  and  over  this  a 
piece  of  the  bark,  while,  at  a  distance,  the  acrid  sap  itself  was 
applied,  doubtless  as  a  counter  irritant.  In  severe  pain  arising 
from  any  cause,  the  juice  was  spread  over  the  painful  area,  or  the 
bark,  rolled  up  into  little  fragments,  was  ignited  and  the  glowing 
ember  applied.  This  was  reserved  for  sufferers  otherwise  strong, 
two  dusky  comrades  holding  the  patient  during  this  administration 
of  the  cautery. 

Another  tree,  however,  of  the  Queensland  and  New  South 
Wales  coastal  scrubs  is  the  rough-leaved  Fig  tree  (Ficus  asjjei-a, 
R.Br.),  which  likewise  yields  an  acrid,  milky  juice,  employed  by 
the  Cleveland  Bay  natives  to  expedite  the  healing  of  certain 
wounds  (James  Morrill). 

The  tannin-containing  kino  of  the  Apple-tree  (Angophora  sp.) 
was  included  in  the  Ngarrabul  pharmacopoeia.  Mr.  J.  F.  Mann  J 
also  speaks  of  the  use  of  the  astringent  sap  of  the  Apple  tree  in 
aboriginal  therapeutics.  In  times  of  drought  the  Yukumbul 
Blacks  procured  water  from  Apple  trees  to  obviate  their  drinking 
impure  natural  water;  as  in  other  parts  of  Australia,  under  the 
stress  of  thirst,  the  natives  derive  water  from  the  roots  of  the 


*  Proc.  Roy.  Soc.  N.S.W.,  Vol.  xvii.,  1883,  p.  107. 

f  Indigenous  Vegetable  Drugs  (Dept.  of  Agriculture  Publications)  Pt.  ii.,  p.  19. 

X  Proc.  Geogr.  Soc.  Aust.,  1st  Session,  1883-4,  Vol.  i.,  p.  49. 


642  NGARRABUL    AND    OTHER    ABORIGINAL    TRIBES, 

Red  Mallee  {Eucalyptus  oleosa,  F.v.M.)  and  the   Coolabah   or 
Dwarf  Box  {E.  mic7'otheca,  F.v.M.). 

The  Red  Gum,  Yarrah  (probably  Eucalyptus  rostrata^  Schlecht.), 
also  found  a  place  in  the  Ngarrabul  materia  medica.  E.  rostrata, 
of  course,  yields  the  Eucalyptus  gum  of  the  British  Pharmacopceia. 
According  to  various  writers.*  the  medicinal  use  of  the  exudations 
of  species  of  Eucalypts  was  of  wide  extent  among  the  Australian 
aborigines. 

The  White  or  Manna  Gum,  Horra  {E.  viminalis,  Labill.)  forms 
one  of  the  most  conspicuous  features  of  the  Glen  Innes  forests. 
Glencoe  and  ]Marowan,  from  the  prevalence  of  this  species,  were 
designated  in  Ngarrabul  Horra-ivill.  In  some  seasons  the  trunks 
are  prettily  tinted  with  patches  of  pink  or  red.  Golden  beetles 
[Anoplognathus),  Psyllid  insects,  ttc,  feed  largel}^  upon  the 
leaves.  The  common  opossum  [Trichosurus  vidj^ecula,  Kerr) 
subsists  to  a  great  extent  upon  the  young  foliage  and  bark  of  the 
smaller  shoots  and  twigs.  The  yield  of  oil  from  the  leaves  is 
about  J%,  consisting  mainl}''  of  pinene,  with  comparatively  little 
eucalyptol.  The  active  principle  (or  principles)  of  Eucalyptus 
oils,  however,  for  therapeutic  purposes — whether  eucalyptol 
(cineol),  peppermint  ketone,  or  other  body — has  yet  to  be 
determined.  In  this  connection  the  admirable  researches  of 
Messrs.  R.  T.  Baker  and  H.  G.  Smith  have  opened  up  an 
enormous  field  for  the  pharmacologist.  The  leaves  of  E.  vimi- 
nalis  were  used  by  the  Ngarrabul  Blacks  topically  for  ophthalmia 
— Narrada  mil  (i.e.,  "bad  eye"),  and  by  internal  administration 
for  diarrhoea.  It  may  have  been  mereh^  a  coincidence  that  a 
captive  opossum  of  mine,  when  deprived  of  these  leaves,  suffered 
from  diarrhoea  with  green  bilious  stools,  but  was  restored  to 
health  when  they  were  again  added  to  his  diet.  Several  authors 
have  observed  the  therapeutic  exhibition  of  Eucalyptus  leaves 


*  Dr.  W.  E.  Roth,  '  Ethnological  Studies  among  the  North-West  Central 
Queensland  Aborigines';  James  Dawson,  'Australian  Aborigines,'  p.  57; 
J.  F.  Mann.  loc.  cit.,  d'C. 


BY    JOHN    MACPHERSON,  643 

b}  the  aborigines  in  different  regions  of  Australia.*  The  tannin- 
containing  bark  of  the  White  Gum  was  employed,  moistened,  by 
the  N'garrabul  Blacks  as  an  outward  application  in  ophthalmia, 
just  as  in  other  parts  of  the  continent  the  bark  of  divers  species 
of  Eucalypts  was  used  for  various  other  purposes. 

To  pass  on  to  the  Animal  Kingdom.  The  fat  of  the  Carpet 
Snake  {Python  spilotes  var.  varieyaia,  Gra}^)  was  in  great  demand 
as  an  emollient  application  for  burns,  or  an  embrocation  for 
rheumatism,  just  as,  for  rheumatic  pains,  the  pristine  dwellers  of 
Tasmania  employed  the  fat  of  the  Mutton  Bird  {Pvffinus  brevi- 
cattdus,  Gould t),  or  the  Australian  settlers  use  Emu  or  Iguana 
oil.  Some  of  my  Yukumbul  informants  averred  that  rheumatism 
was  uncommon  before  the  advent  of  the  white  colonists,  and  the 
consequent  change  in  the  aboriginal  mode  of  life.  An  old  Oban 
native,  when  I  wished  him  to  conduct  me  to  the  ground  of  his 
Bora  {Orba7i-bi,h),  proffered  his  rheumatic  joints  as  an  excuse  for 
declining. 

The  surgical  practice  was  simple.  Slight  wounds  amongst  the 
Yukumbul  people  were  left  untreated  —healing  readily.  In  the 
Ngarrabul  tribe  an  eagle-hawk's  feather  was  placed  upon  a  large 
cut  or  wound  to  close  it;  over  this  some  soft  tea-tree  bark,  the 
whole  being  bound  up  with  a  piece  of  kangaroo-skin  to  keep  it 
warm  and  comfortable.  On  some  of  the  natives  irregular 
cicatrices,  chiefly  about  the  forehead,  gave  evidence  of  the  blows 
of  clubs  [Murrunibn^i  or  Dappirr),  wielded  by  co-tribesmen  in 
altercations,  or  hostile  blacks  in  intertribal  feuds.  One  Oban 
aboriginal  of  great  age,  in  addition  to  such  scars,  had  a  periosteal 
node  upon  his  frontal  bone,  and  great  depressions  in  his  cranial 
vault.  These,  he  said,  were  the  marks  of  quarrels  with  his 
fellows;  but  whether  they  were  depressed  "  pond "  fractures  or 
examples  of  the  absorption  of  the  parietal  bones  which  sometimes 
supervenes  in  advanced  age,  I  cannot  say. 


I 


""■  Edw.  Palmer,  Oleman,  &c.  In  the  National  Dispensatory,  5th  ed., 
p.  627,  it  is  stated  that  the  virtues  of  Eucalyptus  leaves  were  well  known  to 
the  aboriginal  inhabitants  of  Australia. 

t  Bonwick,  *  The  Daily  Life  of  the  Tasmanians,'  p.  89. 


644  NGARRABUL    AND    OTHER    ABORIGINAL    TRIBES, 

Fractures  were  not  uncommon,  the  result  of  accident  or 
encountered  in  war,  from  the  blow  of  the  large  and  powerful  war 
boomerang  {Burr-gun) — the  kind  which  did  not  return  to  the 
thrower.  In  the  Yukumbul  tribe  the  bark  of  a  special  tree, 
Bugaibil,wsi^  used  in  setting  fractures.  Thus,  for  the  forearm,  two 
sheets  of  bark  were  used,  placed  along  the  anterior  and  posterior 
sides  of  the  injured  member;  these  were  secured  by  tying.  The 
dusky  surgeons  said  that  the  juice  of  the  bark  penetrated  to  the 
seat  of  fracture  and  effected  its  healing.  In  the  Bundela  race, 
when  a  limb  was  fractured,  the  old  people  gathered  round  and 
bound  it  up  with  tea-tree  bark  to  keep  it  warm  and  comfortable. 
Around  this  was  fastened  string  made  from  a  scrub-tree  like  the 
Black  Kurrajong,  Yap  par,  of  the  Inland  Plains  {Sterculia  diver  si- 
folia,  G.  Don).  Thus  it  was  secured  until  union  had  taken  place. 
The  bark  of  the  Australian  tea-trees  or  paper-bark  trees 
(Melaleuca,  Gallistemon,  and  Leptosperinum)  lends  itself  admirably 
to  extemporaneous  use  in  setting  broken  bones — placed  outside 
in.  The  inner  layers  are  sufficiently  firm  to  secure  imuiobility  of 
the  fragments,  while  the  soft  exfoliating  outer  layers  form  an 
excellent  padding,  as  recommended  by  Dr.  AVoodward  in  his 
Railway  Ambulance  Handbook.* 

The  bite  of  the  centipede  and  spider,  and  sting  of  the  scorpion, 
were  treated  by  vigorous  sucking  until  the  blood  was  withdrawn, 
and  with  it  the  poison.  The  nip  of  the  powerful  soldier-ant  was 
not  deemed  important  enough  to  require  interference,  whereas 
the  sting  of  the  English  bee  {Boo)  was  considered  highl}^  dangerous 
and  often  fatal.  The  imported  bee  was  very  "  saucy,"  and  should 
be  robbed  of  its  honey  only  by  night  (Bundela). 

Snakes  of  the  most  venomous  kinds  are  common  throughout 
the  whole  district,  but  the  natives  rarely  die  from  the  effects  of 
their  bites.  A  Yukumbul  female  told  me  that  they  were  seldom 
bitten,  and  that  she  had  never  heard  of  a  fatality  arising 
from  such  a  cause.  It  is  said,  however,  that  an  aboriginal  at 
Oban     succumbed    to    the   venom    of    a    tiger    snake,   Erkullah 

*  Third  Edition,  p.  100. 


BY    JOHN    MACPHERSON.  645 

(j^ofechis  scntatiLs,  Peters).  The  Ngarrabul  natives  used  to  cut 
the  skin  at  the  site  of  the  wound  until  it  bled  freely.  Then  all 
the  doctors  sucked  it.  No  ligature  was  applied.  The  Yukumbul 
Blacks,  however,  in  addition  to  sucking,  applied  above  the  wound 
a  ligature  of  the  skin  of  Koohi,  the  opossum.  The  late  Dr. 
George  Bennett,  in  his  'Gatherings  of  a  Naturalist'  (p.  275), 
gives  an  account  of  the  measures  adopted  by  the  Clarence  River 
natives  in  combating  the  effects  of  snake  venom — scarification, 
free  bleeding,  and  keeping  the  patient  running  about  until  the 
effects  of  the  poison  had  passed  away.  The  Ngarrabul  Blacks 
say  that  Bri-pryn,  the  Soldier  Bird  {Myzantha  garrnla,  Lath.), 
finds  snakes,  proclaiming  their  presence  b}'  an  outburst  of  noisy 
clamour. 

In  the  olden  days  poisoned  spears  were  in  use.  The  men 
covered  them  with  the  melted  resin  of  the  Grass  Tree,  Burr-hurr 
{Xanthorrhoea  sp.).  They  were  then  passed  on  to  the  women, 
who  alone  knew  how  to  impregnate  them  with  the  powerful 
poison.  A  wound  with  such  a  spear  was  generally  mortal  unless 
treated  by  a  woman.  Men  were  powerless  to  cope  with  such 
cases.  A  Yukumbul  female  informed  me  that  in  her  tribe  spears 
were  poisoned  with  a  substance  obtained  from  the  mountains.  It 
is  related  of  the  Narrinyeri  tribes  of  South  Australia  that  they 
were  wont  to  poison  a  kind  of  dagger  by  leaving  it  in  a  putrid 
human  corpse  for  some  weeks,  and  then  wrap  up  the  point  of  the 
dagger  in  some  hair  or  feathers  soaked  in  the  fat  of  a  decaying 
corpse.  This  was  termed  Neilyeri  or  the  poison  revenge.*  Poison 
in  any  other  form  was  quite  unknown  to  the  Narrinyeri. f  An 
old  Ngarrabid  tribesman,  on  the  contrary,  was  in  great  dread  of 
poison,  Mittie.  He  said  the  Inverell  blacks  (who  obtained  it 
originally  from  Queensland)  keep  it  in  a  small  vessel.  It  is  like 
a  scent,  and  is  in  use  to  the  present  day.  If  the  possessor  be  at 
enemity   against   anyone,   his   victim's    fate    is    sealed.      In    his 

*Rev.  Geo.  Taplin  in  the  '  Native  Tribes  of  South  Australia,'  p.  29,  et  seq. 
Mr.  C.  Hedley  (Proc.  Eoy.  Soc.  Qsld.,  v.)  speaks  of  the  Port  Curtis  Blacks 
employing  the  milky  juice  of  Exccecaria  agallocha  to  poison  their  spears. 
fTaplin,  loc.  cit. ,  p.  47. 


646  NGARRABUL    AND    OTHER    ABORIGINAL    TRIBES, 

absence  from  camp  some  of  the  poison  is  sprinkled  on  his  rug. 
When  he  returns  to  sleep,  it  is  his  last  eternal  slumber.  He  dies 
"all  puffed  out."  My  informant  could  tell  me  nothing  more 
about  this  lethal  drug.  Even  the  police,  he  said,  are  mystified. 
Current  rumour  certainly  in  this  district  ascribes  the  death  of 
aboriginals  to  some  mysterious  poison  employed  by  hostile  natives. 
It  is  said  that  in  earlier  days  also  the  natives  were  in  great  terror 
of  being  thus  put  to  death  by  revengeful  Blacks. 

As  regards  ritual  or  ceremonial  and  "ornamental"  surgery, 
I  saw  no  natives  with  the  septum  nasi  pierced.  The  Ngarrabul 
Blacks  told  me  that  neither  circumcision  nor  knocking  out  the 
incisor  teeth  was  practised  in  their  tribe,  nor  was  that  remarkable 
rite,  urethrotomy  or  mutilation  of  the  penis,  described  amongst 
other  Australian  people.  Scarification  of  the  body  (erroneously 
termed  "tattooing"),  however,  was  performed  after  attaining 
adult  years.  It  was  entirely  optional,  and  members  of  either  sex 
could  be  so  adorned  if  they  felt  disposed — an  advantage  that 
some  at  least  did  not  avail  themselves  of.  The  form  and  distri- 
bution of  the  lines,  etc.,  made  by  this  operation  differed  materially 
in  different  tribes,  and  travellers  say  that  many  tribes  could  be 
thus  easily  distinguished  by  the  bodily  markings  of  their  mem- 
bers.* I  saw  an  old  Oban  native  with  extensive  cicatrices  upon 
the  skin  of  his  chest  (back  and  front),  but  not  upon  his  abdomen. 
One  vertical  scar  lay  over  the  upper  sternum.  On  either  side  of 
the  chest  in  front,  below  the  level  of  this  one,  were  four  scars; 
those  on  the  left  being  almost  horizontal,  while  those  on  the  right 
were  shorter  and  directed  obliquely  downwards  and  inw^ards, 
thus : — 

I  The  scars  were  all  pale  and  atrophic,  and  neither 

^      upon  these  nor  on  those  resulting  from  injuries  was 

^      ^=1:^     there  any  elevation  of  false  keloid.      In  fact  we 

are  told   that  wounds  require  to   be   subjected  to 

special  measures  to  ensure  the  formation  of  such  cicatrical  over- 


*  Bulmer  in  Biough  Smythe's  'Aborigines  of  Victoria,'  Vol.  i.,  p.  295. 
J.  M.  Davis,  ibid.  Vol.  ii.,  p.  313. 


BY    JOHN    MACPHERSON.  647 

growth—  described  intpv  alios  by  Drs.  Milne  Robertson*  and  W. 
Wyatt.t 

Obstetrics  in  Ngarrabul  were  quite  simple.  The  female  rela- 
tives, sister,  mother  or  grandmother,  looked  after  the  patient  and 
her  baby.  The  men  knew  but  little  about  such  matters,  and  held 
aloof. 

8uch,  then,  was  the  practice  of  medicine  and  surgery  among 
these  primitive  people.  Drunkenness,  with  its  attendant  evils, 
exposure  to  wet  and  the  cold  of  night,  tubercle,  hydatids,  venereal 
disease,  the  diminished  bodily  vigour  resulting  from  occupying  and 
sleeping  in  damp,  dark  and  ill-ventilated  hovels,  incident  upon 
their  civilisation,  have  all  combined  to  destroy  the  tribes,  and 
soon  the  few  remnants  will  be  gathered  to  their  fathers  and  we 
shall  know  them  no  more. 


*  Report  on  certain  Customs  of  the  Aborigines  of  Western  Australia,  p.  6. 
t  Adelaide  and  Encounter  Bay  Tribes  in  '  Native  Tribes  of  South  Austra- 
lia,' p.  163. 


648 


NOTES  ON  AUSTRALIAN  RHOPALOCERA- 
LYC.EyiD.E.     Part  II. 

By  G.  a.  Waterhouse,  B.Sc,  B.E. 

This  Part  contains  observations  on  the  remainder  of  the  new 
Lycaenidi\3  obtained  by  Mr.  R.  E.  Turner  during  the  present 
year  in  North  Queenshind.  I  propose  in  my  next  part  to  deal 
fully  with  each  genus  and  di-aw  up  a  comparison  of  the  species 
placed  in  their  respective  genera. 

PsEUDONOTis,  Druce. 

Ann.  Mag.  Nat.  Hist.  (6),  xii.,  p.  252,  1894. 

This  genus  is  allied  to  Thysonotis,  Hiibn.  (it  has  often  been 
known  as  Danis,  Fabr.).  The  insect  hereunder  described  appears 
to  belong  to  this  genus;  it  is  certainly  unlike  any  other  form 
found  in  Australia. 

PsEUDONOTIS    TURNERI,   n.Sp. 

(J.  Uppersid  e. — Foretviny  with  costa  nearly  straight,  apex 
acute,  outer  margin  nearly  straight,  inner  margin  straight. 
Hindivinc/  with  costa  arched,  outer  margin  slightly  concave  in 
centre,  anal  angle  greatly  produced,  inner  margin  nearly  straight; 
t?vo  tails  are  present  on  the  hindwing.  Both  wings  white  with 
broad  black  margins;  base  black;  the  broad  black  margin  occupy- 
ing almost  the  whole  of  the  cell  of  the  forewing  and  extending  in 
an  even  band  along  the  outer  margins  of  both  wings  to  the  inner 
margin  of  the  hindwing,  widest  near  the  anal  angle.  Between 
the  submedian  anil  first  median,  and  first  and  second  medians 


BY    G.    A.    WATERHOUSE.  649 

the  anal  angle  is  marked  with  two  very  black  conical  spots  which 
are  bounded  externally  with  white  lines  and  internally  with  pale 
blue  crescents.  Along  the  outer  margin  are  faint  white  lines, 
and  above  these  are  a  few  pale  blue  scales.  The  tails  are  nearly 
^  inch  long,  black  tipped  with  white;  and  arise  from  the  sub- 
median  nervure  and  first  median  nervule.  The  pale  blue  scales 
on  the  hindwing  only  appear  so  when  viewed  directly,  but  when 
the  insect  is  held  obliquely  it  will  be  seen  that  the  blue  scales  are 
very  extensive  and  are  of  a  very  brilliant  electric  blue  colour. 
Cilia  black. 

Under  sid  e. — The  scheme  of  colouration  is  as  on  upperside, 
but  the  white  is  replaced  by  cream.  The  marginal  blue  markings 
are  more  extensive  and  do  not  change  colour  when  viewed  in 
different  directions.  These  pale  blue  scales  are  represented  on 
the  forewing  by  a  submarginal  narrow  band,  external  to  which  is 
a  pale  line,  and  in  the  hindwing  by  arches  between  each  of  the 
nervules,  the  legs  of  these  arches  being  continued  down  the 
nervules,  while  between  the  median  nervules  and  submedian 
nervure  are  four  blue  lunules;  the  margin  has  a  distinct  white 
line  running  along  it. 

9.  Costa  more  arched  ;  apex  less  acute  ;  outer  margin  more 
convex  than  in  the  forewing  of  the  male,  and  in  the  hindwing  the 
anal  angle  is  less  produced  and  the  tails  are  shorter.  Otherwise 
as  in  the  male. 

In  the  available  specimens  the  female  is  smaller  than  the  male. 

Expanse,  g  30-35  mm;  9  22-29  mm. 

This  species  is  probably  the  Australian  form  of  F.  danis,  Feld.,"^ 
from  which  it  differs  in  having  much  more  white  on  the  upper- 
side.  Felder's  figure  shows  very  little  white  on  the  upperside, 
while  in  the  species  described  the  white  is  of  larger  extent  to  the 
black.  It  also  resembles  P.  tniJo,  G.  Smith  ct  Kirb}^!  in  general 
appearance,  but  that  species  is  described  and  figured  without,  or 
at  least  with  very  short,  tails  (these  may  have  been  broken  off), 
and  the  upperside  is  without  the  blue  scales  of  the  hindwing. 

*  Reise  Novara,  Lep.  ii.,  p.  240,  11.  273,  t.  30,  figs.  12-13  (1865). 
t  Rhop.  Exot.  Lyceen.  orient.,  xi.,  tigs.  1  and  2,  1897. 


650  AUSTRALIAN  RHOPALOCERA  :  LYC.ENID^'E,  II. , 

It  is  also  allied  to  Myriita  derpiha^  Hew.,"^  especially  on  the 
underside. 

The  larva  is  of  a  brilliant  green,  and  was  found  by  Mr.  R.  E. 
Turner  feeding  on  a  species  of  Orchid  ( Vanda  section).  The  pupa 
is  also  brilliant  green  with  no  markings,  but  a  depression  behind 
the  wings. 

I  am  indebted  to  Mr.  R.  E.  Turner  for  specimens  of  this 
insect  from  Kuranda  near  Cairns. 

Philiris,  Rober. 
Tijdschr.  Ent.  xxxiv.,  p.  317;  Exot.  Schmett.  Theilii.,  p.  273, 1892. 

The  type  of  this  genus  is  Thecla  ilias,  Feld.,  and  as  both  the 
following  species  are  closely  allied  to  that  species,  I  have  placed 
them  in  this  genus,  leaving  aside  the  question  for  the  present  as 
to  whether  it  should  be  sunk  under  Candalides,  Hiibn.,!  the  type 
of  which  is  C.  xanthospilos,  Hiibn. 

Philiris  Kamerung^,  sp.n. 

(^.  Forewing  with  costa  nearly  straight,  apex  acute,  outer 
margin  straight,  inner  margin  straight.  Hindwing  with  outer 
margin  entire. 

Up  per  side. — Foreioing  with  costal  margin  black,  apex 
broadly  black,  outer  margin  black.  The  black  margin  is  bounded 
by  the  subcostal  nervure  until  the  end  of  cell,  then  crosses  the 
end  of  cell  towards  the  outer  margin  and  becomes  less  extensive 
to  the  angle.  The  rest  of  the  wing  purple,  but  viewed  in  some 
directions  blue,  except  a  small  white  patch  in  the  centre  of  the 
wing  external  to  the  cell  in  the  region  of  the  median  nervules. 
Hindwing  with  a  narrow  black  outer  margin,  rest  of  wing  purple 
except  the  apical  angle,  which  is  white  above  the  subcostal 
nervure.     Abdominal  fold  white.     Cilia  white. 


Diurnal  Lepid.     Plate  iii.  b.  (Supplt.),  figs.  101,  102  (cJ),  1878. 
t  Verzeichniss  bekannt.  Schmett.  20  (1816). 


BY    G.    A.    WATERHOUSE.  651 

Underside  pure  silky  white  with  very  faint  black  termina- 
tions to  the  nervules  on  the  outer  margin.  One  small  black  spot 
midway  near  the  inner  margin  of  hindwing. 

9.  Forewing  with  costa  nearly  straight,  apex  pointed,  but  outer 
margin  convex.  Outer  margin  of  hindvnng  more  rounded  than 
in  the  male. 

Upper  side. — For  p.iving  h\3^ck  and  white,  with  a  faint  sub- 
basal  blue  suffusion;  the  black  costal  border  is  wider  than  in  the 
male  and  occupies  all  the  cell,  and  extends  somewhat  along  the 
inner  margin  (in  one  specimen  it  extends  the  whole  length  of  the 
inner  margin),  thus  leaving  an  almost  circular  central  white  spot. 
This  central  white  spot,  which  is  wholly  below  the  cell,  when 
viewed  directly,  is  white,  Avith  the  basal  portion  slightly  blue, 
but  when  viewed  obliquely  has  a  beautiful  pale  silvery -blue  sheen. 
Hindwing  brown  with  apical  angle  white,  the  white  apical  patch 
in  some  specimens  extending  downwards  through  the  cell  to  the 
median  nervule  and  not  having  the  blue  sheen  as  in  the  forewing. 
Cilia  white.     Abdominal  fold  white. 

Underside  as  in  the  male,  but  the  terminations  of  the 
nervules  are  more  marked. 

Expanse,  $  23  mm.;  9  24-26  mm. 

This  insect  is  allied  to  P.  ilias,  Felder,  but  the  white  patches 
in  both  sexes  are  a  ready  means  of  identifying  it. 

Log. — Cairns  District. 

Philiris  Kurand^,  n.sp. 

^.  Foreiving  with  costa  very  straight,  apex  very  acute,  outer 
margin  straight.      Hindwing  with  anal  angle  slightly  produced. 

Uppersid  e. — Foreiuing  dark  purple  with  black  costal  border, 
apex  broadly  black  and  inner  margin  less  broadly  black.  Hind- 
wing  with  costal  border  broadly  black,  outer  margin  narrowly 
black,  the  rest  of  wing  bright  blue.  Abdominal  fold  grey.  Cilia 
white. 

Underside  silky  white,  terminations  of  veins  black,  but  710 
black  spot  near  inner  margin  of  the  hindwing  as  in  P.  KamerungcH 
and  P.  ilias. 


652  AUSTEALIAN  RHOPALOCERA  :  LYC.EXID^E,  II., 

9-  Forewing  with  costa  straight,  apex  not  nearly  so  acute  as 
in  the  male;  outer  margin  convex.  Uindwing  with  outer  margin 
much  more  rounded  than  in  the  male. 

Upperside. — Forewing  with  a  very  even  black  border 
extending  along  costa.  apex  and  outer  margin,  where  it  is  broadest, 
the  rest  of  wing  dull  blue.  Hindiv'ing  as  in  forewing,  with  the 
borders  of  about  the  same  size.  Cilia  white,  the  nervules  of  the 
hind  wing  marked  with  black. 

Underside  as  in  the  male. 

Expanse,  $  26-30  mm.;  9  29-31  mm. 

It  is  noticeable  that  in  the  male  of  this  species  the  forewing 
is  purple  and  the  hindwing  blue,  while  in  P.  ilias  both  wings  are 
purple;  also  it  is  a  larger  insect  than  P.  ilias. 

Log. — Cairns  District. 

Philiris  ilias,  Felder. 

Thecla  ilias,  Felder,  Sitzb.  Akad.  Wiss.  Wien,  Math.  Nat.  01. 
xl.  p.  454,  n,  22  (1860).  Pseudodipsus  ilias,  Miskin,  Syn.  Cat. 
Rhop.  Aust.  p.  67  (1890);  P.  innotatus,  Misk.,  Ent.  Mo.  Mag. 
1874,  p.  165. 

(J.  Foreiving  with  costa  nearly  straight,  apex  acute,  outer 
margin  straight,  inner  margin  straight.  Hindwing  with  outer 
margin  entire. 

Upperside. — Forewing  with  costal  margin  black,  apex 
broadly  black  and  outer  margin  black,  the  rest  of  wing  purple. 
Hindwing  with  outer  margin  black,  the  rest  of  wing  purple. 
Cilia  white. 

Underside  pure  silky  white,  with  a  fine  black  edge  more 
noticeable  at  the  terminations  of  the  nervules.  A  small  black 
spot  near  the  middle  of  inner  margin  of  hindwing. 

9.  Forewing  with  costa  nearly  straight;  apex  pointed,  outer 
margin  very  convex,  outer  margin  of  hindwing  more  rounded 
than  in  the  male. 

Uppersid  e. — Forewing  brown  with  a  pale  blue  area  situated 
between  the  median  and  submedian  nervures  and  extending  from 


BY    G.    A.    WATERHOUSE.  653 

base  to  a  little  beyond  the  middle.  Hindwing  brown,  with  white 
cilia. 

Underside  as  in  the  male.     Expanse,  ^  25  mm. ;  9  25-30 mm. 

Felder's  type  was  from  Amboina.  I  have  not  been  able  to  see 
a  specimen  from  that  locality,  but  Miskin  himself  considered  his 
species  synonymic  with  Felder's,  so  I  allow  Felder's  name  to  stand. 
It  may  be  that  our  insect  is  not  identical  with  the  Amboina 
species,  as  I  find  Mr.  Grose  Smith  has  described  a  species, 
Holochila  nitens*  from  N.  Queensland  If  this  is  the  case,  P. 
innotatus  has  certain  priority,  and  P.  nitens  must  sink  if  it  is,  as 
I  think,  the  same  insect  as  the  one  under  consideration.  My 
specimens  from  Cape  York  agree  very  well  with  the  figure  of  the 
male  of  P.  yiitens,  and  they  are  not  markedly  different  from 
Cairns  or  Brisbane  specimens. 

Loc. — Richmond  River,  N.S.W.  (G.A.W.)  to  Cape  York,  Q. 


*  G.  Smith,  Nov.  Zool.  v.  p.  107  (1898).     G.  Smith  &  Kirby,  Khop.  Exot. 
Lye.  orient.,  1899,  pi.  xx, 
43 


654 


ON  THE  OCCURRENCE   OF   MOXOGRAPTUS  IN   NEW 
SOUTH  WALES. 

By  T.  S.  Hall,  M.A. 

(Communicated  by  John  Mitchell.) 

The  occurrence  of  graptolites  in  the  Silurian  of  Bowning  and 
Yass  has  been  ah-eady  recorded  by  Mr.  John  Mitchell,*  but  the 
record  has  been  since  referred  to  as  doubtful,  and  it  is,  therefore, 
with  much  pleasure  that  I  accepted  Mr.  Mitchell's  offer  to  allow 
me  to  examine  the  specimens  on  which  the  record  was  based. 
They  are  undoubted  examples  of  Monograptus,  a  genus 
which,  it  is  hardly  necessar}^  to  say,  is  confined  to  the 
Silurian  —  that  is,  the  Upper  Silurian  of  the  State 
Geological  Surveys — though  unfortunatel}'  the  condition 
of  the  specimens  does  not  permit  of  specific  naming. 

Monograptus  sp. 

Hydrosome  straight  for  the  greater  part  of  its  length, 

slightly  incurved  proximally.      Maximum  width  about 

1-7  mm.;  length  of  a  slightly  imperfect  specimen  12  mm. 

This  is  nearly  the  total  length,  as  only  a  small  part  of  the 

proximal   end   is  wanting.     There  does  not  seem  to  be 

any  distal  prolongation  of  the  virgula.      The  proximal 

end  of  the  polypary  is  not  visible  in  any  of  the  four 

specimens  which  lie  close  together  on  the  stone.     The 

^^'    ■     thecjie  number  about  10  or  11  in  1  cm.,  and  are  inclined 

to  the  axis  at  3")°.      As  far  as  can  be  seen,  they  are  about  twice  as 

Fig.  1. — Imperfect  specimen  of  Monograptus,  allied  to  M.  dnhlns,  Suess  { x  5). 

*  Proc.  Linn.  8oc.  N.S.  Wales  (2),  Vol.  i.,   1SS6,  p.  577,  and  (2)  Vol.  ill. 
(1888-1889),  p.  150. 


BV    T.    S.    HALL.  655 

long  as  wide.  Apertures  at  right  angles  to  the  axis'of  the  theca, 
and  slightly  everted  at  their  outer  angle.  The  character  of  the 
matrix  does  not  allow  the  presence  or  absence  of  a  spine  to  be 
definitely  asserted. 

The  imperfection  of  the  specimens  in  the  sicular  region  prevents 
their  absolute  identification,  but  they  apparently  belong  to  the 
group  typified  by  M.  duhius,  which  ranges  through  almost  the 
whole  of  the  Lower  Ludlow  and  Wenlock  in  Britain. 

Loc. — Belle  Vale,  Yass. 

Horizon. — Lower  Trilobite  Bed,  Bowning  Series  (  =  Wenlock, 
Upper  Silurian). 


656 


NOTES    AND    EXHIBITS. 


Mr.  Fred.  Turner  exhibited  from  the  Tweed  River  district  of 
New  South  Wales,  three  suspected  poisonous  plants,  viz.: — 
Indigofera  mistralis,  Willd.,  Phyllanthus  Gasstrcemii,  Muell., 
and  Brei/nia  ohlongifolia,  Muell.  These  plants  have  been  sus- 
pected of  causing  red  water  in  cattle,  and  in  addition,  the  firsts 
named  is  thought  to  bring  on  premature  labour  in  stock.  In 
contirmation  of  this  the  Hon.  P.  L.  C.  Shepherd,  M.L.C., 
informed  Mr.  Turner  that  when  he  first  went  to  live  at  Colo  Vale 
he  took  with  him  several  cows  in  calf  and  mares  in  foal,  and  after 
the  animals  had  eaten  of  the  Indigofera  australi^  they  slipped  the 
foetus.  Singular  to  saj",  that  when  the  stock  had  been  some 
time  on  the  estate  they  never  suffered  in  a  similar  way,  although 
the  indigo  still  grew  fairly  plentifully  there.  These  plants  are 
included  in  Turner's  list  of  supposed  poisonous  plants  of  Australia. 
He  also  showed  a  photograph  of  "  Insect  Mimicry,"  taken  direct 
from  the  natural  objects  by  Andre  k,  Sleigh's  special  three-colour 
process;  and  forwarded  to  the  exhibitor  by  Sir  Henr}^  Trueman- 
Wood,  M.  A.,  Secretary,  Society  of  Arts,  London. 

Mr.  Baker  exhibited  specimens  of  the  Symphcos  described  in 
his  paper. 

Mr.  Froggatt  showed  specimens  of  a  lamellicorn  beetle,  Pento- 
don  australifi^  Blackb.,  from  Sackville  Reach,  Ha wkesbury  River, 
where  the  species  of  late  has  done  much  damage  to  maize  seed  in 
the  ground,  as  well  as  to  the  young  corn;  under  ordinar}'  circum- 
stances the  insect  frequents  the  huW  gYa,%s!,  (^Paspaluiii)  of  the  flats, 
but  in  the  absence  of  floods  has  become  a  pest  to  the  farmer.  He 
also  showed  examples  of  another  beetle  of  the  same  family, 
Chiroplatijs  sp.,  which  has  taken  to  devouring  potatoes  and 
tomatoes  in  the  Windsor  and  Richmond  districts. 

Mr.  Stead  narrated  some  recent  experiences  with  a  very  large 
shoal  of  "salmon  "  (Arr-ipis  salar,  Rich.)  in  Port  Hacking,  Avhich 
comprised  prol^abh'  some  hundreds  of  thousands  of  individuals. 


NOTES    AND    EXHIBITS.  657 

Mr.  Hedley  exhibited  specimens  of  polyzoa  and  foraminifera, 
recently  dredged  in  HI  fathoms,  12 J  miles  to  the  east  of  Cape 
Byron,  by  Mr.  Gerald  Halligan,  F.G.S.  The  exhibit  seemed  to 
show  that  a  reef  rock  was  in  process  of  formation  similar  in 
character  to  the  polyzoal  limestone  at  Mount  Gambler,  S.A. 

Mr.  Gurney  showed  two  specimens  of  the  stridulating  moth, 
Hecatesia  fenestrata,  Boisd.,  recently  captured  by  him  at  Narra- 
been ;  and  he  remarked  that  Mr.  Waterhouse  had  also  noticed  it 
at  Waverley. 

Mr.  Fletcher  took  the  opportunity  to  point  out  that  at  the 
Society's  Meeting  on  March  31st,  1897,  he  exhibited  on  behalf 
of  Mr.  C.  T.  Musson,  specimens  of  a  stridulating  Agarista, 
A.  macleayi,  Koch  (Proceedings,  1897,  p.  44)  ;  and  that  recently, 
and  quite  accidentally,  he  had  met  with  a  much  earlier  record  of 
the  stridulating  habit  of  this  species,  in  Trans.  Ent.  Soc.  N.  S. 
Wales,  i.  Proc,  p.  xxvi.  (Nov.  1863). 


658 


DONATIONS    AND    EXCHANGES 

Received  during  the  period  November  28th,  1901, 
to  November  27th,  1902. 

(From  the  respective  Societies,  cjC'c,  unless  otherwise  mentioned.) 


Adelaide. 

Department  of  Mines — 

Record  of  the  Mines  of  S.  Australia  : 

Tarcoola  and  the  N.W.  District.      By  H.   Y.   L.  Brown, 

Government  Geologist. 
Handbook  of  Mining,  with  Maps  (1901). 
Public  Librahy,  Museum,  (fee,  South   Australia — 
Report  of  the  Board  of  Governors  for  1900-01. 

Royal  Society — 

Transactions,  Proceedings  and  Pteport.    xxv.  2  (1901). 
Transactions,    xxvi.  1  (1902). 
Woods  and  Forests  Department — 

Annual  Progress  Report  upon  State  Forest  Administration 
in  S.  AustraHa  for  the  Year  1900-01.  By  W.  Gill,  F.L.S., 
F.R.H.S.,  Conservator  of  Forests. 

Amsterdam. 

KONINKLIJKE    AkADEMIE    VAN    WeTEXSCHAPPEN  — 

Jaarboek,  1895,  1899,  1900. 

Proceedings  of  the  Section  of  Sciences,    iii.  (1901). 

Verhandelingen.  2"«  Sectie.  iv.  7-9  (1895-96);  v.  1-3  (1896); 
vii.  4-6  (1900-01). 

Verslagen  van  de  Zittingen  der  Wis-  en  Natuurkundige 
Afdeeling.    iv.  (1895-96). 

Verslag  van  de  Gewone  Vergaderingen  der  Wis-  en  Natuur- 
kundige Afdeeling.    ix.  (1900-01). 


DONATIONS    AND    EXCHANGES.  659 

Antwerp. 

SOCIETR    ROYALE    DE    GeOGRAPHIE    d'AnVERS  — 

Bulletin,    xxv.  4(1901);  xxvi.  1-2(1902). 

Auckland,  N.Z. 

Auckland  Institute  and  Museum — 
Report  for  1901-02  (1902). 
Transactions  of  the  N.Z.  Institute,    xxxiii.,  xxxiv.  (1901-02). 

Baltimore. 
Johns  Hopkins  University  — 

Hospital  Bulletin,    i.-vii.,  Nos.  1-67,  except  2,  9,  12  and  19 

(1889-96);  xii.,  127-129(1901);  xiii.,  130-138  (1902). 
Hospital  Reports,  i.,  iii.-vii.,  viii.,1-9,  ix.,  x.  1-2(1896-1901). 

Basel. 

JSTaturforschende  Gesellschaft — 

Verhandlungen.    xiii.  2-3  (1901-02);  Anhang  zum  xiii.  Band 
("Zur   Errinerung  an   Tycho   Brahe,  1546-1601");    xiv. 
(1901). 
Namenverzeichness    u.    Sachregister   der    Bande  vi.  bis  xii. 
(1875-90). 

Batavia. 
KoNiNK.  Natuurk.  Vereeniging  in  Nederl.-Indie — 
Natuurkundig  Tijdschrift.    Deel  Ixi.  (1902). 

Berlin. 

Archiv    f.    Naturgeschichte.    1895,  ii.  3  (1897) :  1898,    ii. 
2,  J  (1901):   1899,  ii.  2,  J(1902):    1901,  i.  1,  u.  3(1901);    ii. 
2,  1(1902):   1902,  i.  2(1902). 
Berliner  Entomologischer  Verein— 

Berliner  Entomologische  Zeitschrift.    xxxv.  2  (1890);  xxxvi. 
2(1891);  xxxvii.-xliv.  (1892-99);  xlri.  4  (1901);  xlvii.  1-2 
(1902). 
Gesellschaft  f.  Erdkunde — 

Verhandlungen.    xxviii.  4-10(1901). 
Zeitschrift.    xxxvi.  2-6(1901);  1902,  1-4. 


660  DONATIONS    AND    EXCHANGES. 

Berne. 

Naturforschende  Gesellschaft  in  Bern — 

Mitteilimgen  aus  dem  Jahre  1901.  Nr.  1500-1518(1902). 

Bonn. 

Naturhistorischer  Verein  in  Bonn — 
Verhandlungen.    Iviii.  1-2(1901-02). 

Sitzungsberichte  der  Niederrheinischen  Gesellschaft  f.  Natur- 
und  Heilkunde  zu  Bonn.    1901,  1-2  (1901-02). 

Boston. 

American  Academy  of  Arts  and  Sciences — 

Proceedings,    xxxvi.  29(1901);  xxxvii.  1-20(1901-02). 
Boston  Society  of  Natural  History — 

Proceedings,    xxix.  15-18;  xxx.  1-2(1901). 

Bremen. 

Naturwissenschaftlicher  Yerein — 
Abhandlimgen.    xvii.  1  (1901). 

Brisbane. 

Colonial  Botanist  of  Queensland — 

The  Queensland  Flora.     Part  iv.  [Hygrophyllacece  to  Elceag- 
nace(€,) ;    Part  v.  (^LoranthacecE  to   Lernnacece).      By  F.   M. 
Bailey,  F.L.S.,  Colonial  Botanist  (1901-02). 
Department  of  Agriculture — 

Annual  Report  for  the  Year  1900-01. 

Queensland  Agricultural  Journal,     ix.  6  (1901);  x.  1-6;  xi. 
1-5(1902). 
Geological  Survey — 

Annual  Progress  Report  of    the  Geological  Survey  for  the 

Year  1901  (S.P.  175;  1902). 
Bulletins.    Nos.  13-18  (1901-02). 

Report  on  the  Geology  of  the  Dawson  and  Mackenzie  Rivers, 
with  Special  Reference  to  the  Occurrence  of  Anthracitic 
Coal.     By  B.  Dunstan,  F.G.S.  (S.P.  155;  1901). 
On  Queensland   Mining  and  Milling   Practice.      By  J.  M. 
MacLaren,  B.Sc.  (156;  1901;. 


DOXATIONS    x\ND    EXCHANGES.  661 

Geological  Society,  Brisbane  (continned) — 

On  the  Jordan  Creek  Gold  Field.      By  L.  C.  Ball,  B.E.  (157; 

1901). 
On  the  Hamilton  and  Coen  Gold  Fields.     By  L.  C.  Ball,  B.E. 

(163;  1901). 
On    the    Cardigan,  Queensland,  Tin    Syndicate    Properties, 
Thompson's  Creek,  and  the  Coolgarra  Federal  Tin   Cor- 
poration,  Limited,   Leases,  Herberton   Mineral    District. 
By  W.  E.  Cameron,  B.A.  (165;  1901). 
Fourth   Report   on    the  Gympie  Gold  Field,  having  special 
reference  to  the  Inglewood  Dyke  and  the  Eastern  Leases. 
By  W.  H.  Bands,  A.R.S.M.,  F.G.S.  (166;  1901). 
On  the  Burrum  Coal  Field.     By  W.  H.  Rands.     (Reprint 
of  No.  24:,  with  Corrections  and  Additions.      170;  1901). 
On  the  Coal  Beds  on  Waterpark  Creek,  near  Port  Clinton. 
By  W.  E.  Cameron,  B.A.  (174;  1902). 
HoxME  Secretary's  Department — 

North  Queensland  Ethnography.     Bulletin  No.  3  ("Food:  its 
Search,  Capture  and  Preparation"),  (1901);  No.  4 ("Games, 
Sports  and  Amusements"),  (1902).     By  W.  E.  Roth,  B.  A., 
M.R.C.S.,  kc. 
Royal  Geographical   Society  of  Australasia,  Queensland 
Branch — 
Queensland  Geographical  Journal,    xvii.  (1902). 
One   Separate  :    "  Summary  of    the  Geographical  Work  of 
Mr.  J.  P.  Thompson"  (January,  1901). 
Royal  Society  of  Queensland — 
Proceedings,    xvii.  1  (1902;. 

Brussels. 

Academie  Royale  de  Belgique  — 

Annuaire.    1900,  1901,  1902. 

Bulletin  de  la  Classedes  Sciences.  1899,1900,  1901,  1902,  1-5. 
Societe  Entomologigique  de  Belgique — 

Annales     xlv.  (1901). 

Memoires.    viii.  (1901). 


662  DONATIONS    AND    EXCHANGES. 

Brussels  (continued) — 

SOCiETE  ROYALE  LiNNEENNE  DE  BrUXELLES 

Bulletin,    xxvii.  2  &  4-9  (1901-02). 

SOCIETE  RoYALE  MaLACOLOGIQUE  DE  BeLGIQUB  — 

Annales.    xxxv.  1900(1901). 

Budapest. 

Museum  National  Hongrois — 

Termeszetrajzi  Fiizetek.    xxiv.  1-4(1901). 

Buenos  Aires. 

MusEo  Nacional  de  Buenos  Aires — 
Comunicaciones.    i.  10(1901). 

Buffalo. 

Buffalo  Society  of  Natural  Sciences  — 
Bulletin,    vii.  1  (1901). 

Calcutta. 

Asiatic  Society  of  Bengal — 
Journal.    Ixx.  Part  i.  No.  2  and  Extra  Nos.  1-2(1901-02); 
Part  a.  No.  2  (1902);  Part  Hi.  No.  2  (1902):  Ixxi.  Part  ii. 
1  (1902);  Part  Hi.  1  (1902). 
Proceedings.    1901,  Nos.  ix.-xi.  (Nov,  1901-Feb.  1902);  1902, 
Nos.  i.-v.  (March-Jul}^). 
Geological  Survey  of  India — 
General  Report  for  1901-02. 

PalcTeontologia  Indica.    New  Series,    ii.  1(1902).    T.p  ,  ikc,  to 
Vols.  iv.  of  Series  x.,  iii.  of  Series  xv.,  and  i.  of  Series  xvi. 
(1901-02). 
Memoirs,  xxx.  3-4  (1901);  xxxi.  2-3  (1901);  xxxii.  1-2  (1901); 
xxxiii.  2(1901);  xxxiv.  1  (1901). 

Indian  Museum — 

Annual  Report,  1900-1901. 

*'  An  Account  of  the  Indian  Triaxonia  collected  by  the 
R.I.M.S.S.  Investigator."  By  Prof.  F.  E.  Schulze  (4to., 
1902). 


DONATIONS    AND    EXCHANGES.  663' 

Cambridge. 

Cambridge  Philosophical  Society — 
Proceedings,    xi.  4-6  (1902). 
Transactions,    xix.  2  (1902). 

Cambridge,  Mass. 

American  Naturalist:    xxxv.  419-420  (1901);  xxxvi.  421-429 

(1902).      From  the  Puhlishers. 
Museum  of  Comparative  Zoology  at  Harvard  College — 
Annual  Report  of  the  Keeper  for  1900-01. 
Bulletin,   xxxviii.  (Geological  Series,  v.)  .5-6;  xxxix.  1-3;  xl. 
1-2;  xli.  1  (1901-02). 

Cape  Town, 

Department  of  Agriculture,  Cape  of  Good  Hopk — 

Annual  Report  of  the  Geological  Commission.  1898,  1899, 

1900  (1900-01).      From  the  Director. 
Marine  Investigations  in  South  Africa.    Nos.  6  tfe  10  (1901); 
Report  of  the  Government  Biologist  for  the  Year  1900. 
From  the  Government  Biologist. 
South  African  Museum — 
Annals,    ii.  6-9  (1901-02). 
South  African  Philosophical  Society — 
Transactions,    xi.  4  (1902). 

Cherbourg. 

SOCIETE  NaTIONALE  DES  SCIENCES  NaTURELLES,   ifec. — 

Memoires.    xxxii.  (1901-02). 

Chicago. 

Chicago  Academy  of  Sciences — 

Bulletin,    ii.  3  (1900);  iv.  (Part  i.  of  Nat.  Hist.  Surv.)  (1900). 
Field  Columbian  Museum — 

Anthopological  Series,    ii.  5;  iii.  1-2(1901). 

Geological  Series.    1.9-10(1901). 

Report  Series,    ii.  1  (1901). 

Zoological  Series,    ii.  2;  iii.  4-5  (1901). 


i 


664  DONATIONS    AND    EXCHANGES. 

Christiania. 

Videnskabs-Selskabet  I  Christiania — 
Forhandlinger.    Aar  1900  (1901). 
Ski'ifter.    i.  Math.-naturvid.  Klasse.  1900,  5-7  (1900-01). 

Cincinnati. 

Cincinnati  Society  of  Natural  History — 

Journal,    xx.  1-2  (1901-02). 
Lloyd  Library — 

Bulletin.    No.  2  (1901)  [Reproduction  Series,  No.  2.] ;    No.  3 

(1902)[Mycological  Series,  No.  1.] 
Mycological  Notes.    Nos.  5-8  (1900-01). 

Columbus. 

Ohio  State  University^ 

Thirtieth  Annual  Report  of  the  Board  of  Trustees  for  the 
Year  ending  June  30,  1900.  Parts  1-2  (1900);  Thirty-first 
Annual  Report.    Parts  1-2  (1901). 

Copenhagen. 

Kongelige  Danske  Videnskabehnes  Selskab — 

Oversigt.    1902,  2-3 
Academie  Royale  des  Sciences  et  des  Lettres  de  Danemark- 

Bulletin.    1901,  4-6;   1902,  1. 
Naturhistoriske  Forening — 

Yidenskabelige  Meddelelser  for  Aaret  1901. 
Kjobenhavns  Universitets  Zoologiske  Museum — 

The  Danish  Ingolf -Expedition,    vi.  1  (1902). 

Dublin. 

Royal  Dublin  Society  — 

Economic  Proceedings,    i.  2(1899). 

Scientific  Proceedings.      New  Series,    ix.  2-4  (1900-01). 

Scientific  Transactions.     Second  Series,    vii.  8-13  (1900-01), 

Royal  Irish  Academy — 

Proceedings.     Third  Series,    vi.  3(1901). 
Transactions,    xxxi.  7  ct  12(1899,  1901). 


\ 


DONATIONS    AND    EXCHANGES.  665 

Edinburgh. 

Scottish  Microscopical  Society — 

Proceedings.     Session  1900-01.    in.  2(1901). 

Florence. 

SociETA  Entomologica  Italiana 

Bullettino.    xxxiii.  2-4  (1901-02) 

Frankfurt  am  Main. 

Senckenbergische  Naturforschende  Gesellschaft — 
Abhandlungen.    xx.  3;  xxv.  3;  xxvi.  4(1902). 
Bericht,  1901. 

Frankfart  a.d.  Oder. 

Naturwissenschaftlicher  Verein  des  Reg.-Bez.,  Frankfurt. 
Helios,    xix.  (1902). 

Freiburg,  i.  Br- 

Naturforschende  Gesellschaft — 
Berichte.    xi.  3  (1901);  xii.  (1902). 

Geelong. 

Geelong  Field  Naturalists'  and  Science  Club,  and  Gordon 
College  Amateur  Photographic  Association — 
"  The  Wombat."     Tp.,  etc.,  to  Vols,  i.-iv.,  1895-1899  (1902); 

Vol.v.  Nos.  1-3  (1902). 
Tp.,  &c.,  to  Vols,  i.-vi.  of  the  Geelong  Naturalist,  1891-1898 
(1902). 

Geneva. 

SociETE  DE  Physique  et  d'Histoire  Naturelle — 
Memoires.    xxxiv.  1-2^1902). 

Graz. 

Naturwissenschaftlicher  Verein  f.  Steiermark — 
Mittheilungen.    1894,  1895,  1896,  u.  1901. 

Hague. 

Nederlandsche  Entomologische  Vereeniging — 

Entomologische  Berichten.    Nos.  1-6  (1901-02). 
Tijdschrift  voor  Entomologie.    xliv.  2-4;  xlv.  1-2(1901-02). 


666  DONATIONS    AND    EXCHANGES. 

Halle 

Kaiserliche  Leopoldino-Carolinische  Deutsche  Akademie- 
Geschichte    der   Akademie,   1852-87.     Von  Dr.   P.  W.  Ule 

(1889). 
Geschichte  der  Bibliothek,  &c.      Yon  Dr.  O.  Grulich  (1894). 
Leopoldina.    xxxv.-xxxvi.  (1899-1900). 
Nova  Acta.    Ixxv.-lxxix.  (1899-1901). 

Repertorium    zu    den    Acta    u.   Nova  Acta.     Von   Dr.    A. 
Grasel.    i  -ii.  (1894-1899). 

Hamburg. 

Naturhistorisches  Museum  in  Hamburg — 

Mittheilungen.    xvi.  (1898);  xviii.  (1900).  ' 

Naturwissenschaftlicher  Vehein — 

Abhandlungen.    xvi.  2(1901). 

Verhandlungen.    iii.  Folge.    viii.  u.  ix.  (1900-01). 
Verein  f.  Naturwissenschaftlicher  Unterhaltung— 

Verhandlungen.    xi.,  1898-1900(1901). 

Hamilton,  U.S.A. 

Hamilton  Scientific  Association — 

Journal  and  Proceedings.    No.  xvii.    Session  1900-01. 

Harlem.. 

SOCIETE  HOLLANDAISE   DES  SCIENCES  — 

Archives  Neerlandaises.    Serie  ii.    iv.  4-5  (1901);   vi.  (1901); 
vii.  1-3  (1902). 

Helder. 

Nederlandsche  Dierkundige  Vereeniging — 
Aanwinsten  van  de  Bibliothek,  1901. 
Tijdschrift.     2^«  Serie.      vii.  2-4  (1901-02). 

Helsingfors. 

SociETE  DES  Sciences  de  Finlande — 

Oefversigt.    xliii.  (1900-01). 
SociETAS  PRO  Flora  et  Fauna  Fennica — 

Acta,    xvi.,  xviii.-xx.  (1897-1901). 

Meddelanden.    xxiv.-xxvii.  (1897-1901). 


DONATIONS    AND    EXCHANGES.  667 

Hobart. 

Depaktmknt  of  Mines — 

Progress    of    the    Mineral    Industr}^  of    Tasmania  for    the 

Quarters  ending  30th  September  and  31st  December,  1901, 

31st  March  and  30th  June,  1902. 
•   Report  of  the  Secretary  for  Mines  for  1900-01  (1901). 
Report    on    the    Coalfield    of    Llandaff,    the     Denison     and 

Douglas  Rivers,  on  Deposits  of  Tin  Ore  on  Schouten  Main, 

and  on   Outcrops  of    Quartz  near  Buckland.      By  W.  H. 

Twelvetrees,  Government  Geologist  (1901). 
Report  on   the   Recent   Discovery  of    Cannel  Coal  in  the 

Parish  of    Preolenna  and  upon  the  New  Victory  Copper 

Mine   near   Arthur  River.      B}^  G.    A.  Waller,  Assistant 

Government  Geologist  (1901). 
Report  on  the  Tin  Mines  of  the  Blue  Tier,  Count}'  of  Dorset. 

By  W.  H.  Twelvetrees  (1901). 
Report  on    Coal    Seams    at    Thornedale,   near    Thompson's 

Marshes,  and  the  Jubilee  Colliery,  near  St,  Mary's.      By 

W.  H.  Twelvetrees  (1901^. 
Report  on  the  Occurrence  of    Coal  near  Catamaran  River, 

Recherche  Bay.      By  W.  H.  Twelvetrees  (1902). 
Report  on  the  Tin  Ore  Deposits  of  North  Dundas.      By  G. 

A.  Waller  (1902). 
Report  on  Country  on  the  East  Shore  of  Lake  Sorell,  and 

on    a    Discover}^    of    Coal    near    Oatlands.      B}^  W.    H. 

Twelvetrees  (1902). 
Report  of  the  Ore  Deposits  (other  than  those  of  Tin)  of  Xorth 

Dundas.     By  G.  A.  W^aller  (1902). 
Report  on  the  Coal  Field  in  the  Neighbourhood  of  Recherche 

Bay.     By  W.  H.  Twelvetrees  (1902). 
Reports  on  Deposits  of  Opal  at  Bothwell,  and  an  alleo^ed 

Discovery  of  Gold  at  Hunterston,  on  the  Shannon.      By 

W.  H.  Twelvetrees  (1902). 
Report  on   Gold  and   Coal   at   Port    Cygnet.      By   W.   H. 

Twelvetrees  (1902). 


668  DONATIONS    AND    EXCHANGES. 

Department  of  Mines,  Hobart  (continued) — 

Report  on  Deep  Sinking  at  the  Moonlight-cum-Wonder  Gold 
Mine.  Beaconsfield.     By  W.  H.  Twelvetrees  (1902). 

Report  on  some  Discoveries  of  Copjjer  Ore  in  the  Yieinity 
of  Point  Hibbs.     By  G.  A.  Waller  (1902). 

Report  on  the  Tin  Ore  Deposits  of  Mount  Heemskirk.  'By 
G.  A.  Waller. 

Report  on  the  Den  Hill  Gold  Deposits.     By  W.  H.  Twelve- 
trees  (1902). 
Royal  Society — 

Abstract  of  Proceedings,  April  29th  and  July  8th,  1902. 

Papers  and  Proceedings  for  the  Years  1900-01  (June,  1902). 

Parliamentary  Report  ("  Language  and  Dialects  spoken  by 
the  Aborigines  of  Tasmania."     By  J.  E.  Calder).     (1901). 

Four  Pamphlets — "  Tasmanian  School  of  Forestry  and  Agri- 
culture." By  W.  Heyn  (1902);  "  Tasmanian  Diatomaceae. " 
By  F.  E.  Burbury  (1902);  "  Tasmanian  Timbers."  By  A. 
O.  Green  (1902);  "Notes  on  Unrecorded  and  other 
Minerals  occurring  in  Tasmania."  B}'^  W.  F.  Petterd 
(1902). 

Honolulu,  H.I. 

Bernice  Pauahi  Bishop  Museum — 

Fauna  Hawaiiensis.    i.  3(1901);  ii.  5(1900);  iii.  i.  (1901). 

Memoirs,    i.  4(1902). 

Occasional  Papers,    i.  3-5(1901-02). 

Indianopolis. 

Indiana  Academy  of  Science — 
Proceedings.    1900(1901). 

Jena. 

Medicinish-Naturwissenschaftliche  Gesellschaft  zu  Jena. 
Jenaische   Zeitschrift    fiir    Naturwissenschaft.     xxxvi.   3-4; 
xxxvii.  1  (1902). 

Kyoto- 

Kyoto  Imperial  University — 
Calendar,  1901-02. 


DONATIONS    AND    EXCHANGKS.  669 

Landshut  (Bayern). 

BOTANISCHER  VeREIN  — 

Sechzehnter  Bericht,  1898-1900(1901). 

La  Plata. 

MusEO  DE  La  Plata — - 
Revista.    Tomo  x.  (1902). 
Seccion  Geologica  y  Mineralogica.    Anales.    iii.  (1900). 

Facultad  de  Ciencias  Fisico-Matematicas  dr  la  Universidad 
DE  La  Plata — 
Publicaciones.    No.  1  (1901). 

Leipzig. 

Verein  f.  Erdkunde  zu  Leipzig — 

Mitteilungen.    1900(1901);  1901(1902). 
Wissenschaftliche  Veroffentlichungen.    v.  u.  Atlas  (1901). 

ZOOLOGISCHER  AnZEIGER 

xxiv.  Nos.  655-658;  xxv.  Nos.  659-684  (1901-1902). 

Liege. 

Societe  Geologique  DE  Belgique — 

Annales.    xxviii.  4  (1902);  xxix.  1-3(1901-02). 

Lincoln,  U.S.A. 

American  Microscopical  Society — 

Transactions.    Vols,  xx  -xxii.  (1899-1901). 

London.  / 

Bentham  Trustees  (per  Royal  Botanic  Gardens,  Kew) — 

Hooker's  Icones  Plantarum.  FourthSeries.  viii.  2(May,1902). 
British  Museum  (Nat,  Hist.) — 

Catalogue  of  Fossil  Fishes.    Part  iv.     By  A.  8.  Woodward, 

LL.D.,  F.R.S.,  F.G.S.  (1901). 
Catalogue   of  Lepidoptera  Phalsenie.     Vol.  iii.,  and  Atlas. 

By  Sir  George  F.  Hampson,  Bart.  (1901). 
Hand-List  of  Birds.    Vol.  iii.     By  R.  B.  Sharpe, LL.D. (1901). 
44 


670  DONATIONS    AND    EXCHANGES. 

London  (continued) — 
Entomological  Society — 

Transactions.    1901,  Parts  iv.-v;  1902,  Parts  i.-ii.  (1901-02). 

Geological  Society — 

Geological    Literature    added    to    the    Geological    Society's 

Library  during    the    Year    ended    December   31st,    1901 

(^Sept.,  1902). 
List  of  the  Society.    November,  1902. 
Quarterly  Journal.    Ivii.  4;  Iviii.  1-3(1901-02). 

Linnean  Society — 

Journal.    Botany,    xxxv.  244-245  :  Zoology,    xxviii.  183-185 

(1901-02). 
List  of  the  Society.    1901-02. 

Proceedings.    113th  Session  (Nov.,  1900-June,  1901). 
Transactions.     Botany.     Second   Series,     vi.  2-3  (1901-02)  : 

Zoology,    viii.  5-8(1901-02). 

Royal  Microscopical  Society — 

Journal.    1901.  Part  6;  1902.  Parts  1-4. 
Royal  Society — 

Proceedings.    Ixix.  451-458;  Ixx.  459-467(1901-02). 

Reports  to  the  Evolution  Committee.    Report  i.  (1902). 

Reports    to    the   Malaria   Committee.     Sixth   and   Seventh 
Series  (1902). 

Zoological  Society — 

Abstract  of  Proceedings.    Nov.  2nd  k  19th,  Dec.  3rd  &  17th, 

1901;  Feb.  4th  and  18th,  March  4th,  April  15th,  May  6th, 

June  3rd  &  17th,  1902. 
Proceedings.    1901,  ii.  1-2:   1902,  i.  1. 
Transactions,    xvi.  3-4(1901-02). 

Madras 

Madras  Government  Museum — 
Bulletin,    iv.  2(1901). 


DONATIONS    AND    EXCHANGES.  671 

Manchester. 

CONCHOLOGICAL  SOCIETY  OF  GrEAT  BrITAIN  AND  IRELAND  — 

Journal  of  CoHchology.    x.  5-8  (1902). 

Manchester  Literary  and  Philosophical  Society — 
Memoirs  and  Proceedings,    xlvi.  1-6  (1901-02). 

Manchester  Museum,  Owens  College — 
Publications.    Nos.  32,  35-38  (1901-02). 

Marseilles. 

Musee  d'Histoire  Naturelle — 

Annales.    Tome  vi.    Section  de  Zoologie  (1901). 

Massachusetts. 
Tufts  College — 

Studies.    No.  7  (May,  1902). 

Melbourne. 

Australasian     Association     for     the     Advancement    of 

Science — 
Report  of  the  Eighth  Meeting,  Melbourne,  1900  (1901). 

Australasian  Journal  of  Pharmacy — 

Yol.xvi.  No.  192  (Dec, 1901);  Vol.  xvii.  Nos.  193-203  (Jan.- 
Nov.,  1902).     From  the  Publishers. 

Australasian  Institute  of  Mining  Engineers — 
Proceedings,  Annual  Meeting,  January,  1902. 

,,  Special  Meeting,  March,  1902. 

Transactions,    viii.  1  (1901). 

Australasian  Ornithologists'  Union — 
"The  Emu."   i.  2-3;  ii.  1-2  (1902). 

Department  of  Agriculture — 
Annual  Report.    1900-1901  (1902). 
Bulletin.    Nos.  1-4  (reprinted  from  the  Journal). 
Fungus  Diseases  of  Stone- Fruit  Trees  in  Australia  and  their 

Treatment.     By  D.  Mc Alpine  (1902). 
Journal,    i.  1-9  (Jan.-Sept.,  1902). 


672  DONATIONS    AND    EXCHANGES. 

Melbourne  (continued) — 
Department  of  Mines — 

Annual    Report    of    the    Secretary  for   Mines    and   Water 

Supply  for  the  Year  1901. 
"Indicator"   Series  of    Booklets   on   Gold   Mining.    No.  3. 

The  Creswick  Field  and  its  Mining.    By  William  Bradford. 
Report  on  the  Shamrock,  Shenandoah,  New  Chum  Railway, 

Eureka  Extended,  and    South    Belle  Yue  United   Mines, 

New  Chum  Line  of  Reef,  &c.      By  H.  S.  Whitelaw  (1901). 
Report  on  the  Walhalla  Gold-field.      By  H.  Herman  (1901). 

Field  Naturalists'  Club  of  Yictoria — 

Yietorian  Naturalist,    xviii.  8-12;  xix.  1-7(1901-02) 
List  of  Members,  September,  1902. 
Rules,  April,  1902. 

Public    Library,    Museums,    and    National    Gallery   of 
Yictoria — 
Report  of  the  Trustees  for  1901  (1902). 

Royal  Society  of  Yictoria  — 
Proceedings,    xiv.  2;  xv.  1  (1902). 

University  of  Melbourne — 

Examination    PajDers  :     Annual  (No\.-Dec.,  1901) ;    Final 
Honour,  Degrees,  ifcc.  (IVJarch,  1902);  Matriculation  (Dec, 
1901,  and  May,  1902). 
University  Calendar  for  1902. 

Zoological  and  Acclimatisation  Society  of  Yictoria — 
Thirty-eighth  Annual  Report  (1901). 

Mexico. 

Instituto  Geologico  de  Mexico — 
Boletin.    Num.  15  (1901). 

Michigan. 

State  Board  of  Agriculture  of  the  State  of  Michigan — 
Thirty-Seventh,  Thirty-Eighth,  Thirty-Ninth,   and  Fortieth 
Annual  Reports  (1897-1901). 


DONATIONS    AND    EXCHANGES.  673 

Missoula. 

UNivBRSiry  OF  Montana — 

Bulletin.    No.  3  [Biological  Series.   No.  1]  (1901). 

Montevideo. 

MusEo  Nacional  db  Montevideo — 
Anales.    iii.  21;  iv.  1-2  (1901-02). 

Montreal. 

Mo.vTREAL  Society  op  Natural  History  — 

Canadian  Record  of  Science,    viii.  6-7  (1901-02). 
Royal  Society  of  Canada — 

Proceedings  and  Transactions.    Second  Series.      One  pocket 
of  Maps  for  Vol.  v.  (1899);  vi.  (1900). 

Moscow. 

SociETE  Imperials  des  Natqralistes  — 
Bulletin.    Annee  1902.  Nos.  1-2. 

Munich. 

Konigliche  Bayerische  Akademie  dbr  VVissenschaftbn — 
Sitzungsberichte  der  Math.-physikal.  Classe.  xx.-xxxi.,xxxii. 
1-2(1890-1902). 

Nantes. 

Societe  des  Suibnces  Naturelles  de  l'Ouestde  la  France — 
Bulletin.    2«  Serie.    i.  1-4(1901). 

Table  des  Matieres  de  la  Premiere  Serie  (Tomes  1  ax.,  1891 
a  1900). 

Naples. 

ZOOLOGISCHE  StA'I'ION  ZU   NeAPEL 

Mittheilungen.    xv.  1-3  (1901). 

New  York. 

American  Geographical  Society — 

Bulletin,    xxxiii.  4-5;  xxxiv.  1-3  (1901-02). 
American  Museum  qf  Natural  History — 

Annual  Report  for  the  Year  1901  (1902). 

Bulletin,    xi.  4;  xiv.,  and  Arts.  20-24  separately;  xv.  Parti; 
xvi.  Arts.  1-20;  xvii.  Parts  1-2(1901  02). 

Memoirs,    i.  7;  iv.  3;  v.  1;  vii.  1(1901-02). 


67-i  DOXATIOXS    AND    EXCHANGES. 

New  York   (continued)  — 

New  York  Academy  of  Sciences— 
Annals,    xiv.  1-2(1901-02). 
Memoirs,    ii.  3(1901). 

Odessa- 

SOCIETE  DES  NaTURALISTES  DE  LA  NOUVELLE-RUSSIE — 

Memoires.    xxiv.  1  (1902). 

Ottawa. 

Geological  Survey  of  Canada — 

Catalogue  of  Canadian  Birds.    Part  i.  (1900).    By  J.  Macoun. 
Catalogue  of  the  Marine  Invertebrata  of   Eastern  Canada 

(1901).     By  J.  F.  Whiteaves. 
Contributions  to  Canadian  Paheontology.    ii.  2;  iv.  2  (1900- 
01). 
General  Index  to  the  Reports  of  Progress,  1863-81(1900). 

Oxford. 

Radcliffe  Library,  Oxford  University  Museum — 
Catalogue  of  Books  added  during  1901. 

Padova. 

La    Nuova    Notarisia.    Serie  xiii.    Gennaio,   Aprile,   Luglio» 
Ottobre,  1902.     From  the  Editor,  Dr.  G.  B.  De  Toni. 

Para. 

Museu  Paraense  de  Historia  Naturale  e  Ethnographia  — 
Album  de  Aves  Amazonicas,  organisado  pelo  Dr.  E.  Goeldi. 
PFasciculo,  Estampas  1-12  (1900). 

Paris. 

Journal  de  Conchyliologie.    xlix.  4;  1.  1  (1901-02). 
Museum  d'Histoire  Naturelle — 

Bulletin.    1901,  4-8;  1902,  1-2(1902). 

Nouvelles  Archives.    4°^^  Serie.    ii.  2:  iii.  1  (1900-01). 
Societe  Entomologique  de  France — 

Annales.    Ixix.  1-4(1900). 

Bulletin.    Annee  1900. 


DONATIOXS  AND  EXCHANGES.  675 

Paris  (continued) — 

SOCIETE  ZOOLOGIQUE  DK   FrANCE  — 

Bulletin,    xxvi.  (1901). 
Memoires.    xiv.  (1901). 

Pavia. 

IST[TUTO  BOTAJ^ICO  DELL'  UxIVERSITA  DI  PaVIA  — 

Atti.    ii.  Ser.   Vol.  Settimo  (1902). 

Perth,  W.A. 
Department  op  Agriculture  of  West  Australia — 

Journal,    iv.  5-6;  v.  1-6;  vi.  1-4(1901-02). 
Geological  Survey  op  West  Australia — 
Annual  Progress  Report  for  the  Year  1900. 
Bulletin.    No.  6  (1902). 
Victoria  Public  Library — 
Report  of  the  Committee  for  the  Year  ended  30th  June,  1901. 
Report  by  the  Under  Secretary  for  Lands  for  the  Year  1897. 
Report  by  the  Surveyor  General  for  the  Year  1900. 
"  Western    Australia  and    its    Resources  "  :    printed  under 

instructions  from  the  Minister  of  Lands  (n.d.). 
The  Land  Selector's  Guide  to  the  Crown  Lands  of  Western 
Australia  (1901).     Edited   by  direction  of    the   Minister 
for  Lands. 

Philadelphia. 

Academy  op  Natural  Sciences — 

Proceedings,    liii.  2-3  (1901-02). 
American  Philosophical  Society — 

Proceedings,    xl.  165-167;  xli.  168(1901-02). 

University  op  Pennsylvania— 
Contributions  from  the  Botanical  Laboratory,    ii.  1-2  (1898- 

1901). 
Contributions   from   the   Laboratory  of    Hygiene.    Nos.  1-2 

(1898). 
Contributions  from  the  Zoological  Laboratory,    i.  1  (1893); 
1901  (1902). 


676  DONATIONS    AND    EXCHANGES. 

University  of  Pennsylvania  (continued) — 

Syllabus  of    Lectures  on  the  Vertebrata.     By  E.  D.  Cope, 
Ph.D.  (1898). 
Zoological  Society — 

Thirtieth  Annual  Report  of  the  Board  of  Directors  (1902). 

Pietermaritzburg. 

Geological  Survey  of  Natal  and  Zululand — 

First    Report.      By   W.    Anderson,    Government    Geologist 
(1901). 

Plymouth. 

Marine  Biological  Association  of  the  United  KixVgdom  — 
Journal.     New  Series,    vi.  3(1902). 

Portland,  U.S.A. 

Portland  Society  of  Natural  History — 
Proceedings,    ii.  5  (1901). 

Prague. 

KOENIGLICHE  BOEHMISCHE  GeSELLSCHAFT  DER  WiSSENSCHAFTEN. 

Jahresbericht.    1899,  1900,  1901. 

Prager    Tychoniana.       Gesammelt     von     Prof.     Dr.    F.    J. 

Studnicka  (1901). 
Sitzungsberichte.    1899,  1900,  1901. 
Studie    o    Drazdivosti    Rostlinne     Plasmy.       Dr.     Bohumil 

Nemec(1900). 

St.  John,  New  Brunswick. 

Natural  History  Society  of  New  Brunswick  — 
Bulletin.    Nos.  xviii.  and  xx.  0899-1902). 

St.  Louis. 

Academy  of  Sciences — 

Transactions,    x.  9-11;  xi.  1-5(1901). 
Missouri  Botanical  Garden  — 

Twelfth  Annual  Report  (1901). 

St.  Petersburg. 

Academie  Imperials  des  Sciences — 

Annuaire  du  Musee  Zoologique.    v.  1-2;  vi.  2-4(1900-01). 


DONATIONS    AND    EXCHANGES.  677 

St.   Petp:rsbuug  (confinupd) — 

CoMiTE  Geologique  (Institut  des  Mines) — 

Bibliotheque  Geologique  de  la  Russie,  1897  (1901). 

Bulletins,    xix.  7-10;  xx.  1-6(1900-01). 

Memoires.    xviii.  1-2  (1901). 
Russisch-Kaiserliche  Mineralogische  Gesellschapt. 

Verhandlungen.     Zweite  Serie.    xxxix.  1-2(1901-02). 
SociETE  Entomologique  de  Russie — 

Horae.  T.p.  &c.  to  Tome  xxxiii. ;  xxxv.  3-4  (1902). 

Santiago. 

SOCIETE    SCIENTIFIQUE    DU    ChILI  — 

Actes.    xi.  2-3(1901). 
Siena. 

ISTITUTO    BOTANICO    DELLA    R.    UnIVERSITA    DI    SiENA 

BuUettino.     Anno  Quarto  (1901). 

Stockholm. 

Entomologiska  Foreningen  — 

Entomologisk  Tidskrift.    xxii.  1-4(1901). 
Kongl.  Svenska  Yetenskaps-Akademie  — 

Bihang.    xxvi.  (1900-01). 

Handlingar.    N.F.    xxxiii.-xxxiv.  (1900-01). 

Lefnadsteckningar.    iv.  1-2(1899-1901). 

Stuttgart. 

Verein  p.  vateklaendische  Naturkunde  in  Wuerttemberg — 

Jahreshefte.    Ivii.  (1901). 

Sydney. 

Australian  Museum  — 

Records,    iv.  2,  and  5-7(1901-02). 

Memoir  iv.      Scientific  Results  of  the  Trawling  Expedition  of 

H.M.C.S.  "  Thetis."     Parts  4  and  5  (1902). 
Special    Catalogue.  No.  i.    Nests   and   Eggs  of  Birds   found 
breeding  in  Australia  and  Tasmania.      By  A.  J.  North, 
C.M.Z.S.    Part  2  (April,  1902). 


678  DONATIONS    AND    EXCHANGES. 

Sydney  (continued) — 

Botanic  Gardens  and  Domain,  Sydney — 

Report  of  the  Director  for  the  Year  1901  (1902). 
Department  of  Mines  and  Agriculture — 

Agricultural   Gazette   of    N.S.W.     xii.  12,  Index,  etc.;  xiii. 

1-11  (1901-02). 
Annual  Report  of  the  Department  of  Mines  for  1900  and 

1901. 
Geological   Survey:    Mineral   Resources.    No.  9.    Report  on 

the  Yahval  Gold-Field.      By  E.  C.  Andrews,  B.A.  (1901); 

No.  10.    Report  on  the  Kiandra  Lead.    By  E.  C.  Andrews, 

B.A.  (1901). 
Handbook  to  the  Mining  and  Geological   Museum.     By  G. 

W.  Card,  A.R.S.M.,  &c.  (1902). 
Records.    Vol.  vii.  Pt..  2(1902). 

Government  Statistician — 

Statistics:   Six  States  of  Australia  and  New  Zealand,  1861- 

1900(1901);  1861-1901(1902). 
Wealth    and    Progress    of     New    South     Wales,    1900-01. 
Thirteenth  Issue  (1902). 
Institution  of  Surveyors — 

The  Surveyor,    xiv.  7;  xv.  1-10(1901-02). 

Public  Library  of  New  South  Wales  — 
Report  of  the  Trustees  for  the  Year  1901. 

Royal  Anthopological  Society  of  Australasia — 

"Science  of  Man."    New  Series,    iv.  10-12;  v.  1-9(1901-02). 

Royal  Society  of  New  South  Wales  — 

Abstract  of  Proceedings.    December,  1901;  May-November, 

1902. 
Journal  and  Proceedings,    xxxv.  (1901). 

Technological  Museum — 

A  Research  on  the  Eucalypts,  especially  in  regard  to  their 
Essential  Oils.  By  R.  T.  Baker,  F.L.S.,  and  Henry  G. 
Smith,  F.C.S.  (4to.,  Sydney,  1902). 


DONATIONS    AND    EXCHANGES.  679 

Sydney  (continued) — 
University  of  Sydney  — 
Calendcar,  1902. 

Taiping. 

Perak  Government  Gazette — 

xiv.    30-37  (Oct.-Dec.  1901);  XV.    1-4  (Jan -Feb.  1902).  i^rom 
the  Govemiinent  Secretary. 

Tiflis, 

Botanical  Garden  in  Tiflis  (Caucasus) — 

Fifth  Annual  Report  (1901);  Sixth  Annual  Report.   Part  1 
(1902). 

Tokyo. 

College  of  Science,  Imperial  University  — 

Journal,    xvi.  Parts  1-2,  and  Art.  6  (1901-02);  xvii.  Parts  1-3, 
Arts.  7-9(1901-02). 
Imperial  University  of  Tokyo — 

Calendar,  1901-02. 
Zoological  Society  of  Tokyo — 

Annotationes  Zoologicae  Japonenses.    iv.  2-3(1902). 

Turin. 

Museo  di  Zoologia,  (fee,  della  R.  Universita  di  Torino — 
Bolletino.    xvi.  404-415  (1901). 

Upsal. 

Regia  Societas  Scientiarum  Upsaliknsis — 
Nova  Acta.     Ser.  iii.    xx.  1  (1901). 

Valparaiso. 

Museo  de  Historia  Natural  de  Valparaiso  — 
Revista  Chilena.    v.  11-12;  vi.  1-2(1901-02). 
Memoria  durante  el  Ano  de  1901  (1902). 

Washington,  D-C- 

Bureau  of  American  Ethnology — 

Eighteenth  Annual  Report,  1896-97.    Part  2  (1899). 


680  DONATIONS    AND    EXCHANGES. 

Washington,   D.C.  (continued) — 
Smithsonian  Institution — 

Annual  Report  of  the  Board  of  Regents  for  the  Years  1879, 
1899,  and  1900. 
U.S.  Department  of  Agriculture— 

Bureau  of  Plant  Industry.     Bulletin.    Xos.  1-4,  6-13,  15-19, 

25,  pp.  1-4  (1901-02). 
Circulars.    Nos.  46,  48-51  (June,  1902). 
Division  oj  Botany.    Bulletin.    Nos.  27-29  (1901). 
Contributions   to  U.S.   National    Herbarium,     v.   4;   vii.   3. 

T.p.,  &c.  (1899-1902). 
Division  of  Entomology.     Bulletin.     New  Series.     Nos.   27, 

31-34(1901-02). 
Farmers'  Bulletin.    Nos.  130,  132(1901). 
Technical  Series.    No.  9  (1901). 
U.S.  Geological  Survey — 

Twenty-first  Annual  Report,  1899-1900.    Parts  i.-iv.,  vi.,  <fe 
vi.  con td.  (1900-01). 
U.S.  National  Museum — 

Bulletin.    No.  50,  Part  i.  (1901). 
Proceedings,    xxii.  (1900). 
Washington  Academy  of  Sciences — 

Proceedings,   iii.  pp.371-612(1901);   iv.  pp.  1-560(;1902). 

Wellington. 

New  Zealand  Institute — 

Transactions  and  Proceedings,    xxxiii.;  xxxiv.  (1901-02). 

Private   Donors. 

S.A.S.  Albert  P^,  Prince  de  Monaco — One  Separate:  'La 
Troisieme  Campagne  Scientifique  de  la  Princesse-Alice  ii®' 
(4to.,  Paris,  1902). 

Bailey,  F.  M,,  F.L.S.,  Brisbane — Four  Botanical  Separates  from 
the  Queensland  Agricultural  Journal  (vii.  4-5;  ix.  2  and  4  • 
Svc,  Brisbane,  1900-01). 


I 


DONATIONS    AND    EXCHANGES.  681 

Ckane,  W.  E.,  New  York.     (By  Exchange) — 

"American     Naturalist."     Vol.     ii.     No.     11    (Jan.    1869); 

Vol.  vii.  Nos.  1-3,  5-6  (1873). 
Annual  Report  of  the  Smithsonian  Institution  for  1865,  '67, 

'68,  '69,  '70,  '71,  '73,  '74  (8  Vols.). 
Annual  Report  of  the  U.S.  Department  of  Agriculture  for 

1869,  '70,  '71,  '72,  '73,  '74,  '79,  '80,  '81,  '82,  '83,  '84, 

'86,  '87,  1890,  '91  (15  Vols.). 
Annual  Report  of  the  U.S.  Fish  Commission.     Parts  xi.-xii. 

(1883-84):  Bulletins.     Vols.  i.-v.  and  xiii.-xv.  (1881-85, 

and  1893-95). 
Monographs  of  the  U.S.  Geological  Surve3^     Vols,  ii,,  vi., 

viii.,  and  ix.  (1883-85). 
Proceedings  of  the  American  Association  for  the  Advance- 
ment of  Science.     Vols,  xxxvi.,  xxxviii.,  xxxix.  (1887, 

1888-90), 
Reports   of  the   U.S.   Geological  Survey  of  the  Territories. 

Vols.  vii.  and  ix.  (1876-78). 

French,  C,  F.L  S.,  Melbourne—  Report  of  the  Government 
Entomologist  of  Victoria  for  1900  (8vo.,  Melbourne,  1901). 

Froggatt,  W.  W.,  F.L.S.,  Sydney — Four  Entomological  Sepa- 
rates from  the  Agricultural  Gazette  of  N.S.W.  (being  Mis- 
cellaneous Publications,  Nos.  487,  520,  521,  538;  1901). 

Guthrie,  F.  B.,  F.I.C,  F.C.S.,  Sydney — Eleven  Separates  from 
Agric.  Gaz.  of  N.S.W.  (Misc.  Publications,  Nos.  384,  386, 
409,  427,  461,  482,  529,  561,  579,  584,  587;  1900-02).  One 
Separate:  "Notes  on  Analj^sis  of  Air  from  Coal  Mines." 
(Journ.  and  Proc.  R.  Soc.  N.S.  Wales,    xxxv.,  1902). 

Jack,  Dr.  R.  L.,  London — One  Separate  :  "  Artesian  Water  in 
the  State  of  Queensland.  Australia  "  (Trans.  Victoria  Inst., 
1902). 

Maiden,  J.  H.,  F.L.S.,  &c.,  Sydney — Twelve  Botanical  Separates 
from  the  Agricultural  Gazette  of  N.S.  Wales  (including 
Misc.  Publications,  Nos.  499,  509,  510,  531,  532,  553),  Rept. 


682  DONATIONS    AND    EXCHANGES. 

Aust.  Assoc.  Adv.  Sc.  (Vol.  viii.),  Victorian  Nat.  (Vol.  xviii.), 
Proc.  R.  Soc.  Tasmania,  and  Trans.  R.  Soc.  S.  Australia 
(Vol.  XX vi.) 

Marshall,  C.  E.,  Michigan  State  Agricultural  College  Experi- 
ment Station,  U.S.A.— Bulletins  :  Nos.  139-140,  146-147, 
159,  172-173,  182-184  (1896-1900).  Special  Bulletin:  No.  13 
(1899,  8vo.) 

MussoN,  C.  T.,  Richmond,  N.S.W. — J.  H.  Balfour's  "Introduction 
to  PalEeontological  Botany"  (8yo.,  1872). 

Smith,  E.  A.,  F.Z.S.,  &g. — Four  Conchological  Separates:  from 
Ann.  Mag.  N.H.  (7)  vii.,  (1901);  Journ.  Conch.  1901;  Journ. 
Malacol.  viii.  (1901);  Proc.  Malacol.  Soc.  iv.  (1901). 

Smith,  R.  Grbig,  M.Sc,  Sydney— One  Separate  from  Agricul- 
tural Gazette  of  N.S.  Wales  (Misc.  Publication,  No.  558; 
1902.) 

TiDSWELL,  Dr.  F.,  Sydney — Three  Separates  :  "  Some  Practical 
Aspects  of  the  Plague  at  Sydney"  (Sanitary  Institute  xxi. 
4) :  "  Small  Pox  Possibilities  and  Obligations "  (Journ. 
Sydney  Medical  Soc;  November,  1901):  "Preliminary  Note 
on  the  Serum-Therapy  of  Snake-Bite  "  (Australian  Medical 
Gaz.,  April,  1902). 

Urban,  Ignatius  (Editor) — Symbolte  Antillan?e  seu  Fundamenta 
fbij^Florse   Indise  Occidentalis.      Vol.  ii.     Fasc.  iii.  (8vo.,  Berlin 
1901).      From  the  Editor. 


683 


WEDNESDAY,  MARCH  25th,  1903. 


The  Twenty-Eighth  Annual  General  Meeting  of  the  Society- 
was  held  in  the  Linnean  Hall,  23  Ithaca  Road,  Elizabeth  Bay, 
on  Wednesday  evening,  March  25th,  1903. 

Mr.  J.  H.  Maiden,  F  L.S.,  &g.,  President,  in  the  Chair. 

The  Minutes  of  the  previous  Annual  General  Meeting  were 
read  and  confirmed. 

The  President  delivered  the  Annual  Address. 


PRESIDENTIAL  ADDRESS. 


THE  PRINCIPLES  OF  BOTANICAL  NOMENCLATURE. 

PAGE 

1.  Introductory,  Codes  of  Laws      ...          ...          ...          ...         ...         ...  685 

2.  1737  and  1753        687 

3.  No  Priority  for  Pre-Linnean  Names      ...          ..           ...          ...           ..  689 

4.  Revisions  of  Botanical  Names  must  be  the  work  of  Monographers...  695 

5.  Names  in  MSS.  or  Herbaria       ..           ..           698 

6.  Uniformity  of  Names  of  Botanical  Authors 701 

7.  On  the  Writing  of  Specific  Names         ...          ...          ...          ...         ...  704 

8.  The  Question  of  Altering  Botanical  Names  with  the  View  to  their 

"Improvement"         ...         ...         ...         ...          ...         ...          .  .  708 

9.  The  "  Botany  of  Cook's  Voyage  "         711 


Addendum. —The  De  Vriesian  Species        ...         ...         ..  ...         ...     717 


The  Society's  history  during  the  Session  now  about  to  close  has 
not  been  marked  by  any  events  of  unusual  importance.  Such  a 
statement  happily  does  not  by  any  means  necessarily  imply  that 
satisfactory  progress  has  not  been  made. 


684  president's  address. 

During  the  year  nine  Ordinar}'  Members  were  elected,  and  five 
resigned.  Three  Corresponding  Members  were  elected — Dr.  R. 
Broom,  B.Sc,  C.M.Z.S.,  of  Pearston,  South  Africa,  Mr.  T>. 
Me  Alpine,  of  Melbourne,  and  Mr.  E.  Meyrick,  B.A.,  F.Z.S.,  of 
Marlborough,  England.  One  Ordinary  Member  and  one  Corres- 
ponding Member  were  removed  by  death. 

Mr.  James  B.  Garland,  who  died  on  May  23rd,  1902,  at  the 
early  age  of  31,  became  a  Member  of  the  Society  in  1899.  In 
Nature-study  he  found  a  hobby,  and  in  following  it  up  he  had 
harvested  much  knowledge  of  plants  and  animals  at  firsthand  by 
his  own  patient  observations. 

Mr.  Richard  C.  Rossiter,  who  was  elected  a  Corresponding- 
Member  in  December,  1881,  died  on  16th  January,  1903,  at 
Noumea,  aged  62  years.  He  was  an  enthusiastic  conchologist, 
and,  during  a  residence  of  about  thirty  j^ears  in  Noumea,  he 
succeeded  in  amassing  a  very  fine  collection  of  shells  from  New 
Caledonia  and  contiguous  islands.  In  1881  (Proc.  vi.  817)  Mr. 
Rossiter  communicated  to  this  Society  "A  List  of  the  Cyprseidie 
found  on  the  Coast  of  New  Caledonia  and  Loyalty  Islands," 
comprising  sixty  species,  based  largely  on  his  own  collection. 

Part  i  of  the  Proceedings  for  1901,  and  three  Parts  of  the 
Proceedings  for  1902  were  issued  during  last  year.  All  but  the 
last  few  sheets  of  the  concluding  Part  of  the  Volume  for  1902 
have  been  printed  off,  and  it  is  expected  that  the  Part  will  be 
ready  for  issue  within  the  ensuing  fortnight.  Forty-three  papers, 
including  Part  i.  of  a  "  Census  Muscorum  Australiensis,"  issued 
separately  as  a  Supplement  to  Part  3,  will  be  found  to  be  com- 
prised in  the  Proceedings  for  1902. 

The  additions  to  the  library  during  the  period  November  28th, 
1901,  to  November  27th,  1902,  amount  to  981,  received  by  gift 
or  exchange  from  181  Societies,  Institutions,  &c.,  and  fourteen 
Individuals.  The  complete  list  for  the  period  mentioned  will  be 
found  at  the  end  of  the  Volume. 

The  Bacteriologist  during  last  year  was  engaged  chiefly  in  con- 
tinuing his  researches  on  the  subject  of  bacteria  in  relation  to  gum- 


president's  address.  685 

formation.  Six  papers  dealing  with  this  or  other  Bacteriological 
subjects  were  contributed,  and  already  have  been  published  and 
distributed.  One  pupil  received  a  full  course  of  laboratory- 
instruction. 

The  Covington  drawings  illustrating  the  Cruise  of  H.M.S. 
*'  Beagle,"  to  which  your  attention  was  invited  in  July  last,  have 
been  framed  and  now  adorn  the  walls  of  the  Hall.  May  they 
help  to  stimulate  a  perennial  interest  in  the  records  of  the  historic 
Voyage  and  what  came  of  it  ! 


1. — Introductory,  Codes  of  Laws. 

It  seems  strange  that  the  principles  which  underlie  the 
bestowal  of  botanical  names  have  not  hitherto  been  brought 
before  this  Society,  at  all  events  except  in  a  casual  manner;  and 
I  venture  to  make  an  attempt  to  supply  this  hiatus. 

Much  of  the  voluminous  correspondence  and  polemic  writings 
on  the  subject  in  European  and  American  publications  are  only 
of  indirect  interest  to  us  because  the  vast  majority  of  plants 
referred  to  are  non-Australian.  I  propose  only  to  touch  upon 
points  that  are  of  special  interest  to  Australian  botanists. 

The  oldest  set  of  Linnean  rules  governing  botanical  nomen- 
clature and  including  many  descriptive  definitions,  are  contained 
in  the  'Philosophia  Botanica'  of  Linnaeus  himself,  a  copy  of  which, 
dated  Vienna,  1755,  is  before  me.  It  remained  in  activ^e  use  for 
at  least  a  century. 

The  principal  code  of  laws  at  present  respected  by  botanists 
throughout  the  world  is  : — 'Lois  de  la  Nomenclature  Botanique 
adoptees  par  le  Congres  international  de  Botanique,  tenu  a  Paris 
en  Aout  1867,'  par  Alph.  de  Candolle  (1867). 

This  is  the  first  and  foremost  of  the  modern  attempts  to  reduce 
the  laws  underlying  the  principles  of  botanical  nomenclature  to 
order,  and  it  has  been  of  incalculable  benefit  to  botanical  science. 

These  "  lois"  have  been  supplemented  by  'Nouvelles  Remarques 
sur  la   nomenclature  botanique'  (1883)  by   the  same  author,  a 
work  which  is  also  indispensable. 
45 


688  president's  address. 

Of  late  years  two  American  codes  have  claimed  attention, 
viz : — 

(1).  The  Rochester  Rules,  promulgated  at  a  meeting  of  the 
American  Association  for  the  Advancement  of  Science,  held  at 
Rochester,  N.Y.,  in  August,  1892. 

(2).  The  Madison  Rules,  promulgated  by  the  Madison  Botani- 
cal Congress,  held  at  Madison,  Wis.,  on  August  23rd  and  24:th, 
1893. 

Our  American  brethren,  with  the  fervour  of  their  nation,  have 
in  certain  points  of  nomenclature  been  more  sweeping  in  their 
changes  than  have  European  botanists,  and  it  will  be  sufficient 
for  my  present  purpose  to  draw  attention  to  the  able  criticisms 
of  those  codes  to  be  found  in  the  'Journal  of  Botany'  (London)  and 
in  the  'Botanical  Gazette'  (Chicago). 

Then  come  the  original  Berlin  Rules  for  nomenclature.  They 
are  14  in  number,  and  were  promulgated  in  1897"^  under  the 
authority  mainly  of  the  staff  of  the  Royal  Botanic  Garden  and 
Museum  of  that  city.  They  have  been  generousl}'-  criticised  by 
Mr.  J.  Britten,!  and  they  certainly  command  the  respectful 
attention  of  botanists  all  over  the  world. 

They  are  supplemented  by  '  Zusatze  zu  den  Berliner  Nomen- 
clatur-Regeln,'t  promulgated  last  year. 

Then  we  have  the  International  Congress  of  Botan}^,  held  at 
the  Paris  Exhibition  from  1st  to  10th  October,  1900,  and  at 
which  I  was  present.  The  question  of  nomenclature  was  there 
iscussed,  and  the  "Bureau  Permanent"  appointed  by  the  Con- 
gress have  invited  me  to  contribute  to  the  preliminary  agenda- 
paper  and  to  be  present  at  the  adjourned  Congress  to  be  held  in 
Vienna  in  1905.  The  work  in  regard  to  Botanical  Nomenclature 
at  the  Paris  Exhibition  was  only  of  a  preliminary  character,  but 
the  work  has  been  taken  up  with  such  enthusiasm  that  I  do  not 
doubt  that  results  important  to  the  whole  botanical  world  will  be 

*  Notizblatt  des  K.  bot.  Gart.  Berlin,  1897,  248. 

t  Journ.  Bot.  xxxv.  305. 

tBeiblatt  zu  den  Botanischen  Jahrbiiehern  Nr.  70  (Berlin),  xxxi.  p.  24. 


president's  address.  687 

achieved  at  Vienna.  The  way  is  gradually  being  paved  for  the 
establishment  of  a  "Tribunal  of  Nomenclature,"  whose  decisions 
no  botanists  may  afford  to  disregard. 

2.— 1737  AND  1753. 

Genera. — It  is  generally  agreed  that  1737  is  the  earliest  date 
for  accepted  generic  names,  that  being  the  year  of  publication  of 
Linnaeus'  'Genera  Plantarum.'  Amongst  many  other  authorita- 
tive statements,  see  the  Journal  of  Botany.* 

The  Berlin  Rule  is,  however,  to  date  both  species  and  genera, 
from  the  year  1752,  the  date  of  publication  of  the  4th  Ed.  of 
the  '  Genera  Plantarum.'  "  After  A.  de  Candolle  had  proposed 
to  take  the  year  1737  as  the  starting  point  of  the  priority  of 
genera,  many  botanists  had  acknowledged  it.  But  we  think  that 
the  turning-point  from  the  ancient  botany  to  our  modern  science 
rests  in  the  introduction  of  the  binomial  nomenclature. 
Before  1752,  the  scientific  position  of  Linnaeus  is  not  superior 
to  Tournefort,  Rivinus,  and  many  other  botanists  who  often  had 
described  and  segregated  the  genera  more  exactly  than  he  did." 
(Note  to  Berlin  Rule  1). 

The  following  is  a  list  of  genera  from   Mueller's  '  Census,'  in 
which  the  compiler  quotes  Linnaeus  (1735)  : — 

C  eratophylluini.  Nepenthes. 

Z ygophyllnm.  Ixora. 

Andrachne  Coffea. 

Cardiospermum.  Strych7ios. 

Sagiyia.  Agrostis. 

Tetragonia. 
Species. — It  is  generally  accepted  that  1753  is  the  earliest  date 
for  specific  names,  that  being  the  year  of  publication  of  Linnaeus' 
'Species  Plantarum.'     The  Berlin  Rule  is  in  favour  of  this  date 
also. 

O.  Kuntze  proposed  173^  as  the  date  for  the  earliest  name, 
that    being    the    date    of    Linnaeus'    '  Systema   Naturae,'  but    B. 

*  e.g.  Journ.  Bot.  xxvi.  260. 


688  PRESIDENT  S  ADDRESS. 

Daydon  Jackson  shows'^  that  this  date  is  too  earh\  At  the  same 
place  will  be  found  a  preliminary  criticism  of  Kuntze's  '  Revisio 
Generum  Plantarum,'  a  monument  alike  to  the  author's  industry 
and  reckless  desire  for  alteration. 

Subsequently  Kuntze  points  out  the  advantages  of  1737  as  a 
starting-point  of  botanical  nomenclature,!  in  which  he  shows  the 
number  of  generic  names  that  need  not  be  suppressed  if  1737  be 
taken  and  not  1735  (the  date  of  Linn?eus'  '  Systema ').  The 
editorial  remarks  in  the  Journal  of  Botany  should  be  read,  and  I 
need  not  repeat  them  here. 

At  p.  471  Kuntze  writes  "  a  plea  for  my  1737  proposal,  which, 
with  the  editorial  notes  that  supplement  it,  should  be  carefully 
perused." 

My  own  view  is  that  the  Berlin  proposal  to  adopt  1753  as  the 
earliest  date  both  for  generic  and  specific  names  is  worthy  of 
acceptance.  British  botanists  are  gradually  accepting  it,  and  I 
think  that  in  a  very  few  years  there  will  be  practical  unanimity 
as  regards  this  single  date. 

We  have  no  pre-Linnean  difficulties  in  reojard  to  Australian 
plants  purely  endemic.  With  the  exception  of  a  handful 
collected  by  Dampier  at  the  end  of  the  17th  century,  and  which 
were  treated  as  curiosities,  no  Australian  plants  were  collected 
until  1770,  and  none  were  described  until  a  few  j^ears  later. 
The  difficulties,  if  any,  arise  chiefly  in  regard  to  plants  which 
form  the  Indo-Malayan  element  in  our  vegetation,  the  cosmo- 
politan species  and  the  introduced  weeds. 

We  are,  therefore,  not  so  much  interested  in  this  really  impor- 
tant matter  as  our  brethren  in  other  parts  of  the  world,  and  we 
look  for  our  fellow- workers  in  Europe  and  elsewhere  to  speedily 
come  to  an  agreement  as  regards  the  date  referred  to,  when  I  do 
not  hesitate  to  say  that  we  in  Australia  will  loyally  fall  into  line 
with  them  in  this  respect. 


*  Journ.  Bot.  xxx.  57. 
t  Gaertnerisches  Centralblatt,  Berlin,  1899.    No.  2.     Translated  in  Journ. 

Bot.  xxxviii.  7. 

■^  Journ.  Bot.  xxxviii. 


PRESIDENT  S  ADDRESS.  689 

3. — Xo  Priority  for  pre-Linnean  Names. 

"We  have  made  it  a  rule  in  our  'Genera  Planfcarum'  to  yield 
no  right  of  priority  to  ante-Linnean  names,  i.e.,  those  published 
before  the  adoption  of  the  Linnean  system  of  nomenclature.  If 
we  once  give  this  right  to  Tournefort  or  Rumphius,  there  is  no 
reason  for  not  going  back  to  Bauhin  or  Clusius,  or  even  to  Pliny 
or  Dioscorides,  to  the  utter  confusion  of  all  synonymy.  Linnaeus, 
by  the  establishment  of    the  binomial  nomenclature,   made   an 

epoch  in  the  study  of  systematic  botany.""^ 

The  greater  part  of  this  paper  is  replete  with  valuable  obser- 
vations on  the  principles  which  should  underlie  the  nomenclature 
of  plants. 

There  is  one  book  to  which  Australian  botanists  are  much 
indebted,  but  which,  I  regret  to  say,  is  to  some  extent  marred  by 
the  recognition  it  gives  to  pre-Linnean  names.  I  allude  to 
Mueller's  '  Census  of  Australian  Plants,'  and  I  trust  that  his 
very  regrettable  example  in  this  respect  will  not  be  followed. 
Following  is  the  list  of  pre-Linnean  genera  adopted  by  him,  and 
the  authors'  names  should  all  be  suppressed  and  that  of  Linnaeus 
substituted,  for  they  are  allf  to  be  found  in  his  'Genera'  or  'Species 
Plantarum.' 

1.  L'Obel  (I576)i. 

Polygonum. 
L'Obel  (L581). 
Tribuliis,  also  the  species  terrestris. 

2.  DoDONiEus  (1583)  or  Dodoens  (spelt  both  ways  in  'Census'), 
the  former  being  the  Latinized  form. 

Amarantus.  Galiwrn. 

Asperula.  Vincetoxicuyn, 

*  Bentham,  G.,  'Notes  on  Euphorbiaeeae,'  Journ.  Linn.  Soc.  Bot.  xvii.  193 
(1879). 

t  Possibly  with  one  or  two  exceptions  that  may  have  slipped  my  search. 

X  I  content  myself  with  giving  the  date  of  each  work,  the  titles  of  which 
may  be  obtained  in  Mueller's  '  Census.' 


690 


PRESIDENT  S  ADDRESS. 


3.  Dalechamps  (1587). 

Abrus. 

4.  L'Ecluse  (1605). 

3fusa. 

5.  Reneaulme  (1611). 

Erythrcea. 

6.  C.  Bauhix  (1623). 

Costus. 

7.  Ray  (1682). 

Colocasia. 

8.  Hermann  (1687). 

Crotalaria. 

9.  Plukenet  (1692). 

Helicteres. 

10.  TOURNEFORT  (1700). 

Anemone. 

Ranunculus. 

NymphcEa. 

Nelumho. 

Fapaver. 

Capparis. 

Cardamine. 

Sisymbrium. 

Alyssum. 

Lepidiiim. 

Viola. 

Cakile. 

Hypericum. 

Polygala. 

Linum. 

Geranium . 

Abutilon. 

Hermannia. 

Corchorus. 

Celtis. 

Ficus. 


Scutellaria. 

Passiflora. 

Parietaria. 

Urtica. 

Fagus. 

Euonymus. 

Rhus. 

Plumbago. 

Portidaca. 

Herniaria. 

Atriplex. 

Chenopodium. 

Lotus. 

Glycyrrhiza. 

Phaseolus. 

Tamarindus. 

Acacia. 

Rubus. 

Agrimonia. 

Aristolochia. 

Myrtus. 

Zizyplius. 

Vitis. 


PRESIDENTS  ADDRESS. 


691 


Hydrocotyle. 

Eryngiwni. 

Apium, 

Shun. 

CEnanthe. 

DauGus. 

Visciim. 

Sambucus. 

Cucnmis. 

Momordica. 

Eupatovium. 

Aster. 

Senecio. 

Gentiana. 

Plantacjo. 

Anagallis. 

Lysimachia. 

Samolus. 

Jasminum. 

Olea. 

Ligustruni. 

Convolvulus. 

Cuscuta. 

Solanutn. 

Nicotiana. 

Veronica. 

TOURNEFORT  (1703). 

Salicornia. 
EUeaynus. 
TOUENEFORT  (1706). 

Lavatera. 
11.  Plumier  (1703). 
Hernandia. 
Su7'iana. 
Triumfetta. 
Tragia. 


Euphrasia. 

Orohanche. 

Acianthus. 

Ocimum. 

Mentha. 

Lycojnis. 

Teucrium. 

Verbe7ia. 

Cynoglossum. 

Heliotropiiim. 

Iris. 

Srriilax. 

Asparagus. 

Typha. 

Sparganium. 

Damasonium. 

Juncus. 

Cyperus. 

Hcirpus. 

Fanicuin. 

Oryza. 

Ariindo. 

O]jhioglossum. 

Osmunda. 

Adiantum. 

Polypodium. 

Ximenia. 
Oldenlandia. 

Lujf'a. 

Cup)ania. 
Pisonia. 
Caesalpinia. 
Bauhinia. 


692 


PRESIDENT  S  ADDRESS. 


Dioscorea. 
Commelina. 


Tahernceyyiontana. 
Ruellia. 

12.  RuMPHius  (1704). 

Dammara. 

13.  RiviNUS  (1718). 

Seseli.  Triglochin. 

Lappula.  Alisma. 

14.  Ruppius  (1718). 

Caltha.  Carex. 

Myosotis.  Lycopodium. 

15.  Vaillant  (1719). 

Elephantopus.  Helichrysum. 

Vaillant  (1722). 
Morinda. 
Vaillant  (1727). 
Pilularia. 

16.  RoYEN  (1724). 
Indiyofera. 

17.  LiNDKRN  (1721). 

Limosella. 

18.  MiTCHELi  (1729). 

Montia.  Vallisneria. 

Tillcea.  Salvinia. 

Eugenia. 

19.  Catesby  (1731). 

Xanthoxylum. 

20.  DiLLENIUS  (1719). 
Festuca. 
DiLLENIUS  (1732). 

Uremia.  Bidens. 

Melochia.  Sidei'oxylon. 

Spermacoce. 
DiLLENIUS  (1735). 
Ejjilohium. 
It  will  be  observed   that  Mueller  has  very  largely  recognised 
Tournefort's  genera;  so  did  Linnaeus,  but  it  is  the  practice  now 


PRESIDENTS  ADDRESS. 


693 


to  put  Linn.,  after  the  names  for  reasons  which  are  adequate,  and 
which  need  not  be  repeated  at  this  place.  Mueller  is  inconsistent 
when  he  gives  a  reference  such  as  this  : — "  Luffa,  Tournefort  in 
act.  Acad.  Paris,  84.  t.  2  (1706)  from  Yesling  (1638)."  If  he 
declines  to  abolish  the  authors'  names  of  pre-Linnean  botanists, 
why  draw  a  line  at  Tournefort,  as  in  this  case  1  Why  not,  as  he 
has  done  in  analogous  cases,  write  "  Luffa,  Vesling  (1638)'?" 
This  is  not  the  only  inconsistency  of  this  kind.  He  states  the  case 
according  to  the  generally  accepted  views  of  the  vast  majority  of 
botanists  in  the  entr}^,  "Curcuma,  Linne  gen.  plant.  332  (1737) 
from  Hermann  1687."  But  this  is  the  only  instance  I  can  find, 
and  is  probably  a  slip. 

Then  we  have  the  following  entries,  which  I  have  put  in  tabu- 
lar form  : — 

Species  and  Date.  Genus  and  Date. 

acquatilis,  Ranunculus, 

Dodoens,  1583.  Tournefort,  1700. 


phrag  mites  ^ 

Dodoens,  1583. 

Arundo, 

Tournefort,  1700. 

Avicennce^ 

Gerard,  1597. 

Abut  Hon, 

Tournefort,  1700. 

latifoliuyn, 

C.  Bauhin,  1623. 

Slum, 

Tournefort,  1700. 

indicum, 

J.  Bauhin,  1651. 

Ahutilon, 

Tournefort,  1700. 

ceyijptiaca, 

Miller,  1703. 

Luffa, 

Tournefort,  1706. 

(Enoplia, 

Miller,  1731. 


Zizyphus, 

Tournefort,  1700. 


These  are  cases   which  absolutely  come  within  De  Candolle's 
dictum,  "  Never  make  an  author  say  what  he  has  never  said." 


694  president's  address. 

How  could  Dodoens,  for  example,  call  a  plant  Bantmculus 
acquatilis  when  the  genus  was  not  founded  until  117  years  later  % 
It  is  absurd  on  the  face  of  it.  All  the  names  in  the  table  are 
pre-Linnean. 

Trihulus  terrestris  is  a  name  attributed  in  the  '  Census  '  to 
L'Obel  (1581),  both  as  regards  genus  and  species.  I  give  this 
because  it  is  an  Australian  instance,  but  other  authors  quote 
many  more.  It  is  not  quite  straightforward  to  attempt  to  take 
awa}'  from  Linnteus  the  credit  of  the  binomial  system.  It  is  quite 
true  that  some  of  the  early  writers  occasionall}'  use  binomials,  but 
that  is  a  mere  accident  and  has  no  signiticance.  Sometimes  a 
name  may  consist  of  eight  or  ten  words,  but  oftener  the  names 
are  three-  or  four-worded  or  more,  and  such  adventitious  binomials 
have  nothing  to  do  with  Linnajus'  binomial  S3^stem. 

But  Mueller's  action  in  not  contenting'  himself  with  ooino;  back 
to  the  author  of  binomials,  but  limiting  himself  to  the  sixteenth 
century,  is  eclipsed  by  that  of  Prof.  Edward  L.  Greene,"^  the 
Californian  botanist,  who  in  certain  cases  goes  right  back  to 
antiquity,  fathering  some  of  his  genera  on  Catullus,  Virgil, 
Theophrastus  and  Pliny. 

But  why,  oh  wdi}^  !  stop  at  such  a  latter-day  botanist  as 
Catullus'?  Let  us  explore  the  Egyptian  hieroglyphics,  the  Assyrian 
tile-books,  and  delve  into  the  innermost  recesses  of  Persian 
antiquit}^  for  botanical  names,  and,  when  we  ha^e  found  them, 
let  us  be  sure  thatwe  fit  them  on  to  the  right  plants.  Let  us  not  be 
daunted  by  the  fact  that  of  the  few  plants  referred  to  in  Holy 
Scripture  very  few  are  determinable  with  certainty.  Of  those 
that  are  so  determinable,  why  does  not  Prof.  Greene  consider 
their  Greek  or  Hebrew,  or  Chaldean  equivalents  as  worthy  of 
record  as  those  of  Catullus  ?  Is  botanical  nomenclature  to  be 
kept  in  a  simmering  chaos  until  such  time  as  architologists 
determine  the  names  our  first  parents  assigned  to  plants  in  the 
Garden  of  Eden  1 


American  Nomenclature  Again,'  Journ.  Bot.  xxix.  810. 


prksidknt's  addkess.  695 

4. — Revisions  of  Botanical  Names  must  be  the  wohk  of 
monographeks. 

Asa  Gray  ever  inculcated  caution  in  the  alteration  of  botanical 
names,  and  one  of  his  last  dicta  states,*  "the  business  of  deter- 
mining rightful  names  is  not  so  simple  and  mechanical  as  to 
younger  botanists  it  appears  to  be,  but  is  very  full  of  pitfalls.'"' 

In  a  review  of  Dr.  Taubert's  paper  in  '  Bot.  Centralblatt '  on 
the  nomenclature  of  some  genera  and  species  of  Leguminosse,  the 
editor!  adds  the  judicious  remark  : — "  In  cases  where  a  restitution 
of  an  old  generic  name  has  been  established,  it  has  been  customary 
to  leave  the  specific  combinations  as  a  whole  to  be  dealt  with  by 
a  monographer  of  the  genus,  and  this  has  the  further  advantage 
of  preventing  an  undue  increase  of  synon3'ms,  as  most  mono- 
graphers sink  a  certain  number  of  species,  which  thus  do  not 
need  re-naming." 

The  practice  of  Rudolph  8chlechter  in  this  respect  appeals  to 
common  sense.  In  his  '  Monographie  der  Podochilinse'^  he 
explains  the  necessity  of  merging  Appendicula  in  Podochilus. 
At  p.  64,  he  opens  a  sort  of  suspense  account,  "  Species  mihi  non 
visse  vel  imperfecte  nottie,"  in  wliicli  he  leaves  the  names  as  he 
finds  them.  If  the  species  of  Ajipmdicula  there  referred  to  prove 
to  be  good  species  of  Podocliihis,  they  can  be  named,  but,  until 
that  is  proved,  Schlechter  declines  the  responsibility  of  coining 
new  names  for  "species  non  visa^  vel  imperfecte  not^e," 

Berlin  Rule  2  may  be  important  to  us,  and  reads  "  The  name  of 
a  genus  is,  however,  allowed  to  lapse  if  it  has  not  been  in  general 
use  fifty  years,  counted  from  the  date  of  its  establishment.  If 
the  name  has,  however,  been  revived  in  monographs  or  large 
floras  as  a  result  of  following  the  laws  of  nomenclature  of  the 
year  1(S68,  '  that  name  is  to  hold  good.'  "     This  well-intentioned 


*  Journ.   Bot.   xxxii.   19.      '  Asa    Giay's   last   words   on   Nomenclature. 
Keprinted  from  '  The  Letters  of  Asa  Gray  '  (Macmillan). 
+  Journ.  Bot.  xxix.  .352. 
%  Mem.  de  I'Herbier  Boissier.    No,  21  (1900). 


696  president's  addkkss, 

rule  is,  however,  sometimes  difficult  to  apply  in  practice,  and  it 

will  be  sufficient  to  draw  attention  to  a  judicious  criticism  of  it."^ 

The  question  of  priority  of  names  resolves  itself  into  two  parts. 

1.  Are  the  newly  disinterred  names  actually  the  earliest  given, 
^.e.,  nearest  to  1737  or  1753?  (or  whatever  the  date  or  dates 
finally  agreed  upon). 

2.  Do  they  represent  good  genera  and  species  ?  In  other  words, 
can  they  be  accepted  by  a  competent  monographer  1 

I  deny  absolutely  the  right  of  a  man  to  alter  nomenclature  for 
literary  reasons.  Alteration  must  be  made  with  a  due  sense  of 
resi^onsibility,  and  we  must  be  assured  that  an  author  proposing 
a  change  has  at  least  critically  examined  the  genus  or  species. 

Many  alterations  in  recent  years  are,  as  far  as  I  can  understand, 
purely  literary  and  are  of  that  mechanical  character  which  could 
be  entrusted  to  any  educated  person  having  a  leaning  towards 
botanical  names.  Rather  than  entrust  the  matter  to  such  hands, 
it  would  be  far  better  to  pay  a  respectable  news-cutting  agency  to 
hunt  up  the  records,  the  evidence  obtained  to  be  sifted  and 
weighed  by  competent  authority,  say  a  Botanical  Society. 

We  must  take  all  reasonable  steps  to  prevent  the  resurrection 
of  a  number  of  wortliless  synonyms,  and  these  may  be  further 
duplicated  by  the  addition  of  new  names. 

AVe  do  not  want  botany  turned  into  librarian's  work.  The  best 
systematic  work  is  done  in  the  herbarium  and  garden,  and  the 
botanist  may  actuall}^  waste  his  time  by  spending  too  much  of  it 
in  a  library,  disinterring  long-forgotten  names  like  a  ghoul.  And 
even  if  he  does  produce  a  new-old  name  that  he  offers  for  our 
acceptance,  let  him  put  it  into  a  l^otanical  suspense  account  for  a 
period  in  order  that,  if  we  use  it  at  all,  we  may  have  some 
guarantee  that  we  have  reached  finality,  and  that  the  new-old 
discovery  may  not  be  superseded  by  a  newer-older  find. 

Some  of  the  ingenious  research  expended  on  ascertaining,  from 
circumstantial  evidence,  the  dates  of  publication  of  genera  and 
species  would  be  ludicrous  if  it  were  not  lamentable. 

*  '  The  tifty  yeuis'  limit  in  Nomenclature,'  Journ.  Bot.  xxxvi.  90, 


president's  address.  697 

We  know  that  even  in  the  20th  century,  with  crowds  of  eager 
botanists  and  abundance  of  cheap  ink  and  paper,  we  have  some- 
times difficult}'  in  ascertaining  the  precise  dates  of  publication  of 
genera  and  species.  How  much  is  the  difficulty  enhanced  as  1737 
or  1753  is  approached  !  In  the  18th  century,  for  example,  we 
have  many  instances  of  books  (I  am  not  merely  referring  to 
botanical  ones)  whose  publication  was  delayed  for  various  reasons, 
and  whose  date  of  publication  is  certainly  not  that  of  the 
imprint. 

Some  works  {e-g.,  Beechey's  '  Voyage  ')  have  been  published 
with  a  title  page  at  the  end  of  the  work  {e.g.,  1845),  while  the 
individual  parts  have  been  dated  1843,  4,  etc.  Are  we  certain 
that  we  always  quote  correct  dates  for  the  species  referred  to  in 
this  work  ?     As  a  matter  of  fact  I  know  we  are  not. 

Following  is  a  purely  Australian  example.  The  excessivel}^ 
rare  work,  'Ned.  Kruidk,  Archief,'  which  contains  Miquel's 
valuable  paper  on  Australian  plants,  in  which  he  described  a 
number  of  species  sent  to  him  by  Mueller,  was  originally  published 
in  1856  as  a  very  thin  part;  the  date  quoted  in  the  'Flora 
Australiensis,'  viz.,  1859,  is  that  of  the  volume  (iv.)  containing  the 
part. 

Another  objection  to  disinterring  old  names  simpl}^  because 
they  appear  to  be  older  than  those  in  use  is  illustrated  by  the 
following  : — 

In  a  paper  Mr.  Britten"^  asserts  that  ChIamysporu7n,  Salisbury, 
antedates  Thysanotus,  Robert  Brown;  and  in  a  second  paperf  the 
same  author  suggests  the  suppression  of  some  of  Brown's  names 
in  favour  of  those  of  Salisbury.  At  p.  298,  Bishop  Goodenough 
accuses  Salisbury  of  cribbing  from  Brown,  and  it  is  well  known 
that  Brown  spoke  very  plainly  to  his  contemporaries  about 
Salisbury,  who,  probably  in  part  for  his  failing,  was  a  kind  of 
Ishmael  amongst  his  contemporaries.     Brown  we  know  as  one  ot 

*  '  Hookera  v.  Broclisea,  with  some  remarks  on  Nomenclature,'  Journ.  Bot. 
xxiv.  49. 

t  '  On  the  Nomenclature  of  some  Proteacese,'  Journ.  Bot.  xxiv.  296. 


698  president's  address. 

the  most  eminent  botanists  of  that  or  any  age.  Salisbury's  names 
were  rejected,  those  of  Brown  accepted  by  his  contemporaries. 
A  thief  may  technically  have  priority  by  hurrying  publication, 
but  he  can  only  be  effectively  punished  by  declining  to  accept  his 
names.  At  the  present  day  there  are  so  many  media  of  prompt 
publication  open  to  an  author  that  a  cribber  has  little  opportu- 
nity of  pursuing  a  dishonest  course,  but  a  centnr}^  ago  (and  much 
more  so  fifty  years  previously)  there  were  few  opportunities  of 
publication,  and  the  periods  which  often  elapsed  between  the 
reading  of  a  paper  and  its  publication  appear  to  us  extraordinary. 

5. — Names  in  MSS.   or  Herbaria. 

Art.  50,  '  Lois  de  la  Nomenclature  Botanique,'*  reads  as 
follows  : — "  Les  noms  publies  d'apres  un  document  inedit  tel  qu'un 
her  bier,  une  collection  non  distribuee,  etc.,  sont  precises  par 
I'addition  du  nom  de  I'auteur  qui  publie,  malgre  I'indication  con- 
traire  qu'il  a  pu  donner,  De  meme  les  noms  usites  dans  les 
jardins  sont  precises  par  la  mention  du  premier  auteur  qui  les 
publie. 

"  Dans  de  texte  developj)e,  on  cite  I'herbier,  la  collection,  le 
jardin  (Lam.  ex  Commers.  mss.  in  herb,  par.;  Lindl.  ex  horto 
Lodd.)." 

J.  Britten,  '  A  point  in  Botanical  Nomenclature,'!  shows  that 
some  botanists  ignore  Art.  50,  and  points  out  some  of  the  incon- 
veniences which  arise  from  such  action. 

Berlin  Rule  13  says,  "Manuscript  names  have  not  under  any 
<3ircumstances  a  right  of  consideration,  even  when  they  appear  on 

printed   labels  in  exsiccata For  the  recognition  of 

any  species  a  printed  diagnosis  is  required,  which  may  of  course 
appear  on  an  exsiccata  label." 

My  own  practice  has  hitherto  been  to  follow  a  name  approved 
(by  me)  in  MSS.  or  herbarium,  e.g.,  '■'■Eucalyptus  vhninalis,  Labill. 
Tar.  pedicellaris,  F.v.M." 

*  Alph.  de  Candolle,  1867. 
t  Journ.  Bot.  xx.  53. 


president's  address.  699 

The  name  given  in  italics  should,  according  to  Art.  50,  be 
written  "  Eucalyptus  viminalis,  LabilL,  var.  pedicellai'is,  Deane 
and  Maiden  ex  F.v.M.  mss.  in  herb.  Melb.,"  which  would  of 
course  be  contracted  in  practice,  the  var.  pedicellarls  being  attri- 
buted to  Deane  and  Maiden  instead  of  to  Mueller,  who  originally 
discriminated  it.  Both  methods  are  open  to  objection,  but 
inasmuch  as  the  original  user  of  a  name  may  have  deliberately 
suppressed  it,  it  may  be  an  injustice  to  him  to  revive  it.  On  the 
other  hand,  his  omission  to  revive  the  name  ma}''  be  a  mere  over- 
sight, in  which  case  it  is  an  injustice  to  him  to  omit  to  credit  him 
with  it.  This,  however,  trenches  on  the  question  as  whether  an 
author's  name  should  be  used  at  all,  which  T  have  discussed  at 
p.  703. 

To  show  that  there  is  a  difference  of  opinion  amongst  eminent 
botanists  in  regard  to  dealing  with  herbarium  names,  I  need  only 
quote  the  following  : — 

*'  The  distribution  of  named  specimens  (and,  aj'ortiori,  of  those 
in  sets,  widely  distributed  in  herbaria,  as  were  Sieber's),  where 
and  as  far  as  they  go,  is  held  to  be  tantamount  to  publication." 

*'  So  of  names  and  original  observations  attached  to  specimens 
in  herbaria.  These  names  are  always  attached  antecedently  to 
publications,  and  a  monographer,  having,  as  he  should,  free  access 
to  all  herbaria  within  his  reach,  might  work  a  deal  of  harm  if  he 
did  not  regard  such  names  as  to  him  all  the  same  as  if  already 
published."* 

I  am  afraid  that  if  Sieber's  names  were  suppressed,  or  even  if 
the  authority  for  the  species  names  were  altered  from  "  Sieber  " 
to  "  Bentham,"  etc.,  no  advantage  to  botanical  science  would 
result,  but  much  incon\'enience  and  uncertainty  might  arise.  The 
name  of  another  Australian  collector,  Allan  Cunningham,  will 
occur  in  this  connection.  Some  well  known  plants  which  bear  his 
name  were  never  described  by  him  at  all,  but  as  he  recognised  the 
species  as  new  and  even  named  them,  Schauer  and  others  wrote 

*  Asa  Gray  in  Silliman's  Journal,  March,  1864,  p.  279,  quoted  in  Journ. 
Bot.  V.  83. 


700  president's  address. 

the  descriptions  after  his  death  in  his  name.  I  admit  that  this 
seems  like  an  infringement  of  the  golden  dictum"^  "  neA^er  make  an 
author  say  what  he  has  not  said." 

Viewing  the  matter  afresh,  I  am  of  opinion  that  Art.  50  is 
least  open  to  objection  and  should  be  followed.  At  the  same 
time  I  think  if  it  were  made  too  retrospective  it  would  become  a 
dead  letter. 

In  some  of  my  papers  I  have  taken  cognisance  of  a  few  names 
which  are  unpublished  (or  apparenth^  unpublished)  attached  to 
specimens  collected  b}^  Robert  Brown,  and  some  of  the  older 
botanists,  as  a  matter  of  expediency,  as  many  of  these  specimens 
have  been  widely  distributed  in  herbaria,  and  clues  as  to  the 
identity  of  species,  particularl}^  those  in  an  exceptionally  difficult 
genus,  are  often  valuable. 

In  this  connection,  I  understand  that  some  English  botanists 
of  the  highest  reputation  will  not  quote  names  suspected  to  be 
nomina  nuda:  I  feel  sure  that  this  practice,  which  has  the  most 
praiseworthy  intention  of  reducing  unnecessary  names,  may  be 
carried  too  far.  In  some  cases  names  .ma}'^  be  printed  as  an  inti- 
mation that  further  information  in  regard  to  the  plants  referred 
to  is  desired.  Some  genera  are  so  special  in  the  intricate 
character  of  their  synonymy  that  the  rigid  rules  of  purists  may 
not  always  be  expedient,  as  they  may  actually  hinder  the  advance- 
ment of  knowledge. 

For  exam.ple,  I.  B.  Balfourf  says: — "  In  the  following  list  I  have 
given  all  the  names  of  species  I  have  been  able  to  find,  quoting  the 
authority  and  giving  a  reference  to  where  it  may  be  found.  Of 
many  I  know  nothing  save  the  name  ;  and  it  may  be  taken  for 
granted  that  a  great  number  have  no  claim  to  be  species;  but  I 
have  thought  it  advisable  to  follow  the  plan  of  giving  all  the 
names  at  present  when  seeking  for  information.'' 

Specimens  labelled  in  herbaria  ma}^  be  perfectly  good  species,  and 
it  may  be  desirable  to  resuscitate  these  names  to  prevent  the  use 

*  Alph.  De  Candolle  in  Bull.  Belg.  xiv.  1877,  p.  482. 
t  "  On  the  genus  Pandanus."     Journ.  Linn.  Soc.  Bot.  xvii.  36. 


president's  address.  701 

of  superfluous  ones.  That  eminent  European  botanists  revive 
nomina    ntida    when    it    suits    their    convenience    the    followingf 

o 

example  will  show.  I  think  Mr.  Salmon's  action  is  quite  justifi- 
able, for  he  shows  that  Sullivan  and  Lequesne's  and  Brotherus 
names  are  identical,  which  some  students  may  be  glad  enough 
to  know. 

"  In  the  Kew  Herbarium  there  is  a  moss  labelled  '  Papillaria 
torticuspis,  Broth.,  no  v.  spec.  Birmah.  comm.  L.  Linden;  received 
December,  1893.'  This  name  has  not,  so  far  as  I  can  find,  been 
published.     The  moss  is  typical  A.  Toccooi.^''*'  .... 

We  Australian  botanists  often  pursue  our  work  with  difficulty 
because  the  types  of  so  many  of  our  plants  are  in  Europe,  and 
some  of  the  descriptions  are  not  available  in  Australian  libraries. 
This  causes  us  sometimes,  perhaps,  to  quote  nomina  nuda  in 
ignorance,  and  European  writers  can  sometimes  set  us  right  on 
questions  of  nomenclature  if  they  are  so  disposed.  Any  short- 
comings of  this  character  may  be  looked  upon  quite  leniently,  if 
Only  because,  on  our  part,  we  are  frequently  able  to  correct  the 
descriptions  and  notes  of  non- Australian  workers  on  Australian 
plants,  who  are  usually  unfamiliar  with  some  points  in  regard  to 
the  plants  described  by  them. 

6. — Uniformity  of  Names  of  Botanical  Authors. 

Alphonse  De  Candollef  wrote  a  letter  to  B.  Daydon  Jackson  on 
the  orthography  of  the  names  of  botanical  authors,  which  is 
worthy  of  close  attention.  He  points  out  that  "  Quelques 
botanistes  ont  ajoute  a  leur  nom  un  autre  nom,  de  fantaisie,  par 
exemple : — 

Brisseau  s'est  affuble  du  nom  de  Mirbel 
Proveniat      ,,  ,,  St.  Hilaire 

Palisot  ,,  „  de  Beauvois 

Nees  ,,  ,,  d'Esenbeck," 

which  should  be  "  Mirbel  voyez  Brisseau,"  and  so  on. 

*  '  Bryological  Notes  on  Anojiiodon  Toccoce,  Sulliv.  &  Lesq.'  By  Ernest  S. 
Salmon.     Journ.  Bot.  Aug.  1902,  p.  273. 

t  Journ,  Bot.  xviii.  217. 
46 


702  president's  address. 

It  happens  that  all  these  authors  described  A  ustralian  plants, 
so  that  these  particular  corrections  by  De  Candolle  should  be 
noted. 

It  is  a  matter  for  regret  that  F.  von  Mueller  was  a  law  unto 
himself  in  regard  to  some  names.  For  example,  European 
practice  (including  Bentham  in  the  'Flora  Australiensis')  adopted 
Muell.  Arg.,  and  Hook,  f.,  but  F.  von  Mueller  chose  to  write  J. 
Muell.  and  J.  Hook.  He  was  not  consistent,  for  he  used  Linn.  f. 
The  writing  of  Cunn.  instead  of  A.  Cunn.,  is  a  difference  from  the 
'Flora  Australiensis,'  and  I  am  one  of  those  who  think  that  that 
classic  should  be  followed  as  closely  as  possible  in  details,  for  it  is 
a  model  of  orderly  arrangement.  Of  course  I  do  not  for  a 
moment  advocate  a  slavish  adherence  to  this  or  an}'-  other  work 
in  cases  where  the  progress  of  botanical  knowledge  shows  that 
modifications  and  corrections  are  necessary. 

Mr.  Daydon  Jackson  has  a  useful  paper*  in  which  (p.  82)  he 
quotes  examples  of  vicious  methods  of  contracting  authors'  names 
which  is  well  worthy  of  perusal. 

Art.  xlviii.  (De  Candolle's  'Lois')  says  that  it  is  necessary  to 
publish  the  name  of  an  author  of  a  species,  etc.  He  further  says,t 
"  The  purpose  of  the  citation  of  the  author's  name  is  misunder- 
stood by  some.  This  is  merely  an  abridged  form  of  a  biblio- 
graphical indication,  intended  to  establish,  without  lengthy 
research,  the  date  which  fixes  the  priority  of  a  name." 

Let  me  quote  Bentham's  wordsj  : — "  Were  everj'-one  agreed  as 
to  the  plant  to  be  designated  by  a  particular  name,  the  binomial 
appellations  devised  by  Linnseus  would  be  quite  sufficient  in  all 
cases  when  a  species  is  referred  to  for  comparison.  But  it  so 
frequently  happens  that  different  authors  have  given  the  same 
name  to  different  plants,  that  the  addition  of  a  third  word  (the 
abbreviated  name  of  the  author)  has  become  indispensable  in  some 


*  Journ.  Bot.  xix.  75.     'On  some  recent  tendencies  in  Botanical  Nomen- 
clature.' 

t  Journ.  Bot.  xxvi.  290. 

X  Journ.  Linn.  Soc.  Bot.  xvii.  190  (1879). 


PR BSi dent's  address.  703 

instances  and  advisable  in  most  cases,  to  avoid  uncertainty,  but 
for  no  other  object." 

Authors  of  species  are  only  quoted  as  a  matter  of  convenience, 
and  not  to  reward  authors.  Hence  the  authors'  names  should  be 
very  clearly  indicated,  contractions  being  avoided  if  possible,  as 
uncertainty  in  regard  to  a  name  hinders  and  perhaps  prevents  the 
bibliographical  references  referred  to. 

Where  we  use  contractions  we  should  adopt  the  accepted  con- 
tractions. The  names  of  Australian  botanists  of  this  and  the 
preceding  generation  who  have  not  yet  attained  cosmopolitan 
renown  should  be  quoted  without  abbreviation. 

One  effect  of  recent  changes  of  nomenclature  lias  been  to 
facilitate  the  attribution  of  different  authors  to  the  same  specific 
name.  William  Matthews*  gives  an  extreme  instance  in  the  case 
of  Erodium  moschatum,  a  plant  familiar  to  us,  and  which  is  by 
different  writers  attributed  to  L'Heritier,  Alton,  Willdenow,  and 
Smith. 

Berlin  Rule  7  says,  "  The  author  who  first  named  the  species, 
although  in  another  genus,  is  always  to  be  acknowledged,  and  his 
name  is  therefore  to  be  placed  in  brackets  before  that  of  the  author 
who  has  removed  the  species  into  another  genus,  thus,  Pulsatilla 
pratensis  (L.),  Mill.  If 'an  author  has  himself  placed  his  specieg 
in  another  genus,  the  bracket  is  omitted." 

This  practice  has  been  little  followed  in  Europe,  and  not  at  all 
in  Australia  I  believe.  An  Australian  example  shows  how  it 
works  :  — 

Damasonium  australe,  Salisb.,  in  Trans.  Hort.  Soc.  Lond.  i.  268 
(1812),  in  'Flora  Australiensis  '  and  Mueller's  'Census.' 

In  '  Natiirliche  Pflanzen-Familien,'  Buchenau  changes  the  name 
into  Dmnasonium.  minus,  (R.Br.)  Buchenau,  because  the  plant 
was  first  described  by  R.Br,  as  Actinocarpus  minor,  R.Br.,  and  if 
the  name  of  the  genus  is  changed,  the  specific  name  of  the  man 
who  described  the  plant  first  should  be  preserved,  unless  there  are 
good  reasons  against  it. 

*  Journ.  Bot.  xvi.  260. 


704  PRESIDENT  S  ADDRESS. 

Anothei'  instance  is  : — 

Bentham  ('  Flora  Australiensis')  writes  Brasenia  jyeltata,  Pursh. 

In  '  Nat.  Pfl.  Fam.'  the  name  is  given  as  Brasenia  2)uiyurea, 
(Michx.),  Casp.,  following  Hydropeltis  pur2mrea,  Michx.  (1S03), 
the  first  used  specific  name.  [Mueller,  '  Census,'  writes  Cabomha 
peltata,  F.V.M.,  but  the  'Nat.  Pfl.  Fam.'  shows  that  Cahomha  and 
Brasenia  are  distinct  genera.] 

What  has  been  styled  the  "  Kew  Rule"  is  that  of  consider- 
ino-  as  the  right  name  of  the  plant  that  by  which  it  was  first 
called  in  the  genus  wherein  it  is  placed.  The  Berlin  Rule  recog- 
nises the  first  specific  name  given,  w^hether  the  genus  is  maintained 
or  not.  The  use  of  the  Berlin  Rule  (which  is  popular  in  America) 
has  in  practice  necessitated  the  use  of  two  authors'  names,  viz., 
the  author  who  gave  the  first  specific  name;  this  is  written  first, 
and  is  in  brackets;  then  comes  the  author  of  the  name  which  is 
accepted. 

I  do  trust  our  European  and  American  brethren  will  soon  come 
to  an  agreement  both  as  to  the  adoption  of  the  Kew  or  Berlin 
Rule,  and  also  as  to  the  use  (personally  I  would  like  to  see  the 
bracket-name  abolished  as  cumbersome)  of  the  double-name  or  the 
reverse.  VVe  do  not  wish  to  institute  a  third  or  Australian 
practice,  and  recognise  that  in  these  matters  there  are  larger 
interests  at  stake  and  more  botanists  affected  than  with  us. 

7. On  the  Writing  of  Specific  Names. 

Mr.  Daydon  Jackson's  paper*  gives,  inter  alia,  Linnaeus' 
practice  in  regard  to  the  use  of  capitals  for  specific  names.  He 
quotes  five  examples,  and  Mr.  Jackson's  words  are  in  inverted 
commas. 

{a).   "  An  old  generic  name,  e.g., 

Crassula  Cotyledon^  Haw.,"  itc. 

This  is  a  usage  which  many  botanists  adhere  to  with  some 
tenacity,  and  was  in  recent  years  included  in  the  Berlin  Rules. 


*  *  On  some  recent  tendencies  in  Botanical  Nomenclature.'     Journ.  Bot. 
xix.  75. 


president's  address.  705 

(6).   "  A  native  name,  e.(j.. 

Cassia  Canca,  Cav.,"  &c. 

I  am  inclined  to  think  that  this  should  be  retained.  It  is  but 
another  form  of  («s),  and  whether  the  name  is  a  pagan  or  a 
Christian  one  does  not  matter.  The  special  difficulty  attending 
the  putting  of  native  names  without  a  capital  lies  in  the  fact 
that  some  of  them  have  what  may  be  termed  a  classical  appear- 
ance, and  it  is  desirable  to  prevent  the  waste  of  time  caused  by 
hunting  Latin  and  Greek  dictionaries  for  clues  to  their  etymology. 

(c).  "A  substantive  used  instead  of  an  adjective  (an  uncommon 
case),  e.g., 

Eucalyptus  Globulus,  LabilL,"  etc. 

I  see  no  reason  for  retaining  the  capital  letter  in  this  case.  In 
the  case  of  the  genus  Eucalyptus,  Mueller  named  two  species, 
Leiicoxyloii  and  Doratoxylon;  but  his  use  of  the  capital  letter  was 
never  widely  followed,  nor  was  he  consistent  in  regard  to  this 
form  of  specific  name  in  other  genera. 

At  p.  '2^0  {op.  ciL),  Daydon  Jackson  expresses  the  opinion  that 
the  capital  in  cases  such  as  U.  Globulus  cannot  be  maintained, 
and  gives  instances  of  Linnaeus'  oversight  or  inconsistency  in 
printing  man}^  of  his  specific  names,  showing  that  we  must  not 
assume  that  Linnseus  meant  more  than  he  did  in  printing  the 
names  in  the  way  he  did. 

{d).   "  A  substantive  used  in  the  genitive  case,  e.g., 
Pyrethrum  Halleri,  Willd.,"  etc. 

I  think  the  use  of  the  capital  is  only  desirable  with  proper 
names.     Bentham  and  Mueller  both  use  capitals  in  such  cases. 

(e).   "  A  substantive  used  adjectively  in  commemoration,  e.g., 
Salix  Russelliana,  Sm.,"  tfec. 

As  a  very  general  rule  I  think  botanists  use  {d)  or  (e)  indis- 
criminately to  commemorate  proper  names.  In  some  cases 
euphony  determines  the  form;  in  many  cases  it  does  not  seem 
apparent  what  consideration  has  weighed  with  the  bestower  of 
the  name  whether  he  has  used  the  genitive  or  not.  But  this  is 
certain,  that  either  the  genitive  or  adjectival  form  should  be 
used. 


706  president's  address. 

I  think,  therefore,  some  of  these  old  Linnean  rules  are  some- 
what fanciful  and  of  no  real  value,  simply  adding  to  the  pitfalls  of 
botanical  nomenclature  and  giving  the  literary  person  oppor- 
tunities of  annoying  the  botanist  by  his  unnecessarj'  criticisms 
and  interference. 

The  following  is  taken  from  the  '  Additions  to  the  Berlin  Rules 
of  Nomenclature,'*  where  it  is  laid  down  : — 

"  All  names  of  species  are  to  be  written  small,  except  those 
derived  from  names  of  persons.  (Examples :  Ficus  indica, 
Opuntia  ticus-indica  but  Isachne  Buettneri)." 

Engler  remarks  in  regard  to  this  rule,  "  Though  he  has  signed 
the  rules  it  goes  against  his  feelings  to  write  those  specific  names 
which  are  genitives  of  nouns  with  small  letters.  He  would  prefer 
to  write  Paccinia  Malvacear am,  Aecidiwni  Berheridis,  instead  of 
Puccinia  incdvacearam."  These  genitives  seem  to  be  confined 
to  Mycology  only,  and  while  it  seems  to  go  against  the  grain 
to  write  small  letters  in  these  instances,  I  recommend  the 
adoption  of  the  Berlin  Rule.  One  reason  of  my  adhesion  to 
this  additional  Berlin  Rule  may  seem  comical, — although  I  have 
tried  over  and  over  again,  my  pen  refuses  to  write  muelleri, 
benthami,  and  so  on.  It  hurts  nw  feelings  to  have  a  dis- 
crepancy in  this  case  between  botanical  and  ordinary  literary 
practice. 

The  additional  Berlin  Rule  deals  with  geographical  adjectives, 
which  are  to  be  written  with  small  letters.  Those  who  prefer  to 
write  specific  names  derived  from  names  of  persons  with  capitals 
may  object  to  writing  the  geographical  names  with  small  letters, 
but  in  practice  comparatively  few  do,  and  I  hope  the  Berlin  Rule 
will  be  followed. 

There  seems  to  me  an  actual  difference  in  the  meaning  of 
geographical  and  personal  specific  names.  For  example,  i<iniiatrana 
is  intended  to  convey  that  the  plant  occurs  in  Sumatra;  the  use 
of  the  name  Jonesii  is  frequently  onl}'^  a  compliment  paid  to  a 


*  Op.  fit.,  p.  G86.     This  modifies  No.  8  of  the  original  Berlin  Kules. 


president's  address.  707 

person  of  the  name  of  Jones,  who  may  or  may  not  be  otherwise 
id  entitled  with  the  plant  in  any  way. 

The  following  specific  names  taken  from  the  '  Census  '  show 
that  Mueller  held  different  views: — Novse-Hollandise,  Novse- 
Cambrioe,  Zeylanica,  Africana,  Tasmanica,  Magellanica,  Indica, 
Malabarica,  Malabathricum,  Arnhemicus,  Tannensis,  Carpentarise, 
Abyssinica,  Javanica,  Philippinensis,  Timorensis,  Balonnensis, 
Neo-Cambrica,  Neo-Anglica,  Kiamensi^i,  Aegyptiaca,  Americana, 
Gangeticum,  Truxillensis,  Mexicana,  Asiatica,  Maderaspatana, 
Celebica,  Xova-Zealandiji?,  Peruviana,  Derwentia,  Nepalensis, 
Curassavicum,  Europ£eum,  Malaccensis,  Halepensis,  Tunbridgense, 
Aethiopicum,  Capensis,  Vitiense,  Arcturi,  Americana,  Japonicum, 
Sibirica,  Chinensis,  Moluccensis,  Sumatrana,  and  even  Capri- 
cornica  !  yet  antipoda  ! 

Australis  is  not  a  place, — in  this  continent  it  is  commonly  used 
for  Australia,  but  it  means  South,  and  hence  is  also  used  to  indi- 
cate S.  Europe,  S.  America,  S.  North  America,  and  so  on. 
Mueller  established  the  practice  of  writing  Australis,  Austra- 
lasica,  Australiana, 

Berlin  Rule  11  says,  "it  is  allowable  to  dispense  with  priority 
when  it  is  a  question  of  names  which  have  arisen  through  obvious 
errors  of  geography  on  the  part  of  the  author,  such  as  Asclejnas 
syriaca,  L.  (which  comes  from  the  United  States)." 

At  the  same  time  I  do  not  doubt  that  this  rule  does  not  apply 
in  cases  where  the  specific  name  is  only  partl}^  descriptive  of  the 
range  of  the  plant. 

Following  are  the  remaining  instances  of  the  use  of  capitals 
for  specific  names  in  the  '  Census  '  : — 

Litscea  Hexanthus,  A.  L.  de  Jussieu, 

Cissanipelos  Pareira,  Linn. 

E7'ysimiim  Blennodia,  F.v.M. 

Melia  Azedarach,  Linn. 

Hibiscus  Ahehnoschu^,  Linn. 

Bossicea  Aquifolium,  Benth. 

Acacia  Sentis,  F.v.M. 
,,        Gnidiutn,  Benth. 


708  president's  addrkss. 

Acacia  Oxi/cedrus,  Sieb. 
GreviUea  Aqiti/olium,  Lindl. 

„  Leucadendroii,  A.  Cunn. 

Hakea  Bjnglottis,  Labill. 
Banksia  Rohur,  Cav. 
Dryandra  Serra,  K.Br. 

,,  Arctotidis,  R.Br. 

Coleocoina  Centaurea,  F.v.M. 
Gynurus  Pseudo-China,  DC. 
Solanum  Hystrix,  R.Br. 
Josephina  Imjoeratricis,  Vent. 
Vitex  Lignuvi-Vitre,  A.  Cunn. 
Styphelia  Bpacridis,  F.v.M. 

,,  Oxycedriis,  Labill. 

,,  Bossicea,  F.v.M. 

Cyperus  Eragrostis,  Vahl. 
Fanicum  Crus-Galli,  Linn. 

Some  are  Linnean  names.  Of  course  Linn^us  was  inconsistent 
in  the  use  of  capitals,  or  perhaps  printers  made  mistakes  in  those 
days  just  as  in  ours.  At  all  events  one  can  prove  various  things 
by  laying  too  much  stress  on  the  use  of  capitals  in  Linnaeus' 
'  Species  Plantarum.' 

As  to  the  other  instances,  some  of  the  capitals  indicate  those 
of  well-known  genera.  Those  who  have  the  time  to  waste  may 
ascertain  under  w4iich  of  the  Linnaean  Rules  (if  any)  the  use  of 
capitals  has  come  about  in  some  of  the  other  cases.  My  own  view 
is  that  the  fact  that  the  meanings  of  the  capitals  cannot  be  given 
by  the  average  botanist  on  sight  is  an  argument  for  their 
suppression,  except  in  the  case  of  proper  names. 

8. — The   Question   of   Altering  Botanical  Names  with  the 
VIEW  to  their  "  Improvement." 

Art.  3  of  the  'Lois'  stands  thus: — "Dans  toutes  les  parties 
de  la  nomenclature,  le  principe  essentiel  est  d'eviter  ou  de  repousser 
I'emploi  de  formes  et  de  noms  pouvant  produire  des  erreurs,  des 


president's  address.  709 

equivoques  ou  jetei*  de  la  confusion  dans  la  science.  Apres  cela, 
ce  qu'il  y  a  de  plus  important  est  d'eviter  toute  creation  inutile  de 
noms,  Les  autres  considerations,  telles  que  la  correction  grammati- 
cale  absolue,  la  regidaritc  ou  Teuphonie  des  noms,  un  usage  plus 
ou  moins  repandu,  les  egards  pour  des  personnes,  etc.,  malgre  leur 
importance  incontestable,  sont  relativement  accessoires." 

Names  must  not  be  altered,  except  for  a  grave  reason.  Asa 
Gray  says,^'  '•  Mere  improvement  is  no  warrant  for  alteration. 
Mistakes  may  indeed  be  corrected.  Thus  Nuttall's  genus 
Wisteria,  in  honour  of  Dr.  Wistar,  was  properly  corrected  to 
Wistaria  in  conformity  with  the  rules  that  personal  names  should 
retain  their  orthography  as  nearly  as  possible." 

Mueller  ('  Census  ')  changed  Dicksonia  antarctica  to  D.  Billar- 
dieri,  because  he  said  it  does  not  extend  to  antarctic  regions.  The 
Berlin  Rules  were  not  promulgated  at  the  time,  but  I  think  it 
would  be  considered  as  overstraining  Rule  11. 

Take  another  case,  Fiinhristylis  diphylla,  Vahl,  Enum.  ii.  289 
(1806).  (Syn.  F.  communis,  Kunth,  Enum.  ii.  234  (1837). 
Mueller  accepts  Kunth's  name  {F.  communis)  in  his  '  Census'  in 
spite  of  the  obvious  priority  of  Vahl's  name.  This  seems  to  me 
another  instance  of  Mueller's  habit  of  rejecting  specific  names 
because  the  name  is  badly  chosen.  The  name  diphylla  is  a  very 
bad  one,  and  was  probably  given  by  Vahl  because  his  type  speci- 
mens had  mostly  two  involucral  bracts. 

The  late  R.  D.  Fitzgerald  described  an  orchid  under  the  name 
Dendrohium  fcdcorostrum.  Later  on,  in  publishing  a  plate,  he 
\Yvote  falcorostris.  Mueller  ('Census')  improved  the  name  into 
falconirostre,  which  Fitzgerald  resented,  and  I  think  rightly  so. 
The  correct  name  is  fcdcorostrum,  which  was  the  original  one. 

These  three  Australian  examples  are  perhaps  sufficiently  illus- 
trative. Non- Australian  instances  have  been  quoted  by  the  dozen. 
For  example,  B.    Daydon   Jacksonf  points  out  that  crassinervia 

*  Silliman's  Journ.,  May,  1880,  quoted  in  Journ.  Bot.  xviii.  186. 
t  Journ.  Bot.  xix.  78. 


710  president's  add  HESS. 

may  not  be  structurally  correct  from  a  Latinist's  point  of  view^ 
yet  it  msij  stand. 

And  again,  "This  is  something  worse  than  writing  Potamogeto'n 
lanceolatus,  Sm.,  for  Smith,  considering  the  generic  name  as  neuter, 
wrote  lanceulatum,  and  if  we  use  his  name  we  must  do  so  too."* 

I  hope,  however,  nothing  I  have  said  will  be  interpreted  into 
acquiescence  with  slovenly  methods  of  giving  names.  Let  us 
have  ideals  as  high  as  scientific  men  in  older  parts  of  the  world. 

Let  us  look  upon  the  matter  from  another  aspect.  Consider  our 
common  Gompholohiuni  latifolinm,  Sm.,  which  has  larger  flowers 
than  those  of  G.  (jrandijiorum,  Sm.  Is  grandijioimin,  then,  to 
be  suppressed  ?  If  so,  all  specific  names  indicating  comparisons, 
definite  geographical  localities,  etc.,  etc.,  may  be  suppressed,  and 
botanical  nomenclature  put  into  a  simmer;  for  how  can  any  man 
who  gives  a  species  with  any  descriptive  name  be  certain  that 
at  some  time  or  other,  in  some  part  of  the  world  or  other,  some 
plant  belonging  to  the  same  genus  may  be  discovered  which  might 
be  more  appropriately  described  by  the  specific  name  attached  to 
another  plant?  And  how  about  variation  through  environment. 
Leaves,  for  example,  may  vary  in  an  extraordinary  manner  in 
the  same  species. 

There  are  plenty  of  legitimate  avenues  for  the  energies  of 
archaeologists  and  philologists  without  their  being  permitted  to 
add  new  worries  to  the  life  of  .the  botanical  systematist.  Let  us, 
as  far  as  convenient,  have  precision  in  botanical  names;  this  is 
best  acquired  by  inculcating  caution  on  the  part  of  the  botanist, 
who,  prior  to  the  bestowal  of  a  name,  should  he  feel  diflident  in 
the  matter,  can  usually  obtain  the  advice  of  a  friend  who  knows 
more  about  languages  than  he  does.  The  motto,  '■'' Quieta  non 
movere,'"  is  an  excellent  one  to  offer  to  the  would-be  disturber  of 


*  B.  Daydon  Jackson  in  Journ.  Bot.  xix.  SO. 


PRESIDENT  S  ADDllESS. 


711 


9. — The  Botany  of  Cook's  Voyage. 

The  publication  of  the  'Botany'  of  Cook's  'Voyage,'*  very  sHghtly 
referred  to  in  my  last  address,  deserves  more  extended  notice  at 
this  place,  for  Australians  in  particular  are  much  indebted  to  the 
Trustees  of  the  British  Museum  and  to  Mr.  J.  Britten,  its  editor. 

I  have  tabulated  the  instances  in  which  Mr.  Britben  departs 
from  the  nomenclature  of  the  '  Flora  Australiensis,'  and  the  list 
is  worth}'  of  detailed  study.  The  alterations  are  of  two  kinds, 
those  by  monographers;  the  name  Pleiogynium  Solandri^  Engler, 
may  be  taken  at  random  as  an  instance  of  this. 

Some  of  the  others  appear  to  be  of  the  character  stigmatised  by 
me  as  mere  literar}^  corrections,  and  I  advise  my  fellow  Australians 
not  to  accept  them  until  they  have  been  approved  by  competent 
monographers.  I  have  added  notes  to  some  of  the  names,  but 
they  are  by  no  means  of  an  exhaustive  character. 

The  alterations,  Candollea,  Styj>lielia^  etc.,  accepted  by  Mueller 
will  be  discussed  by  Mr.  Betche  and  myself  in  our  forthcoming 
revised  Census  of  New  iSouth  Wales  Plants. 


No. 


Name  on  Plate. 


Name  in  '  Flora  Australiensis.  ' 


b.Adelioides  deGumheni<,^^v^^k     Adeliopsis  decumhens,  Benth. 

Sol. 
S.Calceolar-ia    enneas2:>e7"ma,    O.     lonldium  suJfruticosuiUjGing. 

Kuntze. 
9.  Calceolaria     filiformit^,     O.         lonidium  filiforme,  F.  Muell. 
Kuntze. 
\O.Maximilianea     yiUivraei,     O.      Cochlospertnum   Gillivraei, 
Kuntze.      Accepted  in  the         Benth. 
'Natiirliche    Pflanzen-Fami- 
lien.' 


"''  "  Illustrations  of  the  Botany  of  Captain  Cook's  Voyage  round  the  World 
in  H.  M.S.  'Endeavour'  in  1768-71,  by  Sir  Joseph  Banks  and  Daniel  Solander, 
with  determinations  by  James  Britten.     Part  i   1900;  Part  ii.  1901." 


712 


PRESIDENT'S  ADDRESS. 


No.  Name  on  Plate. 

13.  PolygaJ  a    lonyifolia,    Poir. 

Doubtless  correct.  Two 
distinct  species  are  also 
given  in  the  Index  Kewen- 

sis. 
X^.Cosmia  quadrivalvis,  Britten. 

19.  Abutilon  indicum,  Sweet,  var. 

aust7'aliense,  Hochreut. 
23.  Triunifetta  suhj^alinata,  Solan- 

der. 
33. Correa  speciosa,  Donn. 
34:. Cot'rea  rubra,  Sm. 

3b.Jamholifera  laevis,  0.  Kuntze. 
Engler  writes  in  'Nat.  Pfl.': 
"  As  the  name  Jamhollfera 
has  not  been  in  use  more 
than  100  years  I  keep  the 
name  Acroriychia,  Forst." 

36.  Micro tneluin  2>ubescens,  Blume. 


4:2.  Dodo  ncea      iiUcosa,     L.,     var. 

laurina. 
Ab.Buchanania  Muelleri,  Engl. 

4:Q.  Pleiogynium   Solandri,    Engl.  \ 
Adopted  in  Engler's  '  Nat.  ' 
Pfl.  Fam.' following  Engler's  ( 
monograph  in  '  DC.   Prod.' ' 
QO.Cracca  reticulata,  O.  Kuntze. 
Taubert  mentions  in  '  Nat.  Pfl. 
species  of  Tephrosia  to  Cracca,  but 
nomenclature. 


Name  in  '  Flora  Australiensis. 
Poly  gala  leptalea,  DC. 


Cala7idrinia  quadrivalvis,  F. 

Muell. 
Abutilon  indicum,  Benth. 

Triumfetta  procumbens, 

Benth. 
Correa  sp)eciosa.  Ait. 
Correa  rubra,  Sm.  (identical 

with  C.  speciosa.  Ait.). 
Acr onychia  laevis,  Forst. 


Micromelump  icbescens,  Blume, 
var.  glabrescens,  Oliv,  (J/. 
glabrescens,  Benth.). 

Dodoncea   viscosa,   (a)  vul- 
garis, Benth. 

Buchanania  angustifolia, 
Benth. 

Oioenia  cerasifera,  F.v.M. 
'  Spondias  acida,  B.  &  S. 
Sjjoiidias  Solaridri,  Benth. 

Tephrosia  reticulata,  Benth. 
Fam.'  that  Kuntze  refers  all 
he  does  not  accept  Kuntze's 


No. 


president's  address.  713 

Name  on  Plate.  Name  in  '  Flora  Australiensis.' 


61.CraccaJi/.ipes,  O.  Kuntze,  var.      Tephi-osia  Jilipes,  Benth. 

lati/'olia. 
62.Cracca purp2trea,Ij.,v£ir.  longi-     Tephrosia   purpurea^  L.  var. 


foHa. 
QS.Sesban  acideatus,  Pers. 
^i.Diphaca  Cochinchinensis, 

Lour.     Accepted  in  '  Nat. 

Pfl.  Fam.' 
65.  Damajjana     conferta^       0. 

Kuntze. 
QS.Caulinia    bimaciclata,     O. 

Kuntze 
69.Cauli7iia  retitsa,  O.  Kuntze. 

70. Caulmia  ruhicunda,  Moench. 
72.Stizolobiu7)i    gigaiifeum,    O. 

Kuntze. 
l^i.Canavali     maintima,     P. 

Thouars. 
76.Vigna     cape7isis,     Walp.,    in 

Index  Kewensis. 
79.  Dolicholus  Cunninghamii, 

Britten. 
81. Cynometr a  ramifiora,  L. 


longi/olia,  Benth. 
Seshania  aculeata,  Pers. 
Ormocarpum  sennoides,  DC. 


Smithia  cortferta,  Sm. 

Hardenhergia  monophylla, 

Benth. 
Hardenhergia  (? )  retusa, 

Benth. 
Keymedya  ruhicunda,  Vent. 
Mucuna  gigantea,  DC, 

Canavalia  ohtusi/olia,  DC. 

In  Index  Kewensis. 
Vigna  iiexillata,  Benth. 


Rhynchosia      Cunninghamii, 

Benth. 
Cynomeira     rainiflora,      var. 
hijuga,  Benth. 
^i. Acacia    Simsii,    A.   Cunn.,  in     Acacia  Simsii,  A.  Cunn. 
Index  Kewensis. 
123. H^Ufuni  calyptrata,  Britt.  Barringtonia  calyptrata.,  Br. 

Niedenzu  gives  the  whole  synomymy  of  Barringtonia  in 
'  Nat.  Pfl.  Fam.,'  but   sticks   to   Barringtonia   though  he 
mentions  Hiittiim,   Adans.  1763.     Huttum  is  rejected  in 
accordance  with  the  Berlin  Rule. 
12  i.Ciomhia  austraJis,  Britt.  Carey  a  arhorea,  var.  (?)  aus- 

tralis,  Benth.  Cai^eya  aus- 
tralis,  F.V.M.,  in  'Nat.  Pfl. 
Fam.' 


714  president's  address. 

No.  Name  on  Plate.  Name  in  '  Flora  Austr.axiensis.' 

125. Rotala  decussata,  DC.  Animaaia  pentandra,  Benth. 

The    section    Rotaht    was    separated  as   a    genus    from 

Ainmania    by  Mueller   in    '  Census/ and  also  in  Engler's 

'Nat.  Pfl.  Fam.' 
\1Q.Ammania  haccifera,  L.  Ammania  iruHca,  Benth. 

Adopted     by    Mueller    in 

'  Census.' 
130. Haliniutn   portidacaMrum,   O.      Sesuvium  portulacastrum,  L. 

Kuntze. 
132. D Idiscus  pi^ocumbens,  F.M.  Trachymeyie  prociunbens, 

Adopted    by    Mueller     in         Benth. 

'  Census.' 
133,  Tr achy mene  erlcoides.,  Sieb.  Siehera  ericoide'^,  Benth. 

Adopted    by    Mueller    in 

'  Census.' 
13 i:.Trachymene     BWardieri,     F.      Siehera  Bidardieri,v3ir.ovata, 

MuelL,    var.     ovata,     F.M.  Benth. 

Adopted    by    Mueller     in 

'  Census.' 
138. Oldenlaridia    rnitrasacmoides,     Hedyotis    mitrasac7noides,    F. 

F.    Muell.       Adopted    by         Muell. 

Mueller  in  '  Census.' 
Ii2. Nelitris  Tinion,  Britt.  Timonius  Rumphii,  DC. 

l^b.Goelosperinutn  decijnens,  Baill.      Morinda  reticidata,  Benth. 

Adopted  by  K.    Schumann 

in  'Nat.  Pfl.  Fam.' 
1 49.  Opercular ia  hyssopi/olia,  Juss.      Opercularia  aspera,  var.  hysso- 

0.  hyssopifolia  is  in  Index         pifolia,  Benth. 

Kew.  a  var.  of  0.  aspera. 
151. Knoxia  stricta,  Gaertn.  Knoxia  corymhosa,  Willd. 

Ib2. Tardavd  marginati,  Britten.       Spermacocemaryinata^^enth.. 
ib3. Shaivia  arguta,  Britt.  Olearia  arguta,  Benth. 

Shawia  is  in  '  Nat.    Pfl.   Fam.,'   a   synonym  of  section 

Astrotricha  of  Olearia. 


PRESIDENT  S  ADDRESS. 


715 


No.  Name  on  Plate. 

lb6.Pteroc(iulo)i     sjihacelatus,      F. 

Muell.    Confirmed  in  '  Nat. 

Pfl.  Fam.' 
\b1  .Pterocaulon    glanrbohsrts,     F. 

IMuell.    Confirmed  in  '  Xat. 

Pfl.  Fam.' 
1  bd.JViebuhria  spilanthoides, 

Britten. 
IQO. Niebuhria  hijlora,  Britt. 
lQ'2.Phacellothrix    claclochaeta,    F. 

Muell.    Confirmed  in  '  Nat. 

Pfl.  Fam.' 
\%D. Elichrysuui  alhicans,  Sieb. 

1 67.  Crossocephalum  Pseudochina, 

O.  Kuntze. 
168. Candollea  sei'i'idata,  Labill. 

16^ .C andollea  alsinoides,  F.  Muell. 
170  A.  Candollea    rotundifolia,     F. 

Muell. 
1  70b.  Candollea uliginosa,  F.  Muell. 
170c.  Candollea    pedunculata,     F. 

Muell. 
182.  Cervicinia  gracilis,  Britt. 
\8LStyphelia  jnnifolia,  Spreng. 
\85.Styphelia  virgata,  Labill. 
186. Sty phelia  ericoides,  Sm. 
187 S.typhelia  rvsci/olia,  Spreng. 
1 88. Sty phelia  leptosper?noides, 

Spreng. 
191.  Woollsia  pu7ig ens,  F.M. 
193. Sideroxylon  Brownii,  F.M. 
19 ii.Miniusops  kauki,  L. 
197.  May epea  axillaris,  F.M. 


Name  in  '  Flora  Australiensis.' 
Mo7ieuteless2)hacelatus,Jun}n\\. 


Monenteles     glandulosus,     F. 
Muell. 

Wedelia      spilanthoides,       F. 

Muell. 
Wedelia  hijlora,  DC. 
Rutidosis  Broivnii,   Benth. 


Heliclirysum  elatum,  A.Cunn. 

In  '  Index  Kewensis.' 
Gynura  Pseudo-China,  DC. 

Stylidium  yyxtminijolium, 

Sw. 
Stylidium  alsinoides,  Br. 
Stylidium  rotundifolium,  Br. 

Stylidium  uliginosum,  Sw. 
Stylidium  pedtinculatum,  Br. 

Wahlenhergia gracilis,  A.  DC. 
Astroloma  pinifolium,  Benth. 
Leucopogon  virgatvs,  Br. 
Leucopogon  ericoides,  Br. 
Leucopogon  ruscifolius,  Br. 
Leucopogon     leptosperm  o  ides, 

Br. 
Lysinema  jnuigeyis,  Br. 
Achras  ohovata,  F.  Muell. 
Miynusops  Browriiayia,  Benth. 
Chionanthus  axillaris,  Br, 


716 

No. 


PRESIDENT  S  ADDRESS. 
Name  on  Plate.  Name  in  'Flora  Australiensis/ 


198. Gi/7iopogon  spicatum,  Britten.     Alyxia  spicata,  Br. 

Confirmed  byK.  Schumann, 

in  'Nat.  Pfl.  Fam.' 
211.  Borraginoides  zei/Ianica,  Trichode^ma  zeylanica,  Br. 

Hiern. 
'il^i.Merreinia  hnstata,  Hallier  f.        Ipomoea  angitstifolia,  Benth. 
Tiie  genus  Merrimia,  Dennst.,  is  admitted  by  Fetter  in 

'  Nat.  Pfl.  Fam.',  not  as  a  syn.  of  Ipomcea  but  alongside  of 

Ipomoea,   "  about   40  species,   mostly  hitherto  placed  with 

Ipomoea  and  Batatas.^' 
2\b.LepistemonFitzalani,^.^l\XQ\\. 

Lepistemon  iirceolatus, 

F.V.M.,  in  'Census." 
21^ .Jacquemontia  multivalvis, 

Hallier  / 
219. Lindernia  subulata,  Br.     Con- 
firmed by  Wittstein  in  'Nat. 

Pfl.  Fam.' 
220.  Raznmovia  hispida,  Britten. 
232. Hyg7'oj)hila    angustifolia,  Br. 

In  'Index  Kewensis,' 
236. Myop>orum  teiiuifoJium, 

Forst.,  in   Mueller's    '  Cen- 
sus.' 
239. Siphonanthus  floribundits, 

Britten.      Siphonanthus    is 

a  section  of    Clerodendron 

in  'Nat.  Pfl.  Fam.' 
2iO.Germanea  australis,    Britten.      Plectranthus  parviflorus, 

Germanea   is   a  section   of         Benth. 

Plectranthus,  in  'Nat.  Pfl. 

Fam.' 


Ipomcea  icrceolata,  Br. 


Convolvidus  midtivalvis, 

Benth. 
Yandellia  subulata,  Benth. 


Centranthera  hispida,  Br. 
Hygrophila  salicifolia, 

Benth. 
Myoporum  acuminatum,  Br. 


Clerodendron  floribundum, 
Br. 


president  s  address.  7  1  7 

Addendum. — The  De  Vriesian  Species. 

I  have  dealt  so  much  with  species  from  the  nomenclature  point 
of  view  that  it  will  be  of  interest  to  refer  to  another  phase  of  the 
species-question.  I  refer  to  the  classical  experiments  of  Prof. 
Hugo  De  Vriese,  undertaken  with  the  view  of  throwing  light  upon 
the  formation  of  species.  When  in  x\msterdam  a  couple  of 
years  ago,  Prof.  De  Vriese  showed  me  very  fully  his  interesting 
experiments  with  plants  of  Oenothera^  and  now  I  offer  a  digest 
by  my  friend,  Mr.  E.  Betche,  of  a  recent  paper,*  giving  an 
account  of  these  remarkable  experiments  and  the  theories  arising 
out  of  them. 

According  to  Darwin,  the  development  of  species  takes  place 
gradually,  by  natural  selection,  but  if  we  consider  that  species 
have  been  proved  to  have  remained  in  an  unaltered  state  for 
many  centuries  (weeds  found  in  Egyptian  mummies  have  been 
found  to  be  exactly  identical  with  species  growing  at  the  present 
day,  after  a  lapse  of  about  4,000  years),  the  time  required  for  the 
development  of  the  present  forms  of  life  is  so  enormous  that 
Darwin  himself  admits  this  to  be  a  weak  point  in  his  theory. 

Kolliker  advances  another  theory  of  the  origin  of  species, 
which  he  claims  overcomes  this  difficulty;  his  theory  is  the 
sudden  development  (mutation  of  the  old  investigators)  of  species, 
as  opposed  to  Darwin's  gradual  development.  According  to  his 
theory,  the  species  lives  for  a  period  in  a  state  of  stability,  then 
the  species  gets  old  and  seems  to  be  no  longer  in  complete 
harmony  witli  the  surrounding  conditions;  it  then  suddenly 
breaks  up  into  numerous  forms — that  is,  a  comparatively  short 
period  of  instability  (mutation)  follows  the  long  period  of 
stability.  The  mother  species  suddenly  develops  a  great  number 
of  variations;  some  of  these  offspring-forms  live  only  a  few  years 
and  die  out  again,  others  develop  into  new  species  and  take 
the   place   of  the   mother-form.      With   youthful  freshness  they 


*  Hugo  De  Vriese,  "  Die  Mutation  und  die  Mutationsperioden  bei  der 
Entstehung  der  Arten  "  Verhandl.  Gesellsch.  Deuts.  Naturf.  und  Aerzte. 
1901. 

46 


718  president's  address. 

live  again  for  a  long  period  in  a  state  of  stability,  till  they  again, 
in  their  turn,  overlive  themselves  and  break  up  again  into  new 
forms.  And  so  the  development  in  Fauna  and  Flora  goes  on 
from  millions  of  years  back  to  the  present  time,  long  periods 
of  stability  always  alternating  with  shorter  mutation-periods. 
According  to  this  theory,  the  species  would  be  again  a  finished 
unit,  not  a  finished  created  species  as  understood  by  the  pre- 
Darwin  naturalists,  but  a  finished  unit  at  least  for  a  certain 
period.  Each  species  has,  like  the  individual,  a  limited  life,  a 
beginning  and  an  end,  and  the  end  of  the  species  is  its  breaking 
up  into  new  progressive  forms.  The  zoologist  Strandfuss  uses 
the  expression  -'explosion"  with  regard  to  the  breaking  up  of  the 
species  into  new  forms,  as  if  the  old  species  "explodes"  into 
numerous  pieces,  some  of  the  pieces  getting  lost,  while  others 
develop  into  new  species.  The  old  botanists  use  the  astronomical 
expression  "  Nebula "  for  extremely  variable  or  "  very  much 
exploded  "  species,  such  as  Hieraciuiii,  Ruhus,  Rusa,  Salix. 

An  interesting  problem  for  investigation  is  to  find  out  the 
species  in  which  the  mutation-period  takes  place  at  the  present 
day.  Prof.  De  Vriese  knows  only  one  plant  which  fulfils  all 
necessary  conditions,  ?.e.,  Oenothevd  Lamarckiana^  a  species  intro- 
duced from  America,  and  now  slowly  spreading  in  several 
European  countries  for  a  period  of  about  a  centur}''.  0.  Lamarck- 
iana  has  the  rare  and  remarkable  property  of  producing  every 
year  new  species  from  seeds.  The  seeds  of  0.  Lamarckiana  are 
sown  in  sufficient  quantities  (care  having  been  taken  to  avoid  all 
possibility  of  cross-fertilization  with  allied  species);  a  number  of 
new  species  will  always  be  found  amongst  the  seedlings.  Prof. 
De  Vriese  enumerates  7  species  thus  obtained  by  him  from  0. 
Lamarckiana,  without  cross-fertilization.  The  species  are : — 
0.  nanella,  lata,  scintilla7is,  oblovc/a,  albida,  gifjas,  and  ruhrinervis. 
These  new  species  difier  onl}'  slightly  from  the  mother  plant,  and 
only  close  observation  teaches  that  new  forms  have  made  their 
appearance,  but  they  are  all  quite  distinct  from  each  other  and 
perfectly  stable  from  the  beginning,  so  that  seeds  of  0.  gigas 
again  produce  0.  giyas. 


president's  address.  719 

Prof.  De  Vriese  concludes  this  part  of  the  paper  with  the 
following  words: — "With  regard  to  the  question  whether  the 
observations  on  Oenothera  hold  good  for  the  development  of  all 
species,  I  do  not  claim  this,  nor  is  it  according  to  my  experience. 
The  theory  I  essaj^ed  to  explain  holds  good  only  for  the  progressive 
development,  not  for  the  numerous  retrogressive  and  side  steps  of 
nature;  generally  speaking  I  maintain  the  progress  in  organic 
life  to  be  intermissive  and  not  continuous." 

The  second  part  of  the  paper  treats  of  the  number  and  length 
of  the  mutation-periods,  but  it  is  too  full  of  speculation  for  me  to 
reproduce  it  here. 

I  fail  to  see  why  the  intermittent  progressive  development  of 
Prof.  De  Vriese's  "  mutation-periods  "  may  not  go  hand  in  hand 
with  Darwin's  gradual  development  theory.  Surel}^  the  slow 
continuous  progress  of  natural  selection  is  not  a  retrogressive 
step  ! 

But  whether  the  progress  is  continuous  or  intermittent,  it  leads 
to  the  same  end.  One  may  compare  the  mutation-theory  to  an 
electric  clock.  The  clocks  are  all  mere  empty  shells  with  dials 
on  two  sides;  they  are  kept  going  by  a  normal  central  clock,  and 
the  hands  are  moved  by  electricity,  not  slowly  and  gradually,  but 
in  jerks  from  minute  to  minute.  The  result  is  the  same  as  the 
gradual  movement  of  the  hands  in  our  watches. 


For  obvious  reasons  I  have  confined  my  attention  to  the  subject 
of  Botanical  Nomenclature.  But  there  is,  I  believe,  a  complemen- 
tary subject,  that  of  Zoological  Nomenclature.  Respecting  this, 
I  am  indebted  to  the  Secretary  for  the  following  brief  statement. 

The  International  Congress  of  Zoologists  was  inaugurated  in 
1889,  and  has  held  five  meetings — at  Paris  in  1889,  at  Moscow  in 
1892,  at  Leyden  in  1895,  at  Cambridge,  England,  in  1898,  and 
at  Berlin  in  1901.  The  Reports  of  three  of  these  Meetings  will 
be  found  in  the  library. 

At  the  First  and  Second  Meetings  the  subject  of  Nomenclature 
received    special   attention,  and  the  Rules  adopted  were  subse- 


720  president's  address. 

quently  reprinted  and  widely  distributed  under  the  auspices  of 
the  Societe  Zoologique  de  France,  in  189o.  In  a  short  preface 
to  the  Rules  so  distributed,  Dr.  Blanchard  says  : — "  Le  caractere 
international  des  Congres  susdits  et  I'adoption  de  ces  Regies  a  la 
presque  unanimite  des  Zoologistes  presents  donne  a  celles-ci  une 
importance  exceptionnelle  :  elles  constituent  desormais  le  code 
auquel  tout  Zoologiste  descripteur  est  tenu  de  se  conformer 
scrupuleusement. " 

It  would  perhaps  be  a  good  thing  if  the  attention  of  Australian 
zoologists  were  drawn  to  these  Rules,  because  in  some  important 
details — such,  for  example,  as  the  use  of  capitals  for  certain  specific 
names — they  are  not  in  agreement  with  the  Stricklandian  Rules 
which  luve  had  many  followers  throughout  the  British  Empire. 

Attention  might  also  be  drawn  to  the  "  Liste  des  Abreviations 
Conventionnelles  des  Noms  d'Auteurs,"  adopted  at  the  Paris 
meeting.  The  method  of  writing  the  familiar  name  of  Sir  William 
Macleay,  viz.,  "Mac-Leay,  W.  (Auteur) — M'Leay  (Abreviation)," 
will  scarcely  meet  with  universal  approbation.  Probably  if  the 
Congress  were  suitably  approached  this  matter,  and  possibly 
others,  might  be  rectified. 

Time  does  not  permit  one  to  go  into  details  on  this  matter  just 
now.  Perhaps  at  one  of  the  Monthly  Meetings  during  the  year 
time  might  be  found  for  a  more  detailed  consideration  of  the 
International  Code. 

On  the  motion  of  Mr.  Henry  Deane  a  cordial  vote  of  thanks 
was  accorded  to  the  President  for  his  interesting  Address. 

The  Hon.  Treasurer,  Mr.  J.  R.  Garland,  presented  the  balance 
sheet  for  the  year  1902,  and  moved  its  adoption,  which  was 
carried.  The  Society's  income  for  the  financial  year  ended 
December  31st,  1902,  was  £1.077  12s.  3d.  ;  the  expenditure 
£1,065  6s.  lOd.;  with  a  credit  balance  of  .£31  8s.  2d.  from  the 
previous  year,  leaving  a  credit  balance  of  £43  13s.  7d.  The 
income  of  the  Bacteriological  Department  was  £545  12s.  6d., 
and  the  expenditure  £485  8s.  8d. ;  with  a  credit  balance  of 
£133  8s.  7d.  from  1901,  leaving  a  credit  balance  of  £194  2s.  5d. 


ANNUAL    MEETING.  721 

On  the  motion  of  the  Hon.  Treasurer  it  was  resolved  that  the 
suspension  of  the  operation  of  Rule  vi.,  providing  for  the  pay- 
ment of  entrance  fees,  should  be  continued  for  all  members 
nominated  or  elected  during  the  year  1903. 

By  the  wish  of  the  Council  Mr.  Garland  intimated  that  before 
the  end  of  the  year  it  was  proposed  to  consider  the  question  of 
the  action  to  be  taken,  after  the  current  year,  in  the  matter  of 
Entrance  Fees,  so  that  a  report  might  be  made  to  the  Society. 

A  motion  relating  to  the  modification  of  the  regulations  for 
the  loan  of  books  from  the  Society's  library,  of  which  notice  had 
been  given,  was  ruled  by  the  President  to  be  out  of  order,  as  he 
considered  the  matter  was  one  for  the  Council  to  deal  with. 

No  nominations  of  other  Candidates  having  been  received,  the 
President  declared  the  following  elections  for  the  current  Session 
to  have  been  duly  made  : — 

President  :  T.  Storie  Dixson,  M.B.,  Ch.M. 

Members  of  Council  (to  fill  six  vacancies) :  Messrs.  Richard 
T.  Baker,  F.L.S.,  W.  W.  Froggatt,  F.L.S.,  Charles  Hedley,  F.L.S., 
A.  H.  S.  Lucas,  M.A.,  B.Sc,  Thomas  Steel,  F.C.S.,  F.L.S.,  Fred. 
Turner,  F.L.S.,  F.R.H.S. 

Auditors  :  Messrs  Duncan  Carson  and  Edward  G.  W.  Palmer, 
J.P. 


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[Printed  off  April  7th,  1903. 


(1902.) 
Names  in  Italics  are  Synonyms. 


PAGE 

Abisara  segecia 370 

Acacia  acinaeea  ...         191,  204 

armata        ....  198,  204,  567,  584 

aspera         191,  204 

Baileyana    ...  198,  202,  204 

binervata     ..,386,388,391,393, 

[395,  396,  399 

Cambagei 563 

cardiophylla  188,  189,  203 

conferta       ...  191,  196,  198,  204 

cultriformis 191,  204 

dealbata  188,  191,  195,  196,  197, 

198,  203,  386,  388, 

564,  567,  573,  575, 

580,  581,  582,  583, 

[584,  588 

decora     188,  191,  195,  203,  204, 

[561 

difformis  188,  189,  191,  195,  196, 

[197,  198,  202,  203,  204 

diffusa    198,  202,  204,  580,  581, 

[583,  584 

discolor       584 

doratoxylon    188,  202,  203,  496, 
[562,  580,  592 

elata  888 

falcata  (?) 567 

flexifolia      ...  195,  196,  204 

hakeoides  188,  189,  191,  195,196, 

[197,  198,  203,  580 

harpophylla  59 

homalophylla  188,  191,  195,  197, 
[203,  562,  563 

implexa       576,  579 

ixiophyUa        188,  189,  191,  195, 

[204 

lanigera       ...  537,  573,  583,  584 

leprosa         537 

linifolia  (?)...  ..        567,583 

lunata  584 

melanoxylon  396,  537,  575,  576, 
[580,  581,  588 


PAGE 

Acacia  microcarpa         ...        191,  204 
montana  191,  197,  202,  203,  204 


obliqua 

... 

191,  204 

obtusata 

'562 

,  580,  584 

Oswaldi 

188 

,  191,  203 

pendula  188,  191, 

197, 

203,  562, 
[563,  590 

penninervis  ..384, 

386, 

387,  388, 

391, 

399, 

403,  404, 
[537,  587 

pubescens    ... 

...     189 

pycnostachya 

... 

...     537 

rhigiophylla    ... 

59 

,  191,  204 

rigens  ...          188 

189 

191,  203 

stenophylla... 

188,  203 

undulifolia  ... 

...     573 

verniciflua  ...  191 

^195 

,  204,  537 

vestita 

567,  583 

Acantholipes  toxica 

...     102 

Acontia  emboloscia 

...       94 

transversa  ... 

...       95 

tripartita     ... 

...     120 

Acontianae 

...       94 

Acrobolbus  unguiculatus 

...     494 

Acronychia  lasvis 

...     536 

Acronycta  ditata 

...      78 

Actinotus  Forsythii 

...       60 

Gibbonsii    ... 

...       60 

Helianthi    ... 

584,  592 

leucocephalus 

...       60 

minor 

...       60 

Adetipa  punctata 

...     319 

Adrapsa  semicircularis 

...      78 

iEnogenes 

...     247 

^thalina 

...     107 

asaphes 

...     108 

Agaricus  diversipes 

...     542 

(Pleurotus)  lividulus 

...     542 

Agarista  macleayi 

., 

..      656 

Agrotinse 

...       79 

Agrotis  enunciatus 

,, 

...       78 

hypochalchis 

...       80 

INDEX. 


PAGE 

PAGE 

Agrotis  subnigra 

...          80 

Argohuccinum    ... 

. 

443,  480 

Alapadna            

...     106 

fasciatmn    ... 

...     443 

pauropis      

...     106 

Argonauta  cornu 

...       23 

Alelimma            

...     123 

Arhopala  wildei.. 

...     370 

Amalthea  conica 

...     601 

Arripis  salar 

, 

...     656 

Amblyanthera 

...       73 

A  sea  laph  u  s  Jlavipes 

..      364 

Amblystoma       

...     559 

sabulosus 

. 

...     363 

Amphilina           

...      50 

siibtrahens  ... 

...     365 

Amphiptyches 

48,53 

Asclepiadeae 

...     541 

urna             

53,  54 

Asclepias  incarnata 

...     378 

Amphistomum  subclavatum  501,  506,  ] 

Ascobacillus  aquatilis  . 

...     144 

[513 

Aspella 

445,  449 

Amydala  femoralis 

...     409 

anceps 

...     445 

Anacardiacese    

...     536 

Aspidiotus  perniciosus 

...     343 

Anachrostis  aphanta    ... 

...     122 

Assiniinea  pagodella    . 

603,  618 

Anatina 

...       17 

Asterolasia  correifolia  . 

55,  56 

Aneura  pinnatifida 

...     493 

var.  mollis 

56,  57 

Angophora  intermedia... 

201,  583 

var.  Muelleri      . 

...       57 

sp 

...     641 

mollis 

55,  56 

Anoplognathus  sp. 

...     642 

Muelleri 

.. 

...       56 

Anthocercis  scabrella   ... 

...       62 

Astralium  aureum 

6 

Anthostomella  danthonias 

...     373 

cucullatum ... 

3 

Aplustrum 

...       16 

fimbriatum... 

2 

3,  4,  5,  6 

brazieri        

16,  29 

var.  cucullatum 

4 

Apollon 444 

449,  479 

var.  squamiferum 

4 

gyrinus         

...     444 

pileolum 

3 

Aquila      

...     446 

squamiferum 

3 

Aquillus 

444,  446 

tentoriforme 

2,  3,  4,  5 

aquatilis 

...     457 

Astrotricha  ledifolia 

...     540 

chemnitzi     

...     457 

Austrodiaphana... 

.. 

...       16 

cutaceiis       

...     444 

Axiocteta 

...     105 

lotorium 

...     457 

oenoplex      .. 

...     106 

retusus 

...     457 

Axiorata 

.. 

...     120 

tranqueharicus 

...     457 

leucozona    ... 

...     121 

tripus           

...     457 

Bacillus  acacias... 

...     398 

Aquilus 

...     446 

citreus 

...     144 

Arachnognatha 

...       91 

levaniformans 

231, 

232,  238, 

Araeopteron       

.-      131 

[234 

,  236,  391 

Araliacese           

...     540 

metarabinus 

...     403 

Aranea  triremis 

27 

Bacterium  acacise  383, 

'393, 

396,  398, 

Araneus  

484,  486 

399, 

400, 

403,  404, 

liberalis 

486,  492 

[405,  407 

Area  zebra          

...       17 

apii 

...       46 

Ardisia  brevipedata 

...     381 

beta^.. 

47 

imperialis    ... 

...     382 

eucalypti  230,  232 

','234 

,  236,  238 

pachyrrhachis 

...     381 

gelatin osum  beta 

...     263 

poranthera 

...     381 

gummis 

...       46 

pseudo-jambosa     ... 

...     381 

luteum 

..     144 

racemosa     

380,  495 

metarabinum   329 

,'403 

,  404,  405 

repandula 

...     381 

mori 

...       47 

solanacea  var.  haplosciadea     381 

pyocyaneum 

...     140 

Argiopidse           

...     486 

sacchari 

"i37 

,  142,  144 

Argiopinae           

...     486 

tracheiphilus 

... 

...       46 

INDEX. 


111. 


Bacterium  vascular  urn 

xylinum 
Baeckea  diffusa... 
Banksia  ericifolia 

marginata  .. 
Berberis  vulgaris 
Bertya  Cunninghamii  ... 
Bifidaria  ... 

macleayi      

Biplex  australasia 

perca 

rubicola 
Biplex     ... 

roHa ... 
Bittacus  australis 

corethrarms 

tipularius    ... 
Blenina  lichenopa 

metachrysa  .. 
Boletus  granulatus 
Bornia  filosa 
Bothynacrum     ... 

storeoides  ... 
Brachycome  scapigera  ... 
Brachylaimus  ... 
Brachyloma  daphnoides 
Brentelina  Bauerleni  ... 
Breynia  oblongifolia  ... 
Bryomima 

elseodes 

ochrochlora 
Bryophila  exquisita 
Bubo  flavipes     ... 
Buccinum 


PAGE 

31,  33,  44, 
[46,  47,  74 

383 

59 

63 

580,  587,  588 

376 

186,  190 

19 

19,  29 

26 

26 

..       26 

444 

444 

358,  .359 

359 

360 

79 

89 

542 

7,  28 

437 

438 

540 

329 


avellana 

brasilianum 

penatum 
Buccinum-tritonis 
Buccitriton 

altus... 


193 
544 
656 
134 
134 
135 

79 
.364 
443 
599 
600 

24 
443 
443 
445 
445 


Bucephalus  505,  512,  513,  514,  515 
polymorphus  512 

Bulimus  carinatus        27 

eximius       ...         ..  ...       27 

zebra  ...          ...         ...       27 

Bulla  f erruginea  27 

ferruginosa 27 

nitida  ...         ...         ...         9 

Bursaria  spinosa  186,  187,  190,  195, 
[197,  202,  203 

Bythoceras  635 


Cabestana 
Csecum    ... 

amputatum 

lilianum 

vertebrale    ... 
Caladenia  clavigera 

coerulea 
Calandria  oryza... 
Ca  I  ca  r  JimhriaUnii 

tentoriforme 
Callistemon 


PAGE 

444,  456 
...  22 
...  17 
603,  618 
...  603 
..  196 
...  186 
...  343 
2 
3 
...     644 


salignus  var.  angustifolia        567 

Callitris  calcarata  186,  190,  193,  195, 

197,  202,  562,  579, 

[580 

robusta    186,  190,  193,  195,  197, 

202,  203,  561,  562,  577, 

[579,  580 

Callocephalon  galeatum  ...     568 

Callorhynchus  antarcticus      ...48,  50 

argenteus    ...         ...         ...       48 

Calythrix  tetragona       186,  190,  193, 

[203,  575,  584 

Camarosporium  dolichi  ...     374 

mali 374 

nigricans     .  374 

Camptorrhinus  ephippiger      ...     409 
Cancellaria  reticulata  ...         ...       27 

Candalides  ...  ..  ...     6-50 

xanthospilos  ...          ...     650 

Canna  indica     ...         ...          ..      375 

Cantharidns  hadius      ...          ...       27 

pieronii         ...         ...         ...       27 

Canthylidia  moribunda  ...       78 

tenuistria    ...         ...         ...       79 

Capnodium  casuarinae 374 

Capparis  Mitchelli         187 

Capulus  602 

australis      600,  601 

danieli         601 

Carabidie  252 

Caradrina  chromoneura  ...       86 

Caradrininse       ...         ...  ...       81 

Carex  gracilis     ...         73 

Carinidea  flmhriata       ...         ...         2 

Caryophyllaeus  ...         ...         ...       50 

mutabilis    ...          ...          ...       54 

Caryophylleai     ..     212 

Cassia  eremophila  186,  190,  193, 197 

Sophera      567 

Cassidaria  gradrata      ...         ...     454 

Cassidea  labiata  ...         ...       27 

Cassinia  leptocephala 61 


iv. 

INDEX. 

PAGE 

PAGE 

Cassinia  sp. 

580 

Cbrysopa  ramburii 

...     369 

Theodori    ...  193, 

194,  195,  197, 

Cbrysopbanus 

...     333 

[202,  203,  573 

cyprotus      

...     333 

Cassis  achatina... 

27 

Cbrysopbrys  australis .. 

...     373 

Cassytha  sp. 

190 

Cbytra     

...     634 

Casuarina  Cambagei 

186,  190,  197, 

Cladina  pycnoclada 

...     371 

[204 

Cladonieae            

...     371 

Cunninghamiana 

203,  204,  561, 

Clanculus  denticulatus 

...       16 

562,  564,  567, 

Clatbrina  retipora 

...     371 

572,  579,  580, 

Clatburella  denseplicata 

...       17 

[581,  582,  5S3 

philomence  .. 

...       17 

distyla 

194 

Cocblodesma      

...       17 

Luehmanni     186, 

190,  193,  197, 

angasi          

...      17 

202, 

203,  204,  541, 

Cocos  capitata 

...     239 

[561 

,  562,  564,  580 

Yatayi          

...     2.39 

paludosa  (?)      193 

,  194,  204,  584 

Collemaceae        

...     371 

quadrivalvis    186, 

197,  202,  203, 

Colubraria          

444,  449 

[204,  580 

granulata 

...     444 

suberosa 

...         ...     194 

maculosa 

443,  444 

torulosa 

.     375 

Columbella  australis     ... 

...    5,6 

Casuarinese 

541 

pavonia       

...     482 

Catada 

124 

semiconvexa 

...    5,  6 

prominens  ... 

124 

Columbellidse     

...     477 

Catepbia 

86 

Compositae          61 

,  241,  540 

didcistriga  .. 

85 

Congeria  lunata 

...  8,  28 

Catoryctis  nono-linea 

249 

Conularia  inornata 

...     495 

subparallela 

.-.     249 

laevigata      

...     495 

truncata 

248 

tasmaniensis 

...     495 

Celsenia 

484,  488 

Conulus  maino 

...       20 

distincta 

484,  488,  492 

Corbula  taitensis 

...       24 

Celastrinese 

586 

Corcobaea           

...     128 

Centetes 

549 

Corgatba  dichionistis  ... 

...     113 

Ceratof  eronia     . . . 

251 

dipyra         

...     113 

Cercaria  armata     499 

,  505;  508,  510 

drosera        

...     114 

cotylocerca  . . . 

500 

figuralis       

...      79 

cystopbora  . . . 

500 

Corula  abebsea 

...     101 

macrocerca... 

505,  514 

dispila          

98,99 

Ceritbium 

606 

eremna        

...     100 

dubium 

343 

mesospila 

...       99 

Charidea  sanguinata 

78 

metableta 

...       98 

Charonia... 

445 

octogesima 

...       97 

Cbiloscypbus  argutus 

493 

stenopbylla 

...     100 

limosus 

493 

Cosmophila  psamathodes 

...     108 

Cbimtera  monstrosa 

...      48,  50,  54 

CossidsB 

...     246 

Ogilbyi 

48,51 

Crioa  acronyctina 

...       96 

Cbione  disjecta... 

28 

leptocbroma 

...       96 

gallinula     ... 

..     596 

nycterina 

...       95 

lagopus 

596 

Criofrontia  sanguinata 

...      78 

Chiroplatys  sp.  ... 

656 

Crossea   ..          

...       12 

Cblamydodera  maculata         ..      237 

biconica       ...           12, 

13,  14,  28 

Cblamys  fenestrata 

495 

gatliffi          

13,28 

Cbolopus  didactylus 

549 

minuta        

...       12 

Cbrysopa  maculipennis            ...     366 

striata          

12,13 

INDEX. 


PAGE 

PAGE 

Cryptandra  Scortechinii 

..       57 

Daviesia  corymbosa,  var. 

195 

Cryptocarya  Gregsoni  . . . 

..     347 

latifolia       

537 

Cryptophaga  confundens 

..     248 

sp 

573 

epigramma 

..     249 

Dendrobium  gracilicaule 

350 

Cryptorhynchides 

..     408 

var.  Howeanum 

350 

Cryptorhynchus      408,  409,  412,  434 

Jonesii         

350 

albicollis 

..     409 

Dendrothripa  baeota    ... 

93 

antares        

409 

clopaea        

92 

australis      

..     409 

Diadoxus  erythrurus    ... 

592 

cariosus       

..     409 

Diaphana  brazieri 

16 

corosus        

..     409 

Diploblasteee      

371 

ephippiger 

..     409 

Diplothemium  maritimum 

239 

femoralis 

..     409 

Dipsacus  sylvestris  torsus 

344 

infiilatus     

..     409 

Discaria  australis 

567 

Lapathi       409,  412 

Discocarpe^       

371 

lithodermus 

..     409 

Distegia  Ledebourii 

593 

longimanus 

..     409 

Distomum  acanthocephalum 

507 

moestus        

.,     409 

duplicatum      500,  501,  505 

,513 

sirius            

..     409 

fasciatum 

506 

solidus         

..     409 

pristiophori              326,  329 

343 

stigmaticus...           409,  410,  411 

tereticolle    ... 

329 

succisus       

.      409 

veliporium 

329 

teter 

..     409 

Distortio. 

454 

verus           409,  410 

Distortrix  anus 

444 

Ctiloceras           

22 

Dnopheropis       

103 

clathratum 

"22,  28 

scotaea         

104 

cyclicum      

22,  28 

Dodonaja     peduncularis     var. 

striatum 

22,  28 

hirsuta 

.. 

57 

Cuciibalus  Behen 

.      212 

pubescens 

58 

Cucumis  sp 

.       194 

sp 190 

203 

Cucurbitaceae 

46 

viscosa         ...  186,  193,  195 

197 

Culama  culiginosa 

..     246 

Dolichos  lablab 

374 

expressa      

..     246 

Drillia 

17 

Cumia      444,  456 

Drimys  aromatica 

380 

Ciirculionidas      

..     408 

dipetala       

380 

Cuscus     ...          

..     546 

Drosera  sp 

197 

Cyaniris  tenella.. 

..     370 

Duboisia  myoporoides  ... 

640 

Cyclostoma         

..     622 

Dymia  sp.           

.. 

495 

Cyclostrema  porcellana 

..       18 

E arias  ochrophylla 

108 

CylindrobuUa  fischeri  ...        604,  618 

Eccleta 

86 

Cymatium       444,  449,  451,  452,  456, 

xuthopanes 

.. 

87 

[457,  458,  471 

Echidna 205,5 

48 

555 

Cymia  woodii 

..     445 

Echidnophaga  ambulans 

. 

205 

Cyperacese          

64,  243 

Echinella            ...  620,  621,  6 

31 

632 

Cyperus  laevigatas 

..       64 

coronaria 6 

31, 

632 

Cypraea  princeps 

25,27 

cumingii 

. 

632 

valentia       

25,  27 

gaidii           

633 

Cytospora  carphosperma 

..     379 

granulata 6 

31, 

632 

Dahlia  capnobela 

..     102 

spinulosa 

632 

Banis       

..     648 

Echinostomum  ...            497,  5 

07 

515 

Danthonia  penicillata  ... 

..     374 

Ecpatia 

85 

Dasypus  viUosus 

,.     549 

Elffiocarpus  sp 2 

"47 

,351 

Dasyurus            

..     205 

Elaeodendron  australe  ... 

. 

536 

INDEX. 


PAGE 

Elusa  oenolopha  ...         ...       84 

Emex  australis  ...         ...         ...     541 

Endodonta  melbournensis       ...     604 
otwayensis  ...         ...        605,  618 

subdepressa  ...        605,  619 

tamarensis 605,619 

Enteles     439 

vicinus         440,  441 

vigorsii         ...         ...        440,  442 

Eopsaltria  georgiana    ...         ...       75 

georgiana 76 

griseogularis  ...     74,  75,  76 

gularis         74,  76 

gularis         ...         ...         ...       75 

leiicogaster ...         ...         ...       75 

leucura        ...         ...         ...       74 

pulverulenta  ...  74,  75 

Epacrideoe  61,  213 

Ephebe  pubescens         ...  ...     371 

Ephebece 371 

Epidromus  ...         ...         ...     449 

buccinum-siilcatum  ...     443 

Epilobium  glabellum 540 

Episparis  chordophoides         ...       78 

Epitripta  123 

acosmopis  ...         ...         ...     123 

Erastria  bryistis  109 

ErastrianiB         108 

Erastroides  lichnomima  ...     110 

Erechtites  mixta  ...  540 

Eremophila  longifolia     187,  193,  197 

Mitchelli     ...  186,  188,  197 

Paisleyi       242 

virgata        ...         ...         ...     242 

Erinaceus  europaeus    ...         ...     549 

Eriostemon  correifoUus  ...       56 

dilformis     186,  190 

myoporoides  580 

Erythraea  australis        575 

Esthlodora         130 

versicolor 1,30 

Eublemma  116,  118 

curvata       ...         ...         ...       79 

eurynipha  ...         ...         ...     117 

glaucochroa  116 

lozostropha  118 

rufipuncta  ...         ...         ...     117 

semirufa      ...         ...         ...       79 

sphragidota     ...    ...  116 

vagusana 78 

Eucalyptus  affinis  195,  198,  202,  203, 
[579,  580,  590 


PAGE 

Eucalyptus  aggregata  580,  581,  582, 
[587,  588,  590 

ambigua      222 

amygdalina     221,  222,  223.  521, 

522,  573,  574,  587, 

[588,  590 

Baueriana       214,  215,  216,  217, 

[529,  530,  538 

var.  conica  216,  217  529, 

Behriana  189,  192,  203,  523,  561 

bicolor     516,  517,  518,  519,  520, 

521,  522,  523,  524,  534, 

[593 

var.  parviflora    ...         ...     520 

Bosistoana  ...  518,  519,  521 

Bridgesiana     198,  200,  201.  202, 

203,  538,  564,  565, 

567,  573,  575,  576, 

577,  579,  580,  581, 

582,  583,  584,  588, 

[589 

calycogona      220,  221,  222,  223, 

[523 

var.  celastroides  ..     222 

var.  gracilis        ...        223,  523 

Cambagei        198,  199,  200,  202, 

203,  564,  567,  572, 

573,  575,  576,  577, 

580,  581,  582,  583, 

[584,  589 

camphora    ...         ...         ...     238 

celastroides 221 

cinerea         225,  227 

cneorifolia 221,  222 

coccifera      ..     586 

conferta       533 

conica     189,  193,  203,  204,  216, 

[538,  561,  564,  579,  580, 

[589 

cordata    225,  226,  584,  585,  586, 

[590 

coriacea  567,  568,  571,  573,  575, 

576,  580,  582,  584,  587, 

[588,  589 

crebra  ..  ..         227,  523 

Dawsoni      ...  527,  529,  535 

dealbata      539 

dives       225,  228,  539,  564,  565, 

566,  567,  571,573,  575, 

576,  580,  581,  582,  584, 

[587,  588,  589 

dumosa     189,  192,  193,  203,  222 

exiniia         206 


INDEX. 


PAGE 

Eucalyptus  fastigata       588,  589,  590 
Fletcheri      ...  216,  217,  529,  538 

foecunda     220,  222 

gamophylla  ..  ..      226 

globulus       585 

goniocalyx 200 

gracilis     220,  221,  222,  223,  224 

var.  breviflora 223 

Gunnii  576 

var.  acervula       215 

var.  rubida         ..  ...     239 

haemastoma    516,  518,  522,  524, 

[568,  570 

var.  micrantha  567,  569,  570, 

571,  573,  580, 

[582,  584,  589 

hemiphloia    ..  216,  218,  562 

var.  albens  196,  198,  200,202, 

203,  523,  525,  535, 

561,  564,  565,  567, 

571,  572,  573,  576, 
577,  579,  580,  582, 

[589 

var.  mierocarpa  216,  218,  523. 

[561,  581 

largiflorens     216,  218,  516,  517, 

518,  519,  520,  521, 

[522.  524 

leucoxylon 199 

Macarthuri 581 

macrocarpa   .         226 

macrorhyncha  196,  198,  202,  203, 
539,  564,  565,  567, 

572,  573,  575,  576, 
579,  580,  581,  582, 

[583,  584,  589 

maculata     577 

maculosa  ...  567,  570,  571,  572, 
[573,  582,  583,  590 

mannifera ..     566 

melanophloia  225,  226,  227,  228, 

[229 

melliodora       189,  193,  195,  196, 

198,  202,  233,  530, 

531,  534,  561,  564, 

565,  567,  571,  572, 

573,  575,  576,  577, 
579,  580,  581,  582, 
[583,  584,  589,  590 

microtheca       226,  227,  228,  523 

Morrisii       539 

obtusiflora  ...  .         ...     221 

ohtusifolia 215 


PAGE 

Eucalyptus  ochrophloia  ...     539 

odorata        ...  224,  522,  523 

var.  floribunda  ...         ...     523 

oleosa  642 

ovalifolia  527,  529,  530, 533,  534, 

[538,  571 

var.  lanceolata     527,  530,  538 

pallens         221 

parviflora  ..  ...  518,  520 
pendula  517,  518,  519,  520,  521 
perforata     ...         ...         ...     222 

pilularis      521,  558 

piperita        ...         207 

polyanthema  215,  529,  531,  532, 

533,  538,  567,  571, 

572,  573,  579,  580, 

581,  582,  583,  584, 

[590 

rar.  glauca         ...        527,  529 

polyanthemos  214,  215,  216,  217, 

218,  520,  527,  529, 

530,531,533,535, 

[538,  593 

polybractea 192,  203 

populifolia  ...189,  192,  193,  203, 

218,  520,  523,  532, 

[592 

pruinosa      ...         ...         ...     226 

pulverulenta    202,  203,  225,  226, 

227,  583,  584,  585, 

[590 

pulvigera     ..  ...         ...     585 

quadrangulata        ..        540,  590 

radiata        540,  574 

regnans       ...         ...         ...     589 

rhomhifolia  ..  ...     215 

Risdoni       225,  228 

Rossii  ..  ...         ...     571 

rostrata  189,  203,  561,  562,  564, 

[579,  580,  589,  642 

rubida     567,  568,  569,  573,  576, 

580,  582,  583,  584,  587, 

[589 

saligna        200 

santalifolia  222 

setosa  226 

sideroxylon     189,  192,  193,  195, 

196,  197,  198,  199, 

202,  203,  579,  580, 

[590 

Sieberiana 570 

Smithii       537 

squamosa 539 


INDEX. 


Eucalyptus  stellulata    564,  567,  580, 

582,  584,  587, 
[588,  589 

stricta  221 

Stuartiana      200,  202,  203,  230, 
[231,  239,  564 

subrotunda 215,216 

tereticornis     193,  195,  196,  198, 
202,  203,  531,  535, 

561,  562,  564,  571, 

572,  573,  576,  577, 
579,  580,  581,  582, 

[583,  584,  589 

var.  dealbata      189,  193,  195, 

196,  198,  202, 

203,  561,  562, 

564,  577,  579, 

[580,  589 

var.  linearis        ...         ...     539 

viminalis  ...  201,  238,  564,  565, 
566,  567,  568,  569, 

573,  575,  576,  580, 
581,  582,  583,  584, 
[587,  588,  589,  642 

i^ar.  pedicellaris  ..         ...     538 

viridis  192,  203 

vitrea  ..  538,  587,  589,  590 

Woollsiana     189.  192,  193,  195, 

196,  198,  199,  200, 

202,  203,  204,  561, 

562,  564,  577,  579, 

[580,  589 
Eudaimonisma  ...         ...         ...     247 

Batchelorella  247 

Eugenia  smithii  ...          ...     378 

sp 378 

Eulejeunea         ..  493 

Eulimella  pulchra         IS 

tricincta      ...          ...         ...       18 

EuphorbiaceiB     ..         ...          ...     541 

Euplexia  callisina         ...          ..        81 

leucostigma  82 

polycmeta  ...          ...         ...       83 

Euroschinus  falcatus   ...        536,  592 

Euspongia  Illawarrre 74 

Eutaxia  empetrifolia    ...         ...     197 

Eutelia  dilatrix 88 

oxylopha     ...         ...         ...       88 

Eutelianse  ...         ...         ...       87 

Excfficaria  agallocha    ...        640,645 

parvifolia    ...         ...         ...     640 

Exithius  cariosus         ..  ...     409 


PAGE 

Exoearpus  aphylla 

190 

cupressiformis 

186,  193, 

195, 

197,  203, 

567, 

[575,  580 

,  587 

stricta 

... 

580 

Falcimala 

123 

Fasciolaria 

480 

Fasciolaria  Jilamentosa 

27 

fusiforrnis  ... 

28 

Feronia 

... 

252 

Feronia  auricollis 



313 

australasice... 

289 

hipinictata  ... 

322 

chalyhea 

254 

,  321 

crfesiis 

264 

gippsiensis  ... 

270 

Ies2ieuri 

... 

325 

opulenta 



313 

philippi 

'  254,  321 

,  322 

plutus 

... 

263 

rujipalpis    . . . 

254 

sairapa 

271 

semiviolacea 

... 

310 

sphodroides 

308 

■•itriaticoUis... 

254 

victoricM 

... 

310 

icilcoxi 

254 

Feronia  (Xotonomu 

s)  chalyhea 

321 

dyscolokles  .. 

309 

molenta 

... 

320 

Feronia  (Omaseus)  mitchelli ... 

289 

occldentalis . . . 

319 

rufpalpis    . . . 

... 

315 

sat  anas 

319 

Feronia  (PeraisJ  hi 

punctata  ... 

321 

lacustris 

320 

montana 

322 

Feronia  ( Pterostich 

lis)  amabilis 

307 

auricoUis     . . . 

312 

comes 

... 

283 

crcesus 

... 

263 

darlingi 

301 

ducalis 

... 

293 

tques 

... 

316 

gippsiensis  ... 

269 

gippslandica 

... 

281 

hunteriensis 

... 

287 

impressicollis 

... 

297 

impressipennis 

var. 

306 

lapeyrousei... 

313 

lesueurii 

... 

325 

mastersi 

297 

opulenta 



312 

INDEX. 


PAGE 

Feronia  ( Pterostichus)  peronii  264 

plutus          263 

purpureo-limbata ...         ..  306 

regalis         ...          ...          ...  252 

resplendens  .          318 

satrapa       ...         ...         ...  270 

semiviolacea           309 

suhvilis        ...         ...         ...  309 

tasmanica  ...         ...         ...  266 

victorke        ..           ...          ...  309 

viridilimbata          ...         ...  300 

viridimarginata     ...         ...  300 

Feronini 252 

Ficus  aspera      ...         ..          ...  641 

Bennettii 239 

Cunninghamii        ...         ...  64 

Henneana  ...         ...         ...  64 

Filices     542 

Fimbriaria  conocephala          ...  493 

Drummondii           ..         ...  494 

Fistulai'ia  serrata          ...         ..  205 

Flammulina  (?)  grenvillei       ...21,  29 

Flata        367 

Fodina  (/Zoriosa   ..         ...         ...  78 

stola 78 

Fossarina            ...         ...        620,  635 

varia            ...         ...         ...  685 

Fossarus 12,13 

Frenchia  casuarinse      ...         ...  592 

Freycinetia  excelsa       ...          ...  381 

Frullaiiia  clavata            ..          ...  493 

Hampeana 494 

megalocarpa           ...         ...  493 

pycnantha  ...         ...         ...  494 

Fungi      ...         ...          ...         ...  542 

Fusanus  acuminatus     186,  190,  193, 

[195,  197,  580 

Fusidae 454 

Fusus  laticostatus          ...         ...  25 

pyrulatus     27 

variegatus  ...          ...         ...  25 

Gadirtha             90 

pulchra        ...         ...         ...  78 

Galeopithecus    ...          549 

Gasterostomum . . .  511,512,514 

Geijera  parviflora            186,  197,  204 

Glenurus 362 

circuiter      362 

erythrocephalus     361 

falsus           361 

fundatus     362 

pulchellus 360,  361 

Glossodia  major            196 


PAGE 

Gmelina  Leichhardtii 62 

Gomphognathus      551,  558,  559,  560 

Gompholobium  Huegelii         ...  582 

Goodenia  ovata..          588 

Gorgonops          ...         ...         ...  560 

Grammodes   excellens             ...  78 

divaricata  ...         ...         ...  78 


minor 

...       78 

pallens 

...       78 

pulcherrima 

...       78 

Grevillea  acuaria 

...     243 

floribunda  ... 

186,  197 

Jamesoniana 

243 

nematophylla 

...     243 

ramosissima 

...     567 

Guttimiium    443,  449,  450,  451,  452, 

[456,  457,  458,  459,  471 

ramda         ..  ..         ...     443 

Gyrina     ...         ...         ...         ...     444 

maculata     ...         ...         ...     444 

Gyrineum  ..         ...         ...     449 

hufonia        ...         ...         ...     444 

giga7iteum  ...         ...         ...     444 

gyrinum      ...         . .  ...     444 

vexillum      ...         ...         ...     443 

Gyrocotyle  ...  48,  49,  50,  53 

nigrosetosa  .  51,  53,  54,  74 

rugosa  48,  50,  51,  52,  53,  54 

urna 48,  50,  51,  52,  53 

Habronestes        ...         ...         ...     485 

Hakea  leucoptera    186,  190,  197,  203 
Halorage^e  ...         ...  59,  537 

Haloragis  serra  ...         ...         ...     537 

Hecatesia  fenestrata     ...         ...     657 

Helices     ...         ...         ...         ...     474 

Helichrysum  apiculatum         . . .     575 
obcordatum...         ...         ...     190 

Helix  porti  ...         ...         ...       21 

rosacea        ...         ...         ...     605 

Helix  beraudi    ...         ...         ...     604 

bisidcata      ...         ...         ...     604 

dandenongensis      ...         ...     605 

mdbournensis         ...         ...     604 

otivayensis  ...         ...         ...     605 

rosacea        ...         ...         ...     605 

■mbangidata  ...         ...     604 

subdepressa...         ...         ...     605 

tamarensis  ..  ...         ...     605 

Helix  (Conulus)  barnardeiisis         21 
darnleyensis  ..  ...       21 

grenvillei     ...         ...         ...       21 

maino  ...         ...         .         20 

nepeanemis . «  20 

reedei  20 


INDEX. 


Hemerobiidse     

Hemerobius  coelivagus 

mimicus 
Hemicentetes 
Henninia  delicata 

semicircularis 
Hermippus  loricatus     ... 
Heterodendron  olesefolium 

[193, 
Heteronota  binoei 
Hexagonia  decipiens     ... 
Hexarchus  griseus 
Hibbertia  stricta 
Hilda 
Himerois... 

thiochroa    

Hipponyx 

aiistrahs 

danieli 
Holocephali 
Holochila  absimilis       .  . 

androdus    ... 

cyprotus 

helentia 

margarita    ... 

purpurea 

stibargentea 
Holocryptis  phasianura 
Holoctula  nitens 
Homarus  vulgaris 
Homodes  erizesta 

riolacea 
Homopterinse     ... 
Hoya  carnosa     ... 
Hyblaea  ibidias 

puera 

synaema 
Hyblaeinae 

Hydatina  tasmanica     ... 
Hydatina  cdhocincta 
Hyela 

argotypa 

thapsina      

Hymenochffita  Mougeottii 
Hypeninse 
Hypera  punctata 
Hyperiosoma 

falcatum      

Hyperlopha  aridela 
Hypoyramma  distincta 
Hypothripa 
Hystrichopsylla  talpae  ... 


PAGE 

358 

368 

367 

549 

78 

78 

484 

186, 

190, 

197 

203 

558 

544 

329 

193 

445 

121 

121 

601 

602 

600 

601 

... 

602 

... 

48 

333 

334 

333, 

334 

333 

343 

333 

343 

370 

333 

333, 

334 

78 

653 

495 

110 

... 

79 

95 

377, 

378 

... 

135 

136 

136 

135 

603 

on 

120 

111 

... 

112 

544 

... 

122 

434 

434 

435 

103 

78 

... 

91 

.. 

205 

Icasma    ... 

minutum     . . . 
Ichthyophis  glutinosus 
lUecobracese 
Imleanga  fluviatilis 
Indigofera  australis 
Inella 
Ingura  cyanodes 

Ino  

Jacksonia  sp. 
Juncus  holoschienus 


207. 


prismatocarpus       207,  210 

Jungermannia  montana 
Keraudrenia 

Hillii 
Knoxia  corymbosa 
Kunzea  opposita 

peduncularis 
Kuphus  arenarius 

polythalamius 
Lacerta 

agilis 
Laches     ... 
Lagena    ... 

toroides 
Lavipusia      444,  446,  448,  45 


nodocostata . . . 
pharcida 
pilearis        ... 

LaurineaB 

Leguminoste 

Leipoa  ocellata  ... 

Lejeunea  (Eulejeunia) 
stipula.. 
elobata 
tasmanica  ... 

Lepidium  ruderale 

Lepidozia  capilligera 
glaucophylla 
gracillima  ... 

Leptospermum  laniger 
myrsinoides 
scoparium  ... 

sp 

Leptosphgeria  cannje 

musarum  ... 
Leucania  abdominalis 

fximata 

irregularis  ... 

labeculis 


PAGE 

90 
91 
559 
541 
79 
656 
...  607 
...  87 
...  607 
...  333 
210,  211, 
[212 


193, 


211, 

[212 

493 

55 

55 

541 

537 

583 

602 

602 

552 

558 

484 

443 

443 

452, 

[456 

463 

465 

444 

63,  347 
..58,  213,  537 

189 

cuspidi- 

493 

493 

...     493 
.      344 

494 

494 

494 

)...  575 
...  190 
...  580 
580,  644 
...  375 
...  375 
...  78 
...  78 
...  79 
...      78 


urn  ('. 


INDEX. 


XI. 


PAGE 

cania /meaWs          

.       78 

melanopasta           

.       81 

polysticha 

80 

sepidchraUs            

.       79 

venalba       

.       78 

cochloridium  paradoxum 

500, 

[501,  513 

Leucopogon  exolasius  .  .  61,  213 

setiiger  ...         ...         ...       62 

Lichenes...         ...         ...         ...     544 

Limnotrochus 631,  634 

Linatella  ...  445,  452,  456,  458 

clngulata     ,.  ...         ...     445 

Liotia      ...         ...         ...         ...       14 

corona  ...         ...  14,  28 

devexa  ..  ...         ...       14 

incidata       ...         ...  14,  28 

Lippistes 23,  24 

cornu  ...         ...         ...       24 

sepai-atista...         ...  ...       24 

Lissanthe  strigosa         ...         ...     190 

Litsea  reticulata  ...         ...       63 

Littorina        620,  621,  622,  623,  624, 

625,  627,  628,  629,  630, 

[633,  634 

littoralis      630 

mauritiana  . . .         ...         ...     630 

scabra  ...         ...         ...     630 

Littorinacea       ...         ...         ...     620 

Littorinidse         620,  621 

Lobelia  purpurascens  ...         ...     592 

Loganiaceas         ...         ...         ...     241 

Lomaria  capensis  var.  Bauerleni     542 
Lophotoma         ...         ...         ...     128 

diagrapha 129 

metabula     ...     129 

LoranthacesB  ...         ...     541 

Loranthus  Bidwilli        ...         ...     541 

linophyllus...         ...         ...     564 

pendulus     ...          ...         ...     564 

sp 218,251 

Lotoriidffi  ...  443,  448,  449,  454 

Lotorium  443,  444,446,  447,  448,449, 

451,  452,  455,  456,  457, 

458,  459,  466,  480,  623, 

[624,  630 

abbotti      454,  466,  468,  473,  483 

segrotum     ...  .  452,  473 

africanum  ...         ...         ...     472 

amictum     ..,     473 

annectans 469 

apenninicum  .         ...     472 

aquatile   460,  472,  476,  477,  483 


PAGE 

Lotorium  australe        ...         474,478 
barthelemyi  ...         472,  481 

bassi         452,  453,  454,  474,  480 

beccarii       472 

bicogei         454 

birostomum  ...         ...     474 

brasilianum  ...         ...     472 

caudatum    ...         ...        453,  473 

chemnitzi   ...         ...        453,  472 

chlorostoma  ...        452,  453 

chlorostomum  460,  472,  483 

chlorostomus  453 

cingulatum      445,  452,  453,  472 
clandestinum    443,  471,  474,  479 

elavator      473,  481 

contabulatum         ...         ...     474 

convolutum    ..  ...  473 

cornutum        450,  460,  470,  471, 
[473,  481,  483 
costatum  443,  457,  460,  470, 

[472,  476,  477,  479 
crispum       ...  452,  453,  472 

cutaceum    ...  444,  450,  472 

cynocephalum         452,  453,  457, 
[458,  473 
cyphus         ..    454,  469,  474,  483 
doliarium      ..  452,  453,  472 

dorsuosum  ...         ...         ...     474 

dunkeri      452,  453,  457,  458,  473 
durbanense...         ...         ...     472 

eburneum        452,  453,  454,  463, 
[473,  483 
elongatum        461,  473,  478,  483 
encausticum  ...        453,  473 

exaratum        444,  452,  453,  458, 
[459,  481 

exile...  473 

femorale      ...  450,  452,  472 

ficoides       • 472 

formosum  ...         ..  ...     481 

fossatum     ...         ...         ...     472 

fraterculum  466,  474,  480 

fusiforme    ...  444,451,481 

gallinago     ...         ...         ...     473 

gemmatum     452,  453,  461,  472, 
[483 

gibbosum 473 

gracile  ...         ...        473,  475 

grandimaculatum  ...         ...     472 

granulatum  ...  ..     474 

intermedium             452,  453,  472 
krebsii         472 


Xll. 


INDEX. 


Lotorium  labiosum 


lignarium 

lineatum 

Icebeccki 

loroisii 

lotor... 


PAGE 

462,  471,  472, 

477,  478,  479, 

[483 

458 

472 

473 

471,  472 
444 


lotorium  444,  452,  453,  457,  472 
mauritianum  ...  ...     473 

minimum 454 

monilifer     ...         ...         ...     473 

moritinctum  ...        473,  478 

mundum     ...  452,  453,  472 

miinsteri     ...         ..  452,  472 

nodiferum      443,  452,  453,  464, 

[476,  481 

nodocostatum  454,  463,  466  473, 

[479,  495 

nodoliratum  ..  ...     474 

olearium      457 

oligostirum     452,  453,  454,  464, 
[466,  469,  473,  483 
orientale      ..  452,  453,  472 

ovoideum    ...  454,  471,  474,  476 
pachycheilos  ...  473 

papillosum  ...         ...          ...     474 

paikinsonianum      448,  451,  454, 

455,  464,  470, 

471,  473,  475, 

[476,  480,  481 

pfeifferianum         ...        458,  473 

pharcidum 465,473 

philomelse   ...452,453,454,457. 

464,  465,  466,  469, 

[473 

pileare     444,  452,  453,  458,  472, 

[476 

poulsenii     ...        •...          ...     472 

protensum  ...454,  466,  468,  474, 
[483 

pyiulum      473  j 

pyrum      444,  452,  453,  457,  458,  j 

[473  j 

quoyi      452,  453,  454,  455,  457, 

459,  463,  472,  473,  477, 

[483 

radiale      454,  467,  471,  473,  483 

retusum       457,  473 

rubeculum  ...         ...         ...     472 

rubicundum  464,  474,  481 
rutilum  450,  462,  472,  477,  483 
sarcostoma 452,  453 


PAGE 

Lotorium  sarcostomum  ...     473 

sauliae  474,  478 

scalariforme  ...         ...     473 

sinense  ...  461,  473,481,  483 
spengleri  452,  453,  458,  462,  472, 
[475,  479,  481,  483 
strangei  ...  462,  472,  478,  483 
subdistortum  452,  453,  454,  468, 
[471,  474 

succinctum 453,  458 

tarbelianum  ...         ...     455 

tenuiliratum  ...         ...     473 

testudinarium        ...         ...     473 

textile  ...  454,  470,  473,  483 

thersites   452,  453,  461,  473,  478, 

[483 

tigrinum      ...         ..         452,  472 

tortirostris       454,  464,  466,  469, 

470,  471,  473,  475, 

[483 

tranquebaricum       452,  453,  472 

trilineatum 452,  473 

tringa  474 

tripus  ...  452,  457,  473 

triton  ...         ...          ...     444 

tritonis   443,  444,  450,  452,  453, 

[474,  475,  476 

tuberosum  443,  453,  46 1 ,  473,  483 

tumulosum     452,  453,  454,  467, 

[474,  483 

variegatum 474 

veliei  ...         ...         ...     472 

verrucosum     452,  458,  454,  462, 

[473,  477,  483 

vespaceum      452,  453,  458,  460, 

461,  465,  473,  475, 

[478,  483 

voigtii  ...         ...         ...     472 

waterhousei    448,  452,  453,  454, 

[462,  472,  475 

woodsi      450,  454,  466,  473,  483 

Luperina  ditata 78 

Luterium  ...         ...         ...     444 

Lutica     484 

Lycaena 370 

nigra  ...         ...          ...     331 

LycEenidse  331 

Lycopodiacege 351,  542 

Lycopodium  cernuum  ...         ...     542 

varium  ..          ...          ...     351 

Lycosuchus         ...         ...         ...     560 

Lymphosporidium  trutt^e        . . .     358 
Lyonsia  reticulata         ...  ..     246 


INDEX. 


PAGE 

Lyonsia  straminea        ...         ...     541 

Macroscelides  proboscideus     ...     549 
Macrosporium  internum         ...     375 
Macrozamia  secunda   ...         ...     580 

Mactra  abbreviata        ...        597,598 

var 599,  618 

parkesiana  ...         ...         ...  8,  28 

pura 598,  599 

pusilla         597 

rufescens     ...         ..  ...     599 

Mactrinula         ...         ...         ..  9 

Madope  curvata  ...         ...       79 

leucocrossa ...         ...         ...       79 

Magusa 84,  85 

Malaisia  tortuosa  88 

Mamestra  nigerrima     ...         ...       84 

Mamestrinte       ...         ...         ...       80 

Manis  brevicaudata      ...         ...     549 

Marapana  nycteropis    ...         ...     128 

Marchantia  cephaloscypha     ...     494 
Marginella  mayii  ...         ...       19 

strangei       ...         ...         ...       18 

subbulbosa  ...         ...         ...       18 

Marsupidium  setulosum  ...     494 

Mastoniaeformis...         ...         ...     607 

Maurilia  iocephala       98 

Mecodina  fluviatilis      79 

Mecoliotia  halligani      ...         ...       15 

spinosa        ...         ...  15,  28 

Megisba 332 

albidisca      332 

malaya        ...  332 

nigra  331,  343 

Melaleuca  parviflora       186,190,191 

sp 644 

uncinata     186,  190 

Melipotis  collustrans     ...         ...       79 

Metachrostis  camptozona        ...     114 
epichroma  ...         ...         ...     114 

punctigera  ...         ...         ...     114 

Metasphaeria  lepidospermEe     . . .     376 
Metyrus  albicollis         ...         ...     409 

Metzgeria  crassicosta 494 

furcata         ...         ...         ...     494 

Micraeschus  diacaustus  ...     115 

oblatarius 79 

Micrococcus  pyogenes  352,  353,  354, 

[356 
Miniopterus    550,  551,  553,  557,  560 

schreibersii..  559 

Minuria  integerrima     241 

Mitra  abbatis 27 

pontificalis 27 


PAGa 

Mitrasacme  distylis      242 

palustris      241 

Modulidae  621,  631 

Modulus 621,631,633 

Monodonta  denticulatus  ...       16 

lupina         ...         ...         ...       16 

Monograptus      ...         ...         ...     654 

dubius         654,655 

sp 654 

Monophx  444,  456 

cornutus      ...         ...  26,  444 

Jonnosus     ...         ...  26,  481 

Monotoca  ledifolia        ...  62,213 

Muehlenbeckia  adpressa  ...     541 

Murex  australis...         ...         ...       26 

denudata     ...         .  ...       26 

palmiferus 26 

Murex     443,  445 

adustus        ...         ...          ...       26 

australis      ...         ...         ...       26 

davator       26,  481 

lotorium      ...         ...         ...     446 

pilearis       ...         ...         ..      476 

tenuispina  ...         ...         ...       27 

Musci      .  ...     544 

Muscicapa  georgiana   ...         ...       75 

gularis         ...         ...  75-,  76 

Myoporinese        242 

Myoporum  deserti  186, 187, 188, 190, 

[193,195,197,203 

platycarpum  ..  ...     188 

Myrina  derpiha...         ...         ...     649 

Myriophyllum  pedunculatum...       59 
Myrmeleon  circuiter     ...         ...     362 

erythrocephalum   ...         ...     361 

falsus  361 

fundatus     362 

pulchellus 360 

Myrsinacese        348 

Myrsine 348,  349 

crassifoUa  ..  ...         ...     349 

platy  stigma  ...         ...     349 

Myrtacese  59,  537 

My  sella   donaciformis  ...  ..         7 

Mytilus  latus  497,  498,  501,  505,  507, 

[508,  514 

Myzantha  garrula         645 

Myzomela  sanguinolenta         ...     237 
Naiades  ...         ...         ...         ..,     474 

Nassa       445 

Nassopsidse        621,  634 

Nassopsis         621,  628,  629,  634,  635 
Natica      624 


XIV. 


INDEX. 


PAGE 

Neptunella         ...         ...         ...     445 

Neiiropates  dyscoloides  ...     309 

pristonychoides        255,  265,  284 

Nicotiana  sp.     ...  ...       74 

Nina        631 

Noctuidge  ...         ...  77,  79 

Noctuinae  ...         ...         ...       95 

Nodaria  acrosema         ...         ...     125 

armatalis     ...         ...         .  78 

ctenucha      124 

Notechis  scutatus  ...         ...     645 

Notonomus     252,  253,  254,  264,  267, 

[269,  300,  324 

accedens  256,  265,  266,  267,  284 

seneomicans    260,  278,  295,  313, 

[315 

amabilis      ...         ...        260,  307 

angustibasis  258,  266,  283,  284, 
[295 
angiistipennis  ...  297,  298 
arthuri  ...  257,  271,  272,  286 
australasisB...  259,  287,  288,  289, 
[291,  292 

bakewelli     258,  276 

bassi  ...  258,  280,  281 

besti 257,273,275 

chalybeus  261,  320,  321,  322,  323 

var.  otwayensis 322 

colossus      260,  314 

croesus        ...  256,  262,  263 

cyaneocinctus         ...        300,  301 

daiiingi       256,  301 

depressipennis  256,  260,  305,  306 
discoderuH  ...         ...        254,  287 

discorimosus  256,  260,  305 

dyscoloides      260,  309,  310,  311, 
[312 
eques        261,  279,  316,  317,  318 
excisipennis  258,  272,  285 

fletcheri      ...  258,  277,  278 

frenchi        256,  261 

froggatti      ...  258,  280,  318 

gippsiensis  ...  257,  269,  272 

gippslandicus  258,  274,  281,  282 
gravis  ...  261,  320,  322 

howitti         257,  275 

incrassatus       259,  290,  291,  292 

ingratus      256,  287 

jervisensis  ...         ..         259,  294 

kershawi      261,  320 

kingi       258,  266,  273,  286,  287, 

[295 

kosciuskianus  257,  266,  267,  271 


PAGE 

Notonomus  lateralis  ...  259,292 
latibasis       ...  259,  298,  302 

leai 259,  291 

lesueuri        ...  261,324,325 

liragerus  ...  259,  294,  295,  296 
macoyi        ...  257,  273,  274 

marginatus...  254,  258,  287,  289 
var.  luculentus  ...  288,  289 
mediosulcatus  253,  261,  319,  320 
miles  ...  261,  323,  324 

molestus      ..  261,  320,  321 

montanus    ...  255,  268,  269 

muelleri      257,  272 

nitidicollis       259,  297,  298,  300, 

[301,  302 
opacicollis  ...  260,  305,  306 

opacistriatis  ..  ...     319 

opulentus   255, 260,  282,  312,  313 
parallelomorpha       255,  312,  313 
peroni         ...  256,  263,  264,  272 
jthiUi'pi        ...         ...         ...     279 

phillipsi      ...  256,  278,  279 

politulus     257,265,266,267,268, 

[287 

pristonychoides      283 

pupureipennis        ...        303,  304 
queenslandicus        259,  296,  302 
rainbowi     ...         ...         257,  268 

resplendens  ...         256,  318 

rufipalpis   ...  261,  276,  315 

rugosicollis...  256,  260,  .304 

satanas        286,  287 

satrapa        ...  257,  270,  271 

simulans     ...  256,  310,  312 

spenceri       261,  317 

sphodroides  260,  308,  309,  310 
striaticollis...  256,  305,  306 

strzleckianus   256,  258,  278,  279, 

[280,  318 
subiridescens  256,  295,  296 

subopacus  259,  301,  302,  305 
sydneyensis  ...        258,  288 

triplogenioides  259,  293,  294 
tubericauda  ...        257,  267 

variicollis  258,  272,  277,  278, 
[283,  286,  287 
victoriensis...  260,  311,  312 

violaceomarginatus        260,  300, 

[303,  304 

violaceus     256 

viridicinctus  297,  298,  300 

viridilimbatus  ...  300,  301 
viridimarginatus  ...          ...     300 


INDEX. 


Notonomuswilcoxi  256,  284,  302,  303 
Nyraphes  myrmelionides         ...     365 


Ochthophora      

89 

sericina        

... 

90 

Ogyris      

334 

abrota          

342 

aenone          

339 

,342 

amaryllis    336,338,339 

,342 

,  343 

genoveva     

341 

hewitsoni        335,  336, 

337, 

338, 

[339 

,  342 

,343 

ianthis         

341 

oroetes      335,  336,  337, 

338, 

339, 

[342 

343 

Olax  stricta         

584 

Olearia  adenophora 

... 

61 

alpicola 

540 

glutescens 

540 

lepidophylla 

'61 

190 

myrsinoides 

540 

pimeloides  ... 

... 

190 

sp 

193 

Omaseus  rufi'paljpis 

... 

316 

Onagrese . .           

540 

Ophiodes  parcimacula  . . . 

... 

78 

parcemacula 

78 

Ophiusa  constricta 

78 

gravata        

... 

78 

parcimacula 

78 

trophidota 

... 

95 

Ophyx  ochroptera 

78 

Orhitus  purpuripennis  . . . 

306 

Orcheia  cyllaria 

79 

Orchideae            

213 

,350 

Ornithopus  purpusillus 

213 

Ornithorhynchus   545,  546, 

547, 

548, 

550,  551, 

553, 

554, 

555,  557, 

558, 

559, 
[560 

Orthozona           

128 

Orycteropus  capensis   ... 

549 

Osmylus  strigatus 

366 

Oxylobium  Pulteneee    .. 

... 

58 

trilobatum  var.  ilicifolium 

537 

Pachylepis          

121 

Paludina..           

622 

,  634 

Panilla  cautophanes     ... 

122 

Parity dia  capistrata 

... 

79 

limbata        

544 

Pareiasaurus      

560 

Parmelia  perforata 

372 

,  544 

perlata         

372 

var.  cetrarioides 

... 

372 

pertusa 

372 

PAGE 

Parmelia  physodes        372 

physodes  var.  mundata     . . .  372 

ParmeliesB          372 

Paspalum  sp,     ...         ..          ...  656 

Passiflora  edulis            377 

Patella 601 

calyptra      ...         602 

cassida        ...         ...         ...  601 

vulgata        630,  634 

Patella  australis           600 

Patellaria  (Patinella)  filicicola  544 

Patersonia          ...         ...         ..  24 

Patinella 544 

Paurophylla       ...         ...         ...  124 

aleuropasta            ...          ...  124 

Peasiella 621,  631,  632,  633 

tantillus      ...         ...         ...  633 

Penicillium  glaucum 207 

Pentamerus  sp 370 

Pentodon  australis        ...          ...  656 

Perameles           ...  205 

Perissops 438 

Persona  ...         ...         ...          ...  444 

anibs...         ...         ...         ...  444 

Personella          ...         ...         ...  445 

septemdentata        ...         ...  445 

Persoonia  augulata       ...         ...  73 

Petermannia  cirrosa 381 

Pezichus 438 

Phascolarctus  cinereus 590 

Phebalium  glandulosum          ...  1£0 

obcordatum            190 

Ralstoni      ...         ...         ...  57 

Philemon  citreogularis 237 

Philiris 650,  651 

ilias 650,652 

kamerungae            ..         650,  651 

kurandse      ...         ...         ...  650 

nitens          ...         ...         ...  653 

Philobrya  crenatulifera           ...  17 

Philotheca  ciliata         190 

Pholidia 242 

Plioma  berberina          ...         ...  376 

passiflorge ...  377 

strobiligena  var.  microspora  377 

Phyllanthus  Gasstrcemii         ...  656 

Gunnii         ...         ...         ...  541 

thymoides  ...         ...         ...  541 

Phylloda 28 

Phyllosticta  asclepiadearum  377,  378 

coprosmse 377 

eriobotryte    ..         ...         ...  377 

grossulariae...         ...          ...  378 

myrticola    ...         ...         ...  378 


XVI. 


INDEX. 


PAGE 
Physcese 372 

Pht/scia  chrysophthalma         ...     372 

Pimelea  ligustrina  var.  glabra        63 

rar.  hypericina  ,.         ...       64 

var.  macrostegia  ...       63 

linifolia       193,  203 

Pinus  insignis 377 

Pittosporum  phillyraeoides     190,  195, 


[561 

,580 

undulatum 

593 

Plagiochila  fasciculata... 

494 

Planispira  zonalis 

.. 

604 

Plantago  coronopus 

592 

Pleospora  asclepiadearum 

i 

577 

,  37S 

diplospora 

.. 

378 

herbarum 

375 

Plesiosaurus       

559 

Pksiotriton        

.. 

445 

Plesiotrochvis     

631 

Pleurotoma  acuta 

28 

densepUcata 

.. 

17 

tigrina         

.. 

28 

Pleurotus            

542 

Plodia  interpunctella    ... 

.. 

343 

Plotheia  rhythmopis     ... 

91 

Poa  serotina       

379 

Poecilodryas       

.. 

75 

cerviniventris 

343 

Poecilus  marginatus 

254 

,287 

Poephila  acuticauda    ... 

208 

aurantiirostris 

208 

Polyporus           

543 

mylittse        

542 

,544 

ovinus         

543 

squamosus 

542 

Polygoneae          

541 

Polypodium  australe  var.  a] 

picola 

542 

Polystictus  trizonatus  ... 

... 

544 

Pomaderris  f erruginea  . . 

.. 

536 

phillyraeoides 

567 

Porismus  strigatus 

... 

366 

Poropterus  lithodermus 

409 

succisus       

409 

Priene     

445 

,449 

riidis            

445 

scaher          

449 

Primula  sinensis 

379 

Prionodon  milberti 

329 

Prionophora  torquesaria 

... 

78 

Prionurus  microlepidotus 

205 

Pristiophorus  cirratus  . . . 

326 

Prodenia 

85 

Prolophota  camptoloma 

127 

PAGE 

Pronophora  tritostlcha  ...     107 

Proscrana  104 

poliophylla 105 

tephropis     104 

Prostanthera  coccinea  . -  ...     190 

ovalifolia 186,  187 

Proteace^  ...         ...  68,243 

Pseudodypsus  ilias       652 

innotatus     ...         ...        652,  653 

Pseudonotis        ...         ...         ...     648 

danis  ...         ...         ...     649 

milo 649 

turner!         648 

Psoralea  eriantha  ...         ...       58 

Psychopsis  coelivagus 368 

insolens       ..  368 

mimica        367,368 

Pteris  falcata      494 

Pterocera  ...         ...         ...     680 

Pterostichus       252 

])hiUipsi      278,  279 

phylarchus 252 

regalis  ...         ...         ...     252 

Pterostylis  mutica         196 

reflexa  580 

Ptilostibes  250 

Ptychognathus  ...         ...         ...     559 

Puffinus  brevicaudus 643 

Pultensea  plumosa         ...         ...       58 

styphelioides  573 

ternata        573 

villosa         537 

Pupa  (Vertigo)  madeayi        ..        19 

Pupina  crossei 20,29 

nitida  20,29 

Pupisoma  ...         ...         ...       21 

circulitum   ...         ...         ...       21 

Purpura 630 

amygdala 599,  600 

pseudamygdala        599,  600,  618 
scabra  ...         ...         ...       26 

Purpura  (CroniaJ  amygdala...     599 
Pycnolejeunea  longidens         ...     494 

Pyralididse  247 

Pyralidinae  247 

Pyrgulina  perspectiva 10,  29 

senex  10,29 

umeralis      11,29 

zea 11,29 

Pyrula  australasia        ...         ...       27 

undulatus    ...         ...         ..        27 

Python  spilotes  var.  variegata       648 
Eadula  Mittenii 494 


INDEX. 


PAGE 

PAGE 

Ifaja  laevis 

329 

Rochefortia  donaciformi 

i        ...  7,28 

Ivamalina  leiodea 

371 

Rubiacca} 

...     541 

Kaiiialincje          

371 

Rulingia  sp 

...       55 

llamularia  graminicola 

379 

Rutaceae 

55,  536 

hordei          

379 

Salmo  iiideus    ... 

...     352 

primulfB       

379 

Salvelinas  fontinalis     .. 

...     356 

KaiieUa  granifera 

26 

Sapindacere        

...       57 

leucostoma  ... 

... 

26 

Saprolegnia  ferax 

...     495 

pulchra       

... 

26 

Sarcopteron  conscripta 

...       78 

Ranellina            

445 

Sarrothripa  metaspilella 

...       92 

madurii      

445 

symmicta 

...       92 

Ranula 

444 

SaiTothripinse    . . 

...       89 

Raiudaria       444,  448,  451,  456 

,457 

Sarticus 

...     252 

lahiata         

444 

Sassia      

445,  472 

llapanea ? 

48 

349 

Scala        

...       12 

crassifolia 

349 

Schoenus  bifidus 

...     245 

myrtillina 349 

350 

Jamesonianus 

...     245 

platystigma            ...        349 

350 

minutulus 

...     244 

llaparna  cyclostigma     . . . 

112 

pedicellatus 

...     244 

nebulosa      

113 

Rodwayanus 

...     243 

Retusa  nitida      

9,28 

trachycarpus 

...     244 

Rhabdotus          

252 

Scleranthus  biflorus 

...     541 

Rhamnacese        

57 

Scleropoides        

...     435 

Rhamnese            

536 

squamicollis 

435,  436 

Rhipidura  dryas 

207 

trianguliferus 

435,  436 

Rhynchocyon     546 

549 

Scleropus  sticticus 

...     436 

cernei          

549 

Scyphiphora  hydrophyla 

Lcea    ...     105 

Rhytida  beiaudi 

604 

Separatista         

..  23,24 

jamesi          

22 

chemnitzii   ... 

...23,24 

franklandiensis 

22 

gabrieli        

...23,24 

Risella    620,  621,  623,  627,  630, 

631, 

gracilenta 

...23,24 

[632,  633 

635 

Grayii 

...       24 

melanostoma 

623 

separatista  ... 

...23,24 

plana            

631 

torcularis 

...23,24 

Risellida3 621 

623 

Se2)ta       444,  446,  448,  ' 

151,  454,  456 

Risellopsis           ...            621,623 

631 

jmrkinsonia 

...       26 

Rissoa  atropurpurea     . . . 

206 

2)arkinsoniana 

444,  448 

australia3  var.  ochroleiica 

206 

ruhecula      

...     448 

cheilostoma 

206 

ruhkunda   

26,  448 

contabulata 

206 

scarlatina    ... 

..      448 

devecta        

206 

spengleri     

...     448 

dissimilis 

206 

triangularis 

...     448 

fiammea      

206 

Sejifidae 

...     448 

frauenfeldi ... 

206 

Septoria  stellariaj 

...     379 

incidata       

206 

Seringia  Hi/Hi 

...       55 

jacksoni 

206 

Seiyida  sipho     

...     602 

nitens           

206 

sulcata 

...     602 

olivacea       

206 

Serpidorbis         

...     603 

salebrosa     

206 

sipho            

...     602 

scrobiculata 

206 

Serpulus 

...     603 

Rissoidse 

200 

sipho 

...     602 

Rissoina  fasciata 

206 

Silene  inflata      

..      212 

hanleyi        

206 

Simplicia  phricozona    ., 

...     125 

JtVlll. 


INDEX. 


PAGE 

Simpidum      443,  449,  451,  452,  456, 
[457,  458,  459,  465,  471 

torosum       ...  443 

Sinusigera  ...       22 

Sipho  mimeticiis  ...         ...     480 

Sirius  badius      ...         ...         ...       17 

Sisura  nana        ...         ...         ..      207 

Sitala  barnarclensis       ...         ...21,29 

darnleyensis  ...         ...21,29 

maino  ..  ...         ..-       20 

nepeanensis  ..  ...20,29 

reedei  20,29 

Solaneae  ...         ...         ...         ...       62 

Solanum  simile  ...         ...         ...     190 

Spekia     635 

Sphserophoron  tenerum  ...     544 

Sphenodon  ...  554,  555,  556 

punctatus    ...         ...        558,  559 

Spodoptera         ...         ...         ...       85 

umbraticula  ...         ...       78 

Stachyohium  Jonesii     ...         ...     350 

Stellaria  media 379 

pungens       ...         ...  ...     575 

Stenanthemum  Scortechinii    ...       57 
Stephanocircus  dasyuri  ...     205 

Stephanopsime 491 

Stephanopsis      484,  491 

altifrons      ...  ...         ...     491 

aspera  491 

obtusifrons 491,492 

Stereulia  acerifolia        ..  ...     578 

diversifolia...  74,  18(),  190,  193, 

202,  203,  562,  564, 

567,  577,  579,  580, 

[644 

rupestris      579 

Sterculiacea'        ...         ...         ..        55 

Sticta  Billardieiii  372 

Freycinetii  ...          ...     372 

orygmaja     ...          ...          ...     372 

Richardii 372 

rubella         372 

Stictete 372 

Storena 484,  485 

ambomaculata  484,  485,  492 

formosa       ...          ...         ...     484 

scintillans  ...  ..  ..     486 

Storenosoma       ...         ...         ...     484 

Storeus 439 

Strepera  graculina         ...         ..      237 

Streplosiphon     ...         ...          ...     480 

poiphyrostoma        ..         ...     4S0 

Strombus  630 


PAGE 

Suphalasca  flavipes 

364,  365 

sabulosa 

363 

subtrahens  ... 

365 

Symmolpis 

...     127 

hyporrhoda 

128 

Sympis  parkeri  . . . 

79 

Symplocaceae 

.348 

Symplocos  Biiuerleni 

594,  595,  656 

candelabrum 

348 

paucistamineus 

594,  595 

spicata 

348,  594,  595 

Stawelii 

348 

Thwaitesii  ... 

...        594,595 

Syntheta 

84 

xylitis 

85 

Tanganyikia 

635 

Tapinocephalus . . . 

560 

Tarache  detrita  . . . 

112 

Tatusia  hybrida 

549 

peba 

549 

Tecoma  australis 

136 

Baileyana    . . . 

381 

Tectarius  ...  620,  622,  631,  632 

bullatus       632 

montroudtri  ...         ...     632 

nodulosus    ...         ...         ...     621 

pagodus       631,632 

spinulosm   ...         ..  ...     632 

tectum-persicum    ...         ..      632 

Teinostoma        ...         ...         ...       14 

involuta       ...          ...          ...13,29 

Tellina  aurea      ...       28 

rostrata       ...         ...  ...       28 

vidiieUa        ...         ...         ...       28 

Tellina  (Phylloda)  aurea  ...       28 

Templetonia  egena        ...         ...     190 

Ternox  ohso/etns  ...         ...     320 

Testudo 559 

Telranthera  reikulala ...         ...       63 

Thai pochares  dividens  . .         ...  •     78 

ptctorora     ...         ...         ...       79 

Thamnophylloblasteie  ..  ...     371 

ThedaiUas         650,651 

Theiosia 250 

astheria        250 

Thelochistes  chrysophthalma...     372 

Thelxinoa  131 

calliscia       ...         ...         132,  133 

canescens    ...          ...         1 3 1 ,  1 32 

epiphracta 131,  132 

Jlavillalis     ...  ...  ...     131 

pleurotypa 132,  133 

Thelymitra  pauciflor a 213 

I  Thermesia  tenthrka      78 


JNDliX. 


PAGE 

PAGE 

Thiaosoma  distinct nm  ... 

488 

Triphoris  maculo.sus     ... 

...     614 

Thomisidi\!          

491 

2)feifferi       

...     616 

Thryi3tomeno  minutiflora 

190 

■scitidus 

...     616 

ThylacodcH  caperatns  .. 

19 

Triplex  denudaia 

...       26 

decmsatus 

603 

flavicunda  ... 

...       26 

sidcatus       

602 

f  rondosa      

...       26 

Thymeleffi           

63 

ruhicunda 

...       26 

Thysanotus  tuberosus  . . . 

575 

Tristoma             

...     606 

Thysonotis          

048 

Triton       444,  445,  446,  447, 

449,451, 

Tiliaceoe 

347 

[456,  471 

Tipasa  macrobela 

126 

abhotti          

..     468 

Titanosuchus     

560 

americanum 

...     477 

Trachylejeunea  elegantissima 

494 

annectans     .. 

...     469 

Tracy  triton        

445 

apenninictim 

...     445 

rincidum 

445 

aquatilis     

...     476 

Trichocolea  tomentella 

494 

australis     

...     478 

Trichosurus  vulpecula... 

558 

642 

hrasilianum 

...    477 

Trichotropis       

1 

2,23 

chlorosfomus 

...     460 

Trigonia... 

... 

454 

clavator 

26,  481 

Trigonotoma  violacea  ... 

253 

corrugatu.s 

...     476 

Triforis 606 

607 

612 

costatus         

...     448 

fasciata 

610 

615 

cyphus         

...     469 

graniferus 

610 

ehurneus      

...     463 

gramdatus 

613 

elongatus     ... 

...     461 

plicata         

606 

exaratus      

26,  460 

sdtulus        

616 

fratercidus 

...     480 

tasmanica 

... 

612 

/usifoi'me    ... 

...       26 

car.          

616 

fimformis 

448,  464 

torquatus     ... 

608 

gemniatus    ... 

...     461 

ustiUaUts     

608 

intermedius 

...     477 

Triphora          606,  607,  60S, 

617 

619 

labiosus 

...     462 

albovittata  ... 

609 

martinianum 

.       477 

ampulla      

615, 

619 

mimeticus 

...     480 

angasi          

610 

612 

nodifer        

...     478 

cinerea         ...            611, 

612 

619 

nodiferum 

...       26 

fasciata        ...            615, 

616, 

619 

nodifervs     

448,  464 

f  estiva         

615 

olJgostiru^   ... 

...     469 

gemmatum 

606 

philomehe 

...     465 

gi-anifera     ...  609,  610, 

616 

619 

pilearis        ...            448 

460,  476 

innotabilis 

608, 

619 

protensiis     ... 

...     468 

kesteveni     

618, 

619 

quoyi           

.      463 

labiata        

617 

radialis 

...     467 

maculosa    ...            614, 

615, 

619 

rubecula      

...     448 

nigrofusca 

611 

619 

sarcostoma 

...     481 

nocturna     

613, 

619 

saidid' 

.      478 

obesula        

618 

sinensis        ...              26 

461,  481 

pfeifferi 

616 

speciosus 

...     479 

regina          

608, 

619 

apengleri      

448,  462 

tasmanica  ...            612, 

616 

619 

strangei       

...     462 

tristoma       

606 

607 

succinctvs    ... 

...     460 

Triphorida^ 

606 

textiUs         

...     470 

Triijliovh            

606 

thersites 

...     461 

angasi         

... 

610 

tortirostris 

...     470 

INDEX. 


. 

PAGE 

Triton  transsylvanicuni 

..      445 

tritonis        

...     444 

tuheroms     

...     461 

tumuloiiu^ 

...     467 

veliei            

...     477 

verrucosus  

...     462 

vespaceiis    ... 

...     460 

vatitus        

.       476 

iroochi 

...     466 

Triton  (AnjohuccinumJ  mime- 

ticus     . , 

...     480 

Tritonium          ...  443,  444 

446,  456 

olearium 

...     460 

rut  Hum        

...     462 

specie-sum 

...     479 

Iritonopsis 

...     445 

suhalveatum 

..      445 

Trochus  conoidalis 

...     633 

Trochus  atropio'ptireus 

...       16 

Jimhriatus     

...    2,  4 

(jeortjianus 

3,  4,  5 

samoen-ns    ... 

16 

squamiferus 

2 

tantUlns       

...     632 

toitoriformis 

3 

urcillei 

...    3,  4 

Trophon 

473,  479 

goldsteini     

...       18 

speciosum 

.      479 

Tropidonotus      

...     559 

natiix          

...     558 

Turbo  helicina 

...       23 

helicoidrs 

...23,24 

■separatisia  ... 

...       24 

Turcia  montrouzieri     ... 

...     632 

Typhis  philippensis 

...     343 

Tyrtceosus           ...            411 

412,  438 

aemulus       

413,  420 

toquus          

413,  432 

assimilis      

414,  426 

biseriatus 

413,  418 

carinatus    ... 

413,  423 

castor 

413,  424 

cinerascens... 

413,  430 

concretus     

...     425 

corpulentus 

413,  429 

cra.ssirostris 

412,  432 

dolosus        

413,  420 

ichthyosomurf 

413,  422 

imitator       

412,  427 

incallidus 

412,  421 

inermis 

412,429 

lateralis       

...     430 

lalerarius    ... 

413,  428 

microthorax     413,  414 

417,  428 

I  Tyrtaeosus  pardalis  414, 

!  pollux  ...  413, 

I  quadratolineatus     . . 

religiosus    ...  414, 

j  urens 

I  ustulatus     ...  413, 

vetustus       ...  413, 

I  Umbelliferjc 
I  Uncinia  debilior 
!  tenella 

I  Uromyces  amygdali 

Urticaceie 
;  Usnea  barbata  var.  ceratina 
j  var.  dasypoga 

1  var.  hirta 

cornuta 
longissima  .. 
trichodea    . . . 
j  Usneeae    ... 
Uvanilla... 
Valsa  ambiens  . . . 
Venus  amygdalam 
australis 
disjecta 
gallinula     ... 
lagopus 
lamellatu     ... 
Verbenaceae 
Vermetus 

arenarius    . . . 
caperatus    . . . 
dentiferus   ... 
gigas 
Veronica  perfoliata 
Vertigo    ... 
Vitex  trifolia     ... 
Voluta  nucleus  ... 
pattersonia  ... 
Westermannia  conchidia 

ocellata 
Xanthorrhoea  sp. 
Xylorycta  heliomacula... 
Xyloryctidai 
Zagira  polycesta 
Zet lies  ch ordo2)hoidcs 

con-scripta   . . . 
Zidora  lodderre  ... 
ZodariidiL' 
Zodariinae 
Zootoca  .. 
Zophochroa 

anthracitis  ... 
j  leucochorda 

melanica     ... 


[Printed  off  April  7th,  1903.  J 


P.L..S.N.5.W.  1908 


PI.  XIX, 


P.L.S.N.S.W  1902 


m^^ 
/%^^^^ 


W.  AH.  del. 


Z7 


PI.  XX. 


P.L.S.N.SW.   I90H 


PI. XXI 


M.r.  del.adnat. 

EUCALYPTUS  PO  LYANTH  EIMOS,  Jc/^<5l/. 


P.L.S  N.S.W.,  1902. 


POLYPORUS  wYLiTT/E,  Coohc  &  Mass.ee. 


P.L.S.N.S.W.  190  2. 


PL.  XXIV 


nJ.c 


^  B.c/e/. 


LACERTA 
HETERONOTA. 


TRICH05URUS 
MINIOPTERUS 


PL.5.N.S.W.  1902. 


PI.  XXV. 


pmx 


R.B  del. 


ICHTHYOPHIS 
SPHENODON 
ZOOTOCA 


AMBLYSTOMA 
PLESiOSAURUS 

TR0PID0N0TU5 


P.L.SN.S.W.  1902 


PI.  XXVI 


R.B.del  "•"'^        '3 

GOMPHOGNATHUS 
PTYCH0GNATHU5 


ORNITHORHYMCHUS 
TE5TUD0 


PL  5.    N.S.W   1902 


^  MddKlgL^Kl® 


^' 


.x4 


,    -filpa 
i  Myall 


^Broken  Hill 

^IveriSn  ^'' 


\    Byrock 
^M-Drysdal 

Muda>l..i, 

■"■''™7°-;- -Dand^o 

^        Bodadah  Nlarruimnt:       'v 

CiljLunnia^^.. — ..,  \Wellin5h)nS       Mudcee 

Double  Reakj-'''  '\Peak  Hillt) 

c  Tvw^  Condobolin  Vmndle^^     Molong  , 


^HillsWn 


WyaIon5^...jj< 


!y 


'p^'x 


ofamundra^ 


;r2<"'§:?^ 


""QGrookwell 


Denillquin 

Rnley 


<?Uerilderie 
bBerrigan    Cu| 


'Cermanron  '    J 
ALBUSy  Av,.^Kiandra'o 


9VervisBay 

rEMANsBAV 


Map  OF       ^^"^©/^fl^ 

Scale  of  Miles 


Cooma 


"ortibala 


Eden  ^JwofotoBAy 
>'  Cape  Howe 


r  /I   S  M  A  /v 


SEA 


Railways  shown  thus . 


Autho 


'rs  route      — 


HECHobmsw.Mt 


PLXXVIII 


RTB.de/ad  nah 


5YMPLOC05    BAUERLENI,  R.TB. 


P.L.S.N.S   W.  1  902. 


^./Yc€^^  ai:/ 


AUSTRALIAN      MOLLUSCA. 


P.L.5.N.S.W.  1902 


PI. XXX 


R  I  SELLA 


H.  L.  Kesteven,  delt. 


P.L.S.N.S.W.  1902. 


^^^aa^f  a'c/ 


AUSTRALIAN     MOLLUSCA. 


PL.SN.  S.W.I  902. 


AUSTRALIAN     MOLLUSCA. 


P.L.S.N. S.W.I  902. 


AUSTRALIAN     MOLLUSCA. 


Issued  August  22nd,  1902. 


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BY    K.    GREIG    SMITH.  145 

DESCKIPTION  OF  PLATE. 

-Balsam  film  from  growth  upon  cane-gelatine.  The  asci  are  shrunken 
by  the  dehydration.  The  cellular  structure  of  some  of  the  asci  can 
be  made  out  from  the  ascus,  which  is  slightly  out  of  focus,  at  the 
bottom  left-hand  corner.  Stained  with  dilute  carbol  fuchsin,  and 
washed  with  alcohol,      x  1000. 

-Flagella  stained  by  the  night-blue  method,      x  1000. 


[Printed  off  August  20th,  1902.] 


PROCEEDINGS,  1902,  PART  I. 


CONTENTS. 

PAGE 

A  Note  on  two  Species  of   Astraliion  from  Port  Jackson.      By  H. 

Leighton  Kesteven     ...         ..  ...         ...         ...         ...         ...  2 

Studies  on  Australian  Mollusca.     Part  vi.      By  C.  Hedley,  F.L.S. 

(Plates  i.-iii.)      •     ...  7 

The  Gummosis  of  the  Sugar-Cane  {Bact.  vascularum,  Cobb).  By  E. 
Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the  Society. 
(Plates  iv.-v.) 31 

On  a  Gyrocotyle  from  Chimcera  Ogilhyi^  and  on  Gyrocotyle  in  general. 
By  W.  A.  Haswell,  M.A.,  D.Sc,  F.R.S.,  Challis  Professor  of 
Biology,  University  of  Sydney.     (Plate  vii.)  48 

Notes  from  the  Botanic  Gardens,  Sydney.     No,  8.     By  J.  H.  Maiden 

and  E.  Betche 55 

Further  Eemarks  upon   the   Mechanism   of    Agglutination.     By  E, 

Gbeig  Smith,  M.  Sc.  ,  Macleay  Bacteriologist  to  the  Society         ...         66 

Note  accompanying  an  exhibit  of  Persoonia  angulata,  R.Br.,  Uncinia 

tenella,  E.Br.,  and  Core.r  yracilh,  E.Br.     By  A.  A.  Hamilton  ...         73 

Note  in  Rectification  of  the  Synonymy  of  Eo2>saltria  yem-giana,  Qnoj& 

Gaim.,  and  E.  gularis,  Quoy  &  Gaim.     By  A.  J.  North,  C.M.Z.S.         74 

New  Genera  and  Species  of   Lepidoptera  belonging  to  the  Family 
Noctuiche.     By  A.  Jefferis  Turner,  M.D.,  F.E.S 77 

An  Ascobaeterium  from  the  Sugar-Cane,  with  Notes  upon  the  Nature 
of  the  Slime  {Bacterium  sacchari,  n.sp.)  By  R.  Greig  Smith, 
M.Sc,  Macleay  Bacteriologist  to  the  Society.     (Plate  vi.)  ...       137 

Elections  and  Announcements       ...30,76 

Notes  and  Exhibit.-, 30,  73 


Issued  October  17th,  1902. 


^^Vol.  XXVII. 
~^^^^>^  Part  2. 


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BY    THOMAS    G.    SLOANE.  325 

NoTONOMUS  LESUEURi,  Casteliiaii. 

Feronia  {PterostichiLs)  lesiieurii,  Casteln.,  Trans.  Roy.  Soc.  Vict. 
1868,  viii.  p.  210;  Fe7\  lesueuri,  Chaud.,  Ann.  Mus.  Civ.  Genova, 
1874,  vi.  p.  596. 

Elongate,  narrow,  siibcylindrical.  Black,  elytra  with  a  l)ronzy 
or  purplish  tinge.  Head  oval,  convex  (3-5  across  eyes)  ;  eyes 
with  orbits  reniform,  depressed.  Prothorax  as  long  as  broad 
(4-5  X  4 '5 5  mm.),  convex,  widest  (not  ampliate)  at  middle,  wider 
across  base  (3 -5  mm.)  than  apex  (3*3  mm.);  sides  lightly  arcuate, 
very  lightly  narrowed  anteriorly,  a  little  more  strongly  so  in  an 
even  curve  posteriorly ;  apex  truncate ;  basal  angles  widely 
rounded;  border  narrow,  not  extending  on  to  the  sides  of  base; 
posterior  marginal  puncture  on  border  at  basal  angle;  median 
line  line,  lateral  basal  impressions  oval.  Elytra  oval  (10  x  5 -3 mm.), 
very  lightly  rounded  on  sides,  convex,  deeply  striate;  interstices 
lightly  convex,  more  strongly  so  towards  apex,  three  inner  ones 
reaching  apex  in  full  width,  third  2-punctate,  eighth  narrow, 
convex,  narrower  than  ninth,  tenth  well  developed  before  apical 
sinuosities,  narrow,  not  long;  humeral  angles  not  marked;  basal 
border  not  raised  at  humeral  angles,  joining  lateral  border  with- 
out any  interruption;  lateral  border  very  narrow,  thickened  but 
not  reflexed  posteriorly.  Intercoxal  declivity  of  prosternum  flat, 
bordered;  of  mesosternum  flat:  metasternal  episterna  elongate. 
Length  16-5-19,  breadth  4-75-5-8  mm. 

Hah. — N.S.W.  :   Burrawang  (Sloane;  common  in  November). 

A  distinct  species,  isolated  by  its  elongate  metasternal  episterna, 
obtuse  basal  angles  of  prothorax,  humeral  angles  of  elytra  not 
dentate,  kc. 


PROCEEDINGS,  1902,  PART  2. 


CONTENTS. 


PAGE 


Preliminary  Note  on  the  Geology  of  the  Queensland  Coast,  with 
References  to  the  Geography  of  the  Queensland  and  N.S.  Wales 
Plateau.     By  E.  C.  Andkews,  B.A.     (With  nine  Text-figures)  ...       146 

Notes  on  the  Botany  of  the  Interior  of  New  South  Wales.     Part  vi. 

From  Marsden  to  Narrandera.    By  R.  H.  Cambage.    (Plates  viii.  x. )     1 86 

Note  on  some  Northern  and  North- Western  Australian  Grass  Finches. 

By  A.  J.  North,  CM. Z.S 207 

Notes  on  Jinicus  /ioZosc//rt';iMs,  R.Br.,  and  J.  priftmatocaiyus^  R.Br.; 

and  on  certain  other  New  South  Wales  Plants.     By  Edwin  Cheel      210 

4.— On  Et(caly2)tiis  Baueriana,  Seha,neY.     By  J.  H.  Maiden 214 

5.— On  Eucalyptus  calycogona,  Turcz.     By  J.  H.  Maiden      220 

On  Eucalyptue  melanophloia,F.v.M.,  Sind  its  cognate  Species.  By 
R.  T.  Baker,  F.L.S.,  Curator  and  Economic  Botanist,  Technolo- 
gical Museum,  Sydney.     (Plate  xi.) 225 

A  Gum  (Levan)  Bacterium  from  a  Saccharine  Exudate  of  Eucalyptui< 
Stuartiana  {Bacterium  eucalypti,  n.sp.).  By  R.  Greig  Smith, 
M.Sc,  Macleay  Bacteriologist  to  the  Society.     (Plate  xii.)  ...       230 

Notes  on  some  hitherto  unrecorded  Species  of  Plants  indigenous  in 
the  State  of  Western  Australia.  By  W.  V.  Fitzgerald,  F.S.Sc, 
Lond.,  F.R.H.S.,  Eng.    {Communicated  by  J.  H.  Maiden,  F.L.S.)      241 

New   Species   of    Queensland   Lepidoptera.     By   Thomas  P.    Lucas, 

L.R.C.P.,Ed.,L.S.A.,Lond.,M.R.C.S.,  Eng 246 

A  Revision  of    the  Genus  Notonomus  (Family  CaraUdcv,  Subfamily 

Feronini).    By  Thomas  G.  Sloane 252 

Elections  and  Announcements       209,  240 

Notes  and  Exhibits 205,237 


Issued  Deceniber  16th,  1902. 


Uo' 


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Part  3. 


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Messrs.  R.  Friedl^ndeb  &  Sohn,  Carlstrasse  11,  Berlin,  N.W, 


^j 


fSi 


NOTICE 


With  the  exception  of  Volumes  I.-VI.  of  the  Proceedings — of 
which  the  Society's  stock  was  totally  destroyed  in  the  Garden 
Palace  Fire — the  Publications  of  the  Linnean  Society  of  N.S.W. 
may  be  obtained  at  the  Society's  Hall,  Ithaca  Road,  Elizabeth 
Bay,  Sydney,  from  Dulau  &  Co.,  37  Soho  Square,  London,  W.,  or 
R.  Friedlander  &  Sohn,  Carlstrasse  11,  Berlin,  N.W.,  at  the  fol- 
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Proceedings  for  1883,  Vol.  VIII.— Part  1,  10s.;  Part  2,  5s.;  Part  3,  7s.; 
Part  4,  8s. 

Proceedings  for  1884,  Vol.  IX.— Part  1,  8s.;  Part  2,  12s.;  Part  3,  £1  5s.; 
Part  4,  £1  5s. 

Proceedings  for  1885,  Vol.  X.— Parti,  12s.;  Part 2,  7s.  6d.;  Part  3,  15s.; 
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Proceedings  for  1886,  Vol.  I.— Part  1,  10s.  6d.;  Part  2,  12s.;  Part  3,  13s.; 
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Proceedings  for  1887,  Vol.  II.— Part  1,  7s.;  Part  2,  8s.;  Part  3,  12s.; 
Part  4,  £1  7s. 

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Proceedings  for  1889,  Vol.  IV.— Part  1,  Us.;  Part  2,  16s.;  Part  3,  19s.; 
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Proceedings  for  1890,  Vol.  V.— Part  1,  lis.;  Part  2,  9s.;  Part  3,9s.; 
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SECOlSriJ     SEIRIES    (continued). 
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10s.  6d. 

Proceedings  for  1900— Part  1,  8s.;  Part  2,  10s.  6d.;  Part  3,  10s.  6d.; 
Part  4,  17s.  6d. 

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17s.  6d. 

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ment, 3s, 

A  reduction  of  20  per  cent,  on  the  above  charges  is  made 
to  Members  of  the  Society. 

The  Macleay  Memorial  Volume  [issued  October  13th,  18931. 
Royal  4to.,  li.  and  308  pages,  with  Portrait,  and  forty-two  plates. 
Price  £3  3s. 

CONTENTS. 

The  Hon.  Sir  William  Macleay,  Kt.,  F.L.S.,  M.L.C.  :  (By  the  Editor). 

Professor     W.     Baldwin     Spencer,     M.A.  :     Contributions     to     our 

Knowledge  of  Ceratodus.     Part  I. — The  Blood  Vessels.     (Plates  i,-v.) 

Prof.  F.  VV.  HuTTON,  F.R.S.  :  The  Pliocene  Mollusca  of  New  Zealand - 
(PI.  VI. -IX.)  —  Prof.  W.  A.  Haswell,  M.A.,  D.Sc.  :  A  Monograph  of  the 
TemnocephaUoi.  (PI.  x.-xv.)  —  Prof.  W.  A.  Harwell,  M.A.,  D.Sc.  :  On 
an  apparently  new  Type  of  the  Platyhelminthes  ( lirematoda  ?)  (PI.  xvi. ) — '■ 
Prof.  T.  Jeffery  Parker,  D.Sc,  F.R.S.,  and  Mias  J.  G.  Rich  :  Observa- 
tions on  the  Myology  of  Paliuurus  Edwardsii,  Hutton.  (PI.  xvii.-xxi)  — 
Prof.  J.  T.  Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B.,  B.Sc  :  Obser- 
vations upon  the  Anatomy  of  the  Muzzle  of  the  Omithorhynchiis.  (PI  xxii 
xxm.)  — Prof.  J.  T.^Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B.',' 
B.Sc.  :    On  the  Peculiar  Rod-like  Tactile  Organs  in  the  Integument  and 

Mucous  Membrane  of  the  Muzzle  of  Ornithprhynchus.     (PI.  xxiv.-xxvi.) 

C.  Hedley,  F.L.S.  :   On  Parmacochlea  Fischeri,  Smith.     (PI.  xxvii.) 

Prof.  R.  Tate,  F.L.S.,  F.G.S.,  &c.  :  On  the  Geographic  Relations  of  the 
Floras  of  Norfolk  and  Lord  Howe  Islands  —  Baron  von  Mueller,,  K.C.M.G., 
F.R.S. :  Notes  on  an  Undescribed  Acacia  from  New  South  Wales.  (PI. 
xxvin.) — Baron  von  Mueller,  K.C.M.G.,  F.R.S.,  and  J.  H.  Maiden, 
F.L.S. ,  F.C.S.  :  Description  of  a  New  Hakea  from  Eastern  New  South 
Wales,  (PI.  xxix. )  —  R.  Etheridge,  Jun.  :  A  Description  of  some  of  the 
Weapons  and  Implements  of  the  Alligator  Tribe,  Port  Essington,  North 
Australia  (PI.  xxx.-xxxv.)  —  N.  A.  Cobb,  Ph.D.  :  New  Nematodes  from 
Fiji  and  Australia.     (PI.  xxxvi.-XLii.) 

Descriptive  Catalogue  of  Australian  Fishes.  By  William 
Macleay,  F.L.S.  [1881].     A  few  copies  only.     Price  £1,  net. 

The  Transactions  of  the  Entomological  Society  of  New 
South  Wales,  2  vols.,  8vo  [Vol.  i.  five  Parts,  1863-66;  Vol.  ii.  five 
Parts,  1869-73 ;  all  published],  price  <£2,  net,  are  also  obtainable, 
but  neither  the  Parts  nor  the  Volumes  are  sold  separately. 


PROCEEDINGS,  1902,  PART  3 


CONTENTS. 


Contributions  to  a  Knowledge  of  Australian  Entozoa.  No.  ii.  On  a 
new  Species  of  Dintomum  from  the  Sawfish  Shark,  Pnstio2)horus 
cirratus.  Lath.  By  S.  J.  Johnston,  B.A.,  B.Sc,  Economic 
Zoologist,  Technological  Museum,  Sydney,     (Plate  xiii.) 

Notes  on  Australian  Uliopalocera,  Lycanida,  Part  i.  By  G.  A. 
Waterhouse,  B.Sc,  B.E.,  F.E.S.     (Plate  xiv.) 

On  a  new  Cryptocanja  from  Lord  Howe  Island,  together  with  Notes 
on  other  Plants  from  that  Island.    By  J.  H.  Maiden.    (Plate  xv.) 

The  Ulcei  Disease  (Black  Ophthalmia?)  of  Rainbow  Trout  f^il/icro- 
cocciLS  pi/ogenesj.  By  R.  Greig  Smith,  M.Sc,  Macleay  Bacteri- 
ologist to  the  Society 

Notes  on  Australian  JiKe?r?'opf6'rfl  and  their  Life-Histories.  By  Walter 
W.  Froggatt,  .^,^ 

Some  Records  of  New-,j^outh  Wales  Mosses.  By  W,  Forsyth. 
[Title  only]         ...       ■;L      ^ 

Census  Muscorum  Australiensium.  A  classified  Catalogue  of  the  Fron- 
dose  Mosses  of  Australia  and  Tasmania,  collate^ltrom  available 
Publications  and  Herbaria  Records,  by  the  Rev.  W.  Walter 
Watts  and  Thomas  Whitelegge,  F.R.M.S.  Part  i.  [Issued 
separately  as  a  Supplement  to  this  Part  of  the  Proceedings] 

Australian  Fangi,  New  or  Unrecorded.  Decades  i. -ii.  By  D. 
Mc Alpine,  Corresponding  Member 

On  a  new  Species  of  Ardisia  from  New  South  Wales.  By  R.  T. 
Baker,  F.L.S.,  Curator,  Technological  Museum,  Sydney.  (Plate 
xvi.) 

The  Bacterial  Origin  of  the  Gums  of  the  Arabin  Group.  By  R. 
Greig  Smith,  M.Sc,  Macleay  Bacteriologist  to  the..^ociety 

Revision  of  the  Australian  Ciirculionidce  belonging  to  the  Subfamily 
Cryptorhynclddes.     Part  v.     By  Arthur  M.  Lea,  F.E  S. 

Notes  on  Prosohranchiata.  No.  1.  Lotorium.  By  H.  Leighton 
Kestkven.     (Plate  xvii.)       ^. 

Descriptions  of  some  new  Araneidce  of  New  South  Wales.  No.  x. 
By  W.  J.  Rainbow,  F.L.S.,  Entomologist  to  the  Australian 
Museum.     (Plate  xviii.)        


Elections  and  Announcements 
Notes  and  Exhibits 


846, 
343, 


326 
331 
347 

,352 

358 
309 

369 
373 

380 
383 
4(is 
443 

484 
372 
370 


Issued  December  16th,  1902. 


Part  3. 

THE 


^3\ 

PEOOEBDINGS 


OF    THE 


LINNEAN   SOCIETY 


OF 


NEV^   SOUTH   VyALES 

FOR  THE  VKAR 

1902. 


SUPPLEMENT   TO   PART   I 


CENSUS  MUSCORUM  AUSTRALIENSIUM. 


A  CLASSIFIED  Catalogue  of  the  Frondose  Mosses  of  Australia 
AND  Tasmania,  collated  from  available  Publications  and 
Herbaria  Records,  by  the  Rev.  W.  Walter  Watts  and 
Thomas  Whitelegge,  F.R.M.S.     Part  I. 


SYDNEY : 
printed  and  published  for  the  society 

BY 

F.  CUNNINGHAME  &  CO.,  146  PITT  STREET, 

AND 

SOLD  BY  THE  SOCIETY. 
[Price,  3/-] 


-A_G!-E3SrTS   lOST   E-CrEOI>E 


-^^^     Messrs.  Dulau  &■  Co.,  37  Soho  Square,  London,  W  ^^a3^ 

^^(^<g  Messrs.  R.  Friedl^nder  &  Sohn,  Carlstrasse  11,  Berlin,  N.W.  jC^^^^\ 


NOTICE 


With  the  exception  of  Volumes  I.-VI.  of  the  Proceedings — of 
which  the  Society's  stock  was  totally  destroyed  in  the  Garden 
Palace  Fire — the  Publications  of  the  Linnean  Society  of  N.S.W. 
may  be  obtained  at  the  Society's  Hall,  Ithaca  Road,  Elizabeth 
Bay,  Sydney,  from  Dulau  &  Co.,  37  Soho  Square,  London,  W.,  or 
R.  Friedlander  &  Sohn,  Carlstrasse  11,  Berlin,  N.W.,  at  the  fol- 
lowing prices  : — 

First   Series. 

Proceedings  for  1882,  Vol.  VIL— Parti,  7s.  6d.;  Part  2,  10s.;  Part  3,  5s.; 
Part  4,  10s. 

Proceedings  for  1883,  Vol.  VIII.— Part  1,  10s.;  Part  2,  5s.;  Part  3,  7s.; 
Part  4,  8s. 

Proceedings  for  1884,  Vol.  IX.— Part  1,  8s.;  Part  2,  r2s.;  Part  3,  £1  5s.; 
Part  4,  £1  5s. 

Proceedings  for  1885,  Vol.  X.— Parti,  12s.;  Part  2,  7s.  6d. ;  Part  3,  15s.; 
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SEOOISTHD     SEIilES- 

Proceedings  for  1886,  Vol.  I.— Part  1,  10s.  6d.;  Part  2,  12s.;  Part  3,  13s.; 
Part  4,  12s.  6d. 

Proceedings  for  1887,  Vol.  II.— Part  1,  7s.;  Part  2,  8s.;  Part  3,  12s.; 
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Proceedings  for  1888,  Vol.  III.— Part  1,  15s.;  Part  2,  £14s.;  Part  3,  £1; 
Part  4,  18s. 

Proceedings  for  1889,  Vol.  IV.— Parti,  lis.;  Part  2,  16s.;  Part  3,  19s.; 
Part  4,  lis. 

Proceedings  for  1890,  Vol.  V.— Part  1,  lis.;  Part  2,  9s.;  Part  3,9s.; 
Part  4,  9s. 

Proceedings  for  1891,  Vol.  VI.— Part  1,  10s.;  Part  2,  9s.  6d.;  Part  3,  17s.; 
Part  4,  7s.  6d. 

Proceedings  for  1892,  Vol.  VII.— Part  1,  6s.  6d.;  Part  2,  4s.  6a.;  Part  3, 
8s.;  Part  4,  8s. 

Proceedings  for  1893,  Vol.  VIII.— Part  1,  5s.;  Part  2,  lis.;  Part  3,  6s.; 
Part  4,  9s. 

Proceedings  for  1894,  Vol.  IX.  —Part  1,  12s.;  Part  2,  12s.;  Part  3,  13s.; 
Part  4,  8s. 

Proceedings  for  1895,  Vol.  X.  — Part  1,  15s.;  Part 2,  8s.  6cl.;  Part  3,  10s., 
Supplement,  Is.  6d.;  Part  4,  12s. 


Proceedings  for  1896— Part  1,  9s.,  Supplement,  2s.  6d.;  Part  2,  6s.  6d.; 
Part  3,  7s.  6d.;  Part  4,  £1  7s.  6d. 

Proceedings  for  1S97— Part  1,  10s.;  Part  2,  8s.  6d. ;  Part  3,  9s.;  Part  4, 
12s.  6d. 

Proceedings  for  1898  .Part  1,  3s.;  Part  2,  6s.;  Part  3,  12s.;  Part  4,  14s. 


SECOISXID     SEKIB3S    (continued). 
Proceedings  for  1899— Part  1,  12s.  6d.;  Part  2,  12s.;  Part  3, 10s.;  Part  4 
10s.  6d. 

Proceedings  for  1900— Part  1,  8s.;  Part  2,  10s.  6d.;  Part  3,  10s.  6d,; 
Part  4,  17s.  6d. 

Proceedings  for  1901 —Part  1,  10s.;  Part  2,  9s.;  Part  3,  5s.;  Part  4, 
17s.  6d. 

Proceedings  for  1902— Fart  1,  7s.;  Part  2,  7s.  6d.;  Part  3,  7s.  6d.,  Supple- 
ment, 3s. 

A  reduction  of  20  per  cent,  on  the  above  charges  is  made 
to  Members  of  the  Society. 

The  Macleay  Memorial  Volume  [issued  October  13th,  18931. 
Royal  4to.,  li.  and  308  pages,  with  Portrait,  and  forty-two  plates. 
Price  £3  3s. 

CONTENTS. 

The  Hon.  Sir  William  Macleay,  Kt.,  F.L.S.,  M.L.C.  :  (By  the  Editor). 

Professor  W.  Baldwin  Spencer,  M.A.  :  Contributions  to  our 
Knowledge  of  Geratodiis.  Part  I. — The  Blood  Vessels.  (Plates  i.-v.)  — 
Prof.  F.  W.  Htjtton,  F.R.S.  :  The  Pliocene  Mollusca  of  New  Zealand. 
(PI.  VI. -IX.)  —  Prof.  W.  A.  Haswell,  M.  A.,  D.Sc.  :  A  Monograph  of  the 
Temnocephaleoi.  (PL  x.-xv.)  —  Prof.  W.  A.  Haswell,  M.A.,  D.Sc.  :  On 
an  apparently  new  Type  of  the  Platyhebninthes  (Trematoda  ?)  (PI.  xvi. ) — 
Prof.  T.  Jeffeby  Parker,  D.Sc,  F.R.S.,  and  Miss  J.  G.  Rich  :  Observa- 
tions on  the  Myology  of  Palmurus  Edioardsii,  Hutton.  (PI.  xvii.-xxi)  — 
Prof.  J.  T.  Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B.,  B.Sc  :  Obser- 
vations upon  the  Anatomy  of  the  Muzzle  of  the  Ornithorhynchus.  (PI.  xxii 
xxm.)  — Prof.  J.  T.  Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B.',' 
B.Sc.  :  On  the  Peculiar  Rod-like  Tactile  Organs  in  the  Integument  and 
Mucous  Membrane  of  the  Muzzle  of  Ornithorhynchus.     (PI.  xxiv.-xxvi.)  — 

C.   Hedley,  F.L.S.  :   On  Farmacochlea  Fischeri,  Smith.     (PI.  xxvii.) 

Prof.  R.  Tate,  F.L.S.,  F.G.S.,"&c.  :  On  the  Geographic  Eelations  of  the 
Floras  of  Norfolk  and  Lord  Howe  Islands  —  Baron  von  Mueller,  K.C.M.G. 
F.R.S. :  Notes  on  an  Undescribed  Acacia  from  New  South  Wales.  (Pi' 
xxvin.) — Baron  von  Mueller,  K.C.M.G.,  F.R.S.,  and  J.  H.  Maiden, 
F.L.S.,  F.C.S.  :  Description  of  a  New  Hakea  from  Eastern  New  South 
Wales.  (PI.  XXIX.)  —  R.  Etheridge,  Jun.  :  A  Description  of  some  of  the 
Weapons  and  Implements  of  the  Alligator  Tribe,  Port  Essington,  North 
Australia  (PI.  xxx.-xxxv.)  —  N.  A.  Cobb,  Ph.D.  :  New  Nematodes  from 
Fiji  and  Australia.     (PI.  xxxvi.-xlii.) 

Descriptive  Catalogue  op  Australian  Fishes.  By  William 
Macleay,  F.L.S.  [1881].     A  few  copies  only.     Price  £\,  net. 

The  Transactions  of  the  Entomological  Society  of  New 
South  Wales,  2  vols.,  8vo  [Vol.  i.  live  Parts,  1863-66;  Vol.  ii.  five 
Parts,  1869-73 ;  all  published],  price  .£2,  net,  are  also  obtainable, 
but  neither  the  Parts  nor  the  Volumes  are  sold  separately. 


Issued  April  9th,  1903. 


XXVII.  ^^ 

Part  4.  ^°  ^ 

THE  ^  i 

PROCEEDINGS 

OF    THE5 

LINNEAN   SOCIETY 


NEW   SOUTH    WALES 

KOR  THK  YEAR 

1902.       V 


PART    IV. 
Containing  Papers  read  in 
SEPTEMBER  (in  part)   to  NOVEMBER 

WITH  FIFTEEN  PLATES. 


SYDNEY : 

PRINTKD    AND    PUBLISJIED    FOR    THE    SOCIETY 
BY 

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AND 

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^a-O-EITTS   IN"  BTJiaOI>B3 


^,^c^>       Messrs.  Dulau  &  Co.,  37  Soho  Square,  London,  W.  '^^^ 

..^^/^Kj,  Messrs.  R.  Friedl^ender  &  Sohn,  Carlstrasse  11,  Berlin,  N.W.  ^^m*Nv^^ 


NOTICE 


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may  be  obtained  at  the  Society's  Hall,  Ithaca  Road,  Elizabeth 
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Proceedings  for  1882,  Vol.  VII.— Part  1,  la.  6d.;  Part  2,  lOs.;  Part  3,  58.; 
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SE3003SraD     SE3RIE3S_ 

Proceedings  for  1886,  Vol.  I.— Part  1,  10s.  6d.;  Part  2, 12s.;  Part  3,  138.; 
Part  4,  12s.  6d. 

s  for  1887,  Vol.  II.— Part  1,  7s.;    Part  2,  8s.;   Part  3,  12s.; 

s  for  1888,  Vol.  III.— Part  1,  15s.;  Part  2,  £14s.;  Part  3,  £1; 

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Proceedmgs  for  1891,  Vol.  VI.— Part  1, 10s.;  Part  2,  9s.  6d.;  Part  3,  17s.; 
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Proceedings  for  1892,  Vol.  VII.— Part  1,  6s.  6d.;  Part  2,  4s.  6d.;  Part  3, 
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Proceedings  for  1893,  Vol.  VIII.— Part  1,  5s.;  Part  2,  lis.;   Part  3,  6s.; 
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Proceedings  for  1896— Part  1,  9s.,  Supplement,  28.  6d.;  Part  2,  68.  6d.; 
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Royal  4to.,  li.  and  308  pages,  with  Portrait,  and  forty-two  plates. 
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CONTENTS. 

The  Hon.  Sir  William  Macleay,  Kt.,  F.L.S.,  M.L.C.  :  (By  the  Editor). 

Professor  W.  Baldwin  Spencer,  M.A.  :  Contributions  to  our 
Knowledge  of  Ceratodus.  Part  I. — The  Blood  Vessels.  (Plates  i.-v.)  — 
Prof.  F.  W.  HuTTON,  F.R.S.  :  The  Pliocene  Mollusca  of  New  Zealand- 
(PI.  VI. -IX.)  —  Prof.  W.  A.  Haswell,  M.A.,  D.Sc.  :  A  Monograph  of  the 
Temnocephaleoi.  (PI.  x.-xv.)  —  Prof.  W.  A.  Haswell,  M.A.,  D.Sc.  :  On 
an  apparently  new  Type  of  the  Platyhelminthes  (Trematoda  ?)  (PI.  xvi. ) — 
Prof.  T.  Jeffeby  Parker,  D.Sc,  F.R.S.,  and  Miss  J.  G.  Rich  :  Observa- 
tions on  the  Myology  of  Palinurus  Edioardsii^  Hutton.  (PI.  xvii.  -xxi),  — 
Prof.  J.  T.  Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B.,  B.Sc  :  Obsei 
vations  upon  the  Anatomy  of  the  Muzzle  of  the  Ornithorhynchus.  (PI.  xxii. 
XXIII.)  — Prof.  J.  T.  Wilson,  M.B.,  Ch.M.,  and  C.  J.  Martin,  M.B., 
B.Sc.  :  On  the  Peculiar  Rod-like  Tactile  Organs  in  the  Integument  and 
Mucous  Membrane  of  the  Muzzle  of  Omithorhynchiis.  (PI.  xxiv.-xxvi.)  — 
C.  Hedley,  F.L.S.  :  On  Parmacochlea  Fischeri,  Smith.  (PI.  xxvil)  — 
Prof.  R.  Tate,  F.L.S.,  F.G.S.,  &c.  :  On  the  Geographic  Relations  of  the 
Floras  of  Norfolk  and  Lord  Howe  Islands  —  Baron  von  Mueller,  K.C.M.G., 
F.R.S. :  Notes  on  an  Undescribed  -4ca«a  from  New  South  Wales.  (PI, 
XXVIII.) — Baron  von  Mueller,  K.C.M.G.,  F.R.S.,  and  J.  H.  Maiden, 
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Weapons  and  Implements  of  the  Alligator  Tribe,  Port  Essington,  North 
Australia  (PI.  xxx.-xxxv.)  —  N.  A.  Cobb,  Ph.D.  :  New  Nematodes  from 
Fiji  and  Australia.     (PI.  xxxyi. -xlii.  ) 

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PROCEEDINGS,  1902,  PART  4. 


CONTENTS. 

PAGE 

Notes  on  some  New  South  Wales  Hepatics.     By  Rev.  W.   Walter 

Watts        493 

On  two  remarkable  Sporocysts  occurring  in  Mytilus  latus,  on  the  Coast 
of  New  Zealand.  By  Professor  W.  A.  Haswell,  M.A.,  D.Sc, 
F.R.S.     (Plates  xix. -XX.)  497 

6. — On  Eucalyptus  bicolor,  A.  Cunn.     By  J.  H.  Maiden  ...         ...     516 

7.  —  On  Eucalyptus  polyanthemos,  Schauer.     By  J.  H.  Maiden.     (Plate 

xxi.)         527 

Contributions  to  a  Knowledge  of  the  Flora  of  Australia.  Part  iv.  By 
R.'T.  Baker,  F.  L.S.,  Curator  and  Economic  Botanist,  Technolo- 
gical Museum,  Sydney,     (Plates  xxii.-xxiii.)...         ...         ...         ...     536 

On  the  Mammalian  and  Reptilian  Vomerine  Bones.  By  R.  Bkoom, 
M.D.,  B.Sc,  C.M.Z.S.,  Corresponding  Member.  (Plates  xxiv.- 
xxvi. )         545 

Notes  on  the  Botany  of  the  Interior  of  New  South  Wales.     Part  vii. — 

From  Forbes  to  Bathurst.     By  R.  H.  Cambage.     (Plate  xxvii. )    ...     561 

On  a  New  Species  of  Symplocos  from  New  South  Wales.  By  R.  T. 
Baker,  F.L.S.,  Curator,  Technological  Museum,  Sydney.  (Plate 
xxviii.)       594 

Studies  on  Australian   MoUusca.     Part  vii.     By  C.    Hedley,  F.L.S. 

(Plates  xxix.,  xxxi.,  xxxii.,  xxxiii.) 596 

Notes  on  Prosobranchiata.       No.  ii.  Littorinacea.       By  H.  Leighton 

Kesteven.     (Plate  XXX.)            620 

Ngarrabul  and  other  Aboriginal  Tribes.      Part  i.  Medical  and  Surgical ' 

Practice.     By  John  MacPherson,  M.A.,  B.Sc,  M.B.,  Ch.M.      ...  637 
Notes    on    Australian    Ehopalocera.     Lycanidce  Part  ii.      By  G.  A. 

Waterhouse,  B.Sc,  B.E 648 

On  the  Occurrence  of  Monograptus  in  New  South  Wales.     By  T.  S. 

Hall,  M. A.     (Communicated  by  John  Mitchell)      654 

Notes  and  Exhibits        495,592,650 

Elections  and  Announcements          593 

Donations  and  Exchanges     ...         ...         ...         ...         ..                      ...  658 

Title-page          i. 

List  of  Contributors  and  Titles  of  Papers iii. 

Contents            ii 

List  of  Plates  ...         ...                     ...         ...         ...         ...         ...         ...  xi. 

Corrigenda        ...         ...         ...         ...         ...         ...         ...         ...         ...  xii. 

List  of  New  Generic  Terms  proposed  in  the  Volume      ...  xii. 

Index  '